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Morphology, Taxonomy, and Biology of Larval Scarabaeoidea
Digitized by the Internet Archive in 2011 with funding from University of Illinois Urbana-Champaign http://www.archive.org/details/morphologytaxono12haye ' / ILLINOIS BIOLOGICAL MONOGRAPHS Volume XII PUBLISHED BY THE UNIVERSITY OF ILLINOIS *, URBANA, ILLINOIS I EDITORIAL COMMITTEE John Theodore Buchholz Fred Wilbur Tanner Charles Zeleny, Chairman S70.S~ XLL '• / IL cop TABLE OF CONTENTS Nos. Pages 1. Morphological Studies of the Genus Cercospora. By Wilhelm Gerhard Solheim 1 2. Morphology, Taxonomy, and Biology of Larval Scarabaeoidea. By William Patrick Hayes 85 3. Sawflies of the Sub-family Dolerinae of America North of Mexico. By Herbert H. Ross 205 4. A Study of Fresh-water Plankton Communities. By Samuel Eddy 321 LIBRARY OF THE UNIVERSITY OF ILLINOIS ILLINOIS BIOLOGICAL MONOGRAPHS Vol. XII April, 1929 No. 2 Editorial Committee Stephen Alfred Forbes Fred Wilbur Tanner Henry Baldwin Ward Published by the University of Illinois under the auspices of the graduate school Distributed June 18. 1930 MORPHOLOGY, TAXONOMY, AND BIOLOGY OF LARVAL SCARABAEOIDEA WITH FIFTEEN PLATES BY WILLIAM PATRICK HAYES Associate Professor of Entomology in the University of Illinois Contribution No. 137 from the Entomological Laboratories of the University of Illinois . T U .V- TABLE OF CONTENTS 7 Introduction Q Economic importance Historical review 11 Taxonomic literature 12 Biological and ecological literature Materials and methods 1%i Acknowledgments Morphology ]* 1 ' The head and its appendages Antennae. 18 Clypeus and labrum ™ 22 EpipharynxEpipharyru Mandibles. Maxillae 37 Hypopharynx <w Labium 40 Thorax and abdomen 40 Segmentation « 41 Setation Radula 41 42 Legs £ Spiracles 43 Anal orifice 44 Organs of stridulation 47 Postembryonic development and biology of the Scarabaeidae Eggs f*' Oviposition preferences 48 Description and length of egg stage 48 Egg burster and hatching Larval development Molting 50 Postembryonic changes ^4 54 Food habits 58 Relative abundance. -
Quick Guide for the Identification Of
Quick Guide for the Identification of Maryland Scarabaeoidea Mallory Hagadorn Dr. Dana L. Price Department of Biological Sciences Salisbury University This document is a pictorial reference of Maryland Scarabaeoidea genera (and sometimes species) that was created to expedite the identification of Maryland Scarabs. Our current understanding of Maryland Scarabs comes from “An Annotated Checklist of the Scarabaeoidea (Coleoptera) of Maryland” (Staines 1984). Staines reported 266 species and subspecies using literature and review of several Maryland Museums. Dr. Price and her research students are currently conducting a bioinventory of Maryland Scarabs that will be used to create a “Taxonomic Guide to the Scarabaeoidea of Maryland”. This will include dichotomous keys to family and species based on historical reports and collections from all 23 counties in Maryland. This document should be cited as: Hagadorn, M.A. and D.L. Price. 2012. Quick Guide for the Identification of Maryland Scarabaeoidea. Salisbury University. Pp. 54. Questions regarding this document should be sent to: Dr. Dana L. Price - [email protected] **All pictures within are linked to their copyright holder. Table of Contents Families of Scarabaeoidea of Maryland……………………………………... 6 Geotrupidae……………………………………………………………………. 7 Subfamily Bolboceratinae……………………………………………… 7 Genus Bolbocerosoma………………………………………… 7 Genus Eucanthus………………………………………………. 7 Subfamily Geotrupinae………………………………………………… 8 Genus Geotrupes………………………………………………. 8 Genus Odonteus...……………………………………………… 9 Glaphyridae.............................................................................................. -
