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VENUS (Jap, Jour. Maluc,) H# Vol. 58, No.4 (l999}: 175-190

New Species of the (: ) from the Collections of

the Yamaguchi Museum and the Hagi City Museum

Shigeo HoRi and Hiroshi FuKuDA

Y'okyoUniversib ] cif Fisheries, 4-5-7, Kbnan, Minato-ku, 7?)k.yo 108-8477 and Department of rvatural History, 7bkyo Metropotitan University, Hktchioji, 7bkyo 192-0397

Abstract: During detailed examination of unidentified specimens of the Pyramidellidae in the

Yamaguchi Museum and the Hagi City Museum, nine hiLherto undescribed species were rec-

ognized, and here described as new: Ondina etachisinoides, Chr:v:sallida stmpa, Trabecuta trun-

catettijbrmis, EgiXina kotoeae, Ettlimella toshikazui, kuraenohamana, 71 gtori- ami.yhimana, Pyrlgisc"s }:oshikoae and foshishigea choshuana. A new , }'bshishigea is

also elected.

Keywords: Pyrnmidellidae, new gcnus, speeies, taxonomy, Hagi City Museum,

Yamaguchi Museum, Yamaguchi Prefecture.new

Introduction

The Yamaguchi Museum and the Hagi City Museum have bQth mude major contributions to malacology in Yamaguchi Prefecture, The Yamaguchi Museum, Yamaguchi City, has in its collec- tiuns a large number of molluscan specimens from all over the Prefecture, including the collection donated by the late Mr, Takusuke Kawamoto, and is said to be one of the most important facilities for overviewing the molluscan fauna ef the Prefecture, Hori & Tsuchlda (1989, 1990), H. Fukuda et al. (1990), and Tsuchida & Hori (1990) carcfully examined and presented revisional reports on the Pyramidellidae in the Yamaguchi Museum, Thc Hagi City Museum is the central institution for malacology in northern Yamaguchi Prefecture. It contains a large number of molluscan spccimens t'rom the region, mainly collected by the ]ate Messrs. Yoshishige Ikeda and Takeichi Tada, and the pyramidellids in its collectiQns were revised in dctail by Hori (1996). Fellowing the examinations of the two museum collections detailed above, many specimens of the Pyramidellidae remained unidcntified. The present authors recently carTied out morphological re-examination of these specimens in detail, with several of them turning eut to belong tQ unde- scribed species. Onc of the present new species (Ondina etachisinoides) is here described with Mr. Hiroshi Yoshizaki, IhonoshD, Yanai, Yamaguchi Prefecture, Japan.

Materialsand methods

All shells were observed and photographed with stereonticroscopes (Nikon SM-Z; WILD M400) and a scanning electronic microscope (JSM-5200LV).

Abbreviations: D: shcll diameter (mm); H: Shell height (mm); HCM: Hagi City Museum, Emukai 525-4, Hagi 758-O041, Yamaguchi Prefecture. Japan; SEM: scanning electronic microscope; YMZ: Yamaguchi Museum (Zoology), Kasuga-ch6, 8-2, Yarnaguchi 753-O073, Yamaguchi Prefecture, Japan.

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Ihxonomy

Superfamily Gray, 1840 Fttmily Pyramidellidae Gray, 1840

Genus Ondina Dc Folin, t870

71}pe speciesi Ondina semiornata De Folin, 1872, by monotypy.

Ondina elachisinoides Hori, H, Fukuda & Yoshizaki, n. sp.

(Figs. 1-3)

71>pe materials: 26 specimens. Holotype YMZ-Mo 6019: H 1.22, D O.70, Kiyogahama Beach, Abu-ch6, Abu-gun, Yamaguchi Pref. (beach drift), Oct. 1, 1992, coll. by S. Hori; two paratypes HCM-Mo 2723: ditto. Dec. 30, l992, coll. by S. Hori; fbur paratypcs HCM-Mo 2724: ca. 10 m A deep, off Kiwado Heki-ch6, Otsu-gun, Yamaguchi Pref. (shelly sand piled in octopus traps), May 2, 1993, coll. by S, Hori; 17 paratypes YMZ-Mo 6020: off Kuwabara, Tokuyama City, Yamaguchi Pref. (shell sand pi]ed in octopus pots), Jan. 3, 1990, cell. by T. Sasaki; two paratypes YMZ-Mo 6021: off 6mishima Island, Ehime Pref., western Japan (sand brought to the Southern Seto Ihonosh6 Marinc Park, ]honosh6, Yanai City, Yamaguchi Pref.), Jan. 13, 1996, by H. Yoshizaki. Description: Shell ovate, moderately thick, lustrous, and translucently white. Protoconch (Fig. 3) of at least 1.5 whorls, diametcr 250 ptm, translucent, smooth; proteconch heterostrophic at an angle ef ca. 160- to teleeconch axis; 50-55% of protoconch subsumed in first teleoconch whorl, with earliest portion completely concealed. Teleoconch of 2.6 whorls, slightly convex-sided, Suture distinct, slightly tabulatc immediately abapically. Growth lines opisthocline, flexed forward (i.e. ventrally). Teleoconch sculpture of thin spirat grooves on abapical portion of each whorl; two grooves en penultimate whorls and ca, 1O on body whorl including base; grooves become feebler toward umbilicul region, Body whorl 74% of shel] height; periphery and base convex. Aperture 45% of shell height, ovate. Co]umella thick, concave. Single columellar fold on posteriormost part of celumella, oblique to co]umellar axis; rclatively smal1 but distinct. Inner lip covered with a thick callus. Outer lip modcrately thick with a shallow sinus at abapicalmost part. Palatal sculpture absent. Umbilicus widely epen.

71ype locality: Kiyogahama Bcach, Abu-ch6, Abu-gun, Yamaguchi Prefecture (34031'44"N, 131020'4S"E).

Etymolog),: Elacrhisina (Latin): a genus of the family Elachishinidac; -oides (I.atin): similar, like. The present species is namcd for its similarity in shell appearance with Etac'hisina (e.g. E. ziczac H. Fukuda & Ekawa, 1997).

Remarksi Ondina etachisinoides resembtes Odvstomia (Jordanula) megembtl},on Saurin (1958: p. 70, pl. 1, fig. 7) from South Vietnam, but differs from the latter in lacking a distinct narrow spi- ral groove on the periphery. Ondina etachisinoides can also be distinguished from Odostomia (J.) sulcatetla Corgan & van Aartsen, 199S (Saurin, 1959: p. 240, pl, 3, fig. 26, as Odostomia (J.) in.fin- sulcata Saurin, 1959) from Victnam by its broader shell (DIHi O.57 in specimen with 2.6 whorls vs. O,4t in specimen with four whorls). The present species is placed in the gcnus Ondina by virtue of its ovate shell, opisthocline growth lines, and spiral grooves on the base, all common features with the type species Ondina

semiornata De Folin, 1872. Here the nominal species belonging to Ondina is recerded from Japan fbr the first time, Although H. Fukuda (1995) recorded a congeneric species from the Ogasawara Islands, it was an unidentified one.

