Contributions on Entomology, International

Volume 5, Number 3, 2004

Annotated catalog of the of the world ()

By Michael Ohl

Associated Publishers 2004

Contributions on Entomology, International Edited by Virendra K. Gupta

Volume 5, Number 3 of the Contributions is an annotated catalog of the family Mantispidae (Neuroptera) of the world. All fossil and extant species described to date are catalogued. Mantispids are distinctive in having raptorial forelegs, and since their general appearance superficially resembles that of praying mantids (Mantodea), they are frequently called mantid flys. The catalog includes 11 fossil and 561 extant species-group names, and 8 fossil and 61 extant genus-group names. Of these, 410 extant species and subspecies assigned to 44 genera are recognized as valid. Data on original and current generic placements, current synonymy (including literature source, type status, type locality, type depository, and taxon distributions) are presented based on an exhaustive examination of the literature pertaining each genus and species. Fifteen new generic combinations, thirteen new species synonyms, four new replacement names, and one new genus synonym are proposed.

Author

Michael Ohl Museum für Naturkunde der Humboldt-Universität, Institut für Systematische Zoologie Invalidenstr. 43 D-10099 Berlin, Germany E-mail: [email protected]

Published November 30, 2004

ISSN: 1080-0745

Associated Publishers P. O. Box 140103 Gainesville, FL 32614-0103, USA Ohl: World Catalog of Mantispidae 131

ANNOTATED CATALOG OF THE MANTISPIDAE OF THE WORLD (NEUROPTERA)

by

MICHAEL OHL

in collaboration with John D. Oswald

TABLE OF CONTENTS

ABSTRACT...... 133 INTRODUCTION...... 134 FORMAT ...... 135 ABBREVIATIONS OF TYPE DEPOSITORIES...... 140 MISCELLANEOUS ANNOTATIONS...... 142 SUMMARY OF NEW NOMENCLATURAL ACTIONS...... 143 ACKNOWLEDGMENTS...... 145 THE CATALOG...... 146 Subfamily Symphrasinae Navás...... 146 Genus Anchieta Navás ...... 146 Genus Plega Navás ...... 147 Genus Trichoscelia Westwood...... 148 Subfamily Drepanicinae Enderlein ...... 151 Genus Ditaxis McLachlan ...... 151 Genus Drepanicus Blanchard in Gay...... 151 Genus Gerstaeckerella Enderlein ...... 152 Genus Theristria Gerstaecker ...... 153 Subfamily Calomantispinae Navás ...... 157 Genus Calomantispa Banks...... 157 Genus Nolima Navás ...... 157 Subfamily Leach...... 159 Genus Asperala Lambkin...... 159 Genus Austroclimaciella Handschin...... 159 Genus Austromantispa Esben-Petersen ...... 161 Genus Buyda Navás...... 161 Genus Campanacella Handschin ...... 162 Genus Campion Navás...... 162 Genus Cercomantispa Handschin...... 164 Genus Enderlein...... 166 Genus Dicromantispa Hoffman in Penny...... 168 Genus Entanoneura Enderlein...... 169 Genus Euclimacia Enderlein...... 170 132 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Genus Eumantispa Okamoto ...... 174 Genus Haematomantispa Hoffman in Penny...... 176 Genus Leptomantispa Hoffman in Penny ...... 176 Genus Madantispa Fraser ...... 177 Genus Mantispa Illiger in Kugelann ...... 177 Genus Mimetispa Handschin ...... 192 Genus Nampista Navás ...... 192 Genus Necyla Navás ...... 193 Genus Orientispa Poivre ...... 194 Genus Paramantispa Williner and Kormilev ...... 196 Genus Paulianella Handschin ...... 196 Genus Perlamantispa Handschin...... 197 Genus Pseudoclimaciella Handschin ...... 199 Genus Rectinerva Handschin...... 202 Genus Sagittalata Handschin ...... 202 Genus Spaminta Lambkin...... 203 Genus Stenomantispa Stitz...... 204 Genus Toolida Lambkin ...... 204 Genus Tuberonotha Handschin ...... 204 Genus Xaviera Lambkin...... 205 Genus Xeromantispa Hoffman in Penny...... 206 Genus Zeugomantispa Hoffman in Penny...... 206 Nomina dubia in Mantispidae ...... 208 Genus Manega Navás...... 208 Genus Nivella Navás...... 208 Mantispidae unplaced to genus ...... 208 Fossil Taxa in Mantispidae ...... 209 Genus † Promantispa Panfilov in Dolin et al...... 209 Genus † Prosagittalata Nel...... 209 Genus † Vectispa Lambkin...... 209 Genus † Whalfera Engel ...... 210 Subfamily † Mesomantispinae Makarkin...... 210 Genus † Mesomantispa Makarkin...... 210 Questionable Fossil Mantispidae ...... 210 Genus † Mantispidiptera Grimaldi ...... 210 Genus † Paraberotha Whalley...... 211 Genus † Retinoberotha Schlüter...... 211 REFERENCES...... 212 Printed Sources...... 212 Internet Sources...... 229 APPENDIX I. SPECIES DISTRIBUTION ...... 231 APPENDIX II. NUMBER OF SPECIES-GROUP NAMES BY AUTHOR...... 232 INDEX ...... 234 ANNOTATIONS ...... 242

Ohl: World Catalog of Mantispidae 133

ABSTRACT

The world fauna of the neuropteran family Mantispidae is cataloged, including all fossil and extant species described to date. The catalog includes 11 fossil and 561 extant species-group names, and 8 fossil and 61 extant genus-group names. Of these, 410 extant species and subspecies assigned to 44 genera are recognized as valid. Two fossil species are assigned to present-day genera, five fossil species in five fossil-only genera are placed in the Mantispidae. Four fossil species in three fossil-only genera and one fossil-only subfamily are listed as dubious Mantispidae. Data on original and current generic placements, current synonymy (including literature source, type status, type locality, type depository, and taxon distributions are presented based on an exhaustive examination of the literature pertaining each genus and species. Fifteen new generic combinations are reported: Austroclimaciella lacolombierei (Navás); Austroclimaciella subfusca (Nakahara); Dicromantispa interrupta (Say); Euclimacia nodosa (Westwood); Gerstaeckerella riedeliana (Fischer von Waldheim); Necyla formosana minor (Stitz); Necyla formosana sumatrana (Stitz); Pseudoclimaciella africana (Esben-Petersen); Pseudoclimaciella africana ragazziana (Navás); Pseudoclimaciella flava (Esben-Petersen); Pseudoclimaciella sanguinea (Navás); Pseudoclimaciella thomensis (Viette); Pseudoclimaciella umbripennis (Navás), Trichoscelia trifasciata (Stitz), and Tuberonotha ferrosa (Navás); thirteen new species synonyms are recognized: Euclimacia sauteri Navás, 1927b = Euclimacia badia Okamoto, 1910; Mantis nana Lichtenstein, 1802 = Mantispa nabota (Olivier, 1797 ["1792"]); Mantispa flaveola Erichson, 1839 = Perlamantispa perla (Pallas, 1772); Mantispa perla var. brunnea Navás, 1906 = Perlamantispa perla (Pallas, 1772); Mantispilla debilis var. nuda Stitz, 1913 = Dicromantispa debilis (Gerstaecker, 1888 ["1887"]); Mantispilla debilis var. nigricornis Stitz, 1913 = Dicromantispa gracilis (Erichson, 1839); Mantispilla debilis var. rugicollis Stitz, 1913 = Dicromantispa gracilis (Erichson, 1839); Necyla uniformis Navás, 1927c: 61 = Zeugomantispa compellens (Walker, 1858); Mantispa virescens Rambur, 1842 = Zeugomantispa viridula (Erichson, 1839); Mantispilla punctata Stitz, 1913 = Zeugomantispa viridula (Erichson, 1839); Mantispilla punctata var. major Stitz, 1913 =; Mantispilla stigmata Stitz, 1913 = Zeugomantispa viridula (Erichson, 1839); and Mantispa viridis Stitz, 1913 = Zeugomantispa viridula (Erichson, 1839); and four new names are proposed: Mantispa ambonensis Ohl for Mantispa basalis (Navás, 1929c), Mantispa coorgensis Ohl for Mantispa femoralis (Banks, 1933), Mantispa paraguayana Ohl for Mantispa nana (Navás, 1912b), and Necyla formosana chiaiensis Ohl for Necyla formosana bella (Kuwayama, 1925). One new genus synonym is recognized: Stenispa Navás = Eumantispa Okamoto. Zeugomantispa viridula (Erichson) is a junior secondary homonym of Z. viridula (Houttuyn in Stoll, 1813) (= Z. minuta (Fabricius)) and is replaced by Z. virescens (Rambur), the next available name for this species. The original generic combination of Plega zikani Navás, 1936b is confirmed, and the previously suggested synonymy of Eumantispa hypogastrica Navás, 1914i with Eumantispa ferruginea Stitz, 1913 is rejected. Mantispid species are distributed across the major zoogeographical regions as follows: Afrotropical 99, Palaearctic 33, Nearctic 20, Neotropical 94, Oriental 121, and Australasian (including New Guinea and adjacent islands) 67.

134 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

INTRODUCTION

The development of global species inventories is an urgent and vitally important task that is a primary step and fundamental activity for any kind of biodiversity research and assessment (Stork and Samways, 1995; Purvis and Hector, 2000; Wilson, 2003). The Mantispidae are a near- cosmopolitan family of Neuroptera with an attractive habitus, interesting behaviors and a relatively small number of species.But, their , phylogeny, and evolution are only fragmentarily understood on a worldwide scale. The present catalog aims to provide a complete global inventory of the Mantispidae in order to stimulate future research on this family. Mantispids are distinctive in having raptorial forelegs that insert at the apical end of a generally elongated prothorax. Their general appearance superficially resembles that of praying mantids (Mantodea), and, thus, mantispids are frequently called mantid flys. Most mantispid species are relatively small (forewing length <15 mm), although the largest have forewing lengths of more than 35 millimeters. The first formally described mantispid, the European (Poda, 1761), was originally placed in the genus Raphidia in the snake-fly order Raphidioptera. This is probably because Raphidioptera and (at least European) Mantispidae superficially resemble each other in having an elongate prothorax and wings with prominent venation and pterostigmata (H. Aspöck, 1999). Pallas (1772) described the first mantispids in the genus Mantis, M. perla and M. pusilla (both now in Perlamantispa), and most of the authors of the late 18th and earliest 19th centuries treated mantispids as members of the dictyopteran order Mantodea. It was not until the work of Erichson (1839) that the neuropterid position of the Mantispidae was finally established (although they were first considered to be most closely related to the Raphidioptera). The history of early mantispid research in the 18th century was presented in great detail by H. Aspöck (1999). Raptorial forelegs are known to occur in two other families in the Neuroptera: the Afrotropical Rhachiberothidae, which are relatively small and which lack an elongate prothorax (U. Aspöck and Mansell, 1994), and two species of the Coniopterygidae from Tanzania and Yemen, respectively, which are otherwise typical representatives of the coniopterygids and are easily differentiated from Mantispidae (Sziráki and Greve, 1996; Sziráki, 1997). The occurrence of raptorial forelegs in a few species of Coniopterygidae is obviously derived within that family. The families Mantispidae and Rhachiberothidae, however, are comprised exclusively of species with raptorial forelegs and may constitute a monophyletic taxon within the "dilarid group" of families, i.e., Dilaridae, Berothidae, Rhachiberothidae, and Mantispidae (e.g., U. Aspöck, 1996, 2002; U. Aspöck et al., 2001). A cladistic analysis of the Rhachiberothidae has shown that the Mantispidae are apparantly the sister group to Berothidae + Rhachiberothidae, which implies independent (parallel) evolution of raptorial forelegs in Mantispidae and Rhachiberothidae (U. Aspöck and Mansell, 1994). Almost simultaneously, Willmann (1990, 1994) argued in favour of a sister-group relationship between the Mantispidae and Rhachiberothidae, with raptorial forelegs as one of their synapomorphies. However, a first molecular analysis of the Neuropterida resulted in a paraphyletic "dilarid group", with Mantispidae as the sister group of Chrysopidae + Hemerobiidae (Haring and U. Aspöck, 2004). Adult mantispids are predaceous, as can be inferred from their raptorial forelegs, and as is well known from biological studies. Immature stages of many mantispids are probably parasites in the nests of aculeate Hymenoptera (New World Symphrasinae) or the egg sacs of spiders (Mantispinae), but field associations have been recorded for only a few species. The development of mantispid larvae has been called hypermetamorphic, with the first instar campodeiform and Ohl: World Catalog of Mantispidae 135

quite mobile, and the last two instars scarabaeiform and relatively immobile. Good accounts of the morphology, behavior, and development of mantispid immature stages can be found in Redborg (1985), Hoffman and Brushwein (1989, 1990, 1992), and Brushwein et al. (1995), and are summarized by U. Aspöck (1996) and Redborg (1998). The taxonomy and systematics of the Mantispidae have not been comprehensively treated for most continents. Ulrike Aspöck and H. Aspöck (1994) and U. Aspöck (1996) revised the European fauna, Lambkin (1986a, b) monographed the Australian species, and Penny (1982a, b) provided a comprehensive generic revision of South American taxa. Other published revisions are only regional or are outdated. Hoffman's comprehensive revision and cladistic analysis of the New World Mantispinae, completed as a Ph.D. dissertation, remains only partially published (Hoffman in Penny, 2002). Catalogs are available for India (Ghosh and Sen, 1977), the Neotropical region (Penny, 1977), Australia (New, 1996), America north of Mexico (Penny et al., 1997), and the Malesian region (New, 2003), but other faunas, particularly the Afrotropical and much of the Southeast Asian faunas, are desperately in need of modern species listings and taxonomic revisions. The last comprehensive listing of the genera and species of the world Mantispidae was presented by Hagen (1866), who listed four genera and 73 species known to him. The purpose of this catalog is to provide a complete list of the genus-group and species- group names of the Mantispidae of the world published before June 2004. Every effort has been made to examine all publications with taxonomically relevant information, so that the catalog reflects the current status of all mantispid taxa. Nomina nuda have been included and are assigned to genus where possible. New combinations and new synonymies are presented where examination of specimens or relevant literature has revealed the need for such actions. Notes on these actions, and ancillary information associated with many other names are given in the Annotations section at the end of this work. Future taxonomic studies will undoubtedly require further changes in the generic placements of some taxa. Many mantispid species have been described primarily on the basis of differences in the details of coloration and wing venation (traits that are known to vary considerably in many species), and distribution. One can infer from the few comprehensive modern revisions available (e.g., Lambkin, 1986a, b; Hoffman in Penny, 2002), that many existing species-group names will eventually be relegated to synonymy when revisions of broad geographical scale are completed for poorly known regions. However, more detailed examination of other character systems, particularly male terminalia, will also undoubtedly lead to the discovery and description of many additional new species.

FORMAT

Taxonomic and Nomenclatural Information

1. Scope: This global catalog includes all known available genus- and species-group names that have been proposed for extant (living) and extinct (fossil) taxa that are currently placed in the family Mantispidae (Insecta: Neuroptera). Certain classes of unavailable names that pertain to the Mantispidae (e.g., nomina nuda, prominent or confusing incorrect spellings, etc.) have also been included. Much of the general format of the catalog has been styled after the dipteran catalogs of N. L. Evenhuis (e.g., Evenhuis, 2002).

136 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

2. Arrangement of taxa: Valid genera are listed alphabetically within subfamilies. Subgenera are not currently used within the Mantispidae. Valid species and subspecies are listed alphabetically within genera. Subspecies entries are cross-referenced to their appropriate containing species. Nominotypical subspecies are not repeated separately from their nominate species entries. Synonymical listings include all known synonymous names, whether available or unavailable, and are presented chronologically. Synonyms are not listed separately in the alphabetical sequence under each genus, though non-synonymous unavailable names are. Three classes of taxa are grouped separately and presented at the end of the main sequence of the catalogue: (1) nomina dubia pertaining to extant mantispids, including extant taxa not placeable to genus, (2) genera composed solely of fossil species [fossil species placed in recent genera are listed with their respective genera], and (3) taxa questionably placed in the Mantispidae.

3. Genus entry format: Genus entries consist of a "genus header", a synonymical listing and a distribution statement. Genus headers are centered on the page and cite the genus-group name (in bold type) and its author(s). The distribution statement, given at the end of each valid genus entry, summarizes the total known distribution of the genus (see "Zoogeographical realm delimitation" for information on recognized zoogeographic realms). The synonymical listing entries present the following information for each genus-group taxon listed: Genus/subgenus name. Author(s) – authorship has been cited in accordance with the relevant articles and recommendations of the Code. Year of publication – where the true year of publication varies from the imprint year, the true year of publication is given first, followed in square brackets by the imprint year (e.g., 1885 ["1884"]). Page number – the page number on which the original description starts; for papers published in serials, the page number cited is that from the continuous pagination of the serial issue, not the separate ("reprint"), where these differ Type species – cited in the original combination and in "correct original spelling" format (i.e., with incorrect original spellings of genus-group and/or species-group names corrected as required by the Code); cited also with appropriate author(s) and year of publication (see also "Type species" below). Designation type – e.g., by original designation, by monotypy, etc.

4. Species entry format: Species-group names: Species entries consist of a "species header", a synonymical listing, and a distribution statement. Species headers cite the species- group name and its author(s) [in parentheses where appropriate] and are shown in bold type for valid names, non-bold type for non-synonyous unavailable names. The distribution statement, given at the end of each valid species entry, summarizes the total known distribution of the species (see "Zoogeographical realm delimitation" for information on recognized zoogeographic realms). The synonymical listing entries present the following information for each species-group taxon listed: Species/subspecies name – cited in the original combination and in "correct original spelling" format (i.e., with incorrect original spellings of genus-group and/or species-group names corrected as required by the Code). Author(s) – authorship has been cited in accordance with the relevant articles and recommendations of the Code. Ohl: World Catalog of Mantispidae 137

Year of publication – where the true year of publication varies from the imprint year, the true year of publication is given first, followed in square brackets by the imprint year (e.g., 1885 ["1884"]). Page number – the page number on which the original description starts; for papers published in serials, the page number cited is that from the continuous pagination of the serial issue, not the separate ("reprint"), where these differ. Verbatim spelling of species/subspecies name (where applicable) – if one or more elements of the verbatim species/subspecies name were altered under the articles of the Code to reach a "correct original spelling", the verbatim spelling(s) of that (those) elements are shown in parentheses, and a comment is provided (where applicable) – on incorrect original spellings (e.g., (as "Entatoneura [sic] Albertisii [sic]"; incorrect original spelling and capitalization)). Sex – of the type specimen(s) as indicated in the original description (if not given, the phrase "sex not indicated" is used). Type category – e.g. holotype, syntypes, etc. (see also "Types and type depositories", below). Sex(s) of type(s) –based on current information, published or unpublished (if not known, "sex unknown" is used). Type locality – generally only the country is cited, but sometimes additional geographical information is provided. Type depository – generally cited as a collection coden (see list under Abbreviations of Type Depositories below). Type fixation source (where applicable) – a literature reference to the source of a valid lectotype designation. Synonymization source – a literature reference to the source of a synonymy, though not necessarily the first; this information is not always provided. Cases of apparent multiple publication of the same name by the same author have been carefully investigated to determine if only one, or multiple, available names were created. Comments relevant to such cases are provided in their individual entries. Cited author and year combinations can be used to find the corresponding entries in the References section. When authors have published more than one article in the same year, the articles are differentiated with with letters suffixed to the year of the citation. If an article appeared or was issued in a year after its imprint year of publication (as given in the respective issue or volume), the imprint year is given in square brackets following the true year of publication. For additional reference citation information, see References below. Authorship by more than one author is listed as in the original work. When the authorship of the name is different from the authorship of the work, the full citation (in plain text) in the respective species section takes the form, e.g., "Hoffman in Penny, 2002". Since in the literature covered here, the authors responsible for the species name have included the descriptions in rather isloted paragraphs or sections, the corresponding citation can be located in the Literature Cited section under that name, e.g., in this case, "Hoffman, 2002".

5. Type species: Most of the information on the type species of genus-group names has been taken from Oswald and Penny (1991), but much of the explanatory information provided in that work has been shortened or transferred to the Annotations section. Type species are cited in this catalog in their original combinations and in "correct original spelling" format (see "Species entry format: Species/subspecies name" above) with original authorship and date, irrespective of their citation and orthography at the time of the establishment of the genus-group name. Where 138 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

information provided by the authors of new genus-group names differs from this format, their information is given in parentheses following the original-combination/correct-original-spelling version of the type species citation, e.g., "Mantispa mozambica Westwood, 1852: 269 (as "Cercomantispa mozambica Westwood 1852"). If the type species of a genus-group name is currently considered invalid, its currently valid name is also shown, placed in square brackets, e.g., "Mantispa decorata Erichson, 1839: 163 [= Mantispa ambusta Erichson]".

6. Types and type depositories: The categories of types employed here are: "holotype", "paratype(s)", "lectotype", "paralectotype(s)", and "syntype(s)". In accordance with the ICZN, published allotypes are automatically given paratype status. The term "holotype" is only used: (1) if the describing author explicitly designated a holotype specimen, (2) if the describing author explicitly stated in the original description that the type series consisted of a single specimen, or (3) if other evidence is available to demonstrate that the type series consisted a single type specimen (e.g., information published by other authors, or information derived from personal investigations of type matters). Where the number of specimens in the type series cannot be definitely determined, and no primary type has subsequently been fixed, the type category is given as "holotype (or syntypes)", even if the describing author's citation of single localities or apparently single measurements seem to imply that the type series consisted of only one specimen. The type category "syntypes" is used when solid evidence exists that the type series consisted of multiple specimens, and no primary type has subsequently been fixed. The type category "lectotype" is used where an apparently valid lectotype has been fixed under the relevant articles of the ICZN. In such cases, a literature reference to the lectotype designation that we believe has validly fixed the primary type is provided following the statement of type depository. Issues involving lost types, transfer of type specimens from one collectionto another, and other miscellaneous information relevant to type category, status, locality, depository, etc. are clarified for selected species in the Annotations section.

7. Nomenclatural actions: Changes in binominal combination, rank, and validity presented for the first time in this catalog are marked in boldface in the respective species sections after the type depository. With regard to junior synonyms, changes in binominal combination and changes in rank are not recorded or annotated. Lectotypes are not designated herein and are deferred to future taxonomic revisions. Several species-group names in the Mantispidae were originally proposed as "varieties". Where the taxonomic status of these names have not been changed and fixed by subsequent authors (e.g., through synonymization or treatment at species rank), and where the original descriptions of these "varieties" do not unambiguously reveal that they were intentionally proposed for infrasubspecific entities, they are automatically given subspecific rank here, in accordance with ICZN Art. 45.6. Species-group taxa have generally been placed to genus according to the most recent publication available to me in which they are cited. Some older mantispid genera have historically been used in much broader (non-monophyletic) senses than they are understood today. New putatively monophyletic genera are regularly being separated from such large collective genera as Climaciella and Mantispa (e.g., by Lambkin, 1986a, b; Hoffman in Penny, 2002). This process will undoubtedly continue in the future as new attempts are made to achieve more objective genus delimitations based on the concept of monophyly. In this catalog, one primary exception of the general acceptance of published generic synonyms is made: H. Aspöck et al. (1980) synonymized Perlamantispa with Mantispa, a synonymy that is not followed here. A Ohl: World Catalog of Mantispidae 139

number of species have been intentionally placed in Perlamantispa, which is based on a concept of overall similarity. However, the underlying information that lead to the concept of Perlamantispa would be obscured if these species were simply relisted in the obviously collective and non-monophyletic genus Mantispa. Thus, Perlamantispa is treated here as a genus distinct from Mantispa, and it remains to be seen if the synonymization suggested by H. Aspöck et al. (1980) prooves to be justified when more non-Palaearctic Perlamantispa species have been studied.

8. Annotations: During the course of compiling this catalog a considerable body of miscellaneous information was developed regarding the validity of a variety of taxa taxa, their type depositories, published discussions on nomenclatural issues, etc. As many of these data will be of interest to readers of this catalog, we have taken the opportunity to record many of these data here. Most of this type of information is recorded in the catalog as a series of "annotations", which are collected near the end of the catalogue and linked to the main text by superscript numbering. This manner of presentation has been implemented to avoid breaking up the main alphabetical sequence of the catalog with, sometimes lengthy, annotation text blocks.

9. Nomina nuda: Where they have been associated in the literature with other valid species, nomina nuda are listed under their current synonym, but are shown without type locality or distributional information. Nomina nuda entries are flagged with the statement "A nomen nudum". Where they have not been associate with other valid species, nomina nuda are simply listed in the main alphabetical sequence under the genera to which they are currently believed to belong.

Treatment of Geographical Information

1. Zoogeographical realm delimitation: The zoogeographical realms used here in the distribution summaries are: Afrotropical, Australasian, Nearctic, Neotropical, Oriental, and Palaearctic. The delimitation of these realms mostly follows Franz and Beier (1970). A notable exception is the recognition here of a slightly expanded Australasian realm that includes Australia and New Guinea and their adjacent islands [Franz and Beier included New Guinea and some of its adjacent islands in the Oriental region]. Boundaries between the Nearctic and Neotropical Regions in Mexico, and the Oriental and Palaearctic Regions in China, follow Evenhuis (2002: 13). The boundaries of the remainder of these realms are based on political country boundaries as this provided a convenient delimitation, though they may change in politically instable regions.

2. Type localities: Type localities are usually stated to country level, though additional information (e.g., state or county) is sometimes given. Where historical type localities could not be assigned to a particular country, all possible countries are provided (e.g., Italian Somaliland = Somalia, Djibouti, or Ethiopia) or a larger geographical region is used. For example, in multi- national islands, such as New Guinea (Papua New Guinea / Indonesia), Borneo (now comprising Sarawak, Sabah, Brunei, and the Indonesian province of Kalimantan), and Hispaniola (Haiti / Dominican Republic), the island name is provided instead of the respective country name. Current and widely used geographical units such as "Caucasus" are used and are not replaced by political areas. In some cases, such area names are followed by supplemental geographical information or interpretations in square brackets (e.g., "Amazonia [probably Brazil]". 140 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Some countries and island groups are broken down into states, provinces, archipelagos, island groups, overseas and other outside dependencies. These "sublocalities" are separated from the country of the type locality by a comma (e.g. "Indonesia, Sulawesi") and are placed in parentheses after the main locality in the Distribution section (e.g., "Distribution: Indonesia (Sulawesi)").

3. Spelling of geographical names: Historical locality names have generally been replaced by contemporary names without further annotation, e.g. Formosa is replaced by Taiwan. When more than one name for the same locality is currently in use internationally, the name used by the local people or in the local language is preferred, e.g., Celebes is replaced by Sulawesi. Due to the colonial history and recent political changes in some African and Southeast Asian countries, the history of the country names is sometimes confusing. A prime example is "Congo", which refers to different political units depending on the date of use. "Congo" is used here as currently in use in the respective country: "Congo" = (Peoples) Republic of Congo, formerly French Congo; "DR Congo" = Democratic Republic of Congo, formerly Belgish Congo or Zaire.

4. Fossil taxa: Fossils are listed with their form of preservation (amber, compression/impression), their geographical type localities according to current political boundaries, and their geological horizon.

5. Distribution: The distributions of all valid genera and species are given as completely as possible based on literature records and data from unpublished information obtained from personal examination of specimens. All distributional information has been verified to the best possible extent (e.g., examination of primary literature sources and inspection of specimen-based label data). Localities flagged by a query are considered questionable for the purposes of this catalog and should be regarded as tentative until verified.

ABBREVIATIONS OF TYPE DEPOSITORIES

Type depositories are provided for primary types only, even when we are aware of paratypes or paralectotypes being distributed in other collections. Where syntypes are known to be deposited in more than one collection, all known collections holding syntypes are cited. Depositories of mantispid types are marked with an exclamation point ("!") when they have been confirmed by personal observation by MO. Indirect evidence for depository information has been accepted if type depositories were confirmed by recent information provided by the respective curator or other persons believed to be reliable. Internet databases and catalogs have been accepted as reliable confirmation of type depositories only if the information provided coincide with information contained in original or subsequent publications regarding the species or with general information on the whereabouts of a particular collection or the material of a particular author. Superscript numbers in the list below refer to additional information in the Annotation section at the end of this work. Institutional names that could not be located unambiguously or whose present status, locality, and/or name are unknown, are abbreviated with an appropriate but arbitray three or four letter acronym based on the original name directly cited from the original literature. The original instituitional name is placed in brackets. Ohl: World Catalog of Mantispidae 141

Most acronyms of collections are take from the and Spider Collections of the World web site. For collections not listed there and for personal collections, we have arbitrarily selected an appropriate acronym.