Scarabs Stlqikwmthlffnyotsieiiec
SCARABS STLQIKWMTHLFFNYOTSIEIIEC Occasional Issue Number 84 Print ISSN 1937-8343 Online ISSN 1937-8351 September, 2017 Notes on the Genus Pachypus (Coleoptera: WITHIN THIS ISSUE Scarabaeidae: Melolonthinae: Pachypodini) Notes on the Genus Pachypus ............................ 1 by Stéphane Le Tirant & René Limoges Ville de Montréal Delbert LaRue ................... 7 Montréal Insectarium 4581 rue Sherbrooke Elephant Dung Beetles ... 9 Montréal, Quebec Canada H1X 2B2 Dave Marqua .................. 16 Email: [email protected] Introduction P. sardiniensis Guerlach, Bazzato, Cillo, 2013 - (Sardinia - endemic). To date, no article or photograph of the Pachypodini tribe has ever The species are very similar, making been published in Scarabs. We identification difficult. There is also thought it would be interesting wide variability within each species. to present an overview of genus Pachypus, along with a few Genus Pachypus has antennae BACK ISSUES spectacular photographs of these with eight segments, five of them Available At These Sites: fascinating beetles. comprising the club. These beetles are usually 12 to 16 mm long. The Coleopterists Society www.coleopsoc.org/de- History males have a deeply excavated fault.asp?Action=Show_ pronotum on the disk. The Resources&ID=Scarabs The Pachypodini tribe was created females, few of which are found by Erichson in 1840 and contains in collections, have no scutellum, University of Nebraska a single genus: Pachypus (Dejean wings or elytra whatsoever. www-museum.unl.edu/ research/entomology/ 1821). Five species have been Scarabs-Newsletter.htm described thus far: Mysterious Biology EDITORS Pachypus caesus Erichson, 1840 - The male and female biology is Rich Cunningham (Italy. Sicily - endemic). fascinating. The male spends much [email protected] P. -
SCARABS Rich Cunningham William B
SCARABS Rich Cunningham William B. Warner Barney Streit 3889 Walnut Avenue 2338 W. Rockwell Court 1 Canton Road #4 Chino, CA 91710 Chandler, AZ 85224 N. Quincy, MA 02171 Occasional Issue Number 12 Kung ang amoy ay mabaho, sila ay darating July, 1995 site in early afternoon, and the A Simple Device for Extracting Pseudocotalpa would not emerge for Scarabaeidae and Other Coleoptera several hours, I began sifting the sand at WITHIN THIS from Sand various locations and ecotones. The most productive of these was below Petalonyx ISSUE.... by Delbert LaRue thurberi (Sandpaper Plant) which was abundant. After about two hours of sift- Many rare and highly sought after Scara- ing, with several breaks in between, I had baeidae can only be collected from cryp- accumulated a nice series of Aegialia, tic habitats amid sand dunes or similiar Diplotaxis, Pseudocotalpa, and a myriad sand deposits by carefully working of “tenebes.” I spent the rest of the after- High Plains Sifter through samples of the substrate (for noon relaxing in the shade, catching up example Aegialia, Psammodius, and on recent literature. After all, I had the some species of Aphodius, to name a peace of mind knowing that if the Museum of Comparative Zoology few). Pseudocotalpa did not emerge (or if I “cunninghamed” and was at the wrong A classic method is to take a shovel of sand dunes), a little effort with my sifter Entomology Supplies sand, drop it into a bucket of water, and had provided a nice series of specimens. let the prevalent quarry float to the top. -
Newsletter of the Biological Survey of Canada