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Genus Chr[ysattida Carpenter, 1856

Txpe species: Chemnitzia communis C. B, Adarns, 1852; by original designation.

tht:ysatlitla stttpa n. sp.

(Figs. 4-8)

71},pe materials: Seven specimens. Holotype YMZ-Mo 3282: H 3.40, D 1.55. Tokuyama Bay, Yamaguchi Pref., eoll. by Takusuke Kawamoto; six paratypes YMZ-Mo 6022: off Kuwabara. Tokuyama City, Yamaguchi Pref. (shelly sand piled in octopus traps), Jan. 3, 1990, coll. by Takenori Sasaki. Description: Shell conical, thick, slightly lustreus, and opaquc white, Protoconch (Figs 6, 7) of ca. 1.5 whor]s, diameter 240 ptm, slightly translucent white, smooth; protoconch heterostrophic at angle of ca, 1 20" to teleoconch axis; ca. 30-35 % of protoconeh subsumed in fi rst teleoconch whorl, with earliest portion slightly exposed (Fig, 6). Teleoconch of 5.5 whorls, almost straight-sided. Suture distinct, slightly tabulate immediately abapically. Growth lines fine, prosocline, Teleoconch sculpture of conspicuous spiral cords, three in number on euch whorl; abapical cord broader than

adapical other two on evcry whorl; adapical twe cords on lst to 3rd teleeconch whorl clearly gran- ulose (Fig. 8); a part of additional cord appears on suprasutural part of 3rd teleoconch whorl and penultimate whorl. Interspaces of spiral cords distinct, width about half of that of adapicalmost cord on every whorl; interspaces sculptured with thin axial lirae. Body whorl 56 % of shell height; periphery convex, Periphcry of body whorl encircled by peripheral and subperipheral spiral cords; peripheral cord as wide as 3rd cord on body whorl; subperipheral cord as wide as 2nd cord on body whorl, Umbilical area with no spiral cerd, Aperture 35 % of she]1 height, ovate. Columella thick,

slightly concave. Single columellar.fold on posteriorniost part of columella; oblique to columellar axis; distinct, moderately developed. Inner lip covered with a thin callus. Outer lip moderately

thick. Palata] sculpture absent. Umbilicus extremely narrow. 7ype tocality: Tokuyama Bay, Yamaguchi Prefecture (about 34"OO'N, 131"45'E), Et);moiogy: Stupa (Sanskrit): a buddhist monument. The present species is named for its partly granulose pyramidellid spire which resembles the buddhist monument. Remarks: Chtlysattida st"pa resembles Miratda gemma (A. Adams, 1863) (Dall & Bartsch, 1906: pp. 356-357, pl. 22, t'ig, 1) from Japan and M. scopulorum (Watson, 1889: p. 485, pl. 31, fig. 5) from Hawaii and northwest Pacific, but it differs from the both in having spiral cords which are granulose only on adapical whorls. The present species is placed in thc genus Chtlysallida, by virtue ofits conical shell with strong axial and spiral ribs, of which the axial ones pass only faintly over the base while the spiral ones remains quite prominent, as in the type species Chtlysallida crommttnis (C. B. Adams, 1852).

Genus 7}"abec"la Monterosato, 1884

Txpe species: Ttahecuta J'eXfreysiana Monterosato, 1884, bymonotypy.

7}rabecula truncatellijbrmis n. sp.

(Figs. 9-12)

Four specimens, Kiyogahama 71ype materials: Holotype YMZ-MQ 6023: H 1.737coll.D O.80. Beach, Abu-ch6, Abu-gun, Yamaguchi Pref. (beach drift),Oct 1, 1992, by S. Hori; three

paratypes YMZ-Mo 6024: ditto. Description: Shell elongate ovo-conical, relativcly thin, ]ustrous, translucent white.Protoconch

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Figs 1-3. 0ndina elachisinoides Hori, H. Fukuda & Yoshizaki, n. sp. X Jk'? I: Si 7tb T :f F }- C2-3: SEM photographs). 1-2: shell of ho]otype (YMZ-Mo. 6019), H 1.22. D O.70. 3: umbilical view of protoconch of onc of paratypeg (YMZ-Mo 6021). Figs 4-8. Chrysallida stupa n. sp, f'y F V7f`}: l- (5-8: SEM photographs). 4-5: shell of holotype (YMZ-Mo 3282), H 3.40, D 1.55. 6-7: protoconch of one of paratypes (YMZ-Mo 6022). 6: api- cal view. The arrow indicates the exposed earliest portion of the protoconch; 7: lateral view. 8: seulpture on sccond whorl surface of holotype.

(Figs 11, 12) of 1.5 whorls, diameter 260 um, translucent, smooth; pretoconch heterostrophic at angle of ca. 120" to teleoconch axis; 30-35 % of protoconch subsumed in first tcleoconch whorl, with a half ef earliest portion exposed (Fig. 11). Teleoconch of 3.3 whorls, convex-sided, Suture well constricted. Growth lines fine, orthocline, Teleoconch sculpture of distinct axial costae: costae

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slightly fiexuous, ca. 22 in numbcr on body whorl. Intercostae slightly wider than costae, with three narrow, distinct spiral ridges at nearly equal intervals. Suprasutural part and periphery of body

whorl encircled by an additional spiral ridge, at which axial costae terminate and connect to each other. Body whorl 58 % of shell hcight; periphery convex. Base with faint, feeble axial costae. Aperture 38 % of shell height, ovate but slightly quadrate. Columella thin, slightly twisted at adapi- cal part, forming an indistinct, feeble columellar fbld. Inner lip coyered with a thin callus, Outer lip thin. Palatal sculpture absent, Umbilicus net open.

71ype locatity. : Kiyogahama Beach, Abu-ch6, Abu-gun, Yamaguchi Prefecture (34031'44"N, l31"29d45,iE). Etymology: 1'runcatetla (Latin): a genus of the family Truncatellidae;formis (Latin) shape. The present species is narned fbr its similarity with young shells of Truncatelta. Remarks: Trabec"ta truncatellijbrmis is similar to a European species, 1'rabecula eximia (Jeffi;eys, 1849) (e.g. van Aartsen, l977: p. 51, p]. 1, fig. 5), but differs from the latter in having less convex-sided whorls and in having a part of the earliest portion of the protoconch clearly exposed. The present species is placed in the genus T?abecula, by virture of its having an ovo-conical shell with constricted sutures, distinct axial costac and slightly folded columel]a, all features of the type species 71rahecula jqfirre},siana Monteresato, 1884,

Genus Egitina Dall & Bartsch, 1906 71}ve species: Parthenia mariella A. Adams, 1860; original designation.

llgilina kotoeae n, sp.

