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THE RESEDACEAE a Taxonomical Revision of the Family (Final Instalment)

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THE RESEDACEAE a Taxonomical Revision of the Family (Final Instalment) 582.684.: MEDEDELINGEN LANDBOUWHOGESCHOOL WAGENINGEN • NEDERLAND • 78-14 (1978) THE RESEDACEAE A taxonomical revision of the family (final instalment) DR. MOUSTAFA S. ABDALLAH Professor of botany, Director, Flora and Phytotaxonomy Researches, Agricultural Research Centre, Ministry of Agriculture, Dokki-Cairo, A.R.Egypt and DR. H. C. D. DE WIT Ordinary professor of botany, Laboratory of Plant Taxonomy and Plant Geography, Agricultural University, Wageningen, The Netherlands Dateo fpublicatio n 30-VIII-1978 H. VEENMAN & ZONEN B.V.-WAGENINGEN - 1978 CONTENTS PREFACE SPECIAL PART (continuation) Reseda L 99 Key to the supraspecific taxa of Reseda 103 Subgenus 1. Neoreseda 104 Subgenus 2. Glaucoreseda 105 Subgenus 3. Luteola 105 Subgenus 4. Reseda 106 Section 1. Reseda 107 Series 1. Reseda 108 Subseries 1. Reseda 108 Subseries 2. Multilaciniata 108 Series 2. Asperula 109 Section 2. Leucoreseda 110 Section 3. Phyteuma 110 Series 3. Phyteuma Ill Subseries 3. Phyteuma 112 Subseries 4. Papulosa 112 Series 4. Crenulata 112 Key to the species and infraspecific taxa of Reseda 114 1. R. alba 123 2. R. alopecuros 146 3. R. alphonsi 149 4. R. amblycarpa 153 5. R. arabica 158 6. R. armena 163 7. R. aucheri 165 8. R. balansae 172 9. R. battandieri 174 10. R. buhseana 179 11. R.bungei 183 12. R. complicata 185 13. R. crystallina 188 14. R. diffusa 190 15. R. duriaeana 192 16. R.elata 195 17. R. ellenbeckii 198 18. R. fruticulosa 200 19. R. germanicopolitana 210 20. R.gilgiana 212 21. R.glauca 214 22. R. globulosa 218 23. R. gredensis 220 24. R. haussknechtii 222 25. R. hemithamnodes 224 26. R. inodora 226 27. R.jacquinii 228 28. R. lanceolata 231 29. R. lutea 235 30. R. luteola 259 31. R. macrobotrys 271 32. R. media 273 33. R. microcarpa 277 34. R. migiurtinorum 279 35. R. muricata 280 36. R.odorata 287 37. R. oligomeroides 296 38. R. orientalis 298 39. R. phyteuma 301 40. R. pruinosa 310 41. R.sa'adae 313 42. R. scoparia 316 43. R. sphenocleoides 318 44. R. stenobotrys 320 45. R. stenostachya 322 46. R. stricta 324 47. R. tefedestica 329 48. R. telephiifolia 330 49. R. tomentosa 332 50. R. tymphaea 334 51. R. undata 337 52. R. urnigera 343 53. R. villosa 348 54. R. virgata 350 55. R. viridis 352 Sesamoides [TOURN.] ORT 354 S. canescens 356 TAXA ET NOMINA REJICIENDA VEL DUBIA 372 SUPPLEMENTARY NOTES 393 INDEX TO PLANT NAMES 394 FIGURES 18-91 PREFACE The first instalment of this revision of the Resedaceae was published on July 7, 1967(p . 