Newsletter of the Biological Survey of Canada Vol. 39(2) December 2020 The Newsletter of the BSC is published twice a year by the Biological Survey of Canada, an incorporated not-for-profit In this issue group devoted to promoting biodiversity science in Canada. From the Editor’s Desk.............2 Membership....................3,15 President’s Report.................4 Feature Article: Sandhill Gold: The Goldsmith BSC Facebook & Twitter..........5 Beetle (Cotalpa lanigera, Contributing to the BSC Scarabaeidae, Coleoptera) in Newsletter.......................5 the Sandhills of Southwestern Highlights of the 2020 AGM...6 Manitoba Request for Specimens...........7 Robert Wrigley & Tim Arendse......19 Carabidae from across North America; Kevin Floate Certain Paraleptophlebia and Rhithrogena (Ephemeroptera) from eastern Canada; Steve Burian Elateridae from across Canada; Project Update Scott Gilmore Project Update Spider Diversity of British Spider Diversity of British Columbia: Columbia: Almost 900 Spe- Almost 900 Species & Still Counting cies and Still Counting Robb Bennett, Darren Copley and Robb Bennett, Darren Copley and Claudia Copley..............................8 Claudia Copley........................8 New Projects 1. Warm & Comfortable within Hollow Stems, Leaf-mines and Galls: Little New Projects known habitats for Entomologists & Botanists to explore, Peter G. Kevan, 1. Warm & Comfortable within Hollow Stems, Leaf- Charlotte Coates, Patricia Nunes mines and Galls: Little known habitats for Entomol- Silva, & Marla Larson...................11 -
(Scarabaeidae: Rutelinae: Rutelini) with a Key to the Larvae of the American Genera of Rutelini
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Papers in Entomology Museum, University of Nebraska State January 2001 Descriptions of the Larvae of Chlorota cincticollis Blanchard and Chasmodia collaris (Blanchard) (Scarabaeidae: Rutelinae: Rutelini) with a Key to the Larvae of the American Genera of Rutelini Mary Liz Jameson University of Nebraska - Lincoln, [email protected] Miguel Angel Moron Instituto de Ecologfa, A.C. (Sector SEP-CON,ACYT) Apdo. Postal 63 Xalapa Veracruz 91000, MEXICO Follow this and additional works at: https://digitalcommons.unl.edu/entomologypapers Part of the Entomology Commons Jameson, Mary Liz and Moron, Miguel Angel, "Descriptions of the Larvae of Chlorota cincticollis Blanchard and Chasmodia collaris (Blanchard) (Scarabaeidae: Rutelinae: Rutelini) with a Key to the Larvae of the American Genera of Rutelini" (2001). Papers in Entomology. 59. https://digitalcommons.unl.edu/entomologypapers/59 This Article is brought to you for free and open access by the Museum, University of Nebraska State at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Papers in Entomology by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. The Coleopterists Bulletin, 55(3):385-396. 2001 DESCRIPTIONSOF THE LARVAEOF CHLOROTACZNCTICOLLZS BLANCHARDAND CHASMODIACOLLARIS (BLANCHARD) (SCARABAEIDAE:RUTELINAE: RUTELINI) WITH A KEY TO THE LARVAEOF THE AMERICANGENERA OF RUTELINI MARYLIZ JAMESON W436 Nebraska Hall University of Nebraska State Museum Lincoln, NE 68588-05 14, U.S.A. mjamesonl @unl.edu AND MIGUELANGELMORON Instituto de Ecologfa, A.C. (Sector SEP-CON,ACYT) Apdo. Postal 63 Xalapa Veracruz 91000, MEXICO [email protected] Abstract Larvae of the genera Chlorota Burmeister and Chasmodia Macleay (Scarabaeidae: Rutelinae: Rutelini) are described for the first time based on the larvae of Chlorota cincticollis Blanchard and Chasmodia collaris (Blanchard). -
Board of Game and Inland Fisheries Meeting Agenda