(Figs 13-16)

BabeUa caelatior (Dall & Bartsch, l906): Fukucla et al,, 1992: pl. 47. no. 329.

7ype materials: 1t specimens. Holotype YMZ-Mo 6025: H 1.53, D O.90. Kiyogahama Beach, Abu-ch6, Abu-gun, Yamaguchi Pref. (beach drift), Dec. 30, 1992, coll. by S. Hori: three paratypes HCM-Mo 6026: ditto: Oct. 1, 1992, coll. by S, Hori; two paratypes YMZ-Mo 2725: ditto. Dec. 30, A 1992, coll. by S. Hori; five paratypes HCM-Mo 2726: ca. Ie m off Kiwado, Heki-ch6, Otsu-gun, Yamaguchi Pref, (shelly sand piled in octopus pots), May 2, 1993, coll. by S. Hori. Description: Shell conical, relatively thick, lustrous, and transluccntly grayish white. Protoconch (Figs 15. 16) of 1,5 whorls, diameter 250 um, translucent, with several minute and indistinct spiral ribs (Fig, 16); protoconch heterostrophic at angle of ca. 120e to teleoconch axis; 30-35 % of protoconch subsumed in first teleoconch whorl, with ca, 113 ofearliest portion exposed (Fig. 16). Teleoconch of four whorls, straight-sided, gradate. Suture distinct, deep, tabulate imme- diatcly abapically. Growth lines fine and prosocline, Teleoconch scuipture of axial costae; proso- clinc, ill-defined, with indistinct border, ca. 18 in number on body whorl. Intercostae as wide as or

slightly wider than costae. Immediately below shoulder, axial costae nodulose with elevated inter-

spaces which rise te connect adapical cnd of axial costae. Supraperipheral spiral keel on abapical

1/3 of each whorl; strong, distinct, connecting with abapical end of axial costae. Subperipheral spi- ral keel just abapical to supraperipberal keel on body whorl; wcaker than supraperipheral keel, Body whorl S9 % of shell height; periphery angulate, Base slightly concave, bearing ca. five extremely minute spiral lirae and feeble growth lines. Aperture 39 % of she]1 height, evate to sub- triangular. Umbilical area sunrounded by an indistinct spiral swelling. Columella thick, strongly

concave. Single columellar fold on posteriormost part of columella; fbld pcrpendicular to culumel- la axis, weak and small. Inner lip covered with a thick callus. Outer lip thin, expanding latero-

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Figs 9-12. 1'rabecula truncateU(formis n. sp. e Er7}Vti"K bf }L Lt (10-12: SEM photographs), Holotype (YMZ-Mo 6023). H 1.73, D O.gO. 9-10: ghell. 1 1-12i protoconch. Il: apical view. The arrow indicatcs the exposed earliest portien of the protoconch; 12i lateral view. Figs 13-16. Egitina kotoeae n. sp. = F Ji"f i: V (14-16: SEM photegraphs). Holotype (YMZ-Mo 6025), H 1,53, D O.90. 13-14: shell, 15-16: protoconch. 15: apical yicw. The arrow indicates the expesed earliest portion of the protoconch; 16: laLeral yiew.

abapically. Palatal sculpture absent. Umbilicus narrow.

7iype tocatity: Kiyogahama Beach, Abu-ch6, Abu-gun, Yamaguchi Prefecture (34031'44"N, 131029'45"E).

Etymology: Thc present species is named after Ms, Kotoe Hori, HagiCity, Yamaguchi

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Prefecture. Remarks: H. Fukuda et al. (1992: pl, 47, no. 329) identified the above new species with a com- mon Japanese species, Babella caelatior (Dall & Bartsch, 1906: pp, 347-348, pl. 17, fig, 9), However, EZ itina kotoeae can be clearly distinguished frem B, caetatior hy the following points: in B. caelatior); 2) axial 1) protoconch axis at an angle of ca. 1400 to teleoconch axis ((Ja. IIOO costae connccted with supraperipheral spiral keel (distinctly separate in B. caelatior); 3) highly ele- 11: vatcd and sharp supraperipheral spiral keel; 4) indistinct border between axial costae and inter- costae. Another common Japanese species Chr:ysaUida qt7iectuosa (Yokoyama, 1927) (e,g. Hori, 1996: p. 146, figs 11-12) can be distinguished from Egilina kotoeae in having a protoconch axis at an angle of ca. 150e to the teleeconch axis, a distinct border between axial costae and intercostae, axial costae which cross the supra- and subperipheral spiral keels, nodulose junctures of the supra-

and subperipheral spiral keels and axial costae, and non-elevated supraperipheral spiral keel. The present species is placed in the genus Egitina by virtue of its haying a conical shcll with strong axial costae between sutures, smooth intercostac, and spiral cords on the base, as in the type species of the genus EgiZina mariella (A. Adams, 1860).

Genus Eulimella Fothes & MacAndrew, 1846

7iMl)e .species: Ettlima macandrei Forbes, l844; by original designation, I・ Eutimetla toshikazui n, sp. (Fig. 17)

Turhonitta (Ptycheulimelta) misella stq?ramisetla Nomura: Kawamoto & Tanabe, 1956, p. 47, no. 620 lmisidentification]. Eulimelta (Eulimelta) sp.: Fukuda et at., 1990, p. 16, pl, 6, fig. 1.

Type materials: Four spccimens, Holotype YMZ-Mo 3299: H 1 1.35, D 2.05. Vicinity of Yanai, Yamaguchi Pref., July 1921, coll. by Nishiyama; three paratypes YMZ-Me 6027 (ex, YMZ-Mo

3299): ditto, Protoconch lost. Description: Shell slender conical, thick, lustreus, and opaque white. ! Teleoconch of 18.5 whorls, straight-sided. Suture distinct, slightly indented; siightly censtricted on earlier whorls. Growth lines fine, slightly opisthocline and feebly flexuous; sometime bearing coarse axial scar. Sculpture absent, Body whorl 20 9I) of shcll height; periphery strongly convex. Aperture 13 % ef shell height, rhombic-quadrate. Columella thick, Single columcllar fbld originat- oblique to ing at posterior part of columella and extending to posterior 115 part of columella; fold columellu axis and extremely feebie, hardly visible in ventral view. Columella and inner lip cov-

ered with a continuous, thick callus, Outer lip moderately thick. Pulatal sculpture absent. Umbilicus narrow. 7).ve tocality: Vicinity of Yanai, Yamaguchi Prefecture (about 33e57'N, 132e08'E). Saba-gun, Etymotog.y: The present species is named after Dr. Toshikazu Fukuda, Tokuji-ch6,

/1/ Yamaguchi Prefecture, "Turbonitia Remarks: Kawamoto & Tanabe (1956) recorded (PtycheulimeUa) miseUa "Tsukaduki-itokakegiri" supramisetta NQmura" with a Japanese name fi;om Vicinity of Yanai, The "Tsukaduki-itokakegiri" present materiaLs were also labeled in Japanese name with the locality of "Vicinity of Yanai", Since no other specimen related to Kawamoto & Tanabe's (1956) record above can be recognized in the Kawamoto Collection in the Yamaguchi Museum, the present mate- "T, riuls arejudged to be the voucher specimens of their record. Thus, (P.) misetta supramisella" in Kawamoto & Tanabe (1956) is here revised to Eulimella toshikaz"i. H. Fukuda et at. (1990)

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once figured one of the same materials as undeterrnined.