ACIU [Agricultural College, Imperial University, Tokyo, Japan.] AMNH American Museum of Natural History, New York, USA. AMSA Australian Museum, Sydney, New South Wales, Australia. ANIC Australian National Insect Collection, Canberra, ACT, Australia.1 ANSP Academy of Natural Sciences, Philadelphia, Pennsylvania, USA. BMNH The Natural History Museum, London, Great Britain. CAA [Amber collection Professor Aftim Acra, Beirut, Lebanon.2] CAS California Academy of Sciences, San Francisco, USA. CHAH [Henry A. Hespenheide collection (see Hoffman, 2002: 257).] CMY [Collection T. Miyake, Japan.] CN Navás collection, partly destroyed (see comment below). Surviving remnants probably in MZBS. CSY [Collection T. Suzuki, Japan.] CUMZ University Museum of Zoology, Cambridge, United Kingdom. CZC [coll. F. de Zayas, Cuba.] DEI Deutsches Entomologisches Institut, Eberswalde, Germany (formerly in Berlin- Dahlem). DZUP Museu de Entomologia, Departamento de Zoologia, Curitiba, Brazil. EIHU Entomological Institute, Hokkaido University, Sapporo, Japan (includes Matsumura collection). EMAU Ernst-Moritz-Arndt-Universität, Greifswald, Germany. ETH Eidgenössische Technische Hochschule, Entomologisches Institut, Zurich, Switzerland. FMNH Finnish Museum of Natural History, Helsinki (= Helsingfors), Finland. FSCA Florida State Collection of , Gainesville, Florida, USA. HNHM Hungarian Natural History Museum, Budapest, Hungary. HUAC Personal collection H. and U Aspöck, Vienna, Austria. IFAN Institut Fondamental de l'Afrique Noire, Dakar, Senegal. INPA Coleccio Sistematica da Entomologia, Manaus, Amazonas, Brazil. ISNB Institut Royal des Sciences Naturelles de Belgique, Bruxelles, Belgium. ITLJ Laboratory of Insect Systematics, National Institute of Agro-Environmental Sciences, Tsukuba, Ibaraki, Japan. IZAS Chinese Academy of Sciences, Institute of Zoology, Beijing, China. KUZC Zoological Collection, Kyushu University, Fukuoka, Japan. MACN Museo Argentino de Ciencias Naturales, Buenos Aires, Argentina. MBBJ Museum Zoologicum Bogoriense, Bogor (= Buitenzorg), Indonesia. MCSN Museo Civico di Storia Naturale "Giacomo Doria", Genova, Italy. MCZ Museum of Comparative Zoology, Cambridge, Massachusetts, USA. MHNG Museum d'Histoire Naturelle, Geneva, Switzerland. MLPA Museo de La Plata, Universidad Nacional de La Plata, Argentina. MNHN Museum National d'Histoire Naturelle, Paris, France. MPEG Museu Paraense Emilio Goeldi, Belem, Para, Brazil. MRAC Musee Royal de l'Afrique Centrale, Tervuren, Belgium. 142 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

MVM Museum of Victoria, Abbotsville, Victoria, Australia. MZBS Museo Zoologia, Barcelona, Spain. MZLU Lund University, Lund, Sweden. MZPW Museum of the Institute of Zoology, Polish Academy of Sciences, Warszawa Poland.3 MZSP Museu de Zoologia da Universidade de São Paulo, São Paulo, Brazil. NELG [Nawa Entomological Laboratory, Gifu, Japan.] NHMB Naturhistorisches Museum, Basel, Switzerland. NHMW Naturhistorisches Museum, Vienna, Austria. NHRS Natuhistoriska Riksmuseet, Stockholm, Sweden. NMBS Naturhistorisches Museum, Bern, Switzerland. NMPC National Museum (Natural History), Prague, Czech Republic. NMWC National Museum of Wales, Cardiff, United Kingdom. OUM University Museum, Oxford, Great Britain. PIMR Palaeontological Institute, Russian Academy of Sciences, Moscow, Russia. PIN Palaeontological Institute, Novosibirsk, Russia. PMNH Peabody Museum of Natural History, Yale University, New Haven, Connecticut, USA. QMB Queensland Museum, Brisbane, Australia. RMNH Nationaal Natuurhistorische Museum ("Naturalis") (formerly Rijksmuseum van Natuurlijke Historie), Leiden, Netherlands. SAM South Australian Museum, Adelaide, South Australia, Australia. SMBA Institutos Cientificos de San Miguel, provincia de Buenos Aires, Argentina. SMFD Senckenberg-Museum, Frankfurt, Germany. UZIU Zoological Museum, Uppsala University, Uppsala, Sweden. WAM Western Australian Museum, Perth, Australia. ZIL Zoological Museum, Academy of Sciences, St. Petersburg, Russia. ZMAN Zoologisch Museum, Universiteit van Amsterdam, Amsterdam, Netherlands. ZMB Museum für Naturkunde der Humboldt-Universität, Berlin, Germany. ZSMC Zoologische Staatssammlung, München, Germany. ZMUC Zoological Museum, University of Copenhagen, Copenhagen, Denmark. ZMUH Zoologisches Institut und Zoologisches Museum, Hamburg, Germany.4 ZMUM Zoological Museum, Moscow, Russia.

MISCELLANEOUS ANNOTATIONS

1. The Navás collection (CN): Many of the types of species described by Navás were retained in his personal collection, which is usually indicated in his publications by "coll. Navás". Despite the fact that a considerable portion of the material in the Navás collection was destroyed after his death (Monserrat, 1985, 1986), some of the types survived in the MZBS. Monserrat (1985) provided a list of putative Navás types in this collection. The type material of mantispids that were published as being held in Navás' collection and that were not listed subsequently by Monserrat (1985) or other authors to be housed in the MZBS or a different collection, are listed here as deposited in "coll. Navás" (under depository acronym "CN").

Ohl: World Catalog of Mantispidae 143

2. Hoffman (1992): This work is an unpublished Ph.D. dissertation and is a taxonomic and systematic revision of the New World Mantispinae. It is one of the rare examples of a modern, comprehensive revision of the Mantispidae of a particular geographic area and is, thus, of high relevance and importance. A large number of nomenclaturally relevant actions have been proposed by Hoffman, such as new species and new synonyms. Parts of this dissertation have recently been made available through the publication by Hoffman in Penny (2002), but the remaining results of the dissertation are not considered to be published and are not recognized as available under the current articles of the ICZN. Consequently, Hoffman's results are not included in this catalog, with the exception of those new names and nomenclatural acts published by Hoffman in Penny (2002), which are included here. Any nomenclatural changes proposed in this catalog that are also in Hoffman's dissertation, have been discovered independently. Data without nomenclatural relevancy (e.g., type depositories, citation details, distribution, etc.), however, have frequently been checked against Hoffman's work. This dissertation was not accessible to MO before January 2004.

3. Fossil Taxa: Species and genera described from fossils are included in the general catalog and are marked with the dagger symbol (†). They are listed together with their form of preservation (amber, compression/impression), their known geographical distribution according to recent zoogeographical realms, and their geological horizon. Fossil genera (i.e., genera containing only fossil species) are listed separately, because they are usually assigned to subfamilies only tentatively, if at all. Fossil species described in present-day genera (Gerstaeckerella asiatica Makarkin and Climaciella henrotayi Nel) are listed under their respective generic names. Paraberotha acra Whalley and Retinoberotha stuermeri Schlüter are listed here as Questionable Mantispidae, because they have been discussed as stem-group representatives of either Mantispidae or Rhachiberothidae. Their actual phylogenetic positions depend on the presumed relationships of these families (Willmann, 1990, 1994; U. Aspöck and Mansell, 1994).

SUMMARY OF NEW NOMENCLATURAL ACTIONS

New generic combinations

Austroclimaciella lacolombierei (Navás) Austroclimaciella subfusca (Nakahara) Dicromantispa interrupta (Say) Euclimacia nodosa (Westwood) Gerstaeckerella riedeliana (Fischer von Waldheim) Necyla formosana minor (Stitz) Necyla formosana sumatrana (Stitz) Pseudoclimaciella africana (Esben-Petersen) Pseudoclimaciella africana ragazziana (Navás) Pseudoclimaciella flava (Esben-Petersen) Pseudoclimaciella sanguinea (Navás) Pseudoclimaciella thomensis (Viette) Pseudoclimaciella umbripennis (Navás) 144 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Trichoscelia trifasciata (Stitz) Tuberonotha ferrosa (Navás)

New species synonyms

Euclimacia sauteri Navás, 1927b = Euclimacia badia Okamoto, 1910 Mantis nana Lichtenstein, 1802 = Mantispa nabota (Olivier, 1797 ["1792"]) Mantispa flaveola Erichson, 1839 = Perlamantispa perla (Pallas, 1772) Mantispa perla var. brunnea Navás, 1906 = Perlamantispa perla (Pallas, 1772) Mantispilla debilis var. nuda Stitz, 1913 = Dicromantispa debilis (Gerstaecker, 1888 ["1887"]) Mantispilla debilis var. nigricornis Stitz, 1913 = Dicromantispa gracilis (Erichson, 1839) Mantispilla debilis var. rugicollis Stitz, 1913 = Dicromantispa gracilis (Erichson, 1839) Mantispa virescens Rambur, 1842 = Zeugomantispa viridula (Erichson, 1839) Mantispilla punctata Stitz, 1913 = Zeugomantispa viridula (Erichson, 1839) Mantispilla punctata var. major Stitz, 1913 = Zeugomantispa viridula (Erichson, 1839) Mantispilla stigmata Stitz, 1913 = Zeugomantispa viridula (Erichson, 1839) Mantispa viridis Stitz, 1913 = Zeugomantispa viridula (Erichson, 1839) Necyla uniformis Navás, 1927c = Zeugomantispa compellens (Walker, 1858)

New genus synonym

Stenispa Navás, 1914i = Eumantispa Okamoto, 1910

Replacement names

Mantispa ambonensis Ohl, n. n. (for Mantispilla basalis Navás, 1929c (now in Mantispa), a junior primary homonym of Mantispilla basalis Navás, 1927c (now in Mantispa)). Mantispa coorgensis Ohl, n. n. (for Mantispa femoralis Banks, 1933; a junior primary homonym of Mantispa femoralis Navás, 1914h). Mantispa paraguayana Ohl, n. n. (for Mantispilla nana Navás, 1912b (now in Mantispa), a junior secondary homonym of Mantispa nana Erichson, 1839. Necyla formosana chiaiensis Ohl, n. n. (for Mantispa formosana bella Kuwayama, 1925 (now in Necyla), a junior primary homonym of Mantispa (Trichoscelia) bella Westwood, 1867 (now in Anchieta).

Zeugomantispa virescens (Rambur, 1842) is the next available name for Zeugomantispa viridula (Erichson, 1839), a junior secondary homonym of Zeugomantispa viridula (Houttuyn in Stoll, 1813).

Revised status

Plega zikani Navás, 1936b (original generic combination confirmed) Eumantispa hypogastrica Navás, 1914i (previously suggested synonymy rejected)

Ohl: World Catalog of Mantispidae 145

ACKNOWLEDGMENTS

This catalog is largely based on the Mantispidae parts of the personal Neuropterida catalogs provided by J. D. Oswald and N. D. Penny, respectively. Moreover, Oswald and Penny shared their personal notes on many species with me and allowed me to benefit from their enormous experience. They also provided copies of historical literature that I could not locate in German libraries. Without their help and generosity, this product could not have achieved the high level of completeness that it hopefully has. I am particularly indebted to John Oswald, who is an impressive source of information on historical literature in Neuropterida. Many other people provided critical assistance throughout this study. I am particularly thankful to H. Aspöck for his notes on pre-1800 publications on Mantispidae in addition to his wonderful work from 1999. Upon my request, Kevin Hoffman sent me a copy of his Ph.D. thesis on the New World Mantispinae, and he also provided his personal bibliographic catalogs of the Mantispidae of the world and copies of a few Navás publications that I could not obtain in Germany. Michael S. Engel informed me about the homonymy of Fera and the status of other fossil mantispids. Reinhard Ehrmann send me details about historical publications that include mantispid description originally placed in Mantidae. Reinhard Gaedike was so kind to lend the entire Mantispidae collection of the DEI to me. Barbara Agabiti provided information on the Paoli collection and the type locality of Perlamantispa perla var. brunnea. X.-k. Yang sent me copies of recent Chinese publications. Vladimir N. Makarkin helped with Russian literature and type depositories. Jes Rust and Arnold Staniczek provided copies of literature on fossil mantispids. The staff of the ZMB library, Martina Rissberger and Judit Gulden, assisted with obtaining copies of historical literature. During the last years, a number of volunteers helped in obtaining literature, particularly Kati Wünsche, Susanne Schiller, and Rebecca Kittel. Several museum curators verified the presence of types of Mantispidae in the Neuropterida collections under their repsonsibility: U. Aspöck (NHMW), P. A. Brown (BMNH), E.-G. Burmeister (ZSMC), J. Constant (ISNB), J J. Hogan (OUM), Legrand (MNHN), H. Mejnol (UZIU), G. A. R. Melo (DZUP), W. A. Nässig (SMFD), and W. Tomaszewska (MZPW). I am especially thankful for the assistance of Isolde Dorandt, the technician of the Neuropterida collection in the ZMB, whose help was important at all stages of the preparation of this catalog. She patiently and repeatedly counted the number of names in this catalog, which resulted in the two tables in Appendix I and II. Final drafts of the manuscript have been critically evaluated and significantly improved by Ulrike and Horst Aspöck and John D. Oswald. 146 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

THE CATALOG

Subfamily Symphrasinae Navás

Genus Anchieta Navás

ANCHIETA Navás, 1909a ["1908-1909"]: 483. Type species: Anchieta nobilis Navás, 1909a ["1908-1909"]: 484 (= Anchieta fumosella (Westwood, 1867: 504)), by monotypy. Anisoptera Schneider, 1843: 32. Type species: Mantispa notha Erichson, 1839: 170 (now in Anchieta), by monotypy. A junior homonym of Anisoptera Berthold, 1827: 409 (in Orthoptera) and Anisoptera Herrich-Schäffer, 1840: 57, 69 (in Hymenoptera). Replaced by Platymantispa Rehn, 1939, and Anisopterana Strand, 1942. Synonymized with Anchieta Navás by Penny, 1982b: 216. Platymantispa Rehn, 1939: 82. Replacement name for Anisoptera Schneider, 1843. Anisopterana Strand, 1942: 389. Unnecessary replacement name for Anisoptera Schneider, 1843, previously replaced by Rehn, 1939. DISTRIBUTION: Neotropical: Brazil to French Guiana. bella (Westwood) Mantispa (Trichoscelia) bella Westwood, 1867: 502, female. Holotype: female, "Amazonia" [probably Brazil], no specific locality (OUM!). Anisoptera romani Esben-Petersen, 1917: 14, male. Holotype: male, Brazil (NHRS). Synonymized with Anchieta bella by Penny, 1977: 37. DISTRIBUTION: Neotropical: Brazil, French Guiana. eurydella (Westwood) Mantispa (Trichoscelia) eurydella Westwood, 1867: 501, female [as Eurydella [sic], incorrect original capitalization]. Holotype: female, "Amazonia" [probably Brazil], no specific locality (OUM!). DISTRIBUTION: Neotropical: Brazil. fumosella (Westwood) Mantispa (Trichoscelia) fumosella Westwood, 1867: 504, male. Holotype (or syntypes): male, "Amazonia" [probably Brazil], no specific locality (OUM!). Anchieta nobilis Navás,1909a ["1908-1909"]: 484, male, female. Lectotype: sex not indicated, Brazil (MNHN). Designated by Penny, 1982a: 420. Synonymized with Anchieta fumosella by Penny, 1982a: 419. DISTRIBUTION: Neotropical: Brazil. notha (Erichson) Mantispa notha Erichson, 1839: 170, male. Holotype: male, Brazil (ZMB!). DISTRIBUTION: Neotropical: Brazil.

Ohl: World Catalog of Mantispidae 147

partheniella (Westwood) Mantispa (Trichoscelia) partheniella Westwood, 1867: 501, male, female [as Partheniella [sic], incorrect original capitalization]. Lectotype: male, "Amazonia" [probably Brazil], no specific locality (OUM!). Designated by Penny, 1982a: 421. DISTRIBUTION: Neotropical: Brazil.

Genus Plega Navás

PLEGA Navás, 1928a ["1927"]: 326. Type species: Symphrasis signata Hagen, 1877: 208, by original designation. DISTRIBUTION: Nearctic and Neotropical: Brazil to southern USA. banksi Rehn Plega banksi Rehn, 1939: 248, male, female. Holotype: male, USA, Arizona (AMNH). DISTRIBUTION: Nearctic: USA (Arizona). beardi Penny Plega beardi Penny, 1982a: 423, male. Holotype: male, Brazil (INPA). DISTRIBUTION: Neotropical: Brazil, Trinidad. dactylota Rehn Plega dactylota Rehn, 1939: 250, male, female. Holotype: male, USA, Arizona (ANSP). Plega dactylota lipanica Rehn, 1939: 254, male, female. Holotype: male, USA, Texas (ANSP). Synonymized with Plega dactylota by Rice, 1987: 342. DISTRIBUTION: Nearctic: USA (Arizona, Texas). duckei Penny Plega duckei Penny, 1982a: 424, male. Holotype: male, Brazil (INPA). DISTRIBUTION: Neotropical: Brazil. fasciatella (Westwood) Mantispa (Trichoscelia) fasciatella Westwood, 1867: 503, female. Holotype (or syntypes): female, Venezuela (OUM!). DISTRIBUTION: Neotropical: Venezuela. fratercula Rehn Plega fratercula Rehn, 1939: 247, male, female. Holotype: male, USA, Arizona (MCZ!). DISTRIBUTION: Nearctic: USA (Arizona). fumosa Linsley and MacSwain Plega fumosa Linsley and MacSwain, 1955: 16, male, female. Holotype: male, Mexico, Michoácan (CAS). DISTRIBUTION: Nearctic: Mexico (Michoácan de Ocampo).

148 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

hagenella (Westwood) Mantispa hagenella Westwood, 1867: 504, male [as Hagenella [sic], incorrect orginal capitalization]. Holotype (or syntypes): male, "Amazonia" [probably Brazil], no specific locality (OUM!). Mantispa cognatella Westwood, 1867: 506, sex not indicated. Holotype: female, Venezuela (OUM!). Synonymized with Plega hagenella by Penny, 1982a: 425. DISTRIBUTION: Neotropical: Bolivia, Brazil, Colombia, Nicaragua, Panama, Trinidad, Venezuela. melitomae Linsley and MacSwain Plega melitomae Linsley and MacSwain, 1955: 15, male, female. Holotype: male, Mexico, Chiapas (CAS). DISTRIBUTION: Neotropical: Mexico (Chiapas). paraensis Penny Plega paraensis Penny, 1982a: 426, female [as paraense [sic], incorrect original termination5]. Holotype: female, Brazil (MPEG). DISTRIBUTION: Neotropical: Brazil. signata (Hagen) Symphrasis signata Hagen, 1877: 208, female. Holotype: female USA, California (MCZ!6). DISTRIBUTION: Nearctic: USA (California); Mexico (Baja California). Neotropical: Costa Rica. variegata Navás Plega variegata Navás, 1928a ["1927"]: 327, male. Holotype (or syntypes): male, Mexico: México (CN). Nearctic and Neotropical: Mexico. yucatanae Parker and Stange Plega yucatanae Parker and Stange, 1965: 606, male, female. Holotype: male, Mexico: Yucatán (CAS). DISTRIBUTION: Neotropical: Mexico (Yucatán). zikani Navás, revised generic combination Plega zikani Navás, 1936b: 722, female. Holotype (or syntype): female, Brazil (DEI!)7. DISTRIBUTION: Neotropical: Brazil.

Genus Trichoscelia Westwood

TRICHOSCELIA Westwood, 1852: 270. Type species: Mantispa (Trichoscelia) fenella Westwood, 1852: 269, by subsequent designation by Enderlein, 1910: 376. Trichoscelia was originally proposed as a subgenus of Mantispa Illiger in Kugelann. Trichoscelis; Hagen, 1859: 408, 1861: 323. Incorrect subsequent spelling of Trichoscelia. Ohl: World Catalog of Mantispidae 149

Symphrasis Hagen, 1877: 208. Type species: Raphidia varia Walker, 1853: 212, by subsequent designation by Enderlein, 1910: 374. Synonymized with Trichoscelia by Penny, 1982b: 214. DISTRIBUTION: Neotropical: Argentina to Mexico. anae Penny Trichoscelia anae Penny, 1982a: 427, male. Holotype: male, Brazil (INPA). DISTRIBUTION: Neotropical: Brazil. banksi Enderlein Trichoscelia banksi Enderlein, 1910: 377, female [as Banksi [sic], incorrect original capitalization]. Holotype: female, Mexico, Chiapas (MZPW!). DISTRIBUTION: Neotropical: Mexico (Chiapas). egella (Westwood) Mantispa (Trichoscelia) egella Westwood, 1867: 502, male. Syntypes8: male, Brazil (BMNH!). Anisoptera amoenula Gerstaecker, 1888 ["1887"]: 119, female. Holotype: female, Brazil (EMAU!). Synonymized with Trichoscelia egella by Penny, 1982a: 429. DISTRIBUTION: Neotropical: Brazil. fenella (Westwood) Mantispa (Trichoscelia) fenella Westwood, 1852: 269, sex not indicated. Holotype: male, Brazil (BMNH!). DISTRIBUTION: Neotropical: Brazil. iridella (Westwood) Mantispa (Trichoscelia) iridella Westwood, 1867: 503, sex not indicated. Lectotype: male, "Amazonia" [probably Brazil], no specific locality (BMNH!). Designated by Penny, 1982a: 431. Mantispa (Trichoscelia) basella Westwood, 1867: 504, male. Holotype: male, "Amazonia" [probably Brazil], no specific locality (OUM!). Synonymized with Trichoscelia iridella by Penny, 1982a: 431. DISTRIBUTION: Neotropical: Brazil, Ecuador, Trinidad. latifascia McLachlan Trichoscelia latifascia McLachlan, 1867 ["1868"]: 255, male. Holotype: male, Brazil (BMNH!). Anisoptera jocosa Gerstaecker, 1888 ["1887"]: 117, female. Holotype (or syntypes): female, Brazil (EMAU9). Synonymized with Trichoscelia latifascia by Penny, 1982a: 433. Symphrasis thaumasta Navás, 1915a: 197, male. Holotype: male, Brazil (NMBS). Synonymized with Trichoscelia latifascia by Penny, 1982a: 433. DISTRIBUTION: Neotropical: Brazil. nassonovi (Navás) Symphrasis nassonovi Navás, 1912a ["1911"]: 537, sex not indicated. Lectotype: male, Peru (ZIL). Designated by Krivokhatsky, 1995: 11. DISTRIBUTION: Neotropical: Peru.

150 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

remipes (Gerstaecker) Anisoptera remipes Gerstaecker, 1888 ["1887"]: 120, female. Holotype (or syntypes): female, Colombia (EMAU!). DISTRIBUTION: Neotropical: Colombia, Costa Rica. santareni (Navás) Symphrasis santareni Navás, 1914h: 25, sex not indicated [as Santareni [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Mexico: Tabasco (BMNH!). DISTRIBUTION: Neotropical: Mexico (Tabasco). sequella (Westwood) Mantispa (Trichoscelia) sequella Westwood, 1867: 503, female. Holotype: female, "Amazonia" [probably Brazil], no specific locality (OUM!). DISTRIBUTION: Neotropical: Brazil, Guyana. tobari (Navás) Symphrasis tobari Navás, 1914h: 26, sex not indicated [as Tobari [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Mexico: Tabasco (BMNH!). DISTRIBUTION: Neotropical: Mexico (Tabasco). trifasciata (Stitz), new combination Symphrasis trifasciata Stitz, 1913: 44, female. Holotype: female, Bolivia (ZMB!)10. DISTRIBUTION: Neotropical: Bolivia, Ecuador. varia (Walker) Raphidia varia Walker, 1853: 212, sex not indicated. Holotype: sex unknown, Brazil (BMNH!). Mantispa myrapetrella Westwood, 1867: 505, male, female [as Myrapetrella [sic], incorrect original capitalization]. Lectotype: male, "in nidis vespae (Myrapetrae scutellaris) Americae meridionalis" (OUM!). Designated by Penny, 1982b: 436. Synonymized with Trichsoscelia varia by McLachlan, 1867 ["1868"]: 261 (in footnote). DISTRIBUTION: Neotropical: Argentina, Brazil, Surinam, Uruguay, Venezuela. Ohl: World Catalog of Mantispidae 151

Subfamily Drepanicinae Enderlein

Genus Ditaxis McLachlan

DITAXIS McLachlan, 1867 ["1868"]: 262. Type species: Mantispa biseriata Westwood, 1852: 263, by monotypy. DISTRIBUTION: Australasian: Australia. biseriata (Westwood) Mantispa biseriata Westwood, 1852: 263, sex not indicated. Holotype: sex unknown, Australia, Queensland (BMNH! OUM?). DISTRIBUTION: Australasian: Australia (Queensland, New South Wales). meridiei Lambkin Ditaxis meridiei Lambkin, 1986a: 37, male, female. Holotype: male, Australia, New South Wales (ANIC). DISTRIBUTION: Australasian: Australia (New South Wales, Victoria).

Genus Drepanicus Blanchard in Gay

DREPANICUS Blanchard in Gay, 1851: 130. Type species: Drepanicus gayi Blanchard in Gay, 1851: 132, by monotypy. Molinella Navás, 1910: 239. Type species: Molinella moulti Navás, 1910: 240, by monotypy. Synonymized with Drepanicus by Banks, 1913: 206. Drepaniscus; Schröder, 1925: 836. Incorrect subsequent spelling of Drepanicus. DISTRIBUTION: Neotropical: Argentina, Chile. chrysopinus Brauer Drepanicus chrysopinus Brauer, 1867a: 284, male. Holotype (or syntype): male, Chile (NHMW11). 12 DISTRIBUTION: Neotropical: Chile. gayi Blanchard in Gay Drepanicus gayi Blanchard in Gay, 1851: 132, pl. 1, Fig. 7, sex not indicated [as Gayi [sic], incorrect original capitalization]. Syntypes: sex unknown, Chile (MNHN?13). DISTRIBUTION: Neotropical: Chile, Argentina. moulti (Navás) Molinella moulti Navás, 1910: 240, sex not indicated. [as Moulti [sic], incorrect original capitalization]. Holoype: sex unknown, Chile (MNHN). DISTRIBUTION: Neotropical: Chile.

152 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

prasinus Esben-Petersen Drepanicus prasinus Esben-Petersen, 1912: 272, female. Holotype: female: "Nd. Cal. [= Nieder- Kalifornien, = Baja California]14" (DEI!). DISTRIBUTION: probably Neotropical: no specific locality. schajovskoyi Williner and Kormilev Drepanicus gayi schajovskoyi Williner and Kormilev, 1958: 17, male, female. Holotype: male, Argentina (SMBA). DISTRIBUTION: Neotropical: Chile, Argentina.

Genus Gerstaeckerella Enderlein

GERSTAECKERELLA Enderlein, 1910: 368. Type species: Gerstaeckerella gigantea Enderlein, 1910: 370, by original designation. Fusa Navás, 1925b: 307. Type species: Fusa pirioni Navás, 1925b: 307, by monotypy. Synonymized with Gerstaeckerella by Penny, 1982b: 212. DISTRIBUTION: Neotropical: Argentina to Colombia, Cuba.

† asiatica Makarkin † Gerstaeckerella asiatica Makarkin, 1990: 65, sex not indicated. Holotype: sex unknown, Kazakhstan, Upper Cretaceous (PIN). DISTRIBUTION: Palaearctic: Kazakhstan (Upper Cretaceous) (compression/impression). chilensis (Hagen) Mantispa chilensis Hagen, 1859: 408, male, female. Syntypes: male, female, Chile (MCZ!).15 Fusa pirioni Navás, 1925b: 307, male [as Pirioni [sic], incorrect original capitalization]. Holotype (or syntypes): male, Chile (depository unknown16). Synonymized with Gerstaeckerella chilensis by Penny, 1982b: 212. DISTRIBUTION: Neotropical: Chile. gigantea Enderlein Gerstaeckerella gigantea Enderlein, 1910: 370, male. Holotype: male, Colombia (MZPW!). Gerstaeckerella salonii Navás, 1912b: 201, sex not indicated [as Salonii [sic], incorrect original capitalization]. Holotype (or syntypes): Paraguay (ZSMC! MZBS according to Monserrat, 1985). Synonymized with Gerstaeckerella gigantea by Williner and Kormilev, 1958: 15. Gerstaeckerella anchietai Navás, 1930e: 69, female. [as Anchietai [sic], incorrect original capitalization]. Holotype: female, Brazil (ZMUH). Synonymized with Gerstaeckerella gigantea by Penny and da Costa, 1983: 634. DISTRIBUTION: Neotropical: Argentina, Brazil, Colombia, Paraguay. implexa Navás Gerstaeckerella implexa Navás, 1932a: 116, male, female. Syntypes: male, female, Chile (ZMUH). DISTRIBUTION: Neotropical: Chile.

Ohl: World Catalog of Mantispidae 153

irrorata (Erichson) Mantispa irrorata Erichson, 1839: 162, sex not indicated. Syntypes: sex unknown, Brazil (ZMB!). DISTRIBUTION: Neotropical: Brazil, Cuba. leonina Navás Gerstaeckerella leonina Navás, 1930f: 302, sex not indicated. Holotype (or syntypes): sex unknown, Chile (ZMUH). DISTRIBUTION: Neotropical: Chile. riedeliana (Fischer von Waldheim), new combination Raphidia riedeliana Fischer von Waldheim, 1834: 329, sex not indicated [as Riedeliana [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Brazil (ZIL or ZMUM).17 DISTRIBUTION: Neotropical: Brazil.

Genus Theristria Gerstaecker

THERISTRIA Gerstaecker, 1885a ["1884"]: 43. Type species: Theristria felina Gerstaecker, 1885a ["1884"]: 44, by subsequent designation by Enderlein, 1910: 368. Veura Navás, 1927d: 27. Type species: Veura debetazi Navás, 1927d: 28 (= Theristria hillieri Navás, 1914i: 98), by monotypy. Synonymized with Theristria by Lambkin, 1986a: 38. Theristriella Esben-Petersen, 1929a: 34. Type species: Theristriella stigma Esben-Petersen, 1929a: 35, by original designation. Synonymized with Theristria by Lambkin, 1986a: 38. DISTRIBUTION: Australasian: Australia. aruntae Lambkin Theristria aruntae Lambkin, 1986a: 57, male, female. Holotype: male, Australia, Northern Territory (ANIC). DISTRIBUTION: Australasian: Australia (Northern Territory, Western Australia). attenuata Lambkin Theristria attenuata Lambkin, 1986a: 72, male, female..Holotype: female, Australia, Western Australia (ANIC). DISTRIBUTION: Australasian: Australia (Western Australia, Northern Territory). basalis Banks Theristria basalis Banks, 1939: 475, male, female. Lectotype: male, Australia, Western Australia (ANIC ex MCZ!). Designated by Lambkin, 1986a: 56.18 DISTRIBUTION: Australasian: Australia (Western Australia).