1-98, tab. 1-17). Now, more than 10year s later the second and final instalment follows; the demands of present-day university democracy are not conducive to voluminous and intricate scientific research and publi­ cations. The work was completed by Dr. ABDALLAH and myself but the msc. needed checking, polishing and editing before it could go to press. Mr. G. BOELEMA did this painstaking and time consuming task very ably, in high spirits and undauntedly. Thanks to his help and to the generosity of the authorities of the University at Wageningen, the revision is now completed and published. Miss L. TH. M. E. VAN DER RIET (now Mrs L. TH. M. E. MAAS) made the drawings, in the same way as she did for the first instalment. Two tables were done by Miss I. ZEWALD (nos 3 and 90). Mr. M. R. MOUTHAAN assisted in the registration and labeling of some 20.000 herbarium specimens. Prof. dr. F. A. STAFLEU and Dr. W. T. STEARN repeatedly advised. To the list of Herbaria are to be added: HEID: Bot. Inst. d. Univ., Berg- heimer Strasse 1, Heidelberg, Germany; MPU: Inst. Bot., Univ. Montpellier, Montpellier, France; TL: Lab. Bot., Fac. Sc, Allée Jules Guesde, Toulouse, France. Dr. ABDALLAH, Mr. BOELEMA and I spent months in trying to decipher col­ lector's names and the localities they noted on their labels. Often we were unable to arrive at a clear conclusion and it was then decided to approach some sort of spelling which seemed the best among several possibilities. It is hoped that the numerous shortcomings in this respect will not be criticized too severely; we tried our best. While this second instalment appears as No. 14 of the Mededelingen Land­ bouwhogeschool 78,bot h instalments, the textual parts and the atlas of figures united, will appear as an issue of Belmontia. H. C. D. DE WIT 5. Reseda LINNAEUS Gen.P Led .5,1754 , p.207 ;Houttuyn ,Nat .Hist .2(8) , 1777, p.722 ;Lamarck , Fl. Fr. 3, 1778, p. 202; Murr., Syst. Veg. ed. 14, 1784, p. 448; Necker, Elem. Bot. 2, 1790, p. 242;Moench , Meth. PI. 1794, p. 57; Murr., Syst.Veg .ed .15 , 1798, p. 369; Willdenow, Sp. PI. 2(2), 1799, p. 876; Brotero, Flora Lus. 2, 1804, p. 305; Lam. et DC, Fl. Fr. ed. 3,4 , 1805, p. 724; DC. in Duby, Bot. Gall. ed. 2, 1, 1828, p. 66; Reichenbach, Fl. Germ. 1830-32, p. 696; Link, Handb.3,1831 ,p . 323; Richter,Code xBot . Linn. 1835, p.462 ;Web be tBerthe ­ lot,Phytog .Canar . 1, May 1837, p. 101 ; Raf., Fl.Teil .3 ,Cent .VIII , 1837(Nov. - Dec), p. 123;Spach , Hist. Nat. Vég.Phan . 7, 1839,p .97 ;Bertoloni , Fl. It. 5, 1842, p. 23; Ledebour, Fl. Ross. 1, 1842, p. 235; Griseb., Spie. Fl. Rum. Bithyn. 1,1843 ,p .240 ; Gren.e tGodr. , Fl. Fr. 1,1848 ,p . 187; Hausm., Fl.Tir . 1851,p . 104;Visiani ,Fl . Dalm. 3,1852 ,p .94 ;Buchena u in Bot.Zeit . 11,1853 , p. 361-372, 377-390, tab. VIII (morphol.); Muell. Arg., Mon. Res. 1857, p.96 ,tab . 1-9, lit.! (eti nNeu eDenkschr . Schw.Ges .Zuer . 16,1858) ;Boissier , Fl.Or .1 , 1867,p . 423; Muell. Arg. in DC, Prodr. 16(2), 1868,p . 555 ;Oliv. , Fl.Trop .Afr . 1,1868 ,p . 103;Lang ei nWillkom m etLange ,Prodr . Fl. Hisp. 3, 1880, p. 889;Arcang. , Comp. Fl. It. 1882,p . 66; Battandier in Batt. &Trab. , Fl.Alg . 1888-90, p.83 ;Terraccian oi nCaruel , Fl.It . 10, 1894,p . 157;Rou ye t Foucaud, Fl.Fr .2,1895 ,p .239 ;Post , Fl.Syr .Palest . Sin. 1896,p . 111 ; Dur.e t Schinz, Consp. Fl. Afr. 1(2), 1897, p. 180; Coste, Fl. Fr. 1, 1901, p. 159; Pirotta, Fl. Col. Erit. in Ann. R. Inst. Bot. Rom. 8(5), 1903,p .259 ; Morstatt, Beitr.Kenntn . Resed.