Eutimella toshikazui somewhat resembles Eulimella paclangensis Thiele (1925: p. 315, pl. 14, figs 26, 27) from Indenesia in general appearance of teleocench, but differs in having a much taller shell with a larger number of teleoconch whorls (H: 11.4 with 18.5 whorls vs. 2,8 wjth nine whorls).

The present species is placed in the genus E"limella, by virtue of its slender conic shelt, smooth surface and lack of a distinct columellar fold, as in the type species Eulimella macandrei (Forbes, 1844).

Genus Tberbonilla Risso, 1826 7ype species: 7)irhonitia costulata Risso, 1826; by subsequent designation of Herrmannsen (1852).

Turbonilla kuraenohamana n. sp. (Figs, 18-21)

Turhonilla (Chemnitzia) datei?. Nomura: Hori & Tsuchida. I989, pp. 66-67. 69 (incl. pl. 1), fig. 3a-b. 7leirbonilla sp. A: Hori, r996, p. I54, fig. 33.

71ype materiats: Five specimens, Holotype HCM-Mo 939 (ex. HCM 451-b): H 5.15, D 1.05. Kuraenohama Beach, Hagi City, Yamuguchi Pref. (beach drift), coll, by Yoshishige Ikeda; three paratypes HCM-Mo 2727 (ex. HCM 451-b): ditto; a single paratype YMZ-Mo 3302: Tokuyama

City, Yamaguchi Prcf., coll. by Eiji Tsuchida.

Description: Shell slender conical, moderately thick, lustrous, and translucent milky white with narrow, pale reddish brown spiral band on middle of each whorl and at periphery of body whorl. Proteconch (Figs 20-21) of ca. 2.5 whorls, diameter 240 um, translucent mjlky white, smooth; pro- toconch heterostrophic at Emgle of 900 to teleoconch axis; 5-10 % of protoconch subsumed in first teleoconch whorl, with earliest portiQn exposed (Fig. 20); umbilicus ot' protoconch closed (Fig. 21). Teleoconch of 11,7 whorls, slightly convex-sided. Suture distinct, constricted and mildly gradate, Teleoconch sculptured with axial costae; 17 in number on body whorl; prominent, nearly orthocline to slightly prosocline; abruptly connate at periphcry. Intercostae ca. twice as wide as costae, well- defined, srnooth. Body whorl 24 Vi} ef shell height; periphery convex. Base bcars only indistinct growth lines. Aperture 13 % of shell hcight, subquadrate, Columella moderately thick, feebly con-

cave, with no visible columellar fold. Inncr lip with no callus. Outer lip moderately thin. Palatal

sculpture absent. Umbilicus not open. 71ype loc:alit}?: Kuraenohamtt Beach, Hagi City, Yamaguchi Prefecture (34"24'25'N, 131e22L 23tE).

Etymology: The present species is named after the type locality, Kuraenohama Beach. Remarks: The holotype and three of the paratypes (HCM-Mo 2727) had originally tbrmed a part of the voucher specimens of 71trboniUa hiradoensis Pilsbry, 1904 in Ikeda (1960: p, 8, no. 325; "Hoso-itokakegiri") 1962: p. 12, no. 264; both as and Ikeda & Tuda (1963: p. 29, no. 656: as 71tthonilta (Pyrgolampros)). Hori & Tsuchida (1989) figured another paratype (YMZ-Mo 3302) " under the name of 7-tzrbonilla (Chemnitzia') datei?. Nomura, 1963", and Hori (1996) cited the helo-

types as undetermined.

7leirbonilla kuraenohamana certainly somewhaL resembles TLtrbonilla datei from northeast HQnshu (Nomura, 1936: p. 58, pl. 8, fig. 60a, b, as 71trbonilla (TLtrboniZla)): however, the former differs from the latter in having 1arger size (H: 5,2 with 1 l.7 whorls vs. 4.0 with eight whorls), slen- derer shell (HID: 5.2 in specimen of 11.7 whorls vs. 3.3 in specimen of eight whorls), and colored

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− Fig .17. Eulintella toshikazui n . sp .ト シ カ ズ ク チ キ レ . Shell of holotype(YMZ Mo 3299), H 11..35, D 2,05 .

・ n sp ク ラ エ ノ ハ マ カ ケ ギ リ − : Figs 18 21.Tt{rhonilla kω raenohamana . . イ ト (19 21 SEM photographs). − − − Holotype 〔HCM Mo 939), II5 .]5, D l.05 . t 8 19: shell .20 21 : protoconch,20 : apical vicw ;21 ; lateral Vlew . マ エ − Fig .22 . Tttrbθ nilla gloriamishimana n . sp .ミ シ ノ サ カ イ ト カ ケ ギ リ . Shell of holotype(YMZ Mo 6028),Hl2 .95,D3 .23 .

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spiral bands.

The present species is placed in the genus 7leirhonitta, by virtue of its slender conical shell with axial costae but with no spiral sculpture, as in the type spccies Titrbonilla tactea (Linnaeus, 1758) (= 7berboniita costulata Risso, 1826).

-tmana Turbonitla glbrinmish n.sp. (Fig. 22)

7ype materiats: Two specimens, Helotype YMZ-Mo 6028: H 12,95, D 3,23, ca. 100 m deep, off Mishima Island, Hagi City, Yamaguchi Pref. (trash fish Qf trawl boat), Aug. 14, 1991, coll. by Tomoyuki Sugimura; a single paratype YMZ-Mo 6029: Off Izu-Oshima Island, off Pacific coast of

central Honshu.