Revised Board of Game and Inland Fisheries 4000 West Broad Street, Board Room Richmond, Virginia 23230 August 14, 2012 9:00am Call to order and welcome, reading of the Mission Statement and Pledge of Allegiance to the Flag. 1. Recognition of Employees and Others 2. Public Comments – Department plan to build a new headquarters under PPEA 3. Public Comments – Non-Agenda Items 4. Approval of July 10, 2012 Board Meeting Minutes 5. Committee Meeting Reports: Wildlife, Boat and Law Enforcement Committee: Mr. Turner, Chairman of the Wildlife, Boat and Law Enforcement Committee, will report on the activities of the August 7, 2012 Committee Meeting. The Committee will recommend the following items to the full Board for final action: Staff Recommendations – Fisheries Regulation Amendments Staff Recommendations – Diversity Regulation Amendments Staff Recommendations – Boating Regulation Amendments Staff Recommendations – 2012-2013 Migratory Waterfowl Seasons and Bag Limits Staff Recommendations – ADA Regulation Agency Land Use Plan Proposed CY2013 Board Meeting Schedule Finance, Audit and Compliance Committee: Mr. Colgate, Chairman of the Finance, Audit and Compliance Committee, will report on the activities of the July 25, 2012 Committee Meeting. The Committee will present the following reports: FY2012 Year-end Financial Summary Internal Audit FY2013 Work Plan - Final Action Education, Planning and Outreach Committee: Ms. Caruso, Chairwoman of the Education, Planning, and Outreach Committee Meeting. Ms. Caruso will announce the next Committee Meeting will be held on October 17, 2012 beginning at 10:00am. 6. Closed Session 7. Director's Report: 8. Chairman's Remarks 9. Additional Business/Comments 10. Next Meeting Date: October 18, 2012 beginning at 9:00am 11. -
A Comparison of Antlions, Bees, Darkling Beetles and Velvet Ants Across Sand Dune and Non-Sand Dune Habitats at Ash Meadows National Wildlife Refuge Nicole F
Utah State University DigitalCommons@USU All Graduate Theses and Dissertations Graduate Studies 5-2014 A Comparison of Antlions, Bees, Darkling Beetles and Velvet Ants Across Sand Dune and Non-Sand Dune Habitats at Ash Meadows National Wildlife Refuge Nicole F. Boehme Utah State University Follow this and additional works at: https://digitalcommons.usu.edu/etd Part of the Biology Commons Recommended Citation Boehme, Nicole F., "A Comparison of Antlions, Bees, Darkling Beetles and Velvet Ants Across Sand Dune and Non-Sand Dune Habitats at Ash Meadows National Wildlife Refuge" (2014). All Graduate Theses and Dissertations. 2078. https://digitalcommons.usu.edu/etd/2078 This Thesis is brought to you for free and open access by the Graduate Studies at DigitalCommons@USU. It has been accepted for inclusion in All Graduate Theses and Dissertations by an authorized administrator of DigitalCommons@USU. For more information, please contact [email protected]. A COMPARISON OF ANTLIONS, BEES, DARKLING BEETLES AND VELVET ANTS ACROSS SAND DUNE AND NON-SAND DUNE HABITATS AT ASH MEADOWS NATIONAL WILDLIFE REFUGE by Nicole F. Boehme A thesis submitted in partial fulfillment of the requirements for the degree of MASTER OF SCIENCE in Biology Approved: _________________________ _________________________ Dr. James P. Pitts Dr. Theresa L. Pitts-Singer Major Professor Committee Member _________________________ _________________________ Dr. Barbara Bentz Dr. Mark McLellan Committee Member Vice President for Research and Dean of the School of Graduate Studies UTAH STATE UNIVERSITY Logan, Utah 2014 ii Copyright © Nicole F. Boehme 2014 All Rights Reserved iii ABSTRACT A Comparison of Antlions, Bees, Darkling Beetles and Velvet ants across Sand Dune and Non-sand Dune Habitats at Ash Meadows National Wildlife Refuge by Nicole F. -