154 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

cardaleae Lambkin Theristria cardaleae Lambkin, 1986a: 58, male, female. Holotype: male, Australia, Northern Territory (ANIC). DISTRIBUTION: Australasian: Australia (Northern Territory). commoni Lambkin Theristria commoni Lambkin, 1986a: 71, male, female. Holotype: female, Australia, Western Australia (ANIC). DISTRIBUTION: Australasian: Australia (Western Australia, Queensland, Northern Territory). delicatula (Westwood) Mantispa delicatula Westwood, 1852: 261, sex not indicated. Lectotype19: sex unknown, Australia, South Australia (OUM!). Designated by Lambkin, 1986a: 62. DISTRIBUTION: Australasian: Australia (Queensland, New South Wales, Australian Capital Territory, Victoria, South Australia). dentata Lambkin Theristria dentata Lambkin, 1986a: 55, male, female. Holotype: male, Australia, Northern Territory (ANIC). DISTRIBUTION: Australasian: Australia (Northern Territory). discolor (Westwood) Mantispa discolor Westwood, 1852: 262, female. Holotype: female, Australia, no specific locality (BMNH!). Theristria eurysticta Gerstaecker, 1885a [1984]: 105, female. Syntypes: female, Australia, New South Wales (EMAU!). Synonymized with Theristria discolor by Lambkin, 1986a: 45. DISTRIBUTION: Australasian: Australia (Queensland, New South Wales, Australian Capital Territory, Victoria). felina Gerstaecker Theristria felina Gerstaecker, 1885a ["1884"]: 44, male, female. Lectotype: male, Australia, Queensland (BMNH! EMAU!). Designated by Lambkin, 1986a: 44. DISTRIBUTION: Australasian: Australia (Queensland, New South Wales, South Australia, Northern Australia). gilva Lambkin Theristria gilva Lambkin, 1986a: 74, male, female. Holotype: male, Australia, Western Australia (ANIC). DISTRIBUTION: Australasian: Australia (Northern Territory, Western Australia). hillieri Navás Theristria hillieri Navás, 1914i: 98, "female" [as Hillieri [sic], incorrect original capitalization]. Holotype: male20, Australia (South Australia) (BMNH!). Ohl: World Catalog of Mantispidae 155

Veura debetazi Navás, 1927d: 28, female [as Debetazi [sic], incorrect original capitalization]. Holotype: female, Australia (Victoria) (MNHN). Synonymized with Theristria hillieri by Lambkin, 1986a: 69. Theristria medialis Banks, 1939: 476, male. Holotype: male, Australia (Western Australia) (ANIC ex MCZ!). Synonymized with Theristria hillieri by Lambkin, 1986a: 69. Theristria delicatula var. minor Banks, 1939: 477, sex not indicated. Holotype: male, Australia (Queensland) (ANIC ex MCZ!). Synonymized with Theristria hillieri by Lambkin, 1986a: 69. DISTRIBUTION: Australasian: Australia (Queensland, New South Wales, Victoria, South Australia, Northern Territory, Western Australia). imperfecta Lambkin Theristria imperfecta Lambkin, 1986a: 47, male, female. Holotype: male, Australia, Victoria (ANIC). DISTRIBUTION: Australasian: Australia (Queensland, New South Wales, Victoria). maculosa Lambkin Theristria maculosa Lambkin, 1986a: 53, male, female. Holotype: male, Australia, Northern Territory (ANIC). DISTRIBUTION: Australasian: Australia (Northern Territory). mouldsorum Lambkin Theristria mouldsorum Lambkin, 1986a: 73, male, female. Holotype: female, Western Australia (AMSA) DISTRIBUTION: Australasian: Australia (Western Australia). nigra Lambkin Theristria nigra Lambkin, 1986a: 59, male, female. Holotype: male, Australia, New South Wales (ANIC). DISTRIBUTION: Australasian: Australia (Queensland, New South Wales, Australian Capital Territory, Victoria, Western Australia). pallida Lambkin Theristria pallida Lambkin, 1986a: 75, male, female. Holotype: male, Australia, Northern Territory (ANIC). DISTRIBUTION: Australasian: Australia (Northern Territory). rieki Lambkin Theristria rieki Lambkin, 1986a: 64, male, female. Holotype: male, Australia, Western Australia (ANIC). DISTRIBUTION: Australasian: Australia (Western Australia). smithersi Lambkin Theristria smithersi Lambkin, 1986a: 66, male, female. Holotype: male, Australia, South Australia (ANIC). DISTRIBUTION: Australasian: Australia (South Australia, Western Australia).

156 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

stigma (Esben-Petersen) Theristriella stigma Esben-Petersen, 1929: 35, female. Holotype: female, Australia, Queensland (QMB). DISTRIBUTION: Australasian: Australia (Queensland, New South Wales, Victoria). stigmalis Banks Theristria stigmalis Banks, 1939: 475, sex not indicated. Holotype, sex unknown, Australia, Queensland (ANIC ex MCZ!). DISTRIBUTION: Australasian: Australia (Queensland, New South Wales). storeyi Lambkin Theristria storeyi Lambkin, 1986a: 50, male, female. Holotype: male, Australia, Queensland (QMB). DISTRIBUTION: Australasian: Australia (Queensland). taroom Lambkin and New Theristria taroom Lambkin and New, 1994: 117, male, female. Holotype: male, Australia, Queensland (QMB). DISTRIBUTION: Australasian: Australia (Queensland). tillyardi Handschin Theristria tillyardi Handschin, 1935: 707, male. Holotype: male, Australia, Northern Territory (NHMB). DISTRIBUTION: Australasian: Australia (Northern Territory, Western Australia). uptoni Lambkin Theristria uptoni Lambkin, 1986a: 67, male, female. Holotype: female, Australia, Northern Territory (ANIC). DISTRIBUTION: Australasian: Australia (Northern Territory, South Australia, Western Australia).

Ohl: World Catalog of Mantispidae 157

Subfamily Calomantispinae Navás

Genus Calomantispa Banks

CALOMANTISPA Banks, 1913: 209. Type species: Calomantispa spectabilis Banks, 1913: 210, by monotypy. DISTRIBUTION: Australasian: Australia. picta Stitz Calomantispa picta Stitz, 1913: 45, female. Holotype: female, Australia, New South Wales (ZMB!). DISTRIBUTION: Australasian: Australia (Australian Capital Territory, New South Wales). spectabilis Banks Calomantispa spectabilis Banks, 1913: 210, male. Syntypes21: male, Australia, Queensland (ANIC ex MCZ). Calomantispa spectabilis var. nigrata Banks, 1913: 210, female. Syntypes21: female, Australia, Queensland (ANIC ex MCZ). Synonymized with Calomantispa spectabilis by Lambkin, 1986b: 4. Calomantispa spectabilis var. maculata Banks, 1913: 210, female. Syntypes21: female, Australia, Queensland (ANIC ex MCZ). Synonymized with Calomantispa spectabilis by Lambkin, 1986b: 4. DISTRIBUTION: Australasian: Australia (Queensland). venusta Lambkin Calomantispa venusta Lambkin, 1986b: 8, male, female. Holotype: male, Australia, New South Wales (AMSA). DISTRIBUTION: Australasian: Australia (Australian Capital Territory, New South Wales, Tasmania, Victoria).

Genus Nolima Navás

NOLIMA Navás, 1914i: 100. Type species: Nolima victor Navás, 1914i: 101, by original designation. Bellarminus Navás, 1914i: 102. Type species: Bellarminus pugnax Navás, 1914i: 103, by monotypy. Synonymized with Nolima by Penny, 1982b: 213. DISTRIBUTION: Nearctic and Neotropical: Costa Rica to southern USA. dine Rehn Nolima dine Rehn, 1939: 261, male, female. Holotype: male, USA: Arizona (CAS). DISTRIBUTION: Nearctic: USA (Arizona, New Mexico).

158 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

infensa Navás Nolima infensa Navás, 1924: 61, sex not indicated [as infensus [sic], incorrect original termination]. Holotype (or syntypes): sex unknown, Costa Rica (MNHN). DISTRIBUTION: Neotropical: Costa Rica. kantsi Rehn Nolima kantsi Rehn, 1939: 260, male. Holotype: male, USA: Texas (CAS). DISTRIBUTION: Nearctic: USA (Texas). pinal Rehn Nolima pinal Rehn, 1939: 257, male. Holotype: male, USA, Arizona (MCZ!). DISTRIBUTION: Nearctic: USA (Arizona). praeliator Navás Nolima praeliator Navás, 1914i: 101, sex not indicated. Syntypes: sex unknown, Mexico, Guerrero (BMNH!). DISTRIBUTION: Nearctic: Mexico (Guerrero). pugnax (Navás) Bellarminus pugnax Navás, 1914i: 103, sex not indicated. Holotype (or syntypes): sex unknown, Guatemala (BMNH!). DISTRIBUTION: Neotropical: Guatemala. victor Navás Nolima victor Navás, 1914i: 101, sex not indicated. Syntypes: sex not indicated, Mexico, Guerrero (BMNH!). DISTRIBUTION: Nearctic: Mexico (Guerrero). Ohl: World Catalog of Mantispidae 159

Subfamily Mantispinae Leach

Genus Asperala Lambkin

ASPERALA Lambkin, 1986b: 39. Type species: Mantispa erythraea Brauer, 1867b: 506, by original designation. DISTRIBUTION: Oriental and Australasian: Indonesia, Australia. erythraea (Brauer) Mantispa erythraea Brauer, 1867b: 506, sex not indicated. Syntypes: sex unknown, Australia, Queensland (ZMUH).22 DISTRIBUTION: Australasian: Australia (Queensland, New South Wales, Northern Territory, Western Australia). Oriental: Indonesia. hemichroa (Navás) Mantispa hemichroa Navás, 1914b: 480, sex not indicated. Holotype: female [according to New, 1996], Australia (BMNH!). DISTRIBUTION: Australasian: Australia (Northern Territory, Queensland, Western Australia).

Genus Austroclimaciella Handschin

AUSTROCLIMACIELLA Handschin, 1961: 287. Type species: Mantispa quadrituberculata Westwood, 1852: 264 (as Mantispa 4-tuberculata), by original designation. DISTRIBUTION: Palaearctic and Oriental: northeastern India to Japan, Philippines, Indonesia. brianti (Navás) Climaciella brianti Navás, 1914i: 92, sex not indicated [as Climiaciella [sic] Brianti [sic], incorrect original spelling and capitalization]. Holotype (or syntypes): sex unknown, India, Sikkim (BMNH!).23 DISTRIBUTION: Oriental: India (Sikkim). habutsuella (Okamoto) Climaciella habutsuella Okamoto, 1910: 542, male. Holotype: male, Okinawa (EIHU). Incorrectly cited as a new species in Okamoto, 1911: 300, which is a German translation of Okamoto, 1910.24 DISTRIBUTION: Oriental: China, Taiwan. Palaearctic: China, Japan. lacolombierei (Navás), new combination Climaciella lacolombierei Navás, 1931b: 3, female [as Lacolombierei [sic], incorrect original capitalization]. Holotype (or syntypes): female, China, Jiangsu (IZAS25). DISTRIBUTION: Palaearctic: China (Jiangsu).

160 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

leopoldi (Lestage) Climaciella leopoldi Lestage, 1934: 154, male [as Leopoldi [sic], incorrect original capitalization]. Holotype: male, Vietnam (ISNB). DISTRIBUTION: Oriental: Vietnam. luzonica (van der Weele) Mantispa quadrituberculata luzonica van der Weele, 1909: 94, male. Holotype: male, Philippines (ETH). Climaciella habutsuella var. fasciata Stitz, 1913: 32, male, female. Syntypes: male, female, Philippines (ZMB!). Synonymized with Austroclimaciella luzonica van der Weele by Handschin, 1961: 291. Mantispa taylori Navás, 1927c: 57, female [as Taylori [sic], incorrect original capitalization]. Holotype: female, Philippines (CN). Synonymized with Austroclimaciella luzonica van der Weele by Handschin, 1961: 290. SUBSPECIES: maculata (Stitz). DISTRIBUTION: Oriental: Philippines. maculata (Stitz) Climaciella habutsuella var. maculata Stitz, 1913: 32, male [incorrectly listed as female by Stitz, 1913]. Syntypes: male, Philippines (ZMB!). A subspecies of Austroclimaciella luzonica van der Weele. DISTRIBUTION: Oriental: Philippines. quadrituberculata (Westwood) Mantispa quadrituberculata Westwood, 1852: 264, sex not indicated [as 4-tuberculata [sic], incorrect original spelling26]. Holotype (or syntypes): sex unknown, India (OUM!). Climaciella miyakei Okamoto, 1910: 541, male [as Miyakei [sic], incorrect original capitalization]. Holotype: male, Japan (EIHU). Synonymized with Climaciella quadrituberculata by Kuwayama, 1962: 379. Incorrectly cited as a new species in Okamoto, 1911: 299, which is a German translation of Okamoto, 1910. Climaciella satsumensis Yazaki, 1927: 361, male, female. Syntypes: male, female, Japan (depository not indicated; EIHU?) Synonymized with Climaciella quadrituberculata by Kuwayama, 1962: 381. Climaciella tanegashimensis Yazaki, 1927: 363, female. Syntypes: female, Japan (depository not indicated; EIHU?). Synonymized with Climaciella quadrituberculata by Kuwayama, 1962: 381. DISTRIBUTION: Oriental: China, India, Indonesia (Java), Taiwan. Palaearctic: China, Japan. subfusca (Nakahara), new combination Climaciella subfusca Nakahara, 1912: 562, female [as Subfusca [sic], incorrect original capitalization]. Holotype: female, Japan (depository not indicated). Incorrectly cited as a new species in Nakahara, 1913: 232, which is an English translation of Nakahara, 1912. Climaciella subflava Nakahara, 1912: 15. A nomen nudum.27 DISTRIBUTION: Palaearctic: Japan.

Ohl: World Catalog of Mantispidae 161

weelei Handschin Austroclimaciella weelei Handschin, 1961: 289, sex not indicated. Holotype: sex unknown, Indonesia (RMNH). DISTRIBUTION: Oriental: Indonesia (Sumba)

Genus Austromantispa Esben-Petersen

AUSTROMANTISPA Esben-Petersen, 1917: 11 (as a subgenus of Mantispa). Type species: Mantispa imbecilla Gerstaecker, 1885a ["1884"]: 41, by original designation. DISTRIBUTION: Oriental and Australasian: Indonesia, Papua New Guinea, Australia. imbecilla (Gerstaecker) Mantispa imbecilla Gerstaecker, 1885a ["1884"]: 41, female. Holotype: female, Queensland (EMAU!). Mantispa pullula Banks, 1910: 104, sex not indicated. Holotype: male [according to Lambkin, 1986b], Australia (ANIC ex MCZ!). Synonymized with Mantispa imbecilla (now in Austromantispa) by Esben-Petersen, 1918: 34. Necyla doddi Navás, 1914d: 651, sex not indicated [as Doddi [sic], incorrect original capitalization]. Holotype: male [according to Lambkin, 1986b], Australia (BMNH!). Synonymized with Mantispa imbecilla (now in Austromantispa) by Esben-Petersen, 1918: 34. DISTRIBUTION: Oriental: Indonesia (Flores, Sumba, Timor). Australasian: Australia (New South Wales, Northern Territory, Queensland, South Australia, Tasmania, Victoria, Western Australia); Papua New Guinea. trevori Lambkin Austromantispa trevori Lambkin, 1986b: 31, male, female. Holotype: male, Australia (QM). DISTRIBUTION: Australasian: Australia (New South Wales, Northern Territory, Queensland, South Australia, Western Australia).

Genus Buyda Navás

BUYDA Navás, 1926c: 87. Type species: Buyda apicata Navás, 1926c: 87, by monotypy. DISTRIBUTION: Neotropical: Argentina to Nicaragua. apicata Navás Buyda apicata Navás, 1926c: 87, female. Holotype: female, type locality not indicated (CN). 28 DISTRIBUTION: Neotropical [or Afrotropical]: no specific locality. phthisica (Gerstaecker) Mantispa phthisica Gerstaecker, 1885a ["1884"]: 35, female. Holotype: female, Brazil (EMAU!). DISTRIBUTION: Neotropical: Argentina, Uruguay, Peru, Brazil, Colombia, Surinam, Costa Rica, Nicaragua.

162 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Genus Campanacella Handschin

CAMPANACELLA Handschin, 1961: 280. Type species: Mantispa hamiltonella Westwood, 1867: 506, by original designation. DISTRIBUTION: Oriental: India, Indonesia, Malaysia. hamiltonella (Westwood) Mantispa hamiltonella Westwood, 1867: 506, sex not indicated [as Hamiltonella [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, India (OUM!). DISTRIBUTION: Oriental: India, Indonesia (Java), Malaysia.

Genus Campion Navás

CAMPION Navás, 1914a: 65. Type species: Campion rubellus Navás, 1914a: 65, by monotypy. Isla Navás, 1914d: 652. Type species: Isla verenda Navás, 1914d: 652 (as verendus [sic]) (= Mantispa australasiae Guerin- Meneville, 1844 (1829-1838): 392), by monotypy. Synonymized with Campion by Lambkin, 1986b: 10.29 DISTRIBUTION: Australasian: Australia. australasiae (Guérin-Méneville) Mantispa australasiae Guérin-Méneville, 1844 ["1829-1838"]: 392, sex not indicated [as Australasiae [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Australia (MNHN30). Mantispa pictiventris Gerstaecker, 1885b ["1884"]: 103, male. Holotype: male, Australia, Queensland (EMAU!). Synonymized with Campion australasiae by Lambkin, 1986b: 15. Isla verenda Navás, 1914d: 652, sex not indicated [as verendus [sic], incorrect original termination]. Holotype: male, Australia, Victoria (BMNH!). Synonymized with Campion australasiae by Lambkin, 1986b: 15. Mantispa tillyardi Esben-Petersen, 1915 ["1914-1915"]: 642, male, female. Lectotype: male, Australia, Western Australia (WAM). Designated by Lambkin, 1986b: 16.31 Synonymized with Campion australasiae by Lambkin, 1986b: 15. DISTRIBUTION: Australasian: Australia (Australian Capital Territory, New South Wales, Northern Territory, Queensland, Western Australia, South Australia, Tasmania, Victoria). callosus Lambkin As Mantispilla crucifera: Navás, 1914a: 61 (one female paralectotype [incorrectly cited as a paratype by Lambkin, 1986b] misidentified, corrected to Campion callosus by Lambkin, 1986b). Campion callosus Lambkin, 1986b: 22, male, female. Holotype: male, Australia, Tasmania (ANIC). DISTRIBUTION: Australasian: Australia (Australian Capital Territory, New South Wales, Queensland, South Australia, Tasmania, Victoria, Western Australia).

Ohl: World Catalog of Mantispidae 163

chrysops (Stitz)32 Mantispa chrysops Stitz, 1913: 27, sex not indicated [abdomen lacking]. Holotype: female [according to Lambkin, 1986b: 54], Australia, Tasmania (ZMB!). A nomen dubium in Campion according to Lambkin, 1986b: 54. DISTRIBUTION: Australasian: Australia (Tasmania). cruciferus (Navás) Mantispa crucifera Navás, 1914a: 61, sex not indicated. Lectotype: male, Australia, Tasmania (OUM!). Designated by Lambkin, 1986b: 22.33 DISTRIBUTION: Australasian: Australia (Australian Capital Territory, New South Wales, Tasmania, Victoria). impressus (Navás) Mantispilla impressa Navás, 1914d: 649, sex not indicated. Holotype (or syntypes): female, Australia, Tasmania (BMNH!). DISTRIBUTION: Australasian: Australia (Australian Capital Territory, New South Wales, South Australia, Tasmania, Victoria). kroombitensis Lambkin and New Campion kroombitensis Lambkin and New, 1994: 118, male. Holotype: male, Australia, Queensland (QM). DISTRIBUTION: Australasian: Australia (Queensland). rubellus Navás Campion rubellus Navás, 1914a: 65, male, female. Lectotype: male, Australia, New South Wales (BMNH). Designated by Lambkin, 1986b: 14. DISTRIBUTION: Australasian: Australia (Australian Capital Territory, New South Wales, Northern Territory, Queensland, South Australia, Tasmania, Victoria, Western Australia). spiniferus Lambkin Campion spiniferus Lambkin, 1986b: 26, male, female. Holotype: male, Western Australia (ANIC). DISTRIBUTION: Australasian: Australia (New South Wales, South Australia, Victoria, Western Australia). tenuistriga (Gerstaecker) Mantispa tenuistriga Gerstaecker, 1885a ["1884"]: 42, female. Holotype: female, Australia, Queensland (EMAU!). Mantispa platycephala Stitz, 1913: 26, female. Holotype: female, Australia, Southern Australia (ZMB!). Synonymized with Campion tenuistriga by Lambkin, 1986b: 18. Mantispilla rubicunda Navás, 1933a: 79, female. Holotype: female, Australia, New South Wales (DEI!). Synonymized with Campion tenuistriga by Lambkin, 1986b: 18. DISTRIBUTION: Australasian: Australia (Australian Capital Territory, New South Wales, Northern Territory, Queensland, South Australia, Tasmania, Victoria, Western Australia).

164 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

vittatus (Guérin-Méneville)34 Mantispa vittata Guérin-Méneville in Duperrey, 1831 ["1830"]: 196, sex not indicated. Holotype (or syntypes): Australia, New South Wales (current depository unknown). A nomen dubium in Campion according to Lambkin, 1986b: 54. DISTRIBUTION: Australasia: Australia (New South Wales, Tasmania).

Genus Cercomantispa Handschin

CERCOMANTISPA Handschin, 1959a: 224. Type species: Mantispa mozambica Westwood, 1852: 269 (as "Cercomantispa mozambica Westwood 1852"), by original designation. DISTRIBUTION: Afrotropical: widespread. condei Poivre Cercomantispa condei Poivre, 1982a ["1981"]: 183, male, female. Holotype: male, Ivory Coast (MNHN). Incorrectly cited as new species by Poivre, 1982b: 9. DISTRIBUTION: Afrotropical: Ivory Coast. decellei Poivre Cercomantispa decellei Poivre, 1982a ["1981"]: 187, male, female. Holotype: male, Ivory Coast (MNHN35). Incorrectly cited as new species by Poivre, 1982b: 10. DISTRIBUTION: Afrotropical: Ivory Coast. finoti (Navás) Necyla finoti Navás, 1915b: 18, male [as Finoti [sic], incorrect original capitalization]. Holotype: male, Madagascar (CN). DISTRIBUTION: Afrotropical: Madagascar. keiseri Handschin Cercomantispa keiseri Handschin, 1963 ["1962"]: 214, sex not indicated. Holotype: sex unknown, Madagascar (NHMB). DISTRIBUTION: Afrotropical: Madagascar. mozambica (Westwood) Mantispa mozambica Westwood, 1852: 269, sex not indicated [as Mozambica [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Mozambique (OUM).36 DISTRIBUTION: Afrotropical: Mozambique. natalica (Navás) Mantispa natalica Navás, 1909a ["1908-1909"]: 476, sex not indicated. Holotype (or syntypes): sex unknown, South Africa (CN). DISTRIBUTION: Afrotropical: South Africa. ndjallai Poivre Cercomantispa ndjallai Poivre, 1981 ["1980"]: 81, male. Holotype: male, Cameroon (MHNG). DISTRIBUTION: Afrotropical: Cameroon.

Ohl: World Catalog of Mantispidae 165

nigricornis (Navás) Necyla nigricornis Navás, 1926a ["1925"]: 460, female. Holotype (or syntypes): female, DR Congo (MRAC). Mantispilla honesta Navás, 1936c: 356, sex not indicated. Holotype (or syntypes), sex unknown, DR Congo (MRAC). Synonymized with Cercomantispa nigricornis by Handschin, 1960a: 238. DISTRIBUTION: Afrotropical: DR Congo. paolina (Navás) Necyla paolina Navás, 1930a: 415, male. Holotype (or syntypes): male, Ethiopia (MCSN37). DISTRIBUTION: Afrotropical: Ethiopia. perparva (Esben-Petersen) Mantispa (Necyla) perparva Esben-Petersen, 1917: 4, male, female. Syntypes: male, female, South Africa (NHRS), DR Congo (ZMUH). Mantispilla striatella Navás, 1934a ["1933"]: 65, male. Syntypes: male, Madagascar (MNHN). Synonymized with Cercomantispa perparva by Handschin, 1963 ["1962"]: 215. Cercomantispa perparva var. obscura Handschin, 1963 ["1962"]: 216, sex not indicated. Syntypes: sex unknown, Madagascar (MRAC). An unavailable name38. DISTRIBUTION: Afrotropical: South Africa, Madagascar, Tanzania. picea (Esben-Petersen) Mantispa (Necyla) picea Esben-Petersen, 1917: 6, female. Holotype: female, South Africa (NHRS). DISTRIBUTION: Afrotropical: Madagascar, South Africa. pulla (Navás) Mantispilla pulla Navás, 1935 ["1934"]: 66, male, female. Syntypes: male, female, Madagascar (MZBS). DISTRIBUTION: Afrotropical: Madagascar. stenostigma (Navás) Necyla stenostigma Navás, 1936c: 358, sex not indicated. Holotype: sex unknown, DR Congo (MRAC). DISTRIBUTION: Afrotropical: DR Congo. tristella (Navás) Mantispilla tristella Navás, 1936c: 358, sex not indicated. Holotype (or syntypes): sex unknown, DR Congo (MRAC). DISTRIBUTION: Afrotropical: DR Congo, Ivory Coast. tristis (Navás) Mantispilla tristis Navás, 1936c: 357, sex not indicated. Holotype: sex unknown, DR Congo (MRAC). DISTRIBUTION: Afrotropical: DR Congo, South Africa, Uganda.

166 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

vulpes (Stitz) Mantispilla vulpes Stitz, 1913: 12, male. Holotype: male, Cameroon (ZMB!39). Necyla cercata Navás, 1915c: 178, male. Holotype: male, DR Congo (MRAC). Synonymized with Cercomantispa vulpes by Handschin, 1959a: 225. Incorrectly cited as a new species by Navás, 1916: 237. Mantispilla simplex Navás, 1936c: 355, sex not indicated. Syntypes: sex unknown, DR Congo (MRAC). Synonymized with Cercomantispa vulpes by Handschin, 1959a: 225. Mantispilla simplicia Fraser, 1952: 59, female. Holotype: female, Madagascar (MNHN). Synonymized with Cercomantispa vulpes by Handschin, 1963 ["1962"]: 216. DISTRIBUTION: Afrotropical: Cameroon, DR Congo, Madagascar, Tanzania, Uganda.

Genus Climaciella Enderlein

CLIMACIELLA Enderlein, 1910: 360. Type species: Mantispa brunnea Say in Keating, 1824: 309, by original designation. Climatiella; Zoological Record (Insecta), 1913: 394. Incorrect subsequent spelling of Climaciella. Nobrega Navás, 1914e: 233. Type species: Nobrega tinctus Navás, 1914e: 233, by monotypy. Synonymized with Climaciella by Penny, 1982b: 220. Climiaciella [sic]; Navás, 1919i: 597. Incorrect subsequent spelling of Climaciella. DISTRIBUTION: Nearctic and Neotropical: Argentina to USA. amapaensis Penny Climaciella amapaensis Penny, 1982a: 450, male. Holotype: male, Brazil (DZUP). DISTRIBUTION: Neotropical: Brazil. brunnea (Say) Mantispa brunnea Say in Keating, 1824: 309, male, female. Syntypes: male, female, USA, Minnesota, Pennsylvania (depository unknown according to Penny et al. 1997: 73). Mantispa varia Erichson, 1839: 161, sex not indicated. Syntypes: sex unknown, Mexico (ZMB! MCZ!). Synonymized with Mantispa brunnea by Hagen 1859: 406. Mantispa denaria Taylor, 1862: 494 [as Denarius [sic], incorrect original capitalization and termination]. Synonymized with by Kevan, 1989: 8. Mantispa burquei Provancher, 1875: 247, sex not indicated [as Burquei [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Canada (depository unknown). Synonymized with Mantispa brunnea by Provancher, 1877: 174. Mantispa brunnea var. occidentis Banks, 1911: 348, sex not indicated. Syntypes: male, USA, Colorado, Nevada, Washington (MCZ!). Synonymized with Climaciella brunnea by Welch and Kondratieff, 1991: 70. Climaciella rubescens Stitz, 1913: 37, female. Holotype: female, Mexico, Tabasco (ZMB!). Synonymized with Climaciella brunnea by Handschin, 1960c: 546. Climaciella rubescens var. unicolor Stitz, 1913: 39, female. Syntypes: females, Mexico, Sierra Mixteca (ZMB!). Synonymized with Climaciella brunnea by Handschin, 1960c: 549. Climaciella rubescens var. laciniata Stitz, 1913: 39, female. Syntypes: female, Mexico, Mexico (ZMB!). Synonymized with Climaciella brunnea by Handschin, 1960c: 549. Ohl: World Catalog of Mantispidae 167

Climaciella brunnea "instabilis morph" Opler, 1981: 166, in figure legend. An unavailable name.40 Climaciella brunnea "canadensis morph" Opler, 1981: 166, in figure legend. An unavailable name.40 Climaciella brunnea "carnifex morph" Opler, 1981: 166, in figure legend. An unavailable name.40 Climaciella brunnea "synoeca morph" Opler, 1981: 166, in figure legend. An unavailable name. 40 Climaciella brunnea "erythrocephalus morph" Opler, 1981: 167, in figure legend. An unavailable name. 40 DISTRIBUTION: Nearctic: USA (widespread), Mexico. Neotropical: Costa Rica, Mexico, Nicaragua, Puerto Rico. cubana Enderlein Climaciella cubana Enderlein, 1910: 362, female. Holotype: female, Cuba (MZPW!). DISTRIBUTION: Neotropical: Cuba. duckei Navás Climaciella duckei Navás, 1915a: 196, sex not indicated. [as Duckei [sic], incorrect original capitalization]. Holotype: sex unknown, Peru (NMBS). DISTRIBUTION: Neotropical: Brazil, Peru.

† henrotayi Nel † Climaciella henrotayi Nel, 1989 ["1988"]: 106, sex unkown. Holotype: sex unknown, France, Upper Oligocene (coll. Henrotay). DISTRIBUTION: Palaearctic: France (Upper Oligocene) (compression/impression). obtusa Hoffman in Penny Climaciella obtusa Hoffman in Penny, 2002: 254, male, female. Holotype: male, Ecuador (FSCA). DISTRIBUTION: Neotropical: Ecuador. personata (Stitz) Euclimacia personata Stitz, 1913: 40, male. Holotype: male, Bolivia (ZMB!). DISTRIBUTION: Neotropical: Bolivia. porosa Hoffman in Penny Climaciella porosa Hoffman in Penny, 2002: 256, male, female. Holotype: male, Costa Rica (CHAH). DISTRIBUTION: Neotropical: Costa Rica. semihyalina (Le Peletier de Saint-Fargeau and Audinet-Serville) Mantispa semihyalina Le Peletier de Saint-Fargeau and Audinet-Serville, 1825: 270, sex not indicated. Holotype (or syntypes): sex unknown, Brazil (MNHN?). Mantispa chalybea Erichson, 1839: 160, sex not indicated. Syntypes: sex unknown, Brazil, Suriname (ZMB! MCZ!). Synonymized with Climaciella semihyalina by Enderlein, 1910: 367. 168 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Mantispa grandis; Burmeister, 1839: 967. Not available with Burmeister as author.41 Synonymized with Climaciella semihyalina by Westwood, 1852: 253. Nobrega tinctus Navás, 1914e: 233, sex not indicated. Holotype: sex unknown, Brazil (BMNH!). Synonymized with Climaciella semihyalina by Penny, 1982a: 453. DISTRIBUTION: Neotropical: Argentina, Uruguay, Bolivia, Peru, Brazil, Costa Rica, French Guiana, Surinam, Mexico.