Description: Shell slender conical, thick, lustrous, and opaque orange but slightly translucent on earliest whorls; tinged with pale yellow on subsutural part, anterior half of base, columella, and anteriormost part of outer lip. Protoconch lost. Teleoconch of 12.7 whorls, feebly convex-sided,

slightly tabulate immediately abapically. Suture distinct, slightly constricted, Teleoconch sculpture of axiai costae and minute spiral grooves. Axia]s 30 in number en body whorl; prominent, feebly flexuous, slightly prosocline but nearly orthocline on body whorl; vanishing at periphery of each whorl, lntercostae nearly as wide as cestae, feebly dcprcssed, bearing sculpture of growth lines and spiral grooves. Costae and intercestae gently continuous to each other, their border ambiguous. Growth lines feebly flexuous, distinct, visible only on intercostae. Spiral groovcs extremely indis- tinct, widely separated, visible only between costac, numbering ca. 20 on body whorl. Body whorl

29 % of shell height; periphery convex. Base cncircled by ca. six extremely indistinct, thin, wide- ly separated spiral ridges. Aperture 17 % of shell height, quadrate, Co]umella thick, obliquely straight. Single celumellar fold en posterier 114-115 part of columella; fbld oblique to columellar

axis, extremely indistinct, visiblc only as a low swelling. Inner lip with a callus. Outer lip fractured.

Palatal sculpture absent. Umbilicus not open.

71ype locality: Off Mishima Island, Hagi City, Yamaguchi Prefecture

131"07'-08'E), ca. 100 m deep.

Etymolog},: Gloria (Latin): glQry, The name is refer to the glory of Mishima Island. Remarks: TurboniUa gtoriamishimana is similar to 71irhonilla gravih' (Laseron, 1951: p. 320, fig. 54, as Chemnitzia) from eastern Australia, but differs in having orange coloration, broader

teleoconch whorls, and narrower axial costae. The present new species is placed in the genus 7Ltrbonitla, by virtue of its slcnder conical shell with axial costae but with no marked spiral sculpture, as in the type species of the genus Turhonilla tactea (Linnaeus, 1758) ( := 71irbonilla cost"lata Risso, 1826),

Genus Pyrgiscus Philippi, 1841

7>ve species: r4fa Philippi, 1826; by subsequent designation byDall & Bartsch in Arnold (1903).Melania

Ib,rgiscus yoshthoae n. sp. (Figs. 23-25)

71ype materials: Two spccimens. Holotype YMZ-Mo 6030: H 4.60, D 2.00;a single paratype YMZ-Mo 6031: both ca. 20 m deep, vicinity ot' Otsushima Island, Tokuyama City, Yamaguchi

Pref. (dredged sand brought to Port Harumi, Tokuyama City), Mar. 18, 1979, coll. by Hiroshi,

Toshikazu, and Yoshiko Fukuda.

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Hori & Fukuda : New Specles of Pyramldellidae丘om Yamaguchi and Hagi Museums 185

・ コ − 一 Figs 23 25. Pyrgiscus yoshikoae n . sp ,ヨ シ イ ト カ ケ ギ リ (24 25 : SEM photographs). Holotype (YMZ − Mo 6030), H 4 .60 , D 2 .00 .23 24 : shell .25 : protoconch . Figs 26 −30. yoshishigea ch θ shuana n . sp .ヨ シ シ ゲ イ ト カ ケ ギ リ (27 .29 ,30 : SEM photographs ), Holotype

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186 VENUS: Vol, 58, No.4 (l999)

Des(:ription: Shell conical, thick, faintly lustrous, opaque milky white. Protoconch (Fig. 25) of at least one whorl, diameter 330 um, slightly translucent white, smooth; protoconch heterostroph- ic at angle of 1350 to teleoconch axis; 50-55 % of protoconch subsumed in first teleoconch whorl, with earliest portion completely concealed, Teleoconch of 6.5 whorls, feebly convex-sided. Suture distinct, slightly constricted. Teleoconch sculpture of prominent axial costae, spiraNirae and grooves, Axials 23 in number on body whorl; prominent, straight, nearly orthocline on first to sec-

ond whorls but becoming opisthocline on later whorls; surface of costae crossed by ca. nine spiral

grooves which divide costae into oval lozenges. On body whorl, axial costae continuous from shoulder to umbilical area, and obsolete between periphery and umbilicus. Width of intercostae nearly half that of costae but occasionally nearly equal to that of costae; adapicalrnost portion of

intercostae elevated level with costae; intercostae with ca. nine raised, distinct spiral lirae. Body

whorl 52 % of shell height; periphery convex. Base encircled by 14 distinct spiral grooves, well- scparatcd around pcriphery buL incised on umbilical area. Aperture 38 % of shell height, teardrop- shaped, Columella thick, nearly straight, Single celumellar fold on posteriormost part of columel-

la; fold eblique to columella axis, a]most obsolete and hardly visible in ventral view. Columella and

inner lip covered with a continuous, thin, broad callus. Outer lip relatively thin. Palatal sculpture

absent. Umbilicus not open. 71}ipe locality: Vicinity of Otsushima Island, Tokuyama City, Yamaguchi Prefecture (about

34"OO', 131043'E), ca, 20 m deep.

Etymotogy: The present species is named after Ms. Yeshiko Fukuda, Tokaji-ch6, Saba-gun,

Yamaguchi Prefecture.

Remarks: l5:ngiscus yoshikoae resembles il},rlgisciZla indenta Laseron (1959: pp. 239-240, fig. 171) t'rom Queensland, Australia, in shell appeurance and sculpture, but differs from the latter in

having axial costae breaked into lozenges.

The present species is allocated in the genus P.ytlgiseus, by virture of its slender conical shell, axial costae extending from thc shoulder to thc base on the body whorl, and spiral sculptures, as in

the type species P.y,giscus rbcfa (Philippi, 1841).

Genus Ybshishigea n. gen.

Tlype species: Ybshishigea choshuana n. sp. Diagnosis: Pyramidellids with slender conic shell bearing a protoconch heterostrophic at angle

of 900 to teleoconch axis, axial costae and smooth intercostae on teleoconch wherl surface, ne vis-

ible celumellar fold, and several spiral ribs over entire inner surftLce of teleoconch whorls.

Description: She]1 slender conica]. Protoconch of ca. 2.5 whorls, smQoth; protoconch het- erostrophic at angle of 90U to teleoconch axis; 5-10 % of protoconch subsumed in first teleoconch whorl, with earliest portion exposed; umbilicus of protoconch deep and wide. Sides of teleoconch

whor]s convex abapically. Suture distinct, well-constricted. Teleoconch sculpture ot' axial costae;

promincnt, nearly orthocline. IntercQstae smDoth. Costae and intercostae gently continuous to each other, their border ambiguous. Body wherl with convex periphery, Base with faint axial costae con- tinuing firom shoulder. Aperture ovate to slightly quadraLe. Columella mederately thick, slightly concave, with no visible fold, Several spiral ribs present over entire inner surface ef teteoconch whorls.

・Et]ymology: The prelient genus is named ttfter the late Mr. Yoshishige Ikeda, former curator of

the Hagi City Museum, in recognition of his significant contributions to the malacology of north-

em Yamaguchi Prefecture.