"White Grubs and Their Allies"
WHITE GRUBS AND THEIR ALLIES A Study of North American Scarabaeoid Larvae NUMBER FOUR : ENTOMOLOGY }``` ` .f -' eta STUDIES IN i, BY PAUL O. RITGHER Corvallis, Oregon OREGON STATE UNIVERSITY PRESS .- OREGON STATE MONOGRAPHS STUDIES IN ENTOMOLOGY JoHN D. LATTIN, Consulting Editor NUMBER ONE A Review of the Genus Eucerceris (Hymenoptera: Sphecidae) By HERMAN A. SCULLEN NUMBER TWO The Scolytoidea of the Northwest: Oregon, Washington, Idaho, and British Columbia By W. J. CHAMBERLAIN NUMBER THREE Stonefíies of the Pacific Northwest By STANLEY G. JEWITT, JR. NUMBER FOUR White Grubs and Their Allies By PAUL O. RITCHER © 1966 Oregon State University Press Library of Congress Catalog Card number: 66 -63008 Printed in the United States of America By the Department of Printing, Oregon State University Author's Acknowledgments THE INFORMATION published in this book represents Mrs. Patricia Vaurie, American Museum of Natural work done over the past thirty years while the History ; Bernard Benesh, Sunbright, Tennessee; E. C. writer was on the staffs of the Kentucky Agricul- Cole, University of Tennessee; W. A. Price, the late tural Experiment Station (1936- 1949), North Carolina H. H. Jewett, L. H. Townsend, and other members of State College (1949- 1952), and Oregon State Univer- the Kentucky Department of Entomology and Botany; sity (1952 -1966). I am especially indebted to the Ken- J. D. Lattin, Louis Gentner, and other entomologists at tucky Agricultural Experiment Station for permission Oregon State University; D. Elmo Hardy, University to reproduce much of the material contained in my Ken- of Hawaii ; W. F. Barr of the University of Idaho; tucky Bulletins 401, 442, 467, 471, 476, 477, 506, and Joe Schuh of Klamath Falls, Oregon; Kenneth Fender 537, which have long been out of print. -
Petition to List Six Sand Dune Beetles Under the U.S. Endangered Species Act
PETITION TO LIST SIX SAND DUNE BEETLES UNDER THE U.S. ENDANGERED SPECIES ACT Giuliani’s Big Dune Scarab Beetle Large Aegialian Scarab Beetle Photos: Richard Rust Petition Submitted to the U.S. Secretary of Interior Acting through the U.S. Fish and Wildlife Service Petitioner: WildEarth Guardians 312 Montezuma Avenue Santa Fe, New Mexico 87501 (505) 988-9126 January 29, 2010 1 I. Summary............................................................................................................................................ 1 II. The Great Basin................................................................................................................................ 1 III. Sand Dunes ..................................................................................................................................... 2 IV. Endangered Species Act and Implementing Regulations............................................................... 2 V. Imperiled Sand Dune Beetles Petitioned for Listing under the Endangered Species Act ............... 3 A. Threats to Sand Dune Beetles ...................................................................................................... 6 B. Imperiled Beetles at Crescent Dunes ........................................................................................... 8 • Crescent Dunes aegialian scarab • Crescent Dunes serican scarab Management of Crescent Dunes C. Imperiled Beetles at Sand Mountain and Blow Sand Mountain................................................ 11 • Hardy’s aegialian scarab -
Appendix 5: Fauna Known to Occur on Fort Drum