Genus Dicromantispa Hoffman in Penny

DICROMANTISPA Hoffman in Penny, 2002: 258. Type species: Mantispa sayi Banks, 1897: 23, by original designation. DISTRIBUTION: Nearctic and Neotropical: Argentina to USA. debilis (Gerstaecker) Mantispa debilis Gerstaecker, 1888 ["1887"]: 114, sex not indicated. Syntypes: sex unknown, Brazil (EMAU!). Mantispilla debilis var. nuda Stitz, 1913: 19, female. Holotype: female, Surinam (ZMB!). New synonym42. DISTRIBUTION: Neotropical: Brazil, Costa Rica. gracilis (Erichson) Mantispa gracilis Erichson, 1839: 169, female. Holotype: female, Brazil (ZMB!).43 Mantispilla debilis var. nigricornis Stitz, 1913: 19, female. Holotype: female, Venezuela (ZMB!). New synonym44. Mantispilla debilis var. rugicollis Stitz, 1913: 19, female. Holotype: female, Venezuela (ZMB!). New synonym45. Mantispa trilineata Navás, 1914e: 230, sex not indicated. Holotype, sex unknown, Brazil (BMNH!). Synonymized with Mantispa gracilis by Penny, 1982a: 439. Mantispa bruchi Navás, 1915d: 134, female [as Bruchi [sic], incorrect original capitalization]. Holotype: female, Argentina (type depository dubious46). Synonymized with Mantispa gracilis by Williner and Kormilev, 1958: 6. Mantispa calceata Navás, 1917: 401, male. Holotype (or syntypes): male, Argentina (CN). Synonymized with Mantispa gracilis by Williner and Kormilev, 1958: 6. Mantispa mista Navás, 1923a: 196, sex not indicated. Holotype: sex unknown, Argentina (MACN). Synonymized with Mantispa gracilis by Williner and Kormilev, 1958: 6. Mantispilla gounellei Navás, 1934b: 16, female. [as Gounellei [sic], incorrect original capitalization]. Holotype: female, Brazil (MNHN). Synonymized with Mantispa gracilis by Penny, 1982a: 439. DISTRIBUTION: Neotropical: Argentina, Brazil, Costa Rica, Venezuela. interrupta (Say), new combination Mantispa interrupta Say, 1825: [unpaginated text to plate 25]. Holotype: sex unknown, USA (Pennsylvania) (depository unknown, holotype probably lost according to Penny et al. 1997: 74). Mantispa aurea Taylor, 1862: 494 [as Aureus [sic], incorrect original capitalization and termination]. Synonymized with Mantispa interrupta by Kevan, 1989: 8. Ohl: World Catalog of Mantispidae 169

Mantispa cincticornis Banks, 1911: 347, sex not indicated. Holotype: female, USA, Texas (MCZ!). Synonymized with Mantispa interrupta by Welch and Kondratieff, 1991: 72. DISTRIBUTION: Nearctic: Canada, USA. sayi (Banks) Mantispa sayi Banks, 1897: 23, sex not indicated. Lectotype: male, USA, Texas (MCZ!). Designated by Hoffman, 1989: 638. Mantispa fuscicornis Banks, 1911: 347, sex not indicated. Lectotype: male, USA, Florida (MCZ!). Designated by Hoffman, 1989: 638. Synonymized with Mantispa sayi by Hoffman, 1989: 638. Mantispa uhleri Banks, 1943: 79, male, female. Holotype: male, USA, Pennsylvania (MCZ!). Synonymized with Mantispa sayi by Hoffman, 1989: 639. DISTRIBUTION: Nearctic: USA. Neotropical: Costa Rica, Cuba, Dominican Republic, Puerto Rico. synapsis Hoffman in Penny Dicromantispa synapsis Hoffman in Penny, 2002: 260, male, female. Holotype: male, Brazil (PMNH). DISTRIBUTION: Neotropical: Brazil.

Genus Entanoneura Enderlein

ENTANONEURA Enderlein, 1910: 358. Type species: Mantispa limbata Gerstaecker, 1885a ["1884"]: 36, by original designation. Entatoneura; Navás, 1914i: 90, 1929c: 381. Incorrect subsequent spelling of Entanoneura. DISTRIBUTION: Neotropical: Argentina to Nicaragua. Afrotropical: Madagascar. Oriental: China, Timor, Myanmar. albertisii Navás Entanoneura albertisii Navás, 1929c: 381, sex not indicated [as Entatoneura [sic] Albertisii [sic], incorrect original spelling and capitalization]. Holotype (or syntypes): sex not indicated, Timor (MCSN).47 DISTRIBUTION: Oriental: Timor. batesella (Westwood) Mantispa batesella Westwood, 1867: 507, male, female [as Batesella [sic], incorrect original capitalization]. Syntypes "Amazonia" (OUM!). Mantispa limbata Gerstaecker, 1885a ["1884"]: 36, female. Holotype: female, Panama (EMAU!). Synonymized with Entanoneura batesella by Hoffman in Penny, 2002: 263. Entanoneura picta Navás, 1914i: 90, sex not indicated [in Entatoneura [sic]]. Holotype (or syntypes): sex unknown, Panama (BMNH!). Synonymized with Mantispa limbata by Esben-Petersen, 1917: 13. Entanoneura [sic!] chopardi Navás, 1933c: 310, female [as Entatoneura [sic] Chopardi [sic], incorrect original spelling and capitalization]. Holotype (or syntypes): female, French Guiana (MNHN). Synonymized with Entanoneura batesella by Penny, 1982a: 448. 170 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Entatoneura [sic!] jocosa Navás, 1933c: 311, female [in Entatoneura [sic]]. Holotype (or syntypes): female, Brazil (MNHN). Synonymized with Entanoneura batesella by Penny, 1982a: 448. Entanoneura similis Handschin, 1960c: 531, male . Holotype48: male, Brazil (NHMB). Synonymized with Entanoneura batesella by Penny, 1982a: 448. DISTRIBUTION: Neotropical: Brazil, Colombia, Costa Rica, French Guiana, Nicaragua, Panama. brunneonigra Handschin Entanoneura brunneonigra Handschin, 1960: 533, sex not indicated. Holotype: sex unknown, Brazil (MZSP). DISTRIBUTION: Neotropical: Brazil. costalis (Erichson) Mantispa costalis Erichson, 1839: 164, sex not indicated. Holotype: female, Brazil (ZMB!). Mantispa areolaris Westwood, 1852: 265, sex not indicated. Holotype (or syntypes): sex unknown, Brazil (OUM!). Synonymized with Mantispa costalis by Handschin, 1960c: 534. DISTRIBUTION: Neotropical: Argentina, Brazil. feae Navás Entatoneura feae Navás, 1929c: 382, male, female [as Entatoneura [sic] Feae [sic], incorrect original capitalization]. Syntypes: male, female, Myanmar (MCSN). Entatoneura feai Navás, 1930c: 24 [as Entatoneura [sic] Feai [sic], incorrect original capitalization]. An unjustified emendation of E. feae Navás, 1929c49. DISTRIBUTION: Oriental: Myanmar. mariae (Navás) Mantispa mariae Navás,1909a ["1908-1909"]: 477, sex not indicated [as Mariae [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Madagascar (CN). DISTRIBUTION: Afrotropical: Madagascar. sinica C.-k. Yang Entanoneura sinica C.-k. Yang, 1999: 133, female. Holotype: female, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian).

Genus Euclimacia Enderlein

EUCLIMACIA Enderlein, 1910: 362. Type species: Euclimacia partita Enderlein, 1910: 366, by original designation. DISTRIBUTION: Palaearctic, Oriental, and Australasian: northeastern India to northern Australia.

Ohl: World Catalog of Mantispidae 171

badia Okamoto Euclimacia badia Okamoto, 1910: 543, male. Holotype: male, Taiwan (EIHU). Incorrectly cited as a new species in Okamoto, 1911: 301, which is a German translation of Okamoto, 1910. Euclimacia sauteri Navás, 1927b: 39, female [as Sauteri [sic], incorrect original capitalization]. Holotype: female, Taiwan (DEI!). New synonym50. DISTRIBUTION: Palaearctic: Japan. Oriental: Taiwan. basiflava Handschin Euclimacia basiflava Handschin, 1961: 273, sex not indicated. Holotype: sex unknown, Malaysia (NHMW!). DISTRIBUTION: Oriental: Malaysia. burmanella (Westwood) Mantispa burmanella Westwood, 1867: 507, sex not indicated [as Burmanella [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Myanmar (BMNH!). DISTRIBUTION: Oriental: Myanmar. celebica Handschin Euclimacia celebica Handschin, 1961: 267, male. Holotype: male, Indonesia, Sulawesi (NHMW!). DISTRIBUTION: Oriental: Indonesia (Sulawesi). cottami Navás Euclimacia cottami Navás, 1914i: 93, sex not indicated. [as Cottami [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, India, Sikkim (BMNH!). DISTRIBUTION: Oriental: India (Sikkim). flavicauda Esben-Petersen Euclimacia flavicauda Esben-Petersen, 1917: 7, sex not indicated. Holotype: sex unknown, Indonesia, Sumatra (ZMUH). DISTRIBUTION: Oriental: Indonesia (Sumatra). flavocincta Stitz Euclimacia flavocincta Stitz, 1913: 42, female. Holotype: female, Solomon Islands (ZMB!). DISTRIBUTION: Australasian: Solomon Islands. fusca Stitz Euclimacia fusca Stitz, 1913: 41, male [incorrectly listed as female by Stitz, 1913]. Holotype: male, Taiwan (ZMB!).51 DISTRIBUTION: Palaearctic: Japan. Oriental: Taiwan. gerstaeckeri Banks Euclimacia gerstaeckeri Banks, 1920: 341, sex not indicated. Holotype (or syntypes): sex unknown, Singapore (MCZ!). DISTRIBUTION: Oriental: Singapore.

172 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

grandis (Guérin-Méneville in Duperrey) Mantispa grandis Guérin-Méneville in Duperrey, 1831 ["1830"]: 196, sex not indicated. Holotype (or syntypes): sex unknown, Indonesia, Ambon (current depository unknown52). Mantispa guerinii Westwood, 1852: 255 [as Guerinii [sic], incorrect original capitalization]. Unnecessary replacement name for Mantispa grandis Guérin-Menéville.53 DISTRIBUTION: Oriental: Indonesia (Ambon, Moluccas). horstaspoecki Ohl Euclimacia horstaspoecki Ohl, 2004: 193, male. Holotype: male, Thailand (ZMB!). DISTRIBUTION: Oriental: Thailand. jacobsoni Handschin Euclimacia jacobsoni Handschin, 1961: 262, sex not indicated. Holotype: sex unknown, Indonesia, Sumatra (ZMAN). DISTRIBUTION: Oriental: Indonesia (Sumatra). metallica Esben-Petersen Euclimacia metallica Esben-Petersen, 1917: 10, female. Holotype: female, Indonesia, Sulawesi (ZMUH). DISTRIBUTION: Oriental: Indonesia (Sulawesi, Sumatra). morosa (Gerstaecker) Mantispa morosa Gerstaecker, 1888 ["1887"]: 149, female. Holotype (or syntypes): female, Philippines, Palawan (EMAU!). DISTRIBUTION: Oriental: Borneo, Philippines (Palawan). nelsoni Navás Euclimacia nelsoni Navás, 1914i: 94, sex not indicated [as Nelsoni [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Sri Lanka (BMNH!). DISTRIBUTION: Oriental: Sri Lanka. nigra Handschin Euclimacia nigra Handschin, 1961: 274, sex not indicated. Holotype: sex unknown, Indonesia, Java (RMNH). DISTRIBUTION: Oriental: Indonesia (Java). nodosa (Westwood), new combination Mantispa nodosa Westwood, 1847 ["1848"]: 70, sex not indicated. Holotype (or syntypes): sex unknown, India, Assam (OUM!).54 DISTRIBUTION: Oriental: India (Assam). nuchalis (Gerstaecker) Mantispa nuchalis Gerstaecker, 1885a ["1884"]: 38, female. Lectotype: male, Australia, Queensland (EMAU! ZMUH according to Esben-Petersen, 1917b: 218). Designated by Lambkin, 1986b: 46. Ohl: World Catalog of Mantispidae 173

Euclimacia flavicostata Esben-Petersen, 1917b: 217, male. Holotype: male, Australia, Queensland (BMNH!). Synonymized with Euclimacia nuchalis by Lambkin, 1986b: 46. DISTRIBUTION: Australasian: Australia (New South Wales, Northern Territory, Queensland). partita Enderlein Euclimacia partita Enderlein, 1910: 366, female. Syntypes: female, Indonesia, Sulawesi (MZPW!). DISTRIBUTION: Oriental: Indonesia (Sulawesi). regina Esben-Petersen Euclimacia regina Esben-Petersen, 1917: 8, female. Holotype: female [locality not indicated, probably Indonesia, Sunda Islands55] (NHRS). DISTRIBUTION: Oriental: Indonesia (Java, Sunda Islands), Singapore. rhombica Navás Euclimacia rhombica Navás, 1914i: 96, sex not indicated. Holotype (or syntypes): sex unknown, Myanmar (BMNH!). DISTRIBUTION: Oriental: Myanmar. rufa Esben-Petersen Euclimacia rufa Esben-Petersen, 1928: 230. Holotype: sex unknown, Indonesia: no specific locality (MBBJ). DISTRIBUTION: Oriental: Indonesia. ruficauda Enderlein Euclimacia ruficauda Enderlein, 1910: 365, female. Holotype: female, Indonesia, Sulawesi (MZPW!). DISTRIBUTION: Oriental: Indonesia (Sulawesi). rufocincta Handschin Euclimacia rufocincta Handschin, 1961: 270, sex not indicated. Holotype: sex unknown, Borneo (RMNH).56 DISTRIBUTION: Oriental: Borneo. superba Lambkin Euclimacia superba Lambkin, 1987: 13, male, female. Holotype: male, Australia, Queensland (ANIC). DISTRIBUTION: Australasian: Australia (Queensland). tagalensis Banks Euclimacia tagalensis Banks, 1914 ["1913"]: 176, sex not indicated. Holotype (or syntypes): Philippines, Luzon (MCZ!). DISTRIBUTION: Palaearctic: Japan. Oriental: Philippines (Luzon).

174 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

torquata Navás Euclimacia torquata Navás, 1914i: 95, sex not indicated. Holotype: sex unknown, Australia, Queensland (BMNH!). DISTRIBUTION: Australasian: Australia (New South Wales, Queensland), New Guinea. triangularis Handschin Euclimacia triangularis Handschin, 1961: 265, sex not indicated. Holotype: sex unknown, Philippines (ZMUC). DISTRIBUTION: Oriental: Philippines. vespiformis Okamoto Euclimacia vespiformis Okamoto, 1910: 543, male. Holotype: male, Taiwan (EIHU). Incorrectly cited as a new species in Okamoto, 1911: 300, which is a German translation of Okamoto, 1910. DISTRIBUTION: Palaearctic: Japan. Oriental: Taiwan. woodhousei Navás Euclimacia woodhousei Navás, 1914i: 94, sex not indicated [as Woodhousei [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, India, Sikkim (BMNH!). DISTRIBUTION: Oriental: India (Sikkim). zonalis Navás Euclimacia zonalis Navás, 1914i: 96, sex not indicated. Holotype (or syntypes): sex unknown, Indonesia, Sulawesi (BMNH!). DISTRIBUTION: Oriental: Indonesia (Sulawesi).

Genus Eumantispa Okamoto

EUMANTISPA Okamoto, 1910: 538. Type species: Mantispa suzukii Okamoto, 1910: 538 (as "Mantispa suzukii Mats. [sic]") (= Mantispa harmandi Navás 1909a ["1908-1909"]: 480), by original designation. Incorrect type species designation by Kuwayama, 1962: 378, of Mantispa harmandi Navás. Incorrectly cited as a new taxon in Okamoto, 1911: 294, which is a German translation of Okamoto, 1910. Stenispa Navás, 1914i: 90. Type species: Eumantispa hypogastrica Navás, 1914i: 89, by monotypy. A junior homonym of Stenispa Baly, 1858: 13 (Insecta: Coleoptera: Chrysomelidae). New synonym57. DISTRIBUTION: Palaearctic, Oriental, and Australasian: Far eastern USSR, Japan, northeastern India to New Guinea. araucariae Handschin Eumantispa araucariae Handschin, 1961: 295, sex not indicated. Syntypes: sex unknown, New Guinea (RMNH). DISTRIBUTION: Australasian: New Guinea.

Ohl: World Catalog of Mantispidae 175

ferruginea Stitz Eumantispa harmandi var. ferruginea Stitz, 1913: 31, male. Holotype: male, Indonesia, Sulawesi (ZMB!). DISTRIBUTION: Oriental: Indonesia (Sulawesi). fuscata Navás Eumantispa fuscata Navás, 1914i: 88, sex not indicated. Holotype (or syntypes): sex unknown, Indonesia, Sulawesi (BMNH!). DISTRIBUTION: Oriental: Indonesia (Sulawesi). fuscicolla C.-k. Yang Eumantispa fuscicolla C.-k. Yang, 1992: 445 (Chinese), 451 (English abstract), male. Holotype: male, China, Yunnan (IZAS). DISTRIBUTION: Oriental: China (Yunnan). harmandi (Navás) Mantispa harmandi Navás, 1909a ["1908-1909"]: 480, male, fermale [as Harmandi [sic], incorrect original capitalization]. Syntypes: male, female, Japan (MNHN). Mantispa nawae Miyake, 1910: 216, female [as Nawae [sic], incorrect original capitalization]. Holotype: female, Japan (NELG). Synonymized with Eumantispa harmandi by Kuwayama, 1925 ["1924-1925"]: 257. Mantispa sasakii Miyake, 1910: 217, female [as Sasakii [sic], incorrect original capitalization] Syntypes: female, Japan (ACIU, CMY). Synonymized with Eumantispa harmandi by Nakahara, 1912: 559. Eumantispa suzukii Okamoto, 1910: 538, male, female. Lectotype: female, Japan (EIHU). Designated by Kuwayama, 1966: 137. Synonymized with Eumantispa harmandi by Nakahara, 1912: 559. Incorrectly cited as a new species in Okamoto, 1911: 295, which is a German translation of Okamoto, 1910. SUBSPECIES: taeniata Stitz. DISTRIBUTION: Palaearctic: Far Eastern USSR, Japan, Sibiria. Oriental: Taiwan, widespread in SE Asia. hypogastrica Navás, revised status Eumantispa hypogastrica Navás, 1914i: 89, sex not indicated. Holotype (or syntypes): sex unknown, Indonesia, Sulawesi (BMNH!).58 DISTRIBUTION: Oriental: Indonesia (Sulawesi). lombokensis Handschin Eumantispa lombokensis Handschin, 1961: 294, sex not indicated. Holotype: sex unknown, Indonesia, Lombok (NHMW!). DISTRIBUTION: Oriental: Indonesia (Lombok). moluccensis Handschin Eumantispa moluccensis Handschin, 1961: 296, sex not indicated. Holotype: sex unknown, Moluccas (RMNH). DISTRIBUTION: Oriental: Moluccas.

176 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

rugicollis (Navás) Mantispa rugicollis Navás, 1905: 54, sex not indicated. Holotype (or syntypes): sex unknown, India, Bengal (NMPC?). DISTRIBUTION: Oriental: India (Assam, Bengal, Sikkim). taeniata Stitz Eumantispa harmandi var. taeniata Stitz, 1913: 31, female. Holotype: female, Papua New Guinea (ZMB!). A subspecies of Eumantispa harmandi (Navás). DISTRIBUTION: Australasian: Papua New Guinea. taiwanensis Kuwayama Eumantispa taiwanensis Kuwayama, 1925: 258, male. Holotype: male, Taiwan (TARI). DISTRIBUTION: Palaearctic: Japan. Oriental: Taiwan. tibetana C.-k. Yang Eumantispa tibetana C.-k. Yang in Huang et al., 1988: 205 (Chinese), 212 (English), male. Holotype: male, China (Xizang) (IZAS). DISTRIBUTION: Palaearctic: China (Xizang).

Genus Haematomantispa Hoffman in Penny

HAEMATOMANTISPA Hoffman in Penny, 2002: 263. Type species: Mantispa nubeculosa Navás, 1933b: 196, by original designation. DISTRIBUTION: Neotropical: Costa Rica. nubeculosa (Navás) Mantispa nubeculosa Navás, 1933b: 196, sex not indicated. Holotype (or syntype): sex unknown, Costa Rica (ZMUH). DISTRIBUTION: Neotropical: Costa Rica.

Genus Leptomantispa Hoffman in Penny

LEPTOMANTISPA Hoffman in Penny, 2002: 264. Type species: Mantispilla pulchella Banks, 1912: 179, by original designation. DISTRIBUTION: Neotropical and Nearctic: Brazil to Canada. antillesensis Hoffman in Reeves and Robinson Leptomantispa antillesensis of Hoffman in Reeves and Robinson, 1999: 461. An unavailable name59. chaos Hoffman in Penny Leptomantispa chaos Hoffman in Penny, 2002: 266, male, female. Holotype: male, Panama (MCZ). DISTRIBUTION: Neotropical: Brazil, French Guiana, Venezuela, Panama, Guatemala.

Ohl: World Catalog of Mantispidae 177

nymphe Hoffman in Penny Leptomantispa nymphe Hoffman in Penny, 2002: 268, male, female. Holotype: male, Costa Rica (USNM). DISTRIBUTION: Neotropical: French Guiana, Panama, Costa Rica. pulchella (Banks) Mantispilla pulchella Banks, 1912: 179, sex not indicated. Holotype: female, USA, Utah (MCZ!). DISTRIBUTION: Nearctic: USA, Canada. Neotropical: Costa Rica, Cuba, Mexico.

Genus Madantispa Fraser

MADANTISPA Fraser, 1952: 58. Type species: Madantispa seyrigi Fraser, 1952: 58, by original designation. DISTRIBUTION: Afrotropical: Madagascar. minuta Handschin Madantispa minuta Handschin, 1963 ["1962"]: 219, male, female. Holotype: male, Madagascar (MNHN). DISTRIBUTION: Afrotropical: Madagascar. pauliani Handschin Madantispa pauliani Handschin, 1963 ["1962"]: 219, male. Holotype: male, Madagascar (MNHN). DISTRIBUTION: Afrotropical: Madagascar. seyrigi Fraser Madantispa seyrigi Fraser, 1952: 58, female [as Seyrigi [sic], incorrect original capitalization]. Holotype: female, Madagascar (MNHN). DISTRIBUTION: Afrotropical: Madagascar.

Genus Mantispa Illiger in Kugelann

Mantispa Illiger in Kugelann, 1798: 499. Type species: Mantis pagana Fabricius, 1775: 278 (= Raphidia styriaca Poda, 1761: 101), by monotypy. Incorrect type species designation by Kuwayama, 1962: 377, of Raphidia styriaca Poda, 1761: 101 (as "Mantispa styriaca Poda"). Amycla Rafinesque, 1815: 118. Unjustified emendation of Mantispa Illiger in Kugelann, 1798. Amycla was considered an emendation of Mantispa Illiger in Kugelann, 1798, by Neave, 1939, 1: 167. Mantispilla Enderlein, 1910: 346 (as subgenus of Mantispa Illiger in Kugelann, 1798). Type species: Mantispa indica Westwood, 1852: 268, by original designation. Synonymized with Mantispa by Penny, 1982b: 217. 60 DISTRIBUTION: worldwide (except Australia) .

178 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

adelungi Navás Mantispa adelungi Navás, 1912a ["1911"]: 536, male. Lectotype: male, Caucasus area (ZIL). Designated by Krivokhatsky, 1995: 10. DISTRIBUTION: Palaearctic: Caucasus area. agapeta (Navás) Mantispilla agapeta Navás, 1914l: 213, sex not indicated. Holotype: sex unknown, Sri Lanka (BMNH!). DISTRIBUTION: Oriental: Sri Lanka. alicante Banks Mantispa alicante Banks, 1913: 208, male. Holotype: male, India (MCZ!). DISTRIBUTION: Oriental: India. amabilis Gerstaecker Mantispa amabilis Gerstaecker, 1894 ["1893"]: 152, female. Syntypes: females, Indonesia, Java (EMAU!). DISTRIBUTION: Oriental: Indonesia (Flores, Java, Sumatra). ambonensis Ohl, new name Mantispilla basalis Navás, 1929c: 380, female. Holotype (or syntypes): female, Indonesia, Ambon (MCSN). A junior primary and junior secondary homonym of Mantispilla basalis Navás, 1927c (now in Mantispa). Mantispa ambonensis Ohl. New replacement name for Mantispilla basalis Navás, 1929c (now in Mantispa), a junior primary and junior secondary homonym of Mantispilla basalis Navás, 1927c (now in Mantispa).61 DISTRIBUTION: Oriental: Indonesia (Ambon). annulicornis Gerstaecker Mantispa annulicornis Gerstaecker, 1888 ["1887"]: 151, sex not indicated. Syntypes: sex unknown, Indonesia, Java (EMAU). SUBSPECIES: nigricornis (Enderlein), stenoptera (Gerstaecker). DISTRIBUTION: Oriental: Indonesia (Java, Sulawesi, Sumatra), Thailand. aphavexelte U. Aspöck and H. Aspöck Mantispa aphavexelte U. Aspöck and H. Aspöck, 1994: 110, male, female. Holotype: male, Greece (HUAC). DISTRIBUTION: Palaearctic: southern Europe, Turkey, Caucasus area, Kazakhstan, Mongolia, China? ariasi Penny Mantispa ariasi Penny, 1982a: 454, male, female. Holotype: male, Brazil (INPA). DISTRIBUTION: Neotropical: Brazil.

Ohl: World Catalog of Mantispidae 179

axillaris Navás Mantispa axillaris Navás, 1908 ["1907-1908"]: 412, male, female. Syntypes: male, female, Brazil (MNHN, MZBS). DISTRIBUTION: Neotropical: Brazil. azihuna (Stitz) Mantispilla azihuna Stitz, 1913: 7, female. Holotype: female, Taiwan (ZMB!). DISTRIBUTION: Palaearctic: Japan. Oriental: Taiwan. basalis (Navás) Mantispilla basalis Navás, 1927c: 59, female. Holotype (or syntypes): female, Bolivia (CN). DISTRIBUTION: Neotropical: Bolivia. basilei (Navás) Mantispilla basilei Navás, 1930g: 27, female [as Basilei [sic], incorrect original capitalization]. Holotype: female, Ethiopia (MCSN). DISTRIBUTION: Afrotropical: Ethiopia. bicolor (Stitz) Mantispilla bicolor Stitz, 1913: 8, female. Syntypes: female, Vietnam (ZMB!). SUBSPECIES: immaculata (Stitz). DISTRIBUTION: Oriental: Vietnam. boliviana (Navás) Mantispilla boliviana Navás, 1927c: 58, female. Holotype (or syntypes): female, Bolivia (CN). DISTRIBUTION: Neotropical: Bolivia. brevistigma C.-k. Yang Mantispa brevistigma C.-k. Yang, 1999: 134, female. Holotype: female, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian). capeneri Handschin Mantispa capeneri Handschin, 1959a: 207, male, female. Syntypes: male, female, South Africa (MRAC). DISTRIBUTION: Afrotropical: South Africa. castaneipennis Esben-Petersen Mantispa (Mantispilla) castaneipennis Esben-Petersen, 1917: 3, female. Holotype: female, South Africa (ZMUH). DISTRIBUTION: Afrotropical: South Africa. celebensis Enderlein Mantispa (Mantispilla) celebensis Enderlein, 1910: 348, female. Holotype: female, Indonesia, Sulawesi (MZPW!). DISTRIBUTION: Oriental: Indonesia (Sulawesi).

180 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

centenaria Esben-Petersen Mantispa (Mantispilla) centenaria Esben-Petersen, 1917: 2, male. Holotype: male, South Africa (ZMUC). DISTRIBUTION: Afrotropical: South Africa. ceylanica (Stitz) Mantispilla indica var. ceylanica Stitz, 1913: 10, female [as ceylanica [sic], incorrect original spelling]. Syntypes: female, Sri Lanka (ZMB!). A subspecies of Mantispa indica Westwood. DISTRIBUTION: Oriental: Sri Lanka. chlorodes (Navás) Mantispilla chlorodes Navás, 1914h: 25, sex not indicated. Holotype (or syntypes): sex unknown, Panama (BMNH!). DISTRIBUTION: Neotropical: Panama. chlorotica (Navás) Mantispilla chlorotica Navás, 1912: 200, sex not indicated. Holotype (or syntypes): sex unknown, Paraguay (ZSMC!). DISTRIBUTION: Neotropical: Paraguay. completa Banks Mantispa completa Banks, 1920: 340, sex not indicated. Holotype (or syntypes), sex unknown, Philippines, Luzon (MCZ!). DISTRIBUTION: Oriental: Philippines (Luzon). confluens Navás Mantispa confluens Navás, 1914h: 19, sex not indicated. Holotype (or syntypes): sex unknown, Panama (BMNH!). DISTRIBUTION: Neotropical: Panama. coomani (Navás) Mantispilla coomani Navás, 1930c: 12, sex not indicated [as Coomani [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Vietnam (MNHN). DISTRIBUTION: Oriental: Vietnam. coorgensis Ohl, new name Mantispa femoralis Banks, 1933: 2, sex not indicated. Syntypes: sex unknown, India (MCZ!). A junior primary homonym of Mantispa femoralis Navás, 1914h. Mantispa coorgensis Ohl: New replacement name for Mantispa femoralis Banks, a junior primary homonym of Mantispa femoralis Navás, 1914h.62 DISTRIBUTION: Oriental: India. cora Newman Mantispa cora Newman, 1838: 401, sex not indicated. Holotype (or syntypes): sex unknown, India (BMNH!). DISTRIBUTION: Oriental: India. Ohl: World Catalog of Mantispidae 181

cordieri (Navás) Mantispilla cordieri Navás, 1933a: 78, female [as Cordieri [sic], incorrect original capitalization]. Holotype (or syntypes): female, Indonesia, Java (MNHN). DISTRIBUTION: Oriental: Indonesia (Java). crenata Navás Mantispa crenata Navás, 1914i: 83, sex not indicated. Holotype (or syntypes): sex unknown, Sri Lanka (BMNH! CUMZ?). DISTRIBUTION: Oriental: Sri Lanka. decepta Banks Mantispa decepta Banks, 1920: 341, sex not indicated. Holotype (or syntypes): sex unknown, Philippines, Mindanao (MCZ!). DISTRIBUTION: Oriental: Indonesia (Sulawesi), Philippines (Mindanao). decumana Hagen Mantispa decumana Hagen, 1866: 426 [authorship incorrectly attributed to Erichson]. A nomen nudum.63 delicata (Navás) Mantispilla delicata Navás, 1914g: 649, sex not indicated. Holotype (or syntypes): sex unknown, South Africa (BMNH!). DISTRIBUTION: Afrotropical: South Africa. deliciosa (Navás) Mantispilla deliciosa Navás, 1927e ["1927-1928"]: 4, female. Holotype: female, China, Kiangsu (NHRS). DISTRIBUTION: Oriental: China (Kiangsu). diminuta Matsumura Mantispa diminuta Matsumura, 1907: 169. Holotype: male, Tokyo (EIHU). A subspecies of Mantispa japonica McLachlan. Mantispa dimidiata Matsumura, 1908: 36. Lapsus calami. DISTRIBUTION: Palaearctic: Japan. dispersa (Navás) Mantispilla dispersa Navás, 1914g: 648, sex not indicated. Syntypes: sex unknown, Malawi, South Africa, Zimbabwe (BMNH!). A subspecies of Mantispa tenella Erichson according to Handschin, 1959a: 203 (as Mantispa tenella var. dispersa). DISTRIBUTION: Afrotropical: Malawi, South Africa, Zimbabwe. ellenbergeri (Navás) Mantispilla ellenbergeri Navás, 1927c: 60 female [as Ellenbergeri [sic], incorrect original capitalization]. Holotype (or syntypes): female, South Africa (MNHN). DISTRIBUTION: Afrotropical: South Africa.