Remarks: Ybshishigea is closely related to the genera Tderbonitla Risso, 1826 and Nisiturris

Dall & Bartsch, 1906, their species having similar slender conica] shells bearing axial costae on

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Hori & Fukuda: New Species of Pyramidcllidae frem Yamaguchi and Hagi Museums 187

surface with smooth intercostae, and no visib}e columellar fold. Specimens of }'bshishigea never- theless differ from those in both other genera in having spiral ribs on the entire inner surface of the teleoconch whorls which are completely absent in Tberbonilla and Nisiturris. The phylogenetic relationships of Ybshishigea, 7berbonitta, and Nisiturris are unknown; howev- er, unpublished results ef comparative analysis of shell characters with anatomical characters in the several nominal genera (e.g. Odostomia, Chrysallida, Miraida, and others) suggests that pres- ence of internal teleoconch sculpture is possibly phylogenetically significant in the farnily Pyramidellidae (Hori, unpubl.). YZ]shishigea is therefore here established based upon the presence

of internal shell sculpture. Besides Z (:hvshuana, at least the following species are also assigned to loshishigea: Y, vinhi 265, (Saurin, 1959: p. 260, pl. 6, fig, 24, as 7berboniUa) (comb. nov.), l ph"ae (Saurin, 1959: p. pl. 7, fig. 10, as Chemnitzia (Nisiturris)) (comb. nev.), l subdetia (Saurin, 1959: p. 260, pl. 5, fig. 23, 4, as Chemnitzia) as 7)irbonilta) (comb, nov.), and Z tring"ieri (Saurin, 1959: p. 265, pl. 7, fig. (comb, nov.) all described from Vietnam.

Ybshishigea choshuana n. sp.

(Figs. 26-30)

Turbonilla sp. B: Hori, 1996, p. 154, fig. 34.

H 4.90, D 1,05,Off Futaoijima 71ype materiats: Three specimens. Holotype HCM-Mo 2728: Island, Shimoneseki City, Yamaguchi Pref. (dredged sand), Jan. 11, 1987, coll. by S. Hori; a sin- gle paratype YMZ-Mo 6032: ditto; one paratype HCM-Mo 940 (cx. HCM 451-c): Kuraenohama Beach, Hagi City, Yamaguchi Pref. (beach drift), cell. by Yoshishige lkeda. Description: Shell slender conical, moderately thick, lustreus, translucent milky white. Protoconch (Figs 29-30) of ca. 2.5 whorls, diameter 260 ptm, opaque white, smooth; protoconch heterostrophic at angle of 90" to teleoconch axis; 5-10 % of protoconch subsumed in first teleo- ca. 1/7 conch whorl, with earliest ponion exposed; umbilicus of protoconch deep, wide, occupying ca. 113 of protoconch diameter. Teleoconch of 10,S whorls; sides convex at abapical part (Fig. 30). Suture distinct, well-constricted. Growth lines fine, indistinct, and orthocline. Teleoconch sculpture of axial costae; 22 in number on body whorl; costae prominent, nearly orthocline but sometimes Cestae and slightly prosocline. Intercostae 1.5-2 times as wide as costae, well-depressed, smooth. intercostae gently continuous to each other, their border ambiguous. Body whorl 28 % of shell height; periphery convex. Base bears faint axial costae continuing from shoulder. Aperture 16 % of shell height, Qvate and slightly quadrate. Columella moderately thick, slightly concave, with no vis- ible fold. Colurnella and inner lip coyered with a thin, continuous callus, Outer lip modcrately extemally faintlyvis- thick, Four spiral ribs present over entire inner surface of teleoconch whorls, ible through shell (Fig. 28). Umbilicus not open. 71ype tocality: Off Futaoijima Island, Shimonoseki City, Yamaguchi Prefecture (about 34006'N, 130"47'E), of the Etymology: The present species is named after Ch6sha, the ancient province consisting northwestern part of present Yamaguchi Prel'ecture, where includes the type locality and another recognized locality of this species. Remarks: One of the paratypes (HCM-Mo 940) had originally fbrmed a part of the voucher specimens of 11trbonilla hiradoensis Pilsbry, 1904 in Ikeda (1960: p, 8, no, 325; I962: p. 12, no,

"Hoso-itokakegiri") 264; both as and Ikeda & Tada (1963: p, 29, no. 656, as Turbonilla (Pyrgolampros)). It has aiready been figured and treated as undetermined (Hori, 1996). Ybshishigea choshuana is similar to Nisiturris cr:y,staltina (Dall & Bartsch, 1906: pp. 341-342,

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188 VENUS : Vol .58. No .4 (1999 )

17 fig.3 as hr η ∫ pL , , 加 〃a (Nisiturris))from Samoa in shell appearance , but differs in having spi − ’ ra 且ribs ’ on theinnersurface ot the tcleoconch whorls .}〜 choh huana also resembles Z vinhi (Saurin, shell appearance and 1959)in presence of intemal spiral ribs , but is distinguished from the latter in having more deeply constrictcd suture and swollen sides of whorls .

Acknow − ’edgements : The authors here express their graしitude to Mr . Teruhisa Mitoki (YMZ )and Mr . Naoki Higuchi and Ms .Yasuko F. lt6(both HCM ), who obligingiy facilitated eur examjnatlons of museum collections ;and to Dr . Hiroshi Saito(NSMT )who assistcd them with しhe opcration of low−vacuum SEM 、 The authors are also grateful to Mr . Paul Callomon (Elle Scicntific Publications)for his valuable comments on manuscrip し The seniof author (S . H .)was supported fQr this work by the Fujiwara Natural History Foundation.Als〔⊃ the auth ‘〕r H , junior ( , F.)was supported by the JSPS Fcllowships for Young Scientists.

山 口 博 物 館 お よ び 萩 市郷 土 博 物 館 所 蔵 の トウ ガ タ ガ イ科 貝 類 (直腹 足 亜 綱 :異 鰓 上 目) の 新 種

堀 成 夫 ・福 田 宏

要 約

山 口 県 立 山 凵 博 物 館 お よ び 萩 市郷 土 博 物館 所 蔵 の 未 同 定 の ト ウ ガ タ ガ イ 標 本 群 を再 検 討 し た 結 果 , い つ く か の 未 記 載 種 が さ れ た の で と マ 確 認 新 種 し て 記載 す る 。 そ の う ち 1 種 ス ジ キ シ ズ ク ク チ キ レ は 吉 (山 口 , 崎 宏氏 県柳 丿1市 〉 と共 同 で 記 載 命名す る 。

Ondina ・ elachisinoides H 。 ri, Fukuda & Yoshizaki , n . sp . ス ジ マ キ シ ズ ク クチ キ レ (新 種 新 称)

殻 高 1.22mm ,殻 径 O.70 mm で 卵 形 。 成 長線 は 後 傾 し る の 湾 曲す 。 各 層 前 部 に 螺 溝 を 巡 ら す 。 軸 襞 は 小 さ く め は 斜 。模 式産 地 山 口 県 阿 武郡 阿武 町清 ヶ 浜 。 そ の 他 山 口 大 , 県 津 郡 凵置 町黄波 戸 沖 , 同 県 徳 山 市桑 原 沖 ,愛 媛 県 大 三 か ら ベ ム の 島沖 も産 出。 トナ 産 Odostomia (Jordanuta )megembr [yon Sau 血 .