Appendix 5: Fauna Known to Occur on Fort Drum LIST OF FAUNA KNOWN TO OCCUR ON FORT DRUM as of January 2017. Federally listed species are noted with FT (Federal Threatened) and FE (Federal Endangered); state listed species are noted with SSC (Species of Special Concern), ST (State Threatened, and SE (State Endangered); introduced species are noted with I (Introduced). INSECT SPECIES Except where otherwise noted all insect and invertebrate taxonomy based on (1) Arnett, R.H. 2000. American Insects: A Handbook of the Insects of North America North of Mexico, 2nd edition, CRC Press, 1024 pp; (2) Marshall, S.A. 2013. Insects: Their Natural History and Diversity, Firefly Books, Buffalo, NY, 732 pp.; (3) Bugguide.net, 2003-2017, http://www.bugguide.net/node/view/15740, Iowa State University. ORDER EPHEMEROPTERA--Mayflies Taxonomy based on (1) Peckarsky, B.L., P.R. Fraissinet, M.A. Penton, and D.J. Conklin Jr. 1990. Freshwater Macroinvertebrates of Northeastern North America. Cornell University Press. 456 pp; (2) Merritt, R.W., K.W. Cummins, and M.B. Berg 2008. An Introduction to the Aquatic Insects of North America, 4th Edition. Kendall Hunt Publishing. 1158 pp. FAMILY LEPTOPHLEBIIDAE—Pronggillled Mayflies FAMILY BAETIDAE—Small Minnow Mayflies Habrophleboides sp. Acentrella sp. Habrophlebia sp. Acerpenna sp. Leptophlebia sp. Baetis sp. Paraleptophlebia sp. Callibaetis sp. Centroptilum sp. FAMILY CAENIDAE—Small Squaregilled Mayflies Diphetor sp. Brachycercus sp. Heterocloeon sp. Caenis sp. Paracloeodes sp. Plauditus sp. FAMILY EPHEMERELLIDAE—Spiny Crawler Procloeon sp. Mayflies Pseudocentroptiloides sp. Caurinella sp. Pseudocloeon sp. Drunela sp. Ephemerella sp. FAMILY METRETOPODIDAE—Cleftfooted Minnow Eurylophella sp. Mayflies Serratella sp. -
12-Month Finding on a Petition to List Six Sand Dune Beetles As Endangered Or Threatened
Endangered and Threatened Wildlife and Plants; 12-Month Finding on a Petition to List Six Sand Dune Beetles as Endangered or Threatened Literature Cited Andrews, F.G., A.R. Hardy, and D. Giuliani. 1979. The Coleopterous Fauna of Selected California Sand Dunes. California Department of Agriculture. BLM Contract CA-960-1285- 1225-DE00. Available at Bureau of Land Management – California Desert District Office. Moreno Valley, California. 142 pp. Army Map Service [Photo]. 1954. Available at U.S. Geological Survey – Henderson, Nevada. Army Map Service 1948, ID AR1IF0000050077; 1952, ID ARA010900600474; 1954, ID ARA010912015150; USGS 1970, ID N10NAPPW11576035, N10NAPPW02777042 and N10NAPPW11573051; Google Earth 1990, 1996, 1997, 2004, and 2010 Barrow, C.W. 1996. An Ecological Model for Protection of a Dune Ecosystem. Conservation Biology 10(3):888-891. [BLM] U.S. Department of Interior – Bureau of Land Management. 1988. Manual 1613 – Areas of Critical Environmental Concern. Washington, D.C. [BLM] U.S. Department of Interior – Bureau of Land Management. 1990. Environmental Assessment for Commercial Filming and Still Photography at Jean and Eldorado Dry Lakes, Big Dune, and Red Rock Canyon. Las Vegas Field Office, Las Vegas, Nevada. EA # NV054-90-43. [Service] U.S. Department of Interior – U.S. Fish and Wildlife Service [Memo]. 1992. Beetles from California and Nevada. Chris Nagano, Staff Entomologist. Sacramento Field Office, Sacramento, California. [BLM] U.S. Department of Interior – Bureau of Land Management [Memo]. 1992. Conversations with Richard Rust. Jeanie Cole, Wildlife Biologist. Las Vegas Field Office, Las Vegas, Nevada. [BLM] U.S. Department of Interior – Bureau of Land Management. 1997. Approved Tonopah Resource Management Plan and Record of Decision.