The Vegetation of the Mt Maungatawhiri Area, Coromandel Peninsula
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TANE 21, 1975 THE VEGETATION OF THE MT MAUNGATAWHIRI AREA, COROMANDEL PENINSULA by A.E. Wright* SUMMARY The major vegetation types for the area surrounding Mt Maungatawhiri, Coromandel Peninsula, are discussed, and their distribution is mapped. A species list of 253 vascular plants is presented. The vegetation is dominantly coastal and ranges from estuarine and rocky shore to mature rata {Metrosideros robusta) forest 300m above sea level. Much of the area has been extensively modified by man through farming, accidental burning and selective milling, although pockets of forest apparendy undisturbed by man do exist on the eastern slopes of Mt Maungatawhiri itself. INTRODUCTION These observations were made during the AUFC Scientific Camp to Wharekaho Beach, Whitianga (17-24 August, 1974). It is hoped that they will provide adequate information for future comparisons between the floras of the larger offshore islands of the Coromandel Peninsula and those of the mainland. Therefore, particular emphasis has been placed on the indigenous flora, and those adventive species which play an important part in each vegetation type. Each major vegetation grouping is briefly discussed mentioning important and unusual species, and its distribution shown in Figure 1, together with the typical localities later described. The lichen flora of this area has been studied in detail by Hayward et al.s VEGETATION Estuarine: The estuary of the Akeake River curves back behind Wharekaho Beach to form an area of low silty banks. The plant cover of these banks varies with the frequency of inundation by salt water. The main bank is largely covered by clumps of Juncus maritimus var. australiensis and Scirpus nodusus with a few shrubby Plagianthus divaricatus and manuka (Leptospermum scoparium). Lower areas are sparsely covered in the creeping grass Cynodon dactylon. Coastal forest and shore: Most cliff areas retain a largely natural vegetation similar to that in other areas of the Coromandel Peninsula. Tree species include pohutakawa {Metrosideros excelsa), karaka (Corynocarpus laevigatus), taupata (Coprosma repens), kawakawa (Macropiper excelsum), and kohekohe (Dysoxylum spectabile) with a groundcover of flax (Phormium tenax), Astelia banksii and the herbs Peperomia urvilleana, Apium australe and Disphyma australe. Bare soil on the foreshore provides a habitat for sowthistles (Sonchus spp.), catsear (Hypochaeris radicata), and Oxalis sp., with the natives NZ Convolvulus (Calystegia tuguriorum), Mercury Bay weed (Dichondra repens) and piripiri (Acaena anserinifolia) spreading back into the pastures. Many of the *Department of Geology, University of Auckland 3 Fig. 1. Map of the area surrounding Mt Maungatawhiri showing distribution of major vegetation types and location of the typical localities described in the introduction to the species list. stream valleys running inland through pasture have remnant coastal forest in their headwaters. Large puriri, (Vitex lucens) kohekohe and karaka formed the canopy, with almost all ground cover (except occasional tangles of supplejack — Ripogonum scandens) eaten out by cattle and sheep. The orchid Drymoanthus adversus was found in bud several times, epiphytic on karaka trees. Summit ridge of Mt Maungatawhiri: Between the main peak of Mt Maungata• whiri and the slightly lower knoll to the southsoutheast (see Fig. 1) runs a fairly sharp ridge dropping very steeply to the east and sloping down more gently to the west. The two high points appear to have been burnt off within the last twenty years and the trees on the flat area surrounding the trig have been cut down much more recently. The fallen branches have provided shelter for dense regeneration of shrubs 2 to 3m tall, for example pohutakawa and mapou (Myrsine australis). Between the knolls, the ridge proper has a mature canopy dominated by mahoe {Melicytus ramiflorus), puriri and karaka with few shrub species beneath them. Towards the south, the ground becomes more rocky, and ferns are the most common ground dwelling plants. The necklace fern (Asplenium flabellifolium), velvet fern (Ctenitis velutina) and Arthropteris tenella were all found only in this area. Arthropteris covers almost all boulders, roots and tree trunks over a large area. It exhibits the usual distinct difference between small, sterile, terrestrial fronds forming a dense covering over the substrate, and larger (up to 30cm) fertile fronds epiphytic on tree trunks. Where the canopy is thinner, the native grass Oplismenus imbecilis is abundant. Mature rata forest: Although some tracts have been selectively milled, a few areas appear to be in their virgin state (no old stumps or adventive species were seen). Many of the stream valleys to the west of Mt Maungatawhiri contain very dense forest with a wealth of ferns and shrubs. Filmy ferns are especially common, and over several hundred metres of one stream Hymenophyllum flexuosum is the dominant filmy fern, always rupestral. Treeferns, predomin• antly mamaku {Cyathea medullaris) are frequent in stream valleys and