182 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

elpidica (Navás) Mantispilla elpidica Navás, 1914b: 88, sex not indicated. Holotype (or syntypes): sex unknown, South Africa (BMNH!). DISTRIBUTION: Afrotropical: South Africa. enderleini Banks Mantispa enderleini Banks, 1914 ["1913"]: 176, sex not indicated. Holotype (or syntypes): sex unknown, Philippines, Luzon (MCZ!). DISTRIBUTION: Oriental: Philippines (Luzon). femoralis Navás Mantispa femoralis Navás, 1914h: 25, sex not indicated. Holotype (or syntypes): sex unknown, Mexico, Orizaba (BMNH!). DISTRIBUTION: Nearctic: Mexico (Orizaba). fenestralis Navás Mantispa fenestralis Navás, 1914i: 84, sex not indicated. Holotype (or syntypes): sex unknown, Sri Lanka (CUMZ). DISTRIBUTION: Oriental: Sri Lanka. finoti Navás Mantispa finoti Navás, 1909a ["1908-1909"]: 476, sex not indicated [as Finoti [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Madagascar (CN). DISTRIBUTION: Afrotropical: Madgascar. flavicauda (Navás) Mantispilla flavicauda Navás, 1914i: 86, male. Holotype (or syntypes): male, Mexico, Guerrero (BMNH!). DISTRIBUTION: Nearctic: Mexico (Guerrero). flavinota Handschin Mantispa flavinota Handschin, 1963 ["1962"]: 221, sex not indicated. Holotype: sex unknown, Madagascar (NHMB). DISTRIBUTION: Afrotropical: Madgascar. florealis Hagen Mantispa florealis Hagen, 1866: 426. A nomen nudum.64 floridana Banks Mantispa floridana Banks, 1897: 23, sex not indicated. Holotype: sex unknown, USA, Florida (MCZ). DISTRIBUTION: Nearctic: USA (Florida). frontalis (Navás) Mantispilla frontalis Navás, 1914e: 232, male. Holotype (or syntypes): male, Paraguay (ZSMC!). DISTRIBUTION: Neotropical: Paraguay.

Ohl: World Catalog of Mantispidae 183

fuliginosa Loew Mantispa fuliginosa Loew in Hagen, 1859: 405, sex not indicated. Holotype (or syntypes): sex unknown, Sudan (SMFD65). DISTRIBUTION: Afrotropical: Sudan. fuscipennis Erichson Mantispa fuscipennis Erichson, 1839: 168, sex not indicated. Holotype: male, South Africa (ZMB). DISTRIBUTION: Afrotropical: South Africa, Tanzania. gillavryna (Navás) Mantispilla gillavryna Navás, 1926b: 6, female. Holotype: female, Surinam (CN). DISTRIBUTION: Neotropical: Surinam. gradata (Navás) Mantispilla gradata Navás, 1926c: 86, female. Holotype (or syntypes): female, Brazil (CN). DISTRIBUTION: Neotropical: Brazil. greeni Banks Mantispa greeni Banks, 1913: 209, female. Holotype: female, Sri Lanka (MCZ!). DISTRIBUTION: Oriental: Sri Lanka. guttula Fairmaire in Thompson Mantispa guttula Fairmaire in Thomson, 1858: 261, sex not indicated. Holotype (or syntypes): sex unknown, Gabon (depository unknown). DISTRIBUTION: Afrotropical: Gabon. haematina (Navás) Mantispilla haematina Navás, 1914a: 62, female. Holotype: female, Zimbabwe (OUM!). DISTRIBUTION: Afrotropical: Zimbabwe. haugi Navás Mantispa haugi Navás, 1909a ["1908-1909"]: 475, sex not indicated [as Haugi [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Congo (MNHN). DISTRIBUTION: Afrotropical: Congo. icterica Pictet Mantispa perla var. icterica Pictet, 1865: 58, sex not indicated. Holotype: female, Spain (MCZ!).66 DISTRIBUTION: Palaearctic: Iberian Peninsula. immaculata (Stitz) Mantispilla bicolor var immaculata Stitz, 1913: 9, sex not indicated. Holotype: sex unknown, Vietnam (ZMB!). A subspecies of Mantispa bicolor (Stitz). DISTRIBUTION: Oriental: Vietnam.

184 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

indica Westwood Mantispa indica Westwood, 1852: 268, sex not indicated [as Indica [sic], incorrect original capitalization]. Syntypes: sex unknown, "in India orientali, Calcutta, Nepalia" (BMNH! OUM!). Mantispa torquilla Hagen, 1858b: 482. A nomen nudum.67 SUBSPECIES: ceylanica (Stitz), spilonota Banks. DISTRIBUTION: Oriental: China (Kiangsu, Shanghai), Nepal, Sri Lanka, Taiwan. iridipennis Guérin-Méneville Mantispa gracilis Rambur, 1842: 433, sex not indicated. Holotype (or syntypes): sex unknown, Colombia (ISNB). A junior primary homonym of Mantispa gracilis Erichson, 1839. Mantispa iridipennis Guérin-Méneville, 1844 ["1829-1838"]: 392, sex not indicated. Holotype (or syntypes): sex unknown, Colombia (current depository unknown). Questionable synonym of Mantispa gracilis Rambur by Westwood, 1852: 256. DISTRIBUTION: Neotropical: Colombia. januaria (Navás) Entanoneura januaria Navás, 1936b: 721, female [in Entatoneura [sic]]. Holotype (or syntypes): female, Brazil (DEI! MZBS according to Monserrat, 1985). DISTRIBUTION: Neotropical: Brazil. japonica McLachlan Mantispa japonica McLachlan, 1875a: 178, male. Holotype: male, Japan (BMNH! NMWC?68). SUBSPECIES: diminuta Matsumura. DISTRIBUTION: Palaearctic: Japan, Korea, "Far Eastern USSR". javanica Westwood Mantispa javanica Westwood, 1852: 267, sex not indicated [as Javanica [sic], incorrect original capitalization]. Holotype (or syntypes): Indonesia, Java (BMNH!69). DISTRIBUTION: Oriental: Indonesia (Java). latifrons Enderlein Mantispa (Mantispa) latifrons Enderlein, 1910: 355, male. Holotype: male, Brazil (MZPW!). DISTRIBUTION: Neotropical: Brazil, Mexico. lineaticollis Enderlein Mantispa (Mantispilla) lineaticollis Enderlein, 1910: 348, male. Holotype: male, Brazil (MZPW!). DISTRIBUTION: Neotropical: Brazil. lineolata Westwood Mantispa lineolata Westwood, 1852: 267, sex not indicated. Holotype (or syntypes): sex unknown, Nepal (BMNH!). DISTRIBUTION: Oriental: Indonesia (Java), Nepal, India.

Ohl: World Catalog of Mantispidae 185

lobata Navás Mantispa perla var. lobata Navás, 1912a ["1911"]: 535, sex not indicated. Lectotype: female, Transcaspia (ZIL). Designated by Krivokhatsky, 1995: 10. 70 DISTRIBUTION: Palaearctic: Central Asia. loveni (Navás) Mantispilla loveni Navás, 1928b: 37, sex not indicated [as Loveni [sic], incorrect original capitalization]. Syntypes: sex unknown, "British East Africa71" (NHRS). DISTRIBUTION: Afrotropical: Tanzania, Kenya, or Uganda; Guinea-Bissau. luederwaldti Enderlein Mantispa (Mantispilla) luederwaldti Enderlein, 1910: 353, male [as Lüderwaldti [sic], incorrect original capitalization and German umlaut]. Holotype: male, Brazil (MZPW!). DISTRIBUTION: Neotropical: Brazil. lurida Walker Mantispa lurida Walker, 1860: 181, sex not indicated. Holotype (or syntypes): sex unknown, no locality indicated (BMNH!). DISTRIBUTION: unknown. lutea (Stitz) Mantispilla lutea Stitz, 1913: 17, female. Holotype: female, Ethiopia (ZMB). DISTRIBUTION: Afrotropical: Ethiopia. luzonensis Navás Mantispa luzonica Navás, 1909a (April) ["1908-1909"]: 479, sex not indicated. Holotype (or syntypes): sex unknown, Philippines, Luzon (MNHN). A junior primary homonym of Mantispa quadrituberculata luzonica van der Weele, 1909 (25 January) (now in Austroclimaciella). Mantispa luzonensis Navás, 1909b: 150. Replacement name for Mantispa luzonica Navás, 1909a ["1908-1909"], a junior primary homonym of Mantispa quadrituberculata luzonica van der Weele, 1909 (now in Austroclimaciella). DISTRIBUTION: Oriental: Philippines (Luzon), Indonesia (Sumatra). maindroni Navás Mantispa maindroni Navás, 1909a ["1908-1909"]: 478, sex not indicated [as Maindroni [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, India (MNHN). DISTRIBUTION: Oriental: India. mandarina Navás Mantispa mandarina Navás, 1914l: 213, sex not indicated. Holotype: female [according to information in U. Aspöck and H. Aspöck, 1994: 107], northern China (BMNH!). 72 DISTRIBUTION: Palaearctic: northern China.

186 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

marshalli (Navás) Mantispilla marshalli Navás, 1914g: 650, sex not indicated [as Marshalli [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Zimbabwe (BMNH!). DISTRIBUTION: Afrotropical: Zimbabwe. maynei (Navás) Mantispilla maynei Navás, 1929d: 107, sex not indicated [as Maynei [sic], incorrect original capitalization]. Syntypes: sex unknown, DR Congo (MRAC).73 DISTRIBUTION: Afrotropical: DR Congo. meadewaldina (Navás) Mantispilla meadewaldina Navás, 1914g: 650, sex not indicated. Holotype (or syntypes): sex unknown, Nigeria (BMNH). DISTRIBUTION: Afrotropical: Nigeria. melanocera (Navás) Mantispilla melanocera Navás, 1913b: 502, sex not indicated. Holotype (or syntypes): sex unknown, Papua New Guinea (BMNH! CUMZ?). DISTRIBUTION: Australasian: Papua New Guinea. militaris (Navás) Mantispilla militaris Navás, 1914i: 87, sex not indicated. Holotype (or syntypes): sex unknown, Sri Lanka (BMNH! CUMZ?). DISTRIBUTION: Oriental: Sri Lanka. moesta Hagen Mantispa moesta Hagen: 1861: 210, female. Holotype (or syntypes): female, USA, Tennessee (MHNG?74). DISTRIBUTION: Nearctic: USA (Tennessee). moluccensis Banks Mantispa moluccensis Banks, 1913: 208, male. Holotype: male, Indonesia, Ambon (MCZ!). DISTRIBUTION: Oriental: Indonesia (Ambon). moucheti (Navás) Mantispilla moucheti Navás, 1925c: 572, female [as Moucheti [sic], incorrect original capitalization]. Holotype (or syntypes): female, DR Congo (MRAC). DISTRIBUTION: Afrotropical: DR Congo, South Africa, Tanzania. moulti Navás Mantispa moulti Navás, 1909a ["1908-1909"]: 481, male, female [as Moulti [sic], incorrect original capitalization]. Lectotype: male, French Guiana (MNHN). Designated by Penny, 1982a: 442. DISTRIBUTION: Neotropical: Brazil, French Guiana.

Ohl: World Catalog of Mantispidae 187

mutata Motschulsky Mantispa mutata Motschulsky , 1863: 9 [authorship incorrectly attributed to Walker]. A nomen nudum.75 nana Erichson Mantispa nana Erichson, 1839: 169, sex not indicated. Syntypes: female, Sudan (ZMB! MCZ!).76 Necyla bonhourei Navás, 1922: 397, sex not indicated [as Bonhourei [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Djibouti (MNHN). Synonymized with M. nana by Handschin, 1959a: 198. DISTRIBUTION: Afrotropical: Eritrea, Congo, Djibouti, South Africa, Sudan, Upper Volta. nanyukina Navás Mantispilla nanyukina Navás, 1933d: 216, female. Holotype (or syntypes): female, Kenya (MNHN according to original description, MZBS according to Monserrat, 1985). DISTRIBUTION: Afrotropical: Kenya. navasi Handschin Mantispilla umbripennis Navás, 1914f: 376, female. Holotype (or syntypes): female, DR Congo (MRAC). As Mantispa umbripennis a junior secondary homonym of Mantispa umbripennis Walker, 1860. Mantispa navasi Handschin, 1960a: 184. Replacement name for Mantispa umbripennis (Navás 1914f), a junior secondary homonym of Mantispa umbripennis Walker, 1860. DISTRIBUTION: Afrotropical: DR Congo. negusa Navás Mantispa negusa Navás, 1914g: 647, sex not indicated. Holotype: sex unknown, Ethiopia (BMNH!). DISTRIBUTION: Afrotropical: Ethiopia. neotropica Navás Mantispa neotropica Navás, 1933c: 309, female. Holotype (or syntypes): female, French Guiana (MNHN). DISTRIBUTION: Neotropical: French Guiana. neptunica Navás Mantispa neptunica Navás, 1914l: 212, sex not indicated. Holotype (or syntypes): sex unknown, Malaysia (BMNH!). DISTRIBUTION: Oriental: Malaysia. newmani Banks Mantispa newmani Banks, 1920: 340, sex not indicated. Holotype (or syntypes), sex unknown, Borneo (MCZ!). DISTRIBUTION: Oriental: Borneo.

188 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

nigricornis Enderlein Mantispa annulicornis var. nigricornis Enderlein, 1910: 356, female. Holotype: female, Indonesia, Sulawesi (MZPW!). A subspecies of Mantispa annulicornis Gerstaecker. DISTRIBUTION: Oriental: Indonesia (Sulawesi). nubila (Stitz) Mantispilla nubila Stitz, 1913: 15, female. Holotype: female, Cameroon (ZMB!). DISTRIBUTION: Afrotropical: Cameroon. obscurata (Navás) Mantispilla obscurata Navás, 1914i: 85, sex not indicated. Syntypes: sex unknown, Sri Lanka (BMNH! CUMZ?). DISTRIBUTION: Oriental: Sri Lanka. pallescens Stitz Mantispa pallescens Stitz, 1913: 22, female. Holotype: female, Borneo (ZMB!). DISTRIBUTION: Oriental: Borneo. paraguayana Ohl, new name Mantispilla nana Navás, 1912b: 201, sex not indicated. Holotype (or syntypes): sex unknown, Paraguay (ZSMC!). A junior secondary homonym of Mantispa nana Erichson, 1839. Mantispa paraguayana Ohl. New replacement name for Mantispilla nana Navás, 1912b (now in Mantispa), a junior secondary homonym of Mantispa nana Erichson, 1839.77 DISTRIBUTION: Neotropical: Paraguay. parvula Penny Mantispa parvula Penny, 1982a: 458, male, female. Holotype: male, Brazil (INPA). DISTRIBUTION: Neotropical: Brazil. pasteuri Navás Mantispa pasteuri Navás, 1909a ["1908-1909"]: 482, male [as Pasteuri [sic], incorrect original capitalization]. Holotype (or syntypes): male, New Guinea (MNHN). DISTRIBUTION: Australasian: New Guinea. pehlkei Enderlein Mantispa (Mantispilla) pehlkei Enderlein, 1910: 351, female, unknown sex [as Pehlkei [sic], incorrect original capitalization]. Syntypes: female, unknown sex, Colombia (probably lost78). DISTRIBUTION: Neotropical: Colombia. phaeonota Navás Mantispa phaeonota Navás, 1933d: 215, male. Holotype (or syntype): male, Kenya [probably erroneous]79 (MNHN? MZBS). DISTRIBUTION: Afrotropical: Madagascar (see annotation above).

Ohl: World Catalog of Mantispidae 189

plicicollis Handschin Mantispa plicicollis Handschin, 1935: 704, sex not indicated. Holotype: sex unknown, Indonesia, Timor (NHMB). DISTRIBUTION: Oriental: Indonesia (Timor). pygmaea (Stitz) Mantispilla pygmaea Stitz, 1913: 16, sex not indicated. Holotype: sex unknown [abdomen lacking], Tanzania (ZMB!). DISTRIBUTION: Afrotropical: Tanzania. radialis (Navás) Mantispilla radialis Navás, 1929b: 53, female. Holotype: female, China, Shanghai (CN). DISTRIBUTION: Oriental: China (Shanghai). radiata (Navás) Mantispilla radiata Navás, 1914d: 649, sex not indicated. Holotype (or syntypes): sex unknown, New Guinea (BMNH!). DISTRIBUTION: Australasia: New Guinea. rimata (Navás) Mantispilla rimata Navás, 1929e: 322, sex not indicated. Holotype (or syntypes): sex unknown, Brazil (ZMUH). DISTRIBUTION: Neotropical: Brazil. salana (Navás) Mantispilla salana Navás, 1931a ["1930"]: 89, female. Holotype: female, India, Maharashtra (CN?). DISTRIBUTION: Oriental: India (Maharashtra). scabricollis McLachlan Mantispa scabricollis McLachlan, 1875b: 13, male, female. Lectotype: male, [an Uzbekistanian enclave in Kyrgyzstan] (ZMUM). Designated by U. Aspöck and H. Aspöck, 1994: 103. Mantispilla minuta Kozhanchikov, 1949: 357, male, female. Lectotype: male: Tajikistan (ZIL). Designated by Krivokhatsky, 1995: 10. Synonymized with Mantispa scabricollis by Zakharenko, 1987: 623. [Genus not indicated] caucasica of Handschin, 1959: 190. An unavailable name80 DISTRIBUTION: Palaearctic: Eastern Greece, Turkey, Caucasus area, Iraq, Iran, Afghanistan, Tajikistan, Turkmenistan. schoutedeni (Navás) Mantispilla schoutedeni Navás, 1929d: 106, sex not indicated [as Schoutedeni [sic], incorrect original capitalization]. Holotype (or syntypes): sex not indicated, DR Congo (MRAC). DISTRIBUTION: Afrotropical: DR Congo.

190 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

simplex Stitz Mantispa simplex Stitz, 1913: 23, male. Holotype: male, Indonesia, Sulawesi (ZMB!). DISTRIBUTION: Oriental: Indonesia (Sulawesi). spilonota Banks Mantispa indica var. spilonota Banks, 1913: 209, male. Holotype: male, Sri Lanka (MCZ!). A subspecies of Mantispa indica Westwood. DISTRIBUTION: Oriental: Sri Lanka, Taiwan. stenoptera Gerstaecker Mantispa stenoptera Gerstaecker, 1888 ["1887"]: 115, female. Holotype (or syntypes): female, Papua New Guinea (EMAU!). A subspecies of Mantispa annulicornis Gerstaecker according to van der Weele (1909: 91). DISTRIBUTION: Australasian: Papua New Guinea. styriaca (Poda) Raphidia styriaca Poda, 1761: 101, sex not indicated [as Styriaca [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Austria (depository unknown81). Raphidia mantispa Scopoli, 1763: 272, sex not indicated [as Raphidia Mantispa [sic]; incorrect original capitalization]. Holotype (or syntypes): sex unknown, "Carniolia". Synonymized with Mantis pagana Fabricius (now in Mantispa) by Olivier, 1792: 631.82 Mantis pagana Fabricius, 1775: 278, sex not indicated. Holotype (or syntypes): sex unknown, France (depository unknown83). Synonymized with Mantispa styriaca by H. Aspöck et al., 1980: 183. Mantispa hauseri Poivre, 1982b: 11, female. Holotype: female, Croatia (MHNG). Synonymized by U. Aspöck and H. Aspöck, 1994: 108. Mantispa kononenkoi Makarkin, 1985: 620, male, female. Holotype: male, Russia (ZIL). Synonymized with Mantispa styriaca by Makarkin, 1990: 38. DISTRIBUTION: Palaearctic: widespread in Eurasia south of 50°N, Morocco. subcostalis Navás Mantispa subcostalis Navás, 1929a ["1928"]: 115, female. Holotype: female, Brazil (DEI!). DISTRIBUTION: Neotropical: Brazil. taina (Alayo) Mantispilla taina Alayo, 1968: 13, sex not indicated. Syntypes: sex unknown, Cuba (CZC). DISTRIBUTION: Neotropical: Cuba. tenella Erichson Mantispa tenella Erichson, 1839: 169, sex not indicated. Syntypes: sex unknown, South Africa (ZMB!). Mantispilla tenera Navás, 1914j: 40, sex not indicated. Holotype (or syntypes): sex unknown, Kenya (MNHN). Synonymized with Mantispa tenella by Handschin, 1959a: 203. Mantispa axillata Navás, 1936c: 354: sex not indicated. Holotype (or syntypes): sex unknown, DR Congo (MRAC). Synonymized with Mantispa tenella by Handschin, 1959a: 203. Ohl: World Catalog of Mantispidae 191

SUBSPECIES: dispersa (Navás). DISTRIBUTION: Afrotropical: DR Congo, Ivory Coast, Kenya, Nigeria, "Italian Somaliland " (Somalia, Djibouti, or Ethiopia), South Africa, Tanzania. tessmanni (Stitz) Mantispilla tessmanni Stitz, 1913: 13, male. Holotype: male, Equatorial Guinea (ZMB!84). DISTRIBUTION: Afrotropical: DR Congo, Equatorial Guinea. tonkinensis Navás Mantispa tonkinensis Navás, 1930c: 11, male. Holotype (or syntypes): male, Vietnam (MNHN). DISTRIBUTION: Oriental: Vietnam. transversa (Stitz) Mantispilla transversa Stitz, 1913: 5, female. Holotype: female, Taiwan (ZMB!). DISTRIBUTION: Oriental Japan (Ryukyu Islands), Taiwan. umbripennis Walker Mantispa umbripennis Walker, 1860: 181, sex not indicated. Holotype (or syntypes): sex unknown, South Africa (BMNH!). DISTRIBUTION: Afrotropical: South Africa, Tanzania. usambarica (Fraser) Mantispilla usambarica Fraser, 1955: 21 [authorship incorrectly attributed to Westwood]. A nomen nudum.85 variolosa Navás Mantispa variolosa Navás, 1914d: 648, sex not indicated. Holotype (or syntypes): sex unknown "Oceanía"(BMNH!). 86 DISTRIBUTION: ? Australasia: no specific locality. venulosa (Navás) Mantispilla venulosa Navás, 1914i: 85, sex not indicated. Holotype (or syntypes): sex not indicated, Central America87 (BMNH!). DISTRIBUTION: Neotropical: Central America. verruculata (Navás) Mantispilla verruculata Navás, 1914j: 41, sex not indicated. Holotype (or syntypes): sex unknown, "British East Africa"88 (MNHN). 89 DISTRIBUTION: Afrotropical : DR Congo, Kenya, South Africa, Uganda. zayasi (Alayo) Mantispilla zayasi Alayo, 1968: 13, sex not indicated. Syntypes: sex unknown, Cuba (CZC). DISTRIBUTION: Neotropical: Cuba. zonaria Navás Mantispa zonaria Navás, 1925c: 571, male. Holotype: male, DR Congo (BMNH! MRAC). DISTRIBUTION: Afrotropical: Cameroon, DR Kongo, Ivory Coast, Tanzania. 192 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

zonata Navás Mantispa zonata Navás, 1923b: 21, sex not indicated. Holotype (or syntypes): sex unknown, Gaboon (MNHN). DISTRIBUTION: Afrotropical: Gaboon.

Genus Mimetispa Handschin

MIMETISPA Handschin, 1961: 276. Type species: Mantispa simulatrix McLachlan, 1900: 127, by original designation. DISTRIBUTION: Oriental: Borneo, Indonesia. simulatrix (McLachlan) Mantispa simulatrix McLachlan, 1900: 127, female. Holotype: female, Borneo (OUM!90). Mantispa mimetica Sharp, 1901 ["1900"]: 338 [authorship incorrectly attributed to McLachlan, 1900]. A nomen nudum.91 Euclimacia ferdinandi Navás, 1928c: 137, male [as Ferdinandi [sic], incorrect original capitalization]. Holotype (or syntypes): male, Borneo (ZMUH). Synonymized with Mimetispa simulatrix by Handschin, 1961: 277.92 DISTRIBUTION: Oriental: Borneo, Indonesia (Java).

Genus Nampista Navás

NAMPISTA Navás, 1914i: 97. Type species: Nampista speciosa Navás, 1914i: 98 (= Mantispa auriventris Guerin-Meneville), by monotypy. Forciada Kozhanchikov, 1949: 355. Type species: Forciada relicta Kozhanchikov, 1949: 356 (= Mantispa auriventris Guerin-Meneville, 1838: 202), by monotypy. Synonymized with Nampista by H. Aspöck et al., 1980: 185. Bucharispa; Martynov , 1936: 437. An unavailable name93. DISTRIBUTION: Palaearctic: Greece to Tajikistan. auriventris (Guérin-Méneville) Mantispa auriventris Guérin-Méneville, 1838: 202, sex not indicated. Holotype (or syntypes): sex unknown, Egypt (depository unknown94). Mantispa apicalis Loew, 1843: 433, sex not indicated. Holotype (or syntype): sex unknown, Greece (MCZ!). Synonymized with Mantispa auriventris by Hagen, 1859: 405. Nampista speciosa Navás, 1914i: 98, sex not indicated. Holotype (or syntypes): sex unknown, Greece (BMNH!). Synonymized with Euclimacia auriventris by Esben-Petersen, 1917: 12. Forciada relicta Kozhanchikov, 1949: 356, male, female. Lectotype: male, Tadzhikistan (ZIL). Designated by Krivokhatsky, 1995: 11. Synonymized with Nampista auriventris by Handschin, 1960c: 155. DISTRIBUTION: Palaearctic: Greece, Egypt, Oman, Uzbekistan, Tajikistan.

Ohl: World Catalog of Mantispidae 193

Genus Necyla Navás

NECYLA Navás, 1913a: 280 [incorrect original spelling as Nicyla]95. Type species: Necyla exigua Navás, 1913a: 281 [as Nicyla [sic] exigua], by monotypy. DISTRIBUTION: Palaearctic and Oriental: Saudi Arabia to Japan, New Guinea. Afrotropical: Uganda. arabica Navás Necyla arabica Navás, 1914l: 214, sex not indicated. Holotype (or syntypes): sex unknown, Saudi Arabia (BMNH!). DISTRIBUTION: Palaearctic: Saudi Arabia. chiaiensis Ohl, new name Mantispa formosana var. bella Kuwayama, 1925: 256, female. Holotype: female, Taiwan (ITLJ?96). A junior primary homonym of Mantispa (Trichoscelia) bella Westwood, 1867 (now in Anchieta). A subspecies of Necyla formosana (Okamoto). Necyla formosana chiaiensis Ohl. New replacement name for Mantispa formosana bella Kuwayama, 1925 (now in Necyla), a junior primary homonym of Mantispa (Trichoscelia) bella Westwood, 1867 (now in Anchieta). 97 DISTRIBUTION: Palaearctic: Japan. Oriental: Taiwan. exigua Navás Necyla exigua Navás, 1913a: 281 [as Nicyla [sic] exigua], sex not indicated. Holotype (or syntypes): sex unknown, Sri Lanka (ZSMC!). DISTRIBUTION: Oriental: Sri Lanka. extrema Navás Necyla extrema Navás, 1914c: 481 male. Holotype: male, New Guinea (BMNH). DISTRIBUTION: Australasian: New Guinea. flavonotata Tjeder Necyla flavonotata Tjeder, 1963 ["1962"]: 122, male, female. Holotype: male, Uganda (MZLU). DISTRIBUTION: Afrotropical: Uganda. formosana (Okamoto) Mantispa (Mantispilla) formosana Okamoto, 1910: 537, female, male [authorship incorrectly attributed to Matsumura]. Lectotype: male, Formosa, Tainan (EIHU). Designated by Kuwayama, 1966: 137. Incorrectly cited as a new species in Okamoto, 1911: 297, which is a German translation of Okamoto, 1910. Mantispilla formosana var. major Stitz, 1913: 7, sex not indicated. Syntypes: male, female, Taiwan (ZMB!).98 Synonymized with Mantispa formosana by Kuwayama, 1925: 255. SUBSPECIES: bella (Kuwayama), minor (Stitz), sumatrana (Stitz). DISTRIBUTION: Oriental: Indonesia (Sulawesi), Taiwan. jucunda Navás Necyla jucunda Navás, 1914i: 87 male. Holotype (or syntypes): male, Sri Lanka (BMNH! CUMZ?). DISTRIBUTION: Oriental: Sri Lanka. 194 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

leopoldi Navás Necyla leopoldi Navás, 1931d: 9, sex not indicated. Holotype: sex unknown, New Guinea (ISNB). DISTRIBUTION: Australasian: New Guinea. minor (Stitz), new combination Mantispilla formosana var. minor Stitz, 1913: 6, male, female. Syntypes: Taiwan (ZMB!). A subspecies of Necyla formosana (Okamoto).99 DISTRIBUTION: Palaearctic: Japan. Oriental: Taiwan. natalensis Navás Necyla natalensis Navás, 1914a: 64, male. Holotype (or syntypes): male, South Africa (OUM!). DISTRIBUTION: Afrotropical: South Africa. orientalis (Esben-Petersen) Mantispa orientalis Esben-Petersen, 1913: 261, male. Holotype: male, Japan (DEI!). Mantispilla nigra Stitz, 1913: 3, female. Holotype: female, Taiwan (ZMB!). Synonymized with Mantispa orientalis by Esben-Petersen, 1917: 13.100 DISTRIBUTION: Palaearctic: Japan. Oriental: Taiwan. pupa Navás Necyla pupa Navás, 1927a: 88, female. Holotype (or syntypes): female, Somalia (MCSN101). DISTRIBUTION: Afrotropical: Somalia. sacra Navás Necyla sacra Navás, 1914a: 63, male. Holotype (or syntypes): male, Israel (OUM!). DISTRIBUTION: Palaearctic: Israel. sumatrana (Stitz), new combination Mantispilla formosana var. sumatrana Stitz, 1913: 7, female. Syntypes: female, Indonesia, Sumatra (ZMB!). A subspecies of Necyla formosana (Okamoto).102 DISTRIBUTION: Oriental: Indonesia (Sumatra). trilineata Navás Necyla trilineata Navás, 1929b: 54, sex not indicated. Holotype, sex unknown, Java (ZMUH). DISTRIBUTION: Oriental: Indonesia (Java).

Genus Orientispa Poivre

ORIENTISPA Poivre, 1984a: 27. Type species: Cercomantispa shirozui Nakahara, 1961: 63 (as "Orientispa shirozui (Nakahara, 1961)"), by original designation. DISTRIBUTION: Palaearctic and Oriental: Sri Lanka to Japan, Indonesia.