1958 と Odostomia (」.) sulcatella Corgan & van Aartsen ,1998 は そ れ ぞ れ ,周 縁 に 明 瞭 な細螺 溝 を もつ こ と 殻 が 細 長 い こ と に よ と , り本種 区 別 さ れ る 。 な お , Ondina ス ジ マ キ シ ズ ク ク チ キ レ 属 (新称 ) の 公 称 種 が U 本 か ら 記録 さ れ る の は 初 め て で あ る 。

・ Chrysallida stupa n . sp . プ ッ トウ ク チ キ レ (新種 新 称 ) 〜 殻 高 3.40mm ,殻径 1.55 mm で 円 錐 形 。各層 に 3 本 の 顕 著 な螺 肋 を 巡 らすが ,初層 第 3 層 の 前 方

2 の 肋 は顆 粒 状 と な る 。体 層 周 縁 か ら に は 2 。 殻底 本 螺肋 が あ る 軸 襞 は 強 く斜 め 。 模 式 産 地 は 山 口 県

徳 山 湾 。 そ の 他 ,同 県 徳 山 市 桑 原 か ら の 沖 も産 出。 本 邦 近 海 産 Miralda gemma (A .Adanls ,1863) ホ ソ ア ラ レ ク チ キ レ お よ び M .scoputorum (Watson ,1889)ナ ガ メ ア ラ レ ク チ キ レ は 下 方 の 層 の 螺 肋 も顆粒状 と な る た め 異 な る 。

・ Trabecula truncatelliformis n . sp . ク ビ キ レ マ ガ イ ク チ キ レ (新種 新 称 )

殻 高 . 殻 .80 mm で い 1 73mm , 径 0 細 長 卵 円錐 形 、,殻 表 の 縦 肋 間 に は 3 本 の 螺 肋 が あ る 。縦 肋 は 縫 合 ・ 上 体 層 周縁 で 螺肋 に よ っ て 互 い に 連 続 さ れ て 止 ま る 。軸 襞 は 不 明 瞭 で 弱 い 。模 式 産地 は 山 口 県 阿 武

ヶ の ・ 郡阿 武町清 浜 。 欧 州 産 τrabecula eximia (Jeffreys,1849)は 側 而 が 強 く膨 ら み ,胎殻 頂が 完 全 に埋 没

す る た め 異 な る 、,

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Hori & Fukuda : Ncw Spccics of Pyramidellidae from Yamaguchi and Hagi Museums 189

Egilina kotoeae n . sp . コ トエ ク チ キ レ

殻 高 1.53mm ,殻 径 O.90 mm で 円錐 形 。 殻 表 の 縦 肋 は 各 層 の 肩 部 直 前 で 結 節 し て 互 い に 結 合 し ,さ

ら に 各 層 の 前 方 に お い て も周 縁 卜螺 肋 に よ っ て 連 結 す る 。 体 層 に は さ ら に 周 縁 ド螺 肋 が あ る 。 軸 襞 は

の 小 さ く水 平 。模 式 産 地 は 山 口 県 阿武 郡 阿 武 町 清 ヶ 浜 。そ 他 , 同 県 大 津 郡 日置 町 黄 波 戸 沖 か ら も産 出。

福 田宏他 (1992 :pl、47, no .329 ) に よ っ て Babelta caetatior (Dall & Bartsch,1906)ク サ ズ リ ク チ キ レ と し

て 図 示 さ れ た 種 は 本種 に 訂正 され る 。 ク サ ズ リク チ キ レ お よ び Chr.v,satlida ‘atfeetuosa (Yekoyama ,1927) ・ ヨ コ ス ジ ギ リ と は 胎 殻 の 角 度 ,縦 肋 の 明 瞭 さ と 螺 肋 と の 交 差 結 合 状 態 ,周 縁 上 螺 肋 の 強 さ に よ っ て 区 別 され る 。

・ Eulimella toshikazui n .sp . ト シ カ ズ ク チ キ レ (新 種 新 称 )

殻 高 11,35mm ,殻 径 2.05 で 細 長 い 円 錐 形 。 厳 線 1塒 に 粗 い 縦 痕 を伴 う。 そ れ 以 夕下の 周彡刻 ・ “ は な い 。軸 襞 は極 め て 弱 く斜 め 。模 式 産 地 は 山 口 県柳 井 付 近 。河 本 田 邊 (1956 ) に よ る 勲 厂δ碗 伽 ” (Pty. cheutimetta ) miseUa supramisetta ッ カ ヅ キ イ ト カ ケ ギ リ ,お よ び 福 山 宏他 (1990 ) に よ る Eutimella

sp ン ア の 巳 ン (Eulimella) .は 本 種 に 訂 正 され る 。 イ ドネ シ 産 Eulimella padan ,gensis ThieL 、1S98 (イ ドネ シ

ア 産 ) は 殻 が 低 く螺 層 数 が 少 な い た め異 な る 。

・ Turbonilla kuraenohamana n . sp . ク ラ エ ノ ハ マ イ ト カ ケ ギ リ (新 種 新 称)

い 1 1 殻 高 5」OInm , 殻径 1.[Omm で細 長 円 錐 形 n 各層 [央 と休 層 周 縁 に 淡赤褐 色螺 溝 を巡 らす 。 殻 表

い い の 縦 肋 は 周 縁 で 彑 に 結 合 す る u 縦 肋 間 は 平 滑 、軸 襞 は な .模 式 産 地 は 山 口 県 萩 市倉 江 ノ 浜 。そ の

・ 他 , 同 県 徳 11[市 か ら も産 出。堀 土 田 (1989 ) に よ る 跏 r わo π f〃α (C ぬθ’η 加 r4 の ゴα f2ご7 Nomura ,1936 , お

よ び 堀 (1996 ) に よ る Turbonilla sp . A は 本種 に 訂 止 さ れ る 。 Turbonilla datei Nomura ,1936 は 小 型 で 太

く,赤 褐 色螺帯 が な い た め 異 な る 。

・ Turbonilta gloriamishimana n . sp . ミ シ マ ノ サ カ エ イ ト カ ケ ギ リ (新種 新称 ) ・ ・ ・ 殻 高 12.95 mm ,殻径 3.23 mm ,細 長 い 円 錐形 で 燈 色 で ,縫 合 下 殻 底 前半 部 軸唇 外唇 前端 は 淡