large stretches of these valleys are choked with supplejack. Large rata are dominant in the drier parts, forming the canopy with common rewarewa (Knightia excelsa) and karaka. Beneath these are several species of Coprosma and the NZ Honeysuckle (Alseuosmia macrophylla). Leptospermum scrub: Along the southern fringe of the mature rata forest is a belt of 20 to 30 year old manuka (Leptospermum scoparium) and kanuka (L. ericoides). Grazing animals, including cattle, have entered the scrub and opened up the shrub layer. Occasional manuka and kanuka have been knocked over, and the greater light penetration has allowed thicker seedling growth. The majority of ground cover is formed by lycopods, the ferns Lindsaea linearis and Phymatodes diversifolium, sundew (Drosera auriculata), mapou seedlings and various adventive weeds such as Hypochaeris, Erigeron and Cirsium vulgare. Further to the northwest, regeneration is at a more advanced stage, and less held back by interference by grazing animals. Consequently, the flora is considerably more diverse. The climbers Clematis paniculata and bush lawyer (Rubus cissoides) are common and there are many healthy saplings of kauri (Agathis australis), rimu (Dacrydium cupressinum) and tanekaha (Phyllocladus trichomanoides) up to 3m tall. It is interesting to note that kauri did not occur in the 5 forest or scrub to the east of Mt Maungatawhiri (apart from one unhealthy tree below the summit of the main hill) — it only became common in the north east i.e. more inland. Pasture: Pasture areas provide the best hunting ground for adventive species. Many such as the barberry (Berberis sp.), wooly mullein (Verbascum thapsus), foxglove (Digitalis purpurea), nettle (Urtica mens) and Californian and Scotch thistles (Cirsium arvense and C. vulgare) occur over most of the area and are a considerable nuisance to the farmers. Around streams blackberry (Rubus fruticosus Agg.) and fleshy weeds such as inkweed (Phytolacca octandra) are common. In the streams grow a few water-weeds such as the floating sweet grass (Glyceria fluitans), watercress (Nasturtium officinale) and duckweed (Lemna sp.). Several large colonies of the fern Hypolepis tenuifolia (the smaller, triangular fronded type) occur in depressions in the pastures and these appear to be untouched by grazing animals just as the adventive noxious weeds are. In one area, the fern Athyrium australe occurs commonly in the damp pasture. Remnants of the forest cover such as ancient puriri and one old ngaio (Myoporum laetum) dot the pastures, as do rotting treefern stumps in several areas (lichen zonation on these stumps has been studied by Hay ward et als). The scrambling pohuehue (Muehlenbeckia complexa) is common in coastal pastures, most noticeably on the summit of the pa (Fig. 1) described by Puch.9 There it forms cushions up to 50cm thick which are cut through by grassed animal trails. Swampy areas: Swamps have formed in several basins to the South and East of Mt Maungatawhiri, particularly where soil cover has slumped off the rock basement to block off a valley. Dominant plants are the raupo (Typha orientalis) and Scirpus lacustris, with the water fern Histiopteris incisa occuring around the margins of the swamps. 6 SPECIES LIST OF VASCULAR PLANTS Both native and adventive species are included. While the coverage of the former was fairly thorough, the adventive flora is incomplete, largely consisting of those species that play an obvious part in the communities already described. The total flora listed comprises 253 species, of which 196 species (78%) are indigenous to New Zealand. 59 of the native species are ferns and fern-allies. Nomenclature and ordering for all native plants except grasses follows Volumes 1 and 2 of 'Flora of New Zealand'.1'7 Native grasses follow Cheeseman.2 Exotic species (marked with asterisk) are named largely from Clapham, Tutin and Warburg (Flora of the British Isles);3 and exotic, non-European grasses from Lambrechtsen.6 Well known common names are given, and where applicable follow 'Standard Common Names for Weeds in New Zealand'.10 Each of the habitats and its community (already described) has been designated a number and a typical locality:8 Loc. 1. Estuary of Akeake River, Mercury Bay N44 223678 0-lm above sea level. Loc. 2. Coastal forest, cliff and shore vegetation, Mercury Bay N44 237678 0-10masl. Loc. 3. Summit ridge of Maungatawhiri Mt, Whitianga N44 232687 approx. 300m asl. Loc. 4. South western slopes of Maungatawhiri Mt, Whitianga (a) mature rata forest N44 228689 160m asl (approx.). (b) Leptospermum fringe N44 230684 120m asl (approx.). Loc. 5 (a) pasture to Southeast of Maungatawhiri Mt. (b) pasture around streams (c) pools in streams in pasture N44 236688 100m asl (approx.). Loc. 6. Swampy areas to Southeast of Maungatawhiri Mt. N44 234688 150m asl (approx.). The area studied centred on the latitude and longitude 36° 47'S 175° 47'E. Beside each species is the number of the habitat(s) in which it is most commonly found. Specimens of all ferns and fern allies are held in the author's Herbarium; specimens of most other plants are held in the Herbarium of the Auckland Institute and Museum.