Ohl: World Catalog of Mantispidae 195

bicolor Poivre Orientispa bicolor Poivre, 1984a: 28, male. Holotype: male, Sri Lanka (ISNB). DISTRIBUTION: Oriental: Sri Lanka. coronata C.-k. Yang Orientispa coronata C.-k. Yang, 1999: 138, female. Holotype: female, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian). flavacoxa C.-k. Yang Orientispa flavacoxa C.-k. Yang, 1999: 139, male. Holotype: male, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian). fujiana C.-k. Yang Orientispa fujiana C.-k. Yang, 1999: 135, male. Holotype: male, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian). longyana C.-k. Yang Orientispa longyana C.-k. Yang, 1999: 136, male. Holotype: male, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian). nigricoxa C.-k. Yang Orientispa nigricoxa C.-k. Yang, 1999: 139, female. Holotype: female, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian). ophryuta C.-k. Yang Orientispa ophryuta C.-k. Yang, 1999: 138, male. Holotype: male, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian). pusilla C.-k. Yang Orientispa pusilla C.-k. Yang, 1999: 136, male. Holotype: male, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian). semifurva C.-k. Yang Orientispa semifurva C.-k. Yang, 1999: 137, male. Holotype: male, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian). shirozui (Nakahara) Cercomantispa shirozui Nakahara, 1961: 63, male, female. Holotype: male, Japan (KUZC). DISTRIBUTION: Oriental: Indonesia, Sri Lanka. Palaearctic: Japan. xuthoraca C.-k. Yang Orientispa xuthoraca C.-k. Yang, 1999: 135, female. Holotype: female, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian).

196 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Genus Paramantispa Williner and Kormilev

PARAMANTISPA Williner and Kormilev, 1958: 10. Type species: Mantispa decorata Erichson, 1839: 163 [= Mantispa ambusta Erichson, 1839: 162], by original designation. Neoclimaciella Handschin, 1960a: 208, 243. A nomen nudum. Synonymized with Paramantispa by Handschin, 1960c: 537.103 DISTRIBUTION: Neotropical: Argentina to Brazil. ambusta (Erichson) Mantispa ambusta Erichson, 1839: 162, female. Holotype: female, Uruguay (ZMB!). Mantispa decorata Erichson, 1839: 163, sex not indicated. Syntypes: male, Brazil (ZMB! HNHM!104). Synonymized with Paramantispa ambusta by Williner and Mariluis, 1979 ["1978"]: 40. DISTRIBUTION: Neotropical: Argentina, Bolivia, Paraguay, Uruguay, Brazil. prolixa (Erichson) Mantispa prolixa Erichson, 1839: 163, female. Syntypes: female, Brazil (ZMB! MCZ!). Mantispa muhni Navás, 1930e: 68, female [as Muhni [sic], incorrect original capitalization]. Holotype (or syntypes): female, Argentina (SMBA105). Synonymized with Paramantispa prolixa by Williner and Kormilev, 1958: 7. DISTRIBUTION: Neotropical: Argentina, Brazil. wagneri (Navás) Mantispa wagneri Navás, 1909a ["1908-1909"]: 481, female [as Wagneri [sic], incorrect original capitalization]. Syntypes: female, Argentina (MNHN). DISTRIBUTION: Neotropical: Argentina, Brazil, Paraguay.

Genus Paulianella Handschin

PAULIANELLA Handschin, 1960a: 210. Type species: Climaciella necopina Navás, 1936a ["1935"]: 102, by original designation. DISTRIBUTION: Afrotropical: Madagascar. necopina (Navás) Climaciella necopina Navás, 1936a ["1935"]: 102, female. Holotype (or syntypes): female, Madagascar (MNHN). DISTRIBUTION: Afrotropical: Madagascar. olsufievi (Navás) Climaciella olsufievi Navás, 1936a ["1935"]: 104, female [as Olsufievi [sic], incorrect original capitalization]. Holotype (or syntypes): Madagascar (MNHN?).106 DISTRIBUTION: Afrotropical: Madagascar.

Ohl: World Catalog of Mantispidae 197

Genus Perlamantispa Handschin

PERLAMANTISPA Handschin, 1960a: 191. Type species: Mantis perla Pallas, 1772: 14 (as "Mantispa perla"), by original designation.107 DISTRIBUTION: Palaearctic: southern Europe to central Asia. Afrotropical: widespread. austroafrica Poivre Perlamantispa austroafrica Poivre, 1984b: 642, female. Holotype: female, "Afrique méridionale", no specific locality (MHNG108). DISTRIBUTION: Afrotropical: "Afrique méridionale". bequaerti (Navás) Mantispilla bequaerti Navás, 1932b: 279, female [as Bequaerti [sic], incorrect original capitalization]. Holotype (or syntypes): female, DR Congo (MRAC). Mantispilla bequaerti var. decolor Navás, 1932b: 280, sex not indicated. Syntypes: sex unknown, DR Congo (MZBS, MRAC). Synonymized with Perlamantispa bequaerti (Navás) by Handschin, 1960a: 197. Mantispilla kibumbana Navás, 1936c: 355, sex not indicated. Holotype (or syntypes): sex unknown, DR Congo (MRAC). Synonymized with Perlamantispa bequaerti (Navás) by Handschin, 1960a: 197. DISTRIBUTION: Afrotropical: DR Congo, South Afrika, Tanzania. dorsalis (Erichson) Mantispa dorsalis Erichson, 1839: 168, female. Syntypes: female, South Africa (ZMB! MCZ!). Mantispilla hemichroa Navás, 1931c: 129, female. Holotype (or syntypes): female, DR Congo (MRAC). Incorrectly considered a junior primary homonym of Mantispa hemichroa Navás, 1914c by Navás, 1932b: 279, and unnecessarily replaced by Mantispilla hypophoea Navás, 1932b. Mantispilla hypophoea Navás, 1932b: 279. An unnecessary replacement name for Mantispilla hemichroa Navás, 1931c, which is incorrectly considered a junior homonym of Mantispa hemichroa Navás, 1914c by Navás, 1932b: 279. Synonymized with Perlamantispa dorsalis by Handschin, 1960a: 196. DISTRIBUTION: Afrotropical: DR Congo, South Afrika. girardi Poivre Perlamantispa girardi Poivre, 1982a ["1981"]: 194, male. Holotype: male, Ivory Coast (IFAN?109). DISTRIBUTION: Afrotropical: Cameroon, Ivory Coast. perla (Pallas) Mantis perla Pallas, 1772: 14, sex not indicated. Holotype (or syntypes): sex unknown, not indicated (SMFD?).110 Mantispa christiana Charpentier, 1825: 93, sex not indicated. Syntypes: male, female, "in Russia europaea" (ZMB!111). Synonymized with Mantispa perla by Erichson, 1839: 167. Mantispa flaveola Erichson, 1839: 168, sex not indicated. Holotype: female [Brazil, Para; obviously mislabeled] (ZMB!). New synonym112. 198 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Mantispa victorii Guérin-Méneville, 1844 ["1829-1838"]: 391, sex not indicated [as Victorii [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Caucasus (MCZ!113). Synonymized with Mantispa perla by Hagen, 1858a: 128. Mantispa perla var. brunnea Navás, 1906: 102, sex not indicated. Holotype: sex not indicated, Italy (CN?). New synonym.114 DISTRIBUTION: Palaearctic: Southern Europe, Turkey, Caucasus area, Kazakhstan, Turkmenistan. pusilla (Pallas) Mantis pusilla Pallas, 1772: 15, sex not indicated. Holotype (or syntypes): sex unknown, South Africa (depository unknown115). Mantis brevicornis De Geer, 1778: 620, pl. 46, fig. 9-10, sex not indicated. Holotype (or syntypes): sex unknown, locality not indicated (NHRS?116). Synonymized with Mantispa pusilla by Burmeister, 1839: 967. DISTRIBUTION: Afrotropical: Uganda, South Africa. similata (Navás) Mantispilla similata Navás, 1922: 396, female. Holotype (or syntypes): female, South Africa (MNHN). DISTRIBUTION: Afrotropical: South Africa. royi Poivre Perlamantispa royi Poivre, 1982a ["1981"]: 191, male (paratype sex not indicated). Holotype: male, Ivory Coast (IFAN109). DISTRIBUTION: Afrotropical: Ivory Coast. tincta (Navás) Mantispilla tincta Navás, 1929d: 107, sex not indicated. Holotype (or syntype): sex not indicated, DR Congo (MRAC). DISTRIBUTION: Afrotropical: DR Congo. vassei (Navás) Mantispa vassei Navás, 1909a ["1908-1909"]: 474, male [as Vassei [sic], incorrect original capitalization]. Holotype: male, Mozambique (MNHN). Mantispa (Mantispilla) lineatifrons Enderlein, 1910: 346, female. Holotype: female, Eritrea (MZPW!). Synonymized with Perlamantispa vassei by Handschin, 1960a: 193. Mantispilla sankitana Navás, 1922: 395, female. Holotype (or syntype): female, Gaboon (MNHN). Synonymized with Perlamantispa vassei by Handschin, 1960a: 193. Mantispilla burgeoni Navás, 1923a: 77, sex not indicated [as Burgeoni [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, DR Congo (MNHN). Probable synonym of Perlamantispa vassei according to Handschin, 1960a: 193. DISTRIBUTION: Afrotropical: DR Congo, Gaboon, Ivory Coast, Mozambique, South Africa.

Ohl: World Catalog of Mantispidae 199

Genus Pseudoclimaciella Handschin

PSEUDOCLIMACIELLA Handschin, 1960a: 207, 210. Type species: Mantispa erichsonii Guérin-Méneville, 1844 ["1829-1838"]: 391 (as "Pseudoclimaciella erichsoni [sic] (Guer. 1844)" on page 207, and as "Climaciella erichsoni [sic] Guer. 1844" on page 210), by original designation. Mantispa erichsonii is a replacement name for Mantispa grandis Erichson, 1839: 164, which is a junior primary homonym of Mantispa grandis Guérin-Méneville, 1831 ["1830"]: 196. DISTRIBUTION: Afrotropical: widespread. africana (Esben-Petersen), new combination Euclimacia africana Esben-Petersen, 1927: 17, sex not indicated. Holotype: sex not indicated, Sudan (FMNH). SUBSPECIES: ragazziana Navás. DISTRIBUTION: Afrotropical: Sudan. alberti (Navás) Climaciella alberti Navás, 1936c: 360, sex not indicated [as Alberti [sic], incorrect original capitalization].. Holotype: sex unknown, DR Congo (MRAC according to original description, MZBS according to Monserrat, 1985: 241). DISTRIBUTION: Afrotropical: DR Congo. apicipennis (Kolbe) Mantispa apicipennis Kolbe, 1897: 36, sex not indicated. Holotype: female, Kenya (ZMB!). Climaciella ornata Stitz, 1913: 35, female. Holotype: female, Togo (ZMB!). Synonymized with Pseudoclimaciella apicipennis by Handschin, 1960a: 225. DISTRIBUTION: Afrotropical: Angola, Congo, DR Congo, Kenya, Ruanda, Togo, Tanzania. cachani Poivre Pseudoclimaciella cachani Poivre, 1982b: 3, female [as cahani [sic], incorrect original spelling117]. Holotype: female, Ivory Coast (MNHN). DISTRIBUTION: Afrotropical: Ivory Coast. congensis (Navás) Entanoneura congensis Navás, 1936c: 359, sex not indicated [as Entatoneura [sic] congensis]. Holotype (or syntypes): sex unknown, DR Congo (MRAC). DISTRIBUTION: Afrotropical: DR Congo. coronata (Stitz) Mantispa tropica var. coronata Stitz, 1913: 29, female. Holotype: female, Cameroon (ZMB!). Mantispa fulvicornis Navás, 1929d: 105, sex not indicated. Holotype (or syntypes): sex unknown, DR Congo (MRAC). Synonymized with Pseudoclimaciella coronata by Handschin, 1960a: 230. DISTRIBUTION: Afrotropical: Cameroon, DR Congo, Uganda.

200 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

elisabethae (Navás) Climaciella elisabethae Navás, 1936c: 360, sex not indicated [as Elisabethae [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, DR Congo (MRAC). Entanoneura machadoi Handschin and Markl, 1955: 71, sex not indicated. Holotype: sex unknown, Angola (NHMB). Synonymized with Pseudoclimaciella elisabethae by Handschin, 1960a: 219. DISTRIBUTION: Afrotropical: Angola, DR Congo, Uganda. erichsonii (Guérin-Méneville) Mantispa grandis Erichson, 1839: 164, female. Holotype: female, South Africa (ZMB!). A junior primary homonym of Mantispa grandis Guérin-Méneville in Duperrey, 1831 [1830]. Mantispa erichsonii Guérin-Méneville, 1844 ["1829-1838"]: 391 [as Erichsonii [sic], incorrect original capitalization]. Replacement name for Mantispa grandis Erichson, 1839, a junior primary homonym of Mantispa grandis Guérin-Méneville in Duperrey, 1831 [1830]. DISTRIBUTION: Afrotropical: Mozambique, South Africa. flava (Esben-Petersen), new combination Euclimacia flava Esben-Petersen, 1917: 9, female. Holotype: female, South Africa (ZMUC). DISTRIBUTION: Afrotropical: South Africa. ivoiriensis Poivre Pseudoclimaciella ivoiriensis Poivre, 1982b: 5, male, female. Holotype: male, Ivory Coast (MNHN). DISTRIBUTION: Afrotropical: Ivory Coast. loanga (Navás) Mantispa loanga Navás, 1909a ["1908-1909"]: 475, sex not indicated. Holotype (or syntypes): sex unknown, DR Congo (MZBS). DISTRIBUTION: Afrotropical: Angola, Cameroon, DR Congo, Ivory Coast, Kenya, South Africa, Tanzania. ragazziana (Navás), new combination Euclimacia africana var. ragazziana Navás, 1929c: 385, sex not indicated. Holotype (or syntypes): sex not indicated, Eritrea (MCSN). A subspecies of Pseudoclimaciella africana (Esben-Petersen). DISTRIBUTION: Afrotropical: Eritrea. rubida (Stitz) Climaciella grandis var. rubida Stitz, 1913: 35, male. Holotype: male, Tanzania (ZMB!). Climaciella occidentalis Paulian, 1957: 77, female. Holotype: female, Madagascar (MNHN). Synonymized with Pseudoclimaciella rubida by Handschin, 1960a: 220. DISTRIBUTION: Afrotropical: Madagascar, Tanzania.

Ohl: World Catalog of Mantispidae 201

sanguinea (Navás), new combination Climaciella sanguinea Navás, 1914g: 652, sex not indicated. Holotype (or syntypes): sex unknown, Sierra Leone (BMNH!). DISTRIBUTION: Afrotropical: Sierra Leone. sarta (Stitz) Climaciella grandis var. sarta Stitz, 1913: 35, female. Holotype: female, Cameroon (ZMB!). DISTRIBUTION: Afrotropical: Cameroon, Madgascar, "West Africa". stitzi Handschin Pseudoclimaciella stitzi Handschin, 1960a: 218, female. Holotype: female, Tanzania (ZMB!). DISTRIBUTION: Afrotropical: Tanzania. timmerhansi (Navás) Climaciella timmerhansi Navás, 1931c: 130, sex not indicated [as Timmerhansi [sic], incorrect original capitalization]. Holotype: male118, DR Congo (MRAC). DISTRIBUTION: Afrotropical: DR Congo. thomensis (Viette), new combination Mantispa thomensis Viette, 1958: 216, female. Holotype: female, São Tomé (ZMB!). DISTRIBUTION: Afrotropical: São Tomé. trivenata (Stitz) Mantispa tropica var. trivenata Stitz, 1913: 29, female. Holotype: female, Malawi (ZMB!). A subspecies of Pseudoclimaciella tropica (Westwood). DISTRIBUTION: Afrotropical: Malawi. tropica (Westwood) Mantispa tropica Westwood, 1852: 265, sex not indicated. Holotype (or syntypes): sex unknown, Gambia (BMNH?; OUM!). SUBSPECIES: trivenata (Stitz). DISTRIBUTION: Afrotropical: Cameroon, DR Congo, Gambia, Ivory Coast, Madagascar, Nigeria, Senegal, South Afrika, Upper Volta, Zimbabwe. umbripennis (Navás), new combination Climaciella umbripennis Navás, 1914g: 651, sex not indicated. Holotype (or syntypes): sex not indicated, Sierra Leone (BMNH!). Climaciella umbra of Paulian, 1957: 256 [authorship incorrectly attributed to Navás, 1936a (1935)]. An incorrect subsequent spelling of Climaciella umbripennis Navás, 1914g.119 DISTRIBUTION: Afrotropical: Sierra Leone.

202 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Genus Rectinerva Handschin

RECTINERVA Handschin, 1959a: 221. Type species: Rectinerva braconidiformis Handschin, 1959a: 221, by original designation. DISTRIBUTION: Afrotropical: DR Congo. braconidiformis Handschin Rectinerva braconidiformis Handschin, 1959a: 221, female. Holotype: female, DR Congo (MRAC). DISTRIBUTION: Afrotropical: DR Congo.

Genus Sagittalata Handschin

SAGITTALATA Handschin, 1959a: 215. Type species: Mantispilla hilaris Navás, 1925c: 573 (as "Sagittalata hilaris (Navás 1924 [sic])"), by original designation. DISTRIBUTION: Palaearctic and Oriental: China. Afrotropical: Ivory Coast to Uganda. asiatica C.-k. Yang Sagittalata asiatica C.-k. Yang, 1999: 134, male. Holotype: male, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian). ata C.-k. Yang Sagittalata ata C.-k. Yang, 1999: 134, female. Holotype: female, China, Fujian (IZAS). DISTRIBUTION: Oriental: China (Fujian). bitschi Poivre Sagittalata bitschi Poivre, 1982a ["1981"]: 181, male, female. Holotype: male, Ivory Coast (MHNG). DISTRIBUTION: Afrotropical: Ivory Coast. delamarei Poivre Sagittalata delamarei Poivre, 1982a ["1981"]: 175, male, female. Holotype: male, Ivory Coast (MNHN). Incorrectly cited as new species by Poivre, 1982b: 10. DISTRIBUTION: Afrotropical: Ivory Coast. griveaudi Poivre Sagittalata griveaudi Poivre, 1982a ["1981"]: 179, female. Holotype: female, Ivory Coast (MNHN). Incorrectly cited as new species by Poivre, 1982b: 10. DISTRIBUTION: Afrotropical: Ivory Coast. hilaris (Navás) Mantispilla hilaris Navás, 1925c: 573, female. Holotype (or syntypes): female, DR Congo (MRAC). DISTRIBUTION: Afrotropical: DR Congo, Ivory Coast, Uganda.

Ohl: World Catalog of Mantispidae 203

jucunda (Navás) Mantispilla jucunda Navás, 1932b: 280, female (male, according to Handschin, 1960a: 186). Holotype (or syntypes): male, DR Congo (MRAC). DISTRIBUTION: Afrotropical: Cameroon, DR Congo. lugubris (Navás) Mantispilla lugubris Navás, 1926a: 459, sex not indicated. Holotype (or syntypes): sex unknown, DR Congo (MRAC). DISTRIBUTION: Afrotropical: Cameroon, DR Congo. semeriai Poivre Sagittalata semeriai Poivre, 1981 ["1980"]: 78, female. Holotype: female, Cameroon (MHNG). DISTRIBUTION: Afrotropical: Cameroon. yuata C.-k. Yang and Peng Sagittalata yuata C.-k. Yang and Peng, 1998: 62, female. Holotype: female, China, Henan (IZAS). DISTRIBUTION: Palaearctic: China (Henan).

Genus Spaminta Lambkin

SPAMINTA Lambkin, 1986b: 48. Type species: Spaminta minjerribae Lambkin, 1986b: 50, by original designation. DISTRIBUTION: Australasian: New Guinea, Australia. minjerribae Lambkin Spaminta minjerribae Lambkin, 1986b: 50, male, female. Holotype: male, Australia, Queensland (QM). DISTRIBUTION: Australasian: Australia (New South Wales, Northern Territory, Queensland, Victoria). pavida (Gerstaecker) Mantispa pavida Gerstaecker, 1885a ["1884"]: 41, female. Holotype: female, Australia, Queensland (EMAU!). Mantispa verticalis Banks, 1920: 339, sex not indicated. Holotype (or syntypes): female [according to New, 1996: 32], Australia, Northern Territory (ANIC ex MCZ!). Synonymized with Spaminta pavida by Lambkin, 1986b: 52. Mantispa strigipes var. rufipes Tillyard, 1925 ["1925-1928"]: 43, sex not indicated. Holotype: male, Australia, Northern Territory (SAM). Synonymized with Spaminta pavida by Lambkin, 1986b: 52. DISTRIBUTION: Australasian: Australia (New South Wales, Northern Territory, Queensland, Western Australia).

204 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

scutellaris (Westwood) Mantispa scutellaris Westwood, 1852: 260, female. Holotype: female, "WWS" [= "Nova Hollandia" according to Westwood, 1852] (BMNH!). A nomen dubium according to Lambkin, 1986b: 56120. DISTRIBUTION: ? Australasia: Australia, New Guinea. strigipes (Westwood) Mantispa strigipes Westwood, 1852: 259, female. Holotype: female, Australia, South Australia (BMNH!). A nomen dubium according to Lambkin, 1986b: 55121. DISTRIBUTION: ? Australasia: Australia, New Guinea.

Genus Stenomantispa Stitz

STENOMANTISPA Stitz, 1913: 47. Type species: Mantispa (Stenomantispa) ilsae Stitz, 1913: 48, by monotypy.122 DISTRIBUTION: Australasian: Papua New Guinea. ilsae (Stitz) Mantispa (Stenomantispa) ilsae Stitz, 1913: 48, female. Holotype: female, Papua New Guinea (ZMB!). DISTRIBUTION: Australasian: Papua New Guinea. reinhardi (Stitz) Mantispa reinhardi Stitz, 1913: 24, female. Holotype: female, Papua New Guinea (ZMB!). DISTRIBUTION: Australasian: Papua New Guinea.

Genus Toolida Lambkin

TOOLIDA Lambkin, 1986b: 36. Type species: Toolida infrequens Lambkin, 1986b: 38, by original designation. DISTRIBUTION: Australasian: Australia. infrequens Lambkin Toolida infrequens Lambkin, 1986b: 38, male, female. Holotype: male, Australia, New South Wales (AM). DISTRIBUTION: Australasian: Australia (Australian Capital Territory, New South Wales, Victoria).

Genus Tuberonotha Handschin

TUBERONOTHA Handschin, 1961: 282. Type species: Mantispa strenua Gerstaecker, 1888 ["1887"]: 150, by original designation. DISTRIBUTION: Palaearctic, Oriental, and Australasia: Japan, India to Australia.

Ohl: World Catalog of Mantispidae 205

bouchardi (Navás) Mantispa bouchardi Navás, 1909a ["1908-1909"]: 478, sex not indicated [as Bouchardi [sic], incorrect original capitalization]. Syntypes: sex unknown, Indonesia, Sumatra (MNHN, MZBS). DISTRIBUTION: Oriental: Indonesia (Sumatra). campioni (Navás) Climaciella campioni Navás, 1914i: 91, sex not indicated [as Campioni [sic], incorrect original capitalization]. Holotype (or syntypes): sex unknown, Singapore (BMNH!). SUBSPECIES: insignis (Navás). DISTRIBUTION: Oriental: India?, Singapore. ferrosa (Navás), new combination Climaciella ferrosa Navás, 1914c: 482, sex not indicated. Holotype (or syntypes): sex unknown, Malaysia (BMNH!). DISTRIBUTION: Oriental: Malaysia. insignis (Navás) Climaciella campioni var. insignis Navás, 1914i: 92, sex not indicated. Holotype (or syntypes): sex unknown, Thailand (BMNH!). A subspecies of Tuberonotha campioni (Navás). DISTRIBUTION: Oriental: Thailand. regia (Navás) Climaciella regia Navás, 1930b: 425, female (indicated in illustration), male (indicated in measurements). Holotype (or syntypes): female, male, Borneo (ZMUH). DISTRIBUTION: Oriental: India, Indonesia (Sumatra), Malaysia, Borneo. strenua (Gerstaecker) Mantispa strenua Gerstaecker 1888 ["1887"]: 150, female. Holotype (or syntypes): female, Indonesia, Java (EMAU!). Mantispa magna Miyake, 1910: 214, female. Syntypes: female, Japan (ACIU, NELG). Synonymized with Tuberonotha strenua by Handschin, 1961: 284. DISTRIBUTION: Australasia: Australia (Queensland), New Guinea. Oriental: Indonesia (Java), Philippines. Palaearctic: Japan.

Genus Xaviera Lambkin

XAVIERA Lambkin, 1986b: 33. Type species: Mantispa manca Gerstaecker, 1885a ["1884"]: 39, by original designation. DISTRIBUTION: Oriental and Australasia: Indonesia to Australia. manca (Gerstaecker) Mantispa manca Gerstaecker, 1885a ["1884"]: 39, female. Holotype: female, Indonesia, Ambon (EMAU!). Mantispilla manca var. annulata Stitz, 1913: 10, female. Holotype: female, New Guinea (ZMB!). Synonymized with Mantispa manca by Esben-Petersen, 1923: 598. 206 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Mantispa manca papuana van der Weele, 1909: 92, male. Holotype: male, New Guinea (RMNH). Synonymized with Mantispa manca by Esben-Petersen, 1923: 598. DISTRIBUTION: Australasia: Australia (Northern Territory, Queensland), New Guinea. Oriental: Indonesia (Ambon).

Genus Xeromantispa Hoffman in Penny

XEROMANTISPA Hoffman in Penny, 2002: 270. Type species: Mantispilla scabrosa Banks, 1912: 179, by original designation. DISTRIBUTION: Neotropical and Nearctic: Costa Rica to southern USA. scabrosa (Banks) Mantispilla scabrosa Banks, 1912: 179. Holotype: female, USA, New Mexico (MCZ!). DISTRIBUTION: Nearctic: USA (Arizona, New Mexico, Texas). Neotropical: Costa Rica, Mexico.

Genus Zeugomantispa Hoffman in Penny

ZEUGOMANTISPA Hoffman in Penny, 2002: 271. Type species: Mantis minuta Fabricius, 1775: 278, by original designation. DISTRIBUTION: Neotropical and Nearctic: Argentina to USA. compellens (Walker) Mantispa compellens Walker, 1860: 181, sex not indicated. Syntypes123: female (only?), Brazil (BMNH!). Necyla uniformis Navás, 1927c: 61, sex not indicated. Holotype (or syntypes): sex not indicated, Guatemala (CN). New synonym124. DISTRIBUTION: Neotropical: southern Brazil to Central Mexico. minuta (Fabricius) Mantis minuta Fabricius, 1775: 278, sex not indicated. Holotype (or syntypes): type locality not indicated, sex unknown (BMNH). Mantis lilliputiana Olivier, 1797 ["1792"]: 640, sex not indicated. [Type status unsettled125]. Surinam. Synonymized with Mantis minuta Fabricius by Lichtenstein, 1796: 81. Synonymized with Mantispa flavomaculata Latreille by Erichson, 1839: 172. Unnecessarily cited as a new synonym with Mantispa minuta (Fabricius) by Penny, 1982a: 459. Mantispa flavomaculata Latreille, 1805: 94, sex not indicated. Surinam [Type status unsettled125]. Synonymized with Mantispa lilliputiana (Olivier) by Erichson, 1839: 172. Unnecessarily cited as a new synonym with Mantispa minuta (Fabricius) by Penny, 1982a: 459. Mantis viridula Houttuyn in Stoll, 1813: 77, sex not indicated. Surinam. [Type status unsettled125]. New synonym. Ohl: World Catalog of Mantispidae 207

Raphidia margaritacea Fischer von Waldheim, 1834: 330, sex not indicated. Holotype (or syntypes): sex unknown, Brazil (type depository unknown126). Synonymized with Mantispa minuta (Fabricius) by Penny, 1982a: 459. Mantispa viridis Walker, 1853: 227 [authorship incorrectly attributed to Newman], sex not indicated. Holotype (or syntypes): sex not indicated, USA, Florida (BMNH!). Synonymized with Zeugomantispa minuta by Hoffman in Penny, 2002: 274. Mantispa gulosa Taylor, 1862: 494 [as gulosus [sic], incorrect original termination]. As a nomen dubium considered synonymous with Mantispa viridis by Penny et al., 1997: 75. Mantispa brevicollis Banks in Baker, 1905: 88. A nomen nudum. Synonymized with Mantispa viridis by Penny et al., 1997: 75. Mantispa pallescens Navás, 1914e: 229, sex not indicated. Holotype: male, Brazil (BMNH!). Synonymized with Mantispa minuta (Fabricius) by Penny, 1982a: 459. A junior primary homonym of Mantispa pallescens Stitz, 1913. Mantispilla flavescens Navás, 1914e: 231, sex not indicated. Holotype (or syntypes): male, Brazil (BMNH!). Synonymized with Mantispa minuta (Fabricius) by Penny, 1982a: 459. Mantispilla trichostigma Navás, 1921: 51, sex not indicated. Holotype: sex unknown, Argentina (MNHN). Synonymized with Mantispa minuta (Fabricius) by Penny, 1982a: 459. Mantispilla viridata Navás, 1924: 59, sex not indicated. Holotype (or syntypes): sex unknown, Costa Rica (MNHN). Synonymized with Zeugomantispa minuta (Fabricius) by Hoffman in Penny, 2002: 274. Mantispilla rubricata Navás, 1924: 60, sex not indicated. Holotype (or syntypes): sex unknown, Costa Rica (MNHN). Synonymized with Zeugomantispa minuta (Fabricius) by Hoffman in Penny, 2002: 274. Mantispilla flavicornis Navás, 1930f: 301, female. Holotype (or syntypes): female, Costa Rica (ZMUH). Synonymized with Zeugomantispa minuta (Fabricius) by Hoffman in Penny, 2002: 274. DISTRIBUTION: Neotropical: Argentina, Uruguay, Peru, Ecuador, Surinam, Venezuela, Costa Rica, Mexico, Hispaniola. Nearctic: USA. virescens (Rambur)127 Mantispa viridula Erichson, 1839: 170, sex not indicated. Syntypes128: sex unknown, Brazil (ZMB! MCZ!). A junior secondary homonym of Mantispa viridula Houttuyn in Stoll, 1813 [= Zeugomantispa minuta (Fabricius)]. Mantispa virescens Rambur, 1842: 433, sex not indicated. Holotype: male [here confirmed], type locality unknown, perhaps South America129 (ISNB!). New synonym130. Mantispilla punctata Stitz, 1913: 20, female. Syntypes: female, Brazil (ZMB!). New synonym. Mantispilla punctata var. major Stitz, 1913: 20, sex not indicated. Holotype131: sex unknown, Brazil (ZMB!).132 New synonym. Mantispilla stigmata Stitz, 1913: 20, male. Holotype: male, Brazil (ZMB!). New synonym. Mantispa viridis Stitz, 1913: 29, female. Syntypes: females, Paraguay (ZMB!).133 New synonym. DISTRIBUTION: Neotropical: Argentina to Central Mexico.

208 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Nomina dubia in Mantispidae

Genus Manega Navás

MANEGA Navás, 1930a: 412. Type species: Manega luddemanni Navás, 1930a: 412, by monotypy.134 DISTRIBUTION: Australasian: Australia. luddemanni Navás Manega luddemanni Navás, 1930a: 412, female [as Luddemanni [sic], incorrect original capitalization]. Holotype: female, Australia, New South Wales (ZMUH). DISTRIBUTION: Australasian: Australia (New South Wales).