(i nFünfstück' sBeitr .Wiss .Bot .5 )190 3; Muschler, Man. Fl. Egypt 1, 1912,p . 438;Briq. , Prodr. Fl. Cors. 2(1), 1913,p . 123;Blatt. , Fl. Aden in Rec. Bot. Surv. Ind. 7, 1914-16, p. 110;Thonn. , Fl. PI. Afr. 1915,p 227(repr . Hist.Nat .Class .27,1962) ;id. , Fl.Arab ,i nRec .Bot . Surv.Ind .8(1) , 1919,p .46 ; Ramis, Bestimm. Fl. Aeg. 1929,p . 97; Dinsmore in Post, Fl. Syr. Palest, et Sin. ed. 2, 1, 1932, p. 137; Bolle in Engl, et Prantl, Nat. Pflz. fam. ed.2 , 17b,1936 ,p .684 ,687 ;Czerniak . in Komarov, Fl. URSS 8, 1939,p .607 ; Merr., Ind. Rafin. 1949, p. 132;Andr. , Fl.PI . Anglo-Egypt.Sud . 1,1950 , p. 69; Cufod. in Bull. Jard. Bot. Brux. 24, Suppl. 1954, p. 159; Keay in Hutch, et Dalz., Fl. W. Trop. Afr. ed. 2, 1(1), 1954, p. 108; Hutch, et Dalz., Fl. West Trop. Afr. ed. 2, 1(1), 1954, p. 108;Täckholm , Stud. Fl. Egypt 1956, p.33 1 Elf. etTaylo r in Hubb.e t Milne-Redh.,Fl .Trop .E .Afr. , Resedaceae, 1958, p 3; Ozenda, Fl. Sah.Sept . Centr. 1958,p .274 ;Clapha m et al., Fl. Br. Isl.ed .2 1962, p. 186; Quézel et Santa, Nouv. Fl. Alg. 1, 1962, p. 438; Yeo in Tutin Heywood et al., Fl. Europ. 1, 1964,p . 346; Rech, fil., Fl. Lowl. Iraq 1964,p 324; Coode in Davis, Fl. Turk. East Aeg. Isis 1, 1965, p. 499; Zohary, Fl Palaest. 1, 1966,p . 331. Luteola[Tournef. ] Webb et Berthelot, Phyt. Can. 1, May 1837, p. 104; Spach, Hist. Nat. Vég. Phan. 7, 1839,p . 103;Fuss , Fl. Transsilv. 1866,p .86 ; Rouye tFouc , Fl. Fr. 2,1895,p .250 . Arkopoda Rafinesque, Fl. Tell. 3, Cent. VIII, 1837 (Nov.-Dec), p. 73; Meded.Landbouwhogeschool Wageningen 78-14 (1978) 99 Merrill, Ind. Rafin. 1949, p. 132. Pectanisia Rafinesque, Fl. Teil. 3, Cent. VIII, 1837 (Nov.-Dec), p. 72; Merrill, I.e. Tereianthes Rafinesque, Fl. Teil. 3, cent. VIII, 1837 (Nov.-Dec), p. 72 ( = Tereianthus Fourr. in Ann. Soc. Linn. Lyon N.S. 16, 1868, p. 342); Merrill, I.e. Resedella Rchb., Handb. 1837, p. 261, nom. Eresda Spach, Hist. Nat. Vég. Phan. 7, 1839, p. 101. Stefaninia Chiovenda, Fl. Somala 1929, p. 77 (= Stephaninia auett.). Annual or perennial herbs or suffruticose, ascending or erect. Stems pithy, as a rule finally hollow (pith disintegrating). Indumentum, if present, pilose (hairs whitish, flattened, wrinkly, blunt) or, more often, con­ sisting of small, conical or mitriform trichomes or papillae. Leaves sometimes rosetted, always partly along the stem, entire or incised, or imparipinnatifid or -sect, decurrent laminar rims and midrib (if sometimes very narrowly and continuous onto the stemribs), rarely petiolate, spirally arranged. Blade (often long) attenuate (and decurrent onto the petiole). Midrib and side- nerves prominent on the lower surface, in lobed leaves excentric. Basal minute dents, perhaps glandular, present in the position of stipules. Flowers zygomorphous, in terminal, simple or branched racemes, solitary in the axil of a bract (which is accompanied at each side by one basal dent), white, cream, yellow or greenish.
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