つ 黄 色 。 殻 表 の 縦 肋 は 各層 の 周 縁 で 消 失 す る 。 縦 肋 間 に は不 明 瞭 な螺 溝 を も 。 軸 襞 は極 め て 不 明 瞭 で ー 斜 め 。模 式 産 地 は 山 口 県 萩 市 見 島沖 。そ の 他 ,東 京 都 伊 豆 大 島 沖 か ら も産 出 。東 オ ス トラ リ ア 産 の

Turbonillα gravis (Laseren ,1951 )は 白色 で 細 長 く,縦 肋 が 幅広 い た め 異 な る 。

・ Pyrgiscus yosh ikeae n . sp . ヨ シ コ イ トカ ケ ギ リ (新 種 新称)

殻 高 4.60mm , 殻 径 2.OO mm で 円 錐形 。殻 表 の 縦 肋 は 螺溝 と交 わ っ て 菱 形 の 粒 に 分 割 され る 。縦 肋

い 間 に は 明瞭 な螺 条 が あ る 。殻 底 に は 弱 縦肋 と明 瞭 な螺 溝 が あ る 。 軸襞 は極 め て 不 明瞭 で 斜 め 。 模 式 ー ン ス ラ ン の の 産地 は 山 口 県 徳 山市 大 津 島付 近 。 ク イ ド 産 P .yrgiscillaindenta Laseron 、tg59 は 縦 肋 表 面

が 円 滑 な た め 異 な る 。

・ Genus Yoshishigea n . gen . ヨ シ シ ゲ イ ト カ ケ ギ リ 属 (新 属 新 称〉

細長 い 円錐 形 . 殻 表 の 縦肋 間 は 平 滑 。 明 瞭 な軸 襞 は な い 。 後成 殻 の 内面 全体 に わ た っ て 数本 の 螺肋

が 走 る 。 属 模 式 種 は Yoshishigea choshuana sp . nov ,ヨ シ シ ゲ イ ト カ ケ ギ リ 。 後 成 殻 内 面 の 彫 刻 の 有 無 は

い ロ い ト ウ ガ タ ガ イ 科貝 類 に お て 系統 的 に 重 要 で あ る ∫能性が 高 (堀 , 未 発 表) た め , 筆者 らは こ の 彫

刻 の 存在 を重 視 し ,敢 え て 本属 を 設立 し た 。

・ Yoshishigea choshuana n . sp . ヨ シ シ ゲ イ トカ ケ ギ リ (新 種 新 称 )

殻 高 4.90・ imn ,殻 径 LO5 で 細 長 い 円錐 形 。殻 表 の 縦 肋 は 殻 底 ま で 舗 す る 。縦 肋 間 は 平滑 。軸

唇 に軸 襞 は 見 られ な V 後 成殻 の 内面 全体 に わ た っ て 4 本 の 螺肋 が 走 る 。 模式 産地 は [[[口 県 下 関 市蓋

井 島 沖 。 そ の 他 ,同 県 萩 市 倉江 ノ 浜 か ら も産 出 。 堀 (1996 ) に よ る Turbonitta sp . B は 本 種 に 訂 正 さ れ

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190 VENUS: Vol. 58, No.4 (1999)

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References

Aartsen, ven. J. J., 1977. European Pyramidellidae: I. Ch,pusallidd. Conchiglie, 13 (3-4): 49-64. Dall, W. H. & Bartsch, P., 1903. Pyramidellidae. In: Arnold. R. The paleontology and stratigraphy of the marine Pliecene and Pleistocene of San Pedro, Califomia, Mem, Caif Acad. Sci, 3: 269-285, pls 1-4, Dall, W. H. & Bartsch, P., 1906, Notes on Japanese, Indopacific, and American Pyramidellidae. Proc. U.S. IVdt. Mus, 30: 321-369, pls 17-26. Fukuda, H., 1995. Marine (Mo]lusca) of the Ogasawara (Bonin) Islands. Part 3: Additional records. Ogasawara Research, (21): 1-142. Fukuda, H., Mashino, K. & Sugimura, T., 1992. A Review of the MoUuscan Fauna of Yamaguchi Prqfecture, Wbstem Jtipan. IO + 100 + xxvi pp., 59 pls. Yamaguchi Museum, Yamaguchi (in Japanese). Fukuda, H., Tsuchida, E., Hori, S., Shikano, Y. & Miteki, T., 1990. Study on the of Yamaguchi - Pref, 1. Revision of the remarkable molluscan shells from Yamaguchi Pref. in Mr. T. Kawamoto's Collectien. (1) Gastropodu. Bull. }2]maguchi Mus., (16)i 1-46 (in Japanese with English summur}J). Herrmannsen, A. N., 1852. Indicis generum Malacozob'rum. Supptementa et corrigenda i-iv + 1-140. Cassel. Hori, S., 1996. A revisien of the family Pyramidellidae (Mellusca: Gastropoda: Heterostropha) from the northem district of Yamaguchi Prefecture deposited in the Hagi City Museum. 71he kiriyagai, 4 (1- 2): 139-162 (in Japanese with English summary). Hori, S. & Tsuchida, E., 1989. Revision of the family Pyramidellidae (Gastropoda: ) in the Yamaguchi Prefectural Muscum (1), The Chirihotan, 20 (3): 6S-70 (in Japancsc). Hori, S. & Tsuchida, E.. 1990. Revision of the fumily Pyramidellidae (Gastropodai Opisthobranchia) in the Yamaguchi Prefectural Museum (2). 111ie Chiribotan, 20 (4): 90-94 (in Japanese). Ikcda, Y., 1960. A Catalogue ofMoll"sks around Hdgi. 1 + 26 pp. Hagi City Museum, Hagi (in Japanese), Ikeda. Y.. 1962. Mtinual of Shell Collecting. 24 pp. Hagi City Museum, Hagi (in Japanese). Ikeda, Y. & Tacla, T., 1 963. A Catategue of Shell-bearing Mollusca of the Nbrthern District of Vbmaguchi Preflacture. 70 pp. Hagi City Museum, Hagi 0n Japanese). Kawamoto, T. & Tanabe, S., 1956, Catalogue ofMottuscan Shetls of lamasuchi Preflacture. 8 + viii + 170 pp, Yamaguchi Museum, Yamaguchi (in Japanese), Laseron, C, F., 1951, The New South Wales Pyrarnidellidae and the genus hathitda, Rec. Aust, Mus,, 22: 298-334.

Laseron, C. F,, 1959. The family Pyramidellidae

years 1873-1876, Rep. Sei. Res, Vby. H, M, S, ChaUenger Zoot,, 15: I-722, pls 1-50.

[Received: October 4, ]999; Accepted: October 30, 1999]

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