Genus Nivella Navás

NIVELLA Navás, 1930a: 413. Type species: Nivella rubella Navás, 1930a: 414, by monotypy.135 DISTRIBUTION: Australasian: Australia. rubella Navás Nivella rubella Navás, 1930a: 414, female. Holotype: female, Australia, New South Wales (ZMUH). DISTRIBUTION: Australasian: Australia (New South Wales).

Mantispidae unplaced to genus

The following species-group names were originally described or subsequently placed in "Mantispa", as it was historically understood, as a collective genus for neuropterans with raptorial forelegs. Since the taxonomic identities of these names cannot be established at the moment, they are herein considered as unplaced nomina dubia in Mantispidae. fausta Thunberg136 Mantis fausta Thunberg, 1784: 63, sex not indicated. Holotype (or syntypes): sex unknown, South Africa (UZIU). DISTRIBUTION: Afrotropical: South Africa. nabota Olivier137 Mantis nabota Olivier, 1797 ["1792"]: 640, sex not indicated. [Type status unsettled]. "Coromandel". Synonymized with Mantispa rufescens by Erichson, 1839: 173. Mantis nana Lichtenstein, 1802: 33, sex not indicated. [Type status unsettled]. "Coromandel". New synonym. Mantispa rufescens Latreille, 1807: 94, sex not indicated. [Type status unsettled]. "India orientalis, Coromandel". DISTRIBUTION: Oriental: India (Coromandel). Ohl: World Catalog of Mantispidae 209

sibirica Gmelin138 Mantis sibirica Gmelin, 1790 ["1788"]: 2055, sex not indicated. Holotype (or syntypes): sex unknown, "Sibirien" (depository unknown). DISTRIBUTION: Palaearctic: Russia (Sibiria).

Fossil Taxa in Mantispidae

The higher taxa listed below are strictly fossil, i.e., with no extant representatives. Fossil species belonging to extant genera are listed with their appropriate genera above.

Genus † Promantispa Panfilov in Dolin et al.

† PROMANTISPA Panfilov in Dolin et al., 1980 (27 January): 92. Type species: † Promantispa similis Panfilov in Dolin et al., 1980: 93, by original designation.139

† similis Panfilov in Dolin et al. † Promantispa similis Panfilov in Dolin et al., 1980: 93, sex not indicated. Holotype: sex unknown, Kazakhstan, Upper Jurassic (PIMR). DISTRIBUTION: Palaearctic: Kazakhstan (Upper Jurassic) (compression/impression).

Genus † Prosagittalata Nel

† PROSAGITTALATA Nel, 1989 ["1988"]: 104. Type species: † Prosagittalata oligocenica Nel, 1989 ["1988"]: 104, by monotypy.

† oligocenica Nel † Prosagittalata oligocenica Nel, 1989 ["1988"]: 104, sex unkown. Holotype: sex unknown, France (Upper Oligocene) (MNHN). DISTRIBUTION: Palaearctic: France (Upper Oligocene) (compression/impression).

Genus † Vectispa Lambkin140

† Promantispa Jarzembowski, 1980 (27 March): 255. Type species: † Mantispa relicta Cockerell, 1921: 477, by original designation. A junior primary homonym of † Promantispa Panfilov in Dolin et al., 1980 (27 January), replaced by † Vectispa Lambkin, 1986a. † VECTISPA Lambkin, 1986a: 16. Replacement name for † Promantispa Jarzembowski, 1980 (27 March), a junior primary homonym of † Promantispa Panfilov in Dolin et al., 1980 (27 January).

210 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

† relicta (Cockerell) † Mantispa relicta Cockerell, 1921: 477, sex not indicated: Holotype: sex unknown, Great Britain, Isle of Wight, Upper Eocene (BMNH). DISTRIBUTION: Palaearctic: Great Britain, Isle of Wight (Early Cretaceous) (amber).

Genus † Whalfera Engel141

† Fera Whalley, 1983 (28 Dec): 230. Type species: † Fera venatrix Whalley, 1983: 230, by original designation. A junior primary homonym of † Fera Hong, 1983 (Sept) (Diptera: Pleciofungivoridae), replaced by † Whalfera Engel, 2004. † Whalfera Engel, 2004: 140. Replacement name for † Fera Whalley, 1983 (28 Dec), a junior primary homonym of † Fera Hong, 1983 (Sept).

† venatrix (Whalley) † Fera venatrix Whalley, 1983: 230, sex not indicated. Holotype: sex unknown, Great Britain, Eocene/Oligocene (BMNH). DISTRIBUTION: Palaearctic: Great Britain (Eocene/Oligocene) (amber).

Subfamily † Mesomantispinae Makarkin

Genus † Mesomantispa Makarkin

† MESOMANTISPA Makarkin, 1997 ["1996"]: 92. Type species: † Mesomantispa sibirica Makarkin, 1997 ["1996"]: 92, by original designation.

† sibirica Makarkin † Mesomantispa sibirica Makarkin, 1997 ["1996"]: 92, sex not indicated: Holotype: sex unknown, Russia, East Sibiria, Lower Cretaceous (PIMR). DISTRIBUTION: Palaearctic: Russia, East Sibiria (Lower Cretaceous) (compression/impression).

Questionable Fossil Mantispidae

Genus † Mantispidiptera Grimaldi

† MANTISPIDIPTERA Grimaldi, 2000: 268. Type species: † Mantispidiptera enigmatica Grimaldi, 2000: 270, by original designation.

† enigmatica Grimaldi † Mantispidiptera enigmatica Grimaldi, 2000: 270, sex not indicated. Holotype: sex unknown, USA, New Jersey, Middle/Late Cretaceous (AMNH). DISTRIBUTION: Palaearctic: USA, New Jersey (Middle/Late Cretaceous) (amber).

Ohl: World Catalog of Mantispidae 211

† henryi Grimaldi † Mantispidiptera henryi Grimaldi, 2000: 270, sex not indicated. Holotype: sex unknown, USA, New Jersey, Middle/Late Cretaceous (AMNH). DISTRIBUTION: Palaearctic: USA, New Jersey (Middle/Late Cretaceous) (amber).

Genus † Paraberotha Whalley

† PARABEROTHA Whalley, 1980: 161. Type species: † Paraberotha acra Whalley, 1980: 162, by original designation.

† acra Whalley † Paraberotha acra Whalley, 1980: 162, "male (?)". Holotype: male (?), Lebanon, Early Cretaceous (CAA). DISTRIBUTION: Palaearctic: Lebanon (Early Cretaceous) (amber).

Genus † Retinoberotha Schlüter

† RETINOBEROTHA Schlüter, 1978: 64. Type species: † Retinoberotha stuermeri Schlüter, 1978, by monotypy.142

† stuermeri Schlüter † Retinoberotha stuermeri Schlüter, 1978:64, sex not indicated. Holotype: sex unknown, France, Upper Cretaceous (ZMB143). DISTRIBUTION: Palaearctic: France (Upper Cretaceous) (amber).

212 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

REFERENCES

Printed Sources

The following bibliography gives full citations to more than 270 works cited in the catalog. Citations are generally taken from John Oswald's Bibliography of Neuropterida (see Internet Sources below), but a concerted effort was made to examine all cited references personally to ensure accurate citation of authorship, date of publication, title and pagination. Special attention was paid to obtaining true dates of publication for the purpose of accurately determining nomenclatural priority. All dating information has been checked against the current version of the Bibliography of Neuropterida, and corrected where necessary. Evidence for the dating of some works, particularly evidence not directly available to me, is provided in brackets after the citation and includes the source of information. For publications that appeared after their imprint date (year), the imprint year as printed in the original publication is given in brackets following the correct year of publication. Both years are cited, where appropriate, in the catalog. Exact dates (day and/or month) are given only if these are necessary for determining the priority of one or more mantispid taxon names or nomenclatural acts. References are cited alphabetically by author surname, then chronologically. When more than one article was published with the same authorship in a particular year, a suffix letter follows the year. Lettered citations are listed alphabetically. An attempt has been made to assign letters sequentially based on publication chronology, but, where exact publication dates were not established, letters were asigned randomly. Illustration and plate numbers are given only if these were printed separately or in separate order from the paginated text, or if these are of nomenclatural importance. Titles of publications are given in full, including secondary titles, and are cited as given on the title page of the original work. Upper and lower case letters are used as given in the title of the original work or as required by traditional language standards. Titles of publications printed in non-Roman characters (e.g., Cyrillic or Chinese) are generally given in transliteration, followed by a translation in brackets. The language of a publication is indicated in brackets at the end of the citation if the primary text language does not use a Latin-derived alphabet. Journal titles are given in full as indicated in the article, or on the wrapper or cover of the respective volume. Volume numbers are given in Arabic numbers, irrespective of whether or not Roman numerals were used in the original work. Issue and part numbers are omitted except where needed to distinguish among of overlapping paginations of such issues or parts. Single volume books are cited with full title, publisher, place(s) of publication and total number of pages. Unpaginated parts of a book are placed in brackets. Annotations are given in brackets after the citation, if necessary. These may include information on dating, duplicate versions or various other data.

Alayo, D. 1968. Los Neurópteros de Cuba. Poeyana Instituto de Biologia (B) 2: 1-127. Apstein, C. 1916. Erscheinungsjahre von Gmelin (Linné), Systema Naturae, ed. 13. Zoologischer Anzeiger 47: 32. Aspöck, H. 1999. Beschreibungen und Abbildungen von Mantispiden in der frühen entomologischen Literatur und Österreichs Beitrag zur Erforschung der Fanghafte (Neuropterida: Neuroptera: Mantispidae). Stapfia 60: 209-244. Ohl: World Catalog of Mantispidae 213

Aspöck, H., Aspöck, U., and H. Hölzel. 1980. Die Neuropteren Europas. Eine zusammenfassende Darstellung der Systematik, Ökologie und Chorologie der Neuropteroidea (Megaloptera, Raphidioptera, Planipennia) Europas. 2 vol. Goecke und Evers, Krefeld, 495 + 355 pp. Aspöck, H., Hölzel, H., and U. Aspöck. 2001. Kommentierter Katalog der Neuropterida (Insecta: Raphidioptera, Megaloptera, Neuroptera) der Westpaläarktis. Denisia 2: 606 pp. Aspöck, U. 1996. Die Mantispiden Europas (Neuropteroidea: Neuroptera: Mantispidae). Pp. 224- 230. In Verhandlungen des 14. Internationalen Symposiums über Entomofaunistik in Mitteleuropa (SIEEC, held 4-9 September 1994 in München). Aspöck, U. 2002. Phylogeny of the Neuropterida (Insecta: Holometabola). Zoologica Scripta 31: 51-55. Aspöck, U., and H. Aspöck. 1994. Zur Nomenklatur der Mantispiden Europas (Insecta: Neuroptera: Mantispidae). Annalen des Naturhistorischen Museums in Wien 96 B: 99-114. Aspöck, U., and M. W. Mansell. 1994. A revision of the family Rhachiberothidae Tjeder, 1959, stat. n. (Neuroptera). Systematic Entomology 19: 181-206. Aspöck, U., Plant, J. D., and H. L. Nemeschkal. 2001. Cladistic analysis of Neuroptera and their systematic position within the Neuropterida (Insecta: Holometabola: Neuropterida: Neuroptera). Systematic Entomology 26: 73-86. Baker, C. F. 1905. Notes on neuropteroid of the Pacific Coast of North America with descriptions of new species by Nathan Banks. Invertebrata Pacifica 1: 85-92. Baly, J. S. 1858. Catalogue of Hispidae in the collection of the British Museum, part. I. British Museum (Natural History), London. 172 pp. Banks, N. 1897. New North American neuropteroid insects. Transactions of the American Entomological Society 24: 21-31. Banks, N. 1910. Some Neuroptera from Australia. Psyche 17: 99-105. Banks, N. 1911. Descriptions of new species of North American neuropteroid insects. Transactions of the American Entomological Society 37: 335-360. Banks, N. 1912. Notes on Nearctic Mantispidae. Proceedings of the Entomological Society of Washington 14: 178-179. Banks, N. 1914 ["1913"]. On a collection of neuropteroid insects from the Philippine Islands. Proceedings of the Entomological Society of Washington 15: 170-180. Banks, N. 1913. Synopses and descriptions of exotic Neuroptera. Transactions of the American Entomological Society 39: 201-242. Banks, N. 1920. New neuropteroid insects. Bulletin of the Museum of Comparative Zoology 64: 297-362. Banks, N. 1933. Entomological investigations on the spike disease of sandal (9). Neuroptera. Indian Forest Records, Entomology Series 18 (6): 1-4. Banks, N. 1939. New genera and species of neuropteroid insects. Bulletin of the Museum of Comparative Zoology 85: 439-504. Banks, N. 1943. New Neuroptera and Trichoptera from the United States. Psyche 50: 74-81. Berthold, A. A. 1827. Natürliche Familien des Thierreichs. Aus dem Französischen. Mit Anmerkungen und Zusätzen von Arnold Adolph Berthold. Weimar, 602 pp. [A German translation of Latreille, P. A. 1825. Familles naturelles du règne , … Paris, 570 pp.] Blanchard, C. É. 1851. Mirmeleonianos. Pp. 119-129 and Rafidianos. Pp. 129-135. In C. Gay (ed.): Historia Fisica y Politica de Chile, Zoologia, Vol. 6. Paris. Boeseman, M., and W. de Ligny. 2004. Martinus Houttuyn (1720-1798) and his contributions to the natural sciences, with emphasis on zoology. Zoologische Verhandlingen Leiden 349: 1- 222. 214 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Boyden, T. C. 1984. Mimicry, predation, and potential pollination by the mantispid Climaciella brunnea var. instabilis (Say) (Neuroptera: Mantispidae). Journal of the New York Entomological Society 91: 508-511. Brauer, F. 1867a. Beitrag zur Kenntniss der Mantispiden-Gattungen. Verhandlungen der Kaiserlich-Königlichen Zoologische-Botanischen Gesellschaft in Wien 17: 281-286. Brauer, F. 1867b. Beschreibung neuer Neuroptera aus dem Museum Godeffroy und Sohn in Hamburg. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 17: 505-512. Brushwein, J. R., Culin, J. D., and K. M. Hoffman. 1995. Development and reproductive behavior of Mantispa viridis Walker (Neuroptera, Mantispidae). Journal of Entomological Science 30: 99-111. Charpentier, T. von. 1825. Horae entomologicae, adjectis tabulis novem coloratis. A. Gosohorsky, Wratislaviae. xvi + 255 pp. Charpentier, T. von. 1844. Bemerkungen zu A. A. H. Lichtensteins Abhandlung über die Mantis- Arten, in den Transactions of the Linneean [sic!] Society. Vol. VI. Lond. 1802. Zeitschrift für die Entomologie (Germar) 5: 272-311. Cockerell, T. D. A. 1921. Fossil arthropods in the British Museum. – VI. Oligocene insects from Gurnet Bay, Isle of Wight. Annals and Magazine of Natural History (9) 7: 453-480. Cowan, C. F. 1971. On Guérin's Iconographie: particularly the insects. Journal of the Society for the Bibliography of Natural History 6: 18-29. De Geer, C. 1778. Memoires pour servir a l'histoire des insectes. Vol. 7. P. Hesselberg, Stockholm. xii + 950 pp. Neuropterida parts: "Mante" [a mantispid], pp. 620-621, pl. 46, figs. 9-10. Ehrmann, R. 2002. Mantodea – Gottesanbeterinnen der Welt. Natur und Tier (NTV), Münster. 519 pp. Enderlein, G. 1910. Klassifikation der Mantispiden nach dem Material des Stettiner Zoologischen Museums. Stettiner Entomologische Zeitung 71: 341-379. Engel, M. S. 2004. Thorny lacewings (Neuroptera: Rhachiberothidae) in Cretaceous amber from Myanmar. Journal of Systematic Palaeontology 2: 137-140. Erichson, W. F. 1839. Beiträge zu einer Monographie von Mantispa, mit einleitenden Betrachtungen über die Ordnungen der Orthopteren und Neuropteren. Zeitschrift für die Entomologie (Germar) 1: 147-173. Esben-Petersen, P. 1912. A few new Neuroptera Planipennia. Entomologische Mitteilungen 1: 267-273. Esben-Petersen, P. 1913. H. Sauter's Formosa-Ausbeute. Planipennia II, Megaloptera and Mecoptera. Entomologische Mitteilungen 2: 222-228, 257-265. Esben-Petersen, P. 1915 ["1914-1915"]. Australian Neuroptera. Part i. Proceedings of the Linnean Society of New South Wales 39: 635-645. [Dating: From contents pages of vol 39: pages 635-640 issued 24 Nov 1914, pages 641-645 issued 26 Feb 1915.] Esben-Petersen, P. 1917. Neue und wenig bekannte Mantispiden. Arkiv för Zoologi 11 (10): 1- 15. Esben-Petersen, P. 1923. Australian Neuroptera. Part v. Proceedings of the Linnean Society of New South Wales 48: 593-600. Esben-Petersen, P. 1927. New and little-known species of Mecoptera and Neuroptera in the zoological museum of Helsingfors. Notulae Entomologicae 7: 13-18. Esben-Petersen, P. 1928. New and little-known Neuroptera from the Dutch East Indies. Treubia 10: 225-230. Ohl: World Catalog of Mantispidae 215

Esben-Petersen, P. 1929. Australian Neuroptera. [Part] VI. Queensland Naturalist 7: 31-35. Evenhuis, N. L. 1997. Litteratura Taxonomica Dipterorum (1758-1930). Vol. I: A-K; vol. 2: L-Z. Backhuys, Leiden, 871 pp. Evenhuis, N. L. 2002. Catalog of the Mythicomyiidae of the World (Insecta: Diptera). Bishop Museum Bulletins in Entomology 10: 85 pp. Fabricius, J. C. 1775. Systema entomologiae, sistens insectorvm classes, ordines, genera, species, adiectis synonymis, locis, descriptionibvs, observationibvs. Flensbvrgi et Lipsiae. 832 pp. Fairmaire, L. M. H. 1858. Ordre Névroptères. Pp. 261-262. In M. J. Thomson (ed.): Archives entomologiques ou recueil contenant des illustrations d'insectes nouveaux ou rares, Paris. Vol. 2. 1 plate [#10] with 2 Neuroptera figures. [Dating: From title page of volume 2.] Fischer von Waldheim, G. 1834. Notice sur quelques Orthoptères et Nevroptères du Brésil. Bulletin de la Société [Impériale] des Naturalistes de Moscou [= Byulleten' Muskovskogo obshchestva Ispytatelei Prirody] 7: 322-330. Franz, H., and M. Beier. 1970. Die geographische Verbreitung der Insekten. Handbuch der Zoologie, IV. Band: Arthropoda. 2. Hälfte: Insecta. Zweite Auflage. 1. Teil: Allgemeines. Beitrag 6. W. de Gruyter, Berlin. 139 pp. Fraser, F. C. 1952. New species of Neuroptera in the Museum National d'Histoire Naturelle, Paris. Revue Française d'Entomologie 19: 55-64. Gerstaecker, A. 1885a ["1884"]. Vier Decaden von Neuropteren aus der Familie Megaloptera Burm. Mitteilungen des Naturwissenschaftlichen Vereins für Neu-Vorpommern und Rügen in Greifswald 16: 1-49. [Dating: From the wrapper and title page of volume 16 and from the wrapper of a separate in the ZMB library.] Gerstaecker, A. 1885b ["1884"]. Zwei fernere decaden Australischer Neuroptera Megaloptera. Mitteilungen des Naturwissenschaftlichen Vereins für Neu-Vorpommern und Rügen in Greifswald 16: 84-116. [Dating: From the wrapper and title page of volume 16 and from the wrapper of a separate in the ZMB library.] Gerstaecker, A. 1888 ["1887"]. Weitere Beiträge zur Artenkenntniss der Neuroptera Megaloptera. Mitteilungen des Naturwissenschaftlichen Vereins für Neu-Vorpommern und Rügen in Greifswald 19: 89-130. [Dating: From wrapper and title page of volume 19.] Gerstaecker, A. 1894 ["1893"]. Ueber neue und weniger gekannte Neuropteren aus der Familie Megaloptera Burm. Mitteilungen des Naturwissenschaftlichen Vereins für Neu- Vorpommern und Rügen in Greifswald 25: 93-173. [Dating: From wrapper and title page of volume 25.] Ghosh, S. K., and S. Sen. 1977. Check-list of Indian Planipennia (order Neuroptera). Records of the Zoological Survey of India 73: 277-326. Gmelin, J. F. 1790 ["1788"]. Caroli a Linne, equatis aurati de stella polari, à systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis à Editio decima tertia [= 13th Edition], aucta, reformata. Tom. 1, Pars IV (pp. 1517-2224) and Pars V (pp. 2225-3020). Georg. Emanuel Beer, Lipsiae. [Dating: From information in Hopkinson (1907) and Apstein (1916). Volume 1, part 4 dated 21 May 1790; volume 1, part 5 dated pre 6 December 1790. Title page to v. 1, pt. 1 dated "1788". Title pages to v. 1, pt. 4 and pt. 5 undated.] Grimaldi, D. 2000. A diverse fauna of Neuropterodea in amber from the Cretaceous of New Jersey. Pp. 259-303. In: D. Grimaldi (ed.): Studies on fossils in amber, with particular reference to the Cretaceous of New Jersey. Backhuys Publishers, Leiden. viii + 498 pp. Guérin-Méneville, F. É. 1831 ["1830"]. Névroptères. Pp. 194-197. In L. I. Duperrey (ed.): Voyage autor du monde, exécuté par ordre du roi, sur la corvette de sa majesté, La Coquille, 216 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

pendant les années 1822, 1823, 1824 et 1825 à par m. L. I. Duperrey, [section] Zoologie, Vol. 2, Pt. 2, div. 1. Crustacés, Arachnides et Insectes, pp. 57-302. Plate 10 with 1 neuropterid figure. [Dating: Sherborn and Woodward (1901:392). Volume 2, part 2, pp. 1- 216 dated 12 Nov. 1831, title page dated 1830.] Guérin-Ménéville, F. É. 1838. Mantispe. Mantispa. Illiger. Magasin de Zoologie (1) 8 (text to pl 202): 1-2. Guérin-Méneville, F. É. 1844 ["1829-1838"]. Iconographie du règne animal de G. Cuvier, ou représentation d'après nature de l'une des espèces les plus remarquables, et souvent non encore figurées, de chaque genre d'animaux. Insectes. Paris. 576 pp. Plates 61-63 with neuropterid figures. [Dating: Cowan (1971): Text dated 7 September 1844. Plates dated as follows: pl. 61 (livraison 34, June 1834), pl. 62 (livraison 16, 5 November 1831), pl. 63 (livraison 45, December 1837). Title page dated 1829-1838.] Hagen, H. A. 1858a. Russlands Neuropteren. Stettiner Entomologische Zeitung 19: 110-134. Hagen, H. A. 1858b. Synopsis der Neuroptera Ceylons [Pars I]. Verhandlungen der Kaiserlich- Königlichen Zoologisch-Botanischen Gesellschaft in Wien 8: 471-488. Hagen, H. A. 1859. Beitrag zur Kenntniss der Neuropteren. Stettiner Entomologische Zeitung 20: 405-412. Hagen, H. A. 1861. Synopsis of the Neuroptera of North America, with a list of the South American species. Smithsonian Miscellaneous Collections 4 (1): xx + 1-347. Hagen, H. A. 1862-1863. Bibliotheca Entomologica. Die Litteratur über das ganze Gebiet der Entomologie bis zum Jahre 1862. 2 Bände. Wilhelm Engelmann, Leipzig. 566 pp. + 512 pp. Hagen, H. A. 1866. Hemerobidarum Synopsis Synonymica. Stettiner Entomologische Zeitung 27: 369-462. Hagen, H. A. 1877. Symphrasis, eine neue Mantispiden-Gattung. Stettiner Entomologische Zeitung 38: 208-211. Handschin, E. 1935. Indo-australische Neuropteren und Mecopteren. Revue Suisse de Zoologie 42: 683-714. Handschin, E. 1959a. Beiträge zu einer Revision der Mantispiden (Neuroptera). I Teil. Mantispiden des Musee Royal du Congo Belge, Tervuren. Revue de Zoologie et de Botanique Africaines 59: 185-227. Handschin, E. 1959b. Mantispa styriaca (Poda 1761). Vierteljahresschrift der Naturforschenden Gesellschaft in Zürich 104: 105-114. Handschin, E. 1960a. Zur Revision süd-amerikanischer Mantispiden. Revue Suisse de Zoologie 67: 523-560. Handschin, E. 1960b. Beiträge zu einer Revision der Mantispiden (Neuroptera). II Teil. Mantispiden des "Musée Royal du Congo Belge", Tervuren. Revue de Zoologie et de Botanique Africaines 62: 181-245. Handschin, E. 1960c. Nampista auriventris (Guérin) 1838 (Neuropt. Planip.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 33: 155-160. Handschin, E. 1961. Beiträge zur Kenntnis der Gattungen Euclimacia, Climaciella und Entanoneura Enderlein 1910 im indo-australischen Faunengebiet. Nova Guinea, Zoology 15: 253-301. Handschin, E. 1963 ["1962"]. Beitrag zur Kenntnis der Neuropterenfauna von Madagascar. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 35: 211-226. Handschin, E., and W. Markl. 1955. Neuropteren aus Angola. Companhia de Diamantes de Angola, Publicaçoes Culturais, Lisboa 27: 65-82. Ohl: World Catalog of Mantispidae 217

Haring, E., and U. Aspöck. 2004. Phylogeny of the Neuropterida: a first molecular approach. Systematic Entomology 29: 415-430. Herrich-Schäffer, G. A. W. 1840. Nomenclator Entomologicus. Verzeichnis der europäischen Insecten; zur Erleichterung des Tauschverkehrs mit Preisen versehen. Heft 2. Regensburg, 244 pp. Hoffman, K. M. 1992. Systematics of the Mantispinae (Neuroptera: Mantispidae) of North, Central, and South America. Unpublished Ph.D. dissertation, Clemson University, Clemson, South Carolina, 501 pp. Hoffman, K. [M.] 2002. Family Mantispidae. Pp. 251-275. In N. D. Penny (ed.): A guide to the lacewings (Neuroptera) of Costa Rica. Proceedings of the California Academy of Sciences 53: 161-457. Hoffman, K. M., and J. R. Brushwein. 1989. Species of spiders (Araneae) associated with the immature stages of Mantispa pulchella (Neuroptera: Mantispidae). Journal of Arachnology 17: 7-14. Hoffman, K. M., and J. R. Brushwein. 1990. Spider (Araneae) taxa associated with the immature stages of Mantispa interrupta (Neuroptera: Mantispidae). Entomological News 101: 23-28. Hoffman, K. M., and J. R. Brushwein. 1992. Descriptions of the larvae and pupae of some North American Mantispinae (Neuroptera: Mantispidae) and development of a system of larval chaetotaxy for Neuroptera. Transactions of the American Entomological Society 118: 159- 196. Hong, Y.-C. 1983. Middle Jurassic fossil insects in North China. Geological Publishing House, Beijing, China. (published September 1983 according to Engel, 2004) Hopkinson, J. 1907. Dates of publication of the separate parts of Gmelin's edition (13th) of the "Systema Naturae" of Linnaeus. Proceedings of the Zoological Society of London 1907: 1035-1037. Horn, W., Kahle, I., Friese, G., and R. Gaedike. 1990. Collectiones entomologicae. Akademie der Lanwirtschaftswissenschaften der Deutschen Demokratischen Republik, Berlin, 2 vol., 573 pp. Horn, W., and S. Schenkling. 1828-1929. Index Litteraturae Entomologicae. Serie I: Die Welt- Literatur über die gesamte Entomologie bis inklusive 1863. 4 vols. Selbstverlag, Berlin. 1426 pp. Houttuyn, M. 1813. Register van de in dit Werk afgebeelde Spoken en wandelende Bladen. Table des spectres et des feuilles ambulantes qui sont representes dans cet ouvrage. Index phasmatum et mantum in hoc opere delineatorum. pp. 76-79. In Stoll, C. Natuurlijke en naar het leven nauwkeurig gekleurde Afbeeldingen en Beschrijvingen der Spoken, wandelende Bladen, Zabel-Springhanen, Krekels, Trek-Springhaanen en Kakkerlakken. In alle vier deelen der Wereld, Europa, Asia, Afrika en America, huishoudende. Bij een verzameld en Beschreven door Caspar Stoll. Representation exactement colorée d'après nature des Spectres ou Phasmes, des Mantes, des Sauterelles, des Grillons, des Criquets et des Blattes. Qui se trouvent dans les quatre parties du monde, l'Europe, l'Asie, l'Afrique et l'Amerique; Rassemblées et décrites par Caspar Stoll. J. C. Sepp et fils, Amsterdam, 79 pp. [detailled pagination and plate numbers cannot be provided here, because no copy of this work is currently accessible to us]. [Hagen (1862-1863: Bd. 2, 199) and Horn and Schenkling (1929: 1193) stated that M. Houttuyn continued the work by Stoll 1787 ["1787- 1790"] until 1815 after Stoll died in 1795. However, 1813 is printed on the title page (which is illustrated in H. Aspöck, 1999: Fig. 44) of the work, and there is no printed evidence for a later publication date (based on the photocopies of some parts of the work available to MO). 218 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Thus, we accept 1813 tentatively as the correct year of publication. Since the complete work by Houttuyn is currently not accessible to us, it difficult to tell the differences to the original work by Stoll 1787 ["1787-1790"]. Obviously, Stoll's complete original text and plates were re-printed in the later work. Houttuyn added at least some plates (25 plates are numbered in Houttuyn's index, but Hagen (1862-1863: Bd. 2, 199) and Horn and Schenkling (1929: 1193) gave a total of 75 plates for the entire work] and the index to the plates. Our dating of this work and our interpretation of the authorship is in agreement with Weidner (1983) and Boeseman & de Ligny (2004). In the index, Houttuyn proposed the new name Mantis viridula for the mantispid named "Le Nain de Suriname" by Stoll (p. 77). See notes for Stoll 1787 ["1787-1790"].] ICZN (International Commission on Zoological Nomenclature). 1999. International Code of Zoological Nomenclature. 4th ed. The International Trust for Zoological Nomenclature, London. 306 pp. Illiger, J. K. W. 1798. Genus Mantispa. p. 499. In J. G. Kugelann: Verzeichnis der Käfer Preussens, entworfen von Johann Gottlieb Kugelann … ausgearbeitet von Johann Karl Wilhelm Illiger. Mit einer Vorrede des Professors und Pagenhofmeisters Helwig in Braunschweig, und dem angehängten Versuche einer natürlichen Ordnungs- und Gattungsfolge der Insekten. Halle. 510 pp. Jarzembowski, E. A. 1980. Fossil insects from the Bembridge Marls, Palaeogene of the Isle of Wight, southern England. Bulletin of the British Museum of Natural History (Geology) 33: 237-292. [Dating: 27 March 1980 as printed on title page of paper.] Kevan, D. K. McE. 1989. Ms. Taylor's walking-sticks: "Queen Bess" and odd "orthopterians" of 1862 (Mantodea, Phasmatodea, Dermaptera, a reduviid hemipteran and mantispid Neuroptera). Proceedings of the Entomological Society of Ontario 120: 1-11. Kolbe, H. J. 1897. Neuropteren – Die Netzflügler. Pp. 1-42. In K. Möbius: Die Thierwelt Deutsch-Ost-Afrikas und der Nachbargebiete. Band IV Wirbellose Thiere, Lieferung 8. Dietrich Reimer, Berlin. Kormilev, N. A. 1958. Notas sobre Mantispidae neotropicales, I (Neuroptera). Revista de la Sociedad Entomologica Argentina 21: 1-18. Kozhanchikov, I. V. 1949. New Mantispidae (Neuroptera) from Tadshikistan. Entomologicheskoe Obozreni 30: 353-358 [Text in Russian]. Krivokhatsky, V. A. 1995. Katalog tipovykh ekzemplyarov kollektsii zoologicheskogo instituta RAN. Nasekomye setchatokrylye (Neuroptera) [= A catalog of the type-specimens in the collection of the Zoological Institute Russian Academy of Sciences. Insecta, Neuroptera.] Zoologicheskii Institut RAN, Sankt-Peterburg [St. Petersburg]. 18 pp. [Text in Russian.] Kuwayama, S. 1925 ["1924-1925"]. Notes on the Japanese Mantispidae, with special reference to the morphological characters. Journal of the College of Agriculture, Hokkaido Imperial University 15: 237-267. [Dating: From footer on p. 237: "Vol. XV, Pt. 4, June 1925".] Lambkin, K. J. 1986a. A revision of the Australian Mantispidae (Insecta: Neuroptera) with a contribution to the classification of the family I. General and Drepanicinae. Australian Journal of Zoology, Supplementary Series 116: 1-142. Lambkin, K. J. 1986b. A revision of the Australian Mantispidae (Insecta: Neuroptera) with a contribution to the classification of the family II. Calomantispinae and Mantispinae. Australian Journal of Zoology, Supplementary Series 117: 1-113. Lambkin, K. J. 1987. The Australian Mantispidae (Neuroptera): Supplementary notes. General and Applied Entomology 19: 11-14. Ohl: World Catalog of Mantispidae 219

Lambkin, K. J., and New, T. R. 1994. Australian Mantispidae (Neuroptera): supplementary notes 2. Journal of the Australian Entomological Society 33: 117-121. Latreille, P. A. 1807. Genera crustaceorum et insectorum secundum ordinem naturalem in familias disposita, iconibus exemplisque plurimis explicata. Vol. 3. Parisiis and Argentorati. 258 pp. Lepechin, I. I. 1774. Tagebuch der Reise durch verschiedene Provinzen des Russischen Reiches in den Jahren 1768 und 1769. Erster Theil. Richter, Altenburg. 331 pp. + Erratum [Digital copy: http://www-gdz.sub.uni-goettingen.de/cgi-bin/digbib.cgi?PPN335109918.] Le Peletier de Saint-Fargeau, A. L. M., and J. G. Audinet-Serville. 1825. Mantispe, Mantispa. Pp. 269-270; plate 95, figs 4-5 [as Raphidie], plate 133, figs 2-3 [as Mante]). In Encyclopédie Méthodique. Histoire Naturelle [1782-1828], 'Insectes.' Vol. 10, Latreille, P. A., Le Peletier de Saint-Fargeau, A. L. M., Audinet-Serville, J. G., and F. E. Guérin[-Menéville]. Paris. [Dating: Sherborn & Woodward (1906): volume 10, pages 1-344 dated 1825 (as given on title page of completed volume 10); plates 66-165 of volume comprising plates of "Crustacés, Arachnides, et Insectes" dated 1792 (1818 is given on title page of completed volume).] Lestage, J. A. 1934. Notes sur l'entomofaune indochinoise. IV. – Une nouvelle Climaciella du Tonkin (Climaciella Leopoldi) et remarques sur le rôle des mantispidés en phytopathologie. Bulletin et Annales de la Société [Royale] Entomologique de Belgique 74: 153-160. Lichtenstein, A. A. H. 1796. Catalogus musei Zoologici ditissimi Hamburgi, d. III. Februar 1796. Auctionis lege distrahendi. Sectio Tertia, continens Insecta. Verzeichnis von höchstseltenen, aus allen Welttheilen mit vieler Mühe und Kosten zusammen gebrachten, auch aus unterschiedlichen Cabinettern, Sammlungen und Auctionen ausgehobenen Naturalien, welche von einem Liebhaber, als Mitglied der Batavischen und verschiedener anderer Naturforschender Gesellschaften gesammlet worden. Dritter Abschnitt, bestehend in wohlerhaltenen, mehrentheils ausländischen und höchstseltenen Insekten, die theils einzeln, theils mehrere zusammen in Schachteln festgesteckt sind, und welche am Mittewochen, den 3ten Februar 1796 und den folgenden Tagen auf dem Eimbeckschen Hause öffentlich verkauft werden sollen durch den Mackler Peter Hinrich Packischefsky Gottl. Friedr. Schniebes, Hamburg. Vol. 3. [xiii] + 222 pp. [ii, Appendix]. [This catalogue was produced to document and advertise the sale at auction of the Holthuisen natural history collection. The auction was originally scheduled for 3 February 1796, but was postponed until 16 May 1797. Lichtensteins' introduction to this catalogue contains the date 1 November 1795, and it appears likely that the first edition of this work appeared in late 1795 or early 1796, to predate to original date of the sale. This edition is customarily dated 1796 from the sale date on its title page. A second edition of this work was issued in 1797 prior to the actual sale date in May of that year. Several mantispids are included on p. 81 of the section "Insect[orum]. Ordo II. Hemiptera ... Mantis", pp. 77-81. The mantispids in this work were previously illustrated by Stoll 1787 ["1787-1790"] (pl. I, II, IV). For additional notes on this auction catalogue see Sherborn (1899) and Weidner (1983).] Lichtenstein, A. A. H. 1802. A dissertation on two natural genera hitherto confounded under the name of Mantis. Transactions of the Linnean Society of London 6: 1-39. Linnaeus, C. 1767. Systema natura per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. 12th Edition. Vol. 1, pt. 2 (pp. 533-1327). Salvii, Holmiae. Linsley, E. G., and J. W. MacSwain. 1955. Two new species of Plega from Mexico (Neuroptera, Mantispidae). Pan-Pacific Entomologist 31: 15-19. 220 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Loew, H. 1843. Bemerkungen über die anatomischen Verhälnisse der Neuropteren. Zeitschrift für die Entomologie (Germar) 4: 423-434. Makarkin, V. N. 1985. K faune setchatokrylykh (Neuroptera) Dalnego Vostoka [= A contribution to the fauna of the Neuroptera from the Far East]. Akademiia nauk SSSR. Zoologicheskii Zhurnal 64: 620-622. [Text in Russian, summary/abstract in English. Transliterated and translated Russian title from V. N. Makarkin, pers. com. to J. D. Oswald]. Makarkin, V. N. 1990. Novye setchatokrylye (Neuroptera) iz verkhnego mela Azii [= New lacewings (Neuroptera) from the Upper Cretaceous of Asia]. Pp. 63-68. In Akimov, I. A. (ed.): Novosti faunistiki i sistematiki. Sbornik nauchnyhk trudov [= News of faunistics and systematics. Collected scientific papers]. Naukova Dumka Publ., Kiev. [Text in Russian.] Makarkin, V. N. 1997 ["1996"]. Fossil Neuroptera of the Lower Cretaceous of Baisa, East Siberia. Part 5. Mantispidae. Russian Entomological Journal 5: 91-93. [Dating: From reverse of front wrapper of vol. 5, no. 1-4: "Printed in May 1987".] Martynov, A. V. 1936. Lacewings (Neuroptera). Pp. 437-438. In Zhivotnyi mir SSSR [= Animal kingdom of the USSR]. Izdatel'stvo Akademii Nauk SSSR, Moscow-Leningrad. Vol. 1. 806 pp. Text in Russian. Matsumura, S. 1907. Konchu Bunruigaku [or Konch-Bunrui-Gaku = Systematic Entomology]. Vol. 1. Keiseisha Co., Tokyo. 336 + 8 [index] pp. [Text in Japanese.] Matsumura, S. 1908. Catalogue of the beneficial insects of Japan. Rokumeikan Co., Tokyo. 174 pp. [Text in Japanese.] McLachlan, R. 1867 ["1868"]. New genera and species, andc., of neuropterous insects and a revision of Mr. F. Walker's British Museum Catalogue of Neuroptera, part ii. (1853), as far as the end of the genus Myrmeleon. Journal of the Linnean Society of London, Zoology 9: 230-281. McLachlan, R. 1875a. A sketch of our present knowledge of the neuropterous fauna of Japan (excluding Odonata and Trichoptera). Transactions of the Entomological Society of London 1875: 167-190. McLachlan, R. 1875b. Neuroptera s. str, Planipennia. Pp. 1-24. In A. P. Fedtschenko (ed.): Reise in Turkestan von Alexis Fedtschenko, auf Veranlassung des General-Gouverneurs von Turkestan, General von Kaufmann, hrsg. von der Gesellschaft der Freunde der Naturwissenschaften in Moskau, Vol. 2. pt. 5. Moskou. [Text in Russian]. McLachlan, R. 1900. A remarkable new mimetic species of Mantispa from Borneo. Entomologist's Monthly Magazine 36: 127-129. Miyake, T. 1910. The Mantispidae of Japan. Journal of the College of Agriculture, Tohoku Imperial University, Sapporo 2: 213-221. Monserrat, V. J. 1985. Lista de los tipos de Mecoptera y Neuroptera (Insecta) de la collección L. Navás, depositados en el Museo de Zoología de Barcelona. Miscellània Zooloògica 9: 233- 243. Monserrat, V. J. 1986. Longinos Navás, his neuropterological work and collection. Pp. 173-176. In J. Gepp, H. Aspöck, and H. Hölzel (eds): Recent Research in Neuropterology. Proceedings of the 2nd International Symposium on Neuropterology. Graz, Austria, 176 pp. Motschulsky, V. von. 1863. Essai d'un catalogue des insectes de l'ile Ceylan. Bulletin de la Société [Impériale] des Naturalistes de Moscou [= Byulleten' Muskovskogo obshchestva Ispytatelei Prirody] 36(3): 1-153. Nakahara, W. 1912. On the Japanese Mantispidae. Konchu Sekai [= Insect World] 16: 12-15. [Text in Japanese]. Ohl: World Catalog of Mantispidae 221

Nakahara, W. 1913. A revision of the Mantispidae of Japan. Annotationes Zoologicae Japonenses 8: 229-237. Nakahara, W. 1961. A new species of the Mantispidae from Japan (Neuroptera). Mushi 35: 63- 66. Navás, L. 1905. Notas zoológicas. VII. Insectos orientales nuevos ó poco conocidos. Boletín de la Sociedad Aragonesa de Ciencias Naturales 4: 49-55. Navás, L. 1906. Notas entomológicas. XIII. Sobre una pequeña colección de Neurópteros de Italia, reunida por el profesor Felipe Silvestri, de Portici. Boletín de la [Real] Sociedad Española de Historia Natural 6: 101-102. Navás, L. 1908 ["1907-1908"]. Neurópteros nuevos. Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 6: 401-423. Navás, L. 1909a ["1908-1909"]. Mantíspidos nuevos [I]. Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 7: 473-485. Navás, L. 1909b. Sobre la Mantispa luzonica (Neur.). Rectificación. Boletín de la Sociedad Aragonesa de Ciencias Naturales 8: 150. Navás, L. 1910. Neurópteros chilenos (1.a serie). Revista Chilena de Historia Natural 14: 235- 241. Navás, L. 1912a ["1911"]. Quelques Mantispides (Insectes Névroptères) du Musée Zoologique de l'Academie Impériale des Sciences de St. Pétersbourg. Akademiia nauk SSSR. Zoologicheskii Muzei. Ezhegodnik [= Annuaire du Musée Zoologique de l'Académie Impériale des Sciences, St. Pétersborg] 16: 535-538. Navás, L. 1912b. Neurópteros nuevos de América [Ia]. Brotéria (Zoológica) 10: 194-202. Navás, L. 1913a. Neuroptera asiatica. I series. Revue Russe d'Entomologie 13: 271-284. Navás, L. 1913b. Mis excursiones por el extranjero en el verano de 1912 (25 julio – 16 septiembre). Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 10: 479- 514. Navás, L. 1914a. Algunos Neurópteros del Museo de Oxford. I serie. Boletín de la Sociedad Aragonesa de Ciencias Naturales 13: 61-68. Navás, L. 1914b. Neuroptera nova africana. I series. Memorie dell'Accademia Pontifica dei Nuovi Lincei, Rome (1) 32: 81-90. Navás, L. 1914c. Neurópteros de Oceanía. Primera [I] serie. Revista de la Real Academia de Ciencias Exactas Fisicas y Naturales de Madrid 12: 464-483. Navás, L. 1914d. Neurópteros de Oceania. Segunda [II] serie. Revista de la Real Academia de Ciencias Exactas Fisicas y Naturales de Madrid 12: 645-653. Navás, L. 1914e. Neurópteros sudamericanos. Primera [I] serie. Brotéria (Zoológica) 12: 45-56, 215-234. Navás, L. 1914f. Notes sur quelques Névroptères du Congo Belge [I]. Revue de Zoologie Africaines 3: 365-377. Navás, L. 1914g. Neuropteros nuevos de Africa. Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 10: 627-653. Navás, L. 1914h. Névroptères nouveaux de l'Amerique du Nord. Iième série. Entomologische Zeitschrift, Frankfurt am Main 28: 18-20, 25-26. Navás, L. 1914i. Mantíspidos nuevos (Segunda [II] serie). Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 11: 83-103. Navás, L. 1914j. Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale (1911-1912). Résultats scientifiques. Insectes Névroptères. I. Planipennia et Mecoptera. Paris. 52 pp. 222 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Navás, L. 1914k. Neurópteros nuevos o poco conocidos (Segunda [II] serie). Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 11: 105-119. Navás, L. 1914l. Neurópteros nuevos o poco conocidos (Tercera [III] serie). Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 11: 193-215. Navás, L. 1915a. Neue Neuropteren. Zweite [II] Serie. Entomologische Mitteilungen 4: 194-202. Navás, L. 1915b. Neuroptera nova africana. IV Series. Memorie dell'Accademia Pontifica dei Nuovi Lincei, Rome (2) 1: 9-19. Navás, L. 1915c. Notes sur quelques Névroptères du Congo Belge. III. Revue de Zoologie Africaines 4: 172-182. Navás, L. 1915d. Neurópteros nuevos o poco conocidos (Sexta [VI] serie). Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 12: 119-136. Navás, L. 1916. Neurópteros nuevos o poco conocidos (Séptima [VII] serie). Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 12: 219-234. Navás, L. 1917. Neurópteros nuevos o poco conocidos (Novena [IX] serie). Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 13: 393-406. Navás, L. 1921. Algunos insectos de Santa Fe (República Argentina). Estudios. Revista Mensual (Academia literaria del Plata, Buenos Aires) 1921: 49-53. Navás, L. 1922. Insectos nuevos o poco conocidos [I]. Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 17: 383-400. Navás, L. 1923a. Estudis sobre Neuròpters (Insectes). Arxius de l'Institute [d'Estudis Catalans, Seccio] de Ciencias, Barcelona 7: 179-203. Navás, L. 1923b. Insecta nova. X Series. Memorie dell'Accademia Pontifica dei Nuovi Lincei, Rome (2) 6: 19-27. Navás, L. 1924. Insectos de la América Central. Brotéria (Zoológica) 21: 55-86. Navás, L. 1925a. Algunos insectos del Museo de París [Iib]. Brotéria (Zoológica) 22: 75-83. Navás, L. 1925b. Insectos neotrópicos. 1.a serie. Revista Chilena de Historia Natural 29: 305-313. Navás, L. 1925c. Névroptères nouveaux. Annales de la Société Scientifique de Bruxelles 44 (pt. 1): 566-573. Navás, L. 1926a ["1925"]. Insectos nuevos o poco conocidos. Verhandlungen des III. Internationalen Entomologen-Kongresses (Zürich 1925) 2: 458-461. Navás, L. 1926b. Algunos insectos del Brasil (3.a serie) [IIIb]. Brotéria (Zoológica) 23: 5-15. Navás, L. 1926c. Insectos exóticos Neurópteros y afines. Brotéria (Zoológica) 23: 79-93. Navás, L. 1927a. Insectos de la Somalia Italiana. Memorie della Società Entomologica Italiana 6: 85-89. Navás, L. 1927b. Neuropteren, Megalopteren, Plecopteren und Trichopteren aus dem Deutsch. Entomolog. Institut (Berlin-Dahlem). III serie. Entomologische Mitteilungen 16: 37-43. Navás, L. 1927c. Veinticinco formas nuevas de insectos. Boletín de la Sociedad Ibérica de Ciencias Naturales 26: 48-75. Navás, L. 1927d. Insectos del Museo de París. 4.a serie. Brotéria (Zoológica) 24: 5-33. Navás, L. 1927e ["1927-1928"]. Névroptères de la Chine. Arkiv för Zoologi 19A(18): 1-5. Navás, L. 1928a ["1927"]. Insectos neotropicos. 3.a serie. Revista Chilena de Historia Natural 31: 316-328. Navás, L. 1928b. Insectos del Museo de Estocolmo. Revista de la Real Academia de Ciencias Exactas Fisicas y Naturales de Madrid (2) 24: 28-39. Navás, L. 1928c. Insectos del Museo de Hamburgo. Primera [I] serie. Boletín de la Sociedad Entomologica de España 11: 59-67, 90-100, 121-138 [Errata: 165]. Ohl: World Catalog of Mantispidae 223

Navás, L. 1929a ["1928"]. Insectos neotropicos. 4.a serie. Revista Chilena de Historia Natural 32: 106-128. [Dating: from wrapper of vol. 32 "Febrero 1929".] Navás, L. 1929b. [Insecta orientalia.] VII Series. Memorie dell'Accademia Pontifica dei Nuovi Lincei, Rome (2) 12: 43-56. Navás, L. 1929c. Insectos exóticos Neurópteros y afines del Museo Civico de Génova. Annali del Museo Civico di Storia Naturale Giacomo Doria, Genoa 53: 354-389. Navás, L. 1929d. Insectes du Congo Belge (Série III). Revue de Zoologie et de Botanique Africaines 18: 92-112. Navás, L. 1929e. Insectos Neurópteros del Museo de Hamburgo. Memorias de la [Real] Sociedad Española de Historia Natural 15: 315-322. Navás, L. 1930a. Insecta nova. Series XV. Memorie dell'Accademia Pontifica dei Nuovi Lincei, Rome (2) 14: 409-418. Navás, L. 1930b. Insecta orientalia. VIII Series. Memorie dell'Accademia Pontifica dei Nuovi Lincei, Rome (2) 14: 419-434. Navás, L. 1930c. Insectos del Museo de París. 5.a série. Brotéria (Zoológica) 26: 5-24. Navás, L. 1930d. Insectos del Museo de París. 6.a série. Brotéria (Zoológica) 26: 120-144. Navás, L. 1930e. Insectos neotropicos. 6.a serie. Revista Chilena de Historia Natural 34: 62-75. Navás, L. 1930f. Insectos neotrópicos. 7.a serie [as "Sexta serie" (sic)]. Revista Chilena de Historia Natural 34: 299-307. Navás, L. 1930g. Spedizione di S. A. R. Il Duca degli Abruzzi alle sorgenti dell'Uebi Scebeli – Risultati Zoologici. Descrizione di una nuova Mantispilla. Annali del Museo Civico di Storia Naturale Giacomo Doria, Genoa 55: 27-28. Navás, L. 1931a ["1930"]. Comunicaciones entomológicas. 13. Insectos de la India. 3.a serie. Revista de la [Real] Academia de Ciencias Exactas Fisico-Quimicas y Naturales de Zaragoza (1) 14: 74-92. Navás, L. 1931b. Névroptères et insectes voisins. Chine et pays environnants. Deuxième [II] série. Notes d'Entomologie Chinoise 1 (7): 1-10. Navás, L. 1931c. Insectes du Congo Belge (Série VI). Revue de Zoologie et de Botanique Africaines 21: 123-144. Navás, L. 1931d. Mantispidae. in Résultats scientifiques du voyage aux Indes Orientales Néerlandaises de LL. AA. RR. le Prince et la Princesse Léopold de Belgique, V. Van Straelen, ed. Mémoires du Musée Royal d'Histoire Naturelle de Belgique, Hors Série 4(2): 7-10. Navás, L. 1932a. Décadas de insectos nuevos. Década 21. Brotéria (Ciências Naturais) 28: 109- 119. Navás, L. 1932b. Insectes du Congo Belge. Série VII. Revue de Zoologie et de Botanique Africaines 22: 269-290. Navás, L. 1933a. Insecta orientalia. XII series. Memorie dell'Accademia Pontifica dei Nuovi Lincei, Rome (2) 17: 75-108. Navás, L. 1933b. Insectos neotropicos. 8.a serie. Revista Chilena de Historia Natural 37: 194-197. Navás, L. 1933c. Insectos suramericanos. Septima [VII] serie. Revista de la Real Academia de Ciencias Exactas Fisicas y Naturales de Madrid 30: 303-314. Navás, L. 1933d. Neurópteros exóticos [1.a serie]. Memorias de la Real Academia de Ciencias y Artes de Barcelona (3) 23: 203-216. Navás, L. 1934a ["1933"]. Comunicaciones entomológicas. 17. Insectos de Madagascar. Primera [I] serie. Revista de la [Real] Academia de Ciencias Exactas Fisico-Quimicas y Naturales de Zaragoza (1) 17: 49-76. 224 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Navás, L. 1934b. Insectos suramericanos. Octava [VIII] serie. Revista de la Real Academia de Ciencias Exactas Fisicas y Naturales de Madrid (2) 31: 9-28. Navás, L. 1935 ["1934"]. Comunicaciones entomológicas. 18. Insectos de Madagascar. Segunda [II] serie. Revista de la [Real] Academia de Ciencias Exactas Fisico-Quimicas y Naturales de Zaragoza 18: 42-74. Navás, L. 1936a ["1935"]. Comunicaciones entomológicas. 19. Insectos de Madagascar. Trecera [III] serie. Revista de la [Real] Academia de Ciencias Exactas Fisico-Quimicas y Naturales de Zaragoza (1) 19: 100-110. Navás, L. 1936b. Insectos del Brasil. 5.a serie. Revista do Museu Paulista, Sao Paulo 20: 721- 734. Navás, L. 1936c. Insectes du Congo Belge. Série IX. Revue de Zoologie et de Botanique Africaines 28: 333-368. Neave, S. A. (ed.) 1939-1975. Nomenclator Zoologicus. 7 vols. Zoological Society of London, London. [Vol. 1 (1939), 957 pp.; Vol. 2 (1939), 1025 pp.; Vol. 3 (1940), 1065 pp.; Vol. 4 (1940), 758 pp.; Vol. 5 (1950), 308 pp.; Vol. 6 (1966), 329 pp., M. A. Edwards and A. T. Hopewood, eds.; Vol. 7 (1975), 374 pp., M. A. Edwards and H. G. Vevers, eds.] Nel, A. 1989 ["1988"]. Deux nouveaux Mantispidae (Planipennia) fossiles de l'Oligocene du sud- est de la France. Neuroptera International 5: 103-109. 5 figures. [Dating: From note in volume 5: 253.] New, T. R. 1996. Neuroptera. Pp. 1-104, 184 (Appendix III), 199-216 (Index). In Wells, A. (ed.). Zoological Catalogue of Australia. Vol. 28. Neuroptera, Strepsiptera, Mecoptera, Siphonaptera. CSIRO Publishing, Melbourne. xiii + 230 pp. New, T. R. 2003. The Neuroptera of Malesia. Fauna Malesiana, vol. 4. Brill, Leiden. viii + 204 pp. Newman, E. 1838. Entomological Notes. Entomological Magazine, London 5: 168-181, 372-402, 483-500. Ohl, M. 2002. Die Neuropterida-Sammlung des Museums für Naturkunde der Humboldt- Universität zu Berlin – Sammler, Forscher, Typen. Galathea 13, Supplement (Tagungsbericht, 6. Treffen deutschsprachiger Neuropterologen): 21-29. Ohl, M. 2004. A new wasp-mimicing species of the genus Euclimacia from Thailand (Neuroptera, Mantispidae). Denisia 13: 193-196. Okamoto, H. 1910. Homposan gitorokwa [= The Mantispidae of Japan]. Dobutsugakkai Zasshi [= Zoological Magazine, Tokyo] 22: 533-544. [Text in Japanese. German translation: Okamoto, 1911.] Okamoto, H. 1911. Beitrag zur Mantispiden-Fauna Japans. Zoologischer Anzeiger 37: 293-302. [German translation of Okamoto, 1910.] Olivier, G. A. 1797 ["1792"]. Mante [= Mantis]. Pp. 616-642. In Encyclopedie Méthodique. Histoire Naturelle, Insectes. Vol. 7. Paris. 8 figures. [Dating: Sherborn and Woodward (1906: 578): pp. 369-827 dated 1797, title page of volume 7 dated "M. DCC. XCII. [1792]".] Opler, P. A. 1981. Polymorphic mimicry of polistine wasps by a neotropical Neuropteran. Biotropica 13: 165-176. Oswald, J. D., and N. D. Penny. 1991. Genus-group names of the Neuroptera, Megaloptera and Raphidioptera of the world. Occasional Papers of the California Academy of Sciences 147: 1-94 Ohl: World Catalog of Mantispidae 225

Pallas, P. S. 1772. [Genus Mantis.] Pp. 12-15. In Spicilegia zoologica, quibus novae imprimis et obscurae animalium species iconibus, descriptionibus atque commentariis illustrantur, cura P. S. Pallas. Tome 1, Fasciculus 9. Berolini. 1 plate with 2 Neuroptera figures. Panfilov, D. V. 1980. Novye predstaviteli setcharokrylykh (Neuroptera) iz yury Karatau [= New representatives of lacewings (Neuroptera) from the Jurassic of Karatau]. Pp. 82-111. In V. G. Dolin, D. V. Panfilov, A. G. Ponomarenko, and L. N. Pritykina. Iskopaemye nasekomye mezozoya [= Fossil insects of the Mesozoic]. Akademiya Nauk Ukrainskoj SSR, Institut Zoologii, Naukova Dumka. Kiev. [Text in Russian. Dating: After information in Lambkin, 1986a: 16 (in footnote): publisher's supply invoice in Lambkin's copy of this work is dated 27 January 1980, which indicates that the book was in existence by that date.] Parker, F. D., and L. A. Stange. 1965. Systematic and biological notes on the tribe Platymantispini (Neuroptera: Mantispidae) and the description of a new species of Plega from Mexico. Canadian Entomologist 97: 604-612. Paulian, R. 1957. Les Mantispidae Malgaches (Neuroptera). Naturaliste Malgache 9: 75-81. Penny, N. D. 1977. Lista de Megaloptera, Neuroptera e Raphidioptera do México, América Central, ilhas Caraíbas e América do Sul. Acta Amazonica 7 (4) (Suplemento): 1-61. [Dating: As indicated in J. D. Oswald's internet Bibliography of Neuropterida, several of N. D. Penny's publications in the Acta Amazonica appear to have been published one or two years later than the year of publication as given on the title page of the respective volume. Oswald states that since he could not find the true dates of publication in the journals themselves, the dates of receipt of the volumes in some libraries might be an approximate indication of its publiction date. However, objective evidence for the delayed publication date is not known. Since none of these publications are critical with respect to priority in Mantispidae, I prefer to accept the year as printed on the title pages tentatively as the correct publication date.] Penny, N. D. 1982a. Neuroptera of the Amazon Basin. Part 6. Mantispidae. Acta Amazonica 12: 415-463. [Dating: See annotation for Penny, 1977.] Penny, N. D. 1982b. Review of the generic level classification of the New World Mantispidae (Neuroptera). Acta Amazonica 12: 209-223. [Dating: See annotation for Penny, 1977.] Penny, N. D. (ed.) 2002. A guide to the lacewings (Neuroptera) of Costa Rica. Proceedings of the California Academy of Sciences 53: 161-457. Penny, N. D., Adams, P. A., and L. A. Stange. 1997. Species catalog of the Neuroptera, Megaloptera, and Raphidioptera of America North of Mexico. Proceedings of the California Academy of Sciences 50: 39-114. Penny, N. D., and C. A. da Costa. 1983. Mantispídeos do Brasil (Neuroptera: Mantispidae). Acta Amazonica 13:601-687. [Dating: See annotation for Penny, 1977.] Pictet, A.-E. 1865. Synopsis des Névroptères d'Espagne. H. Georg, Genèva and J. B. Baillière and F. Savy, Paris. 123 pp. Poda, N. 1761. Insecta musei Graecensis, quae in ordines, genera et species juxta systema naturae Caroli Linnaei digessit. Graecii. 127 pp. Poivre, C. 1981 ["1980"]. Mantispides du Cameroun I. (Neuroptera, Planipennia). Neuroptera International 1: 77-89. Poivre, C. 1982a ["1981"]. Mantispides nouveaux d'Afrique et d'Europe (Neuroptera, Planipennia) (1re partie). Neuroptera International 1: 175-205. Poivre, C. 1982b. Mantispides nouveaux d'Afrique et d'Europe (Neuroptera, Planipennia) (seconde partie). Neuroptera International 2: 3-25. 226 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Poivre, C. 1982c. Les Mantispidés du Muséum d'Histoire naturelle de Genève. II. (Insecta, Planipennia). Revue Suisse de Zoologie 89: 667-672. Poivre, C. 1984a. Les mantispides de l'Institut Royal des Sciences Naturelles de Belgique (Insecta, Planipennia) 1re partie: especes d'Europe, d'Asie et d'Afrique. Neuroptera International 3: 23-32. Poivre, C. 1984b. Les Mantispidés du Muséum d'Histoire naturelle de Genève. III. (Insecta, Planipennia). Revue Suisse de Zoologie 91: 635-646. Provancher, L. 1875. Description de plusieurs insectes nouveaux. Naturaliste Canadien 7: 147- 251. Purvis, A., and A. Hector. 2000. Getting the measure of biodiversity. Nature 405: 212-219. Rafinesque, C. S. 1815. Analyse de la nature ou tableau de l'univers et des corps organisés. Palerme. 224 pp. Rambur, M. P. 1842. Histoire Naturelle des Insectes, Névroptères. Librairie encyclopédique de Roret. Fain et Thunot, Paris. [xviii] + 534 pp. 12 plates. Redborg, K. E. 1985. Spider (Arachnida: Araneae) boarding following egg sac penetration by Mantispa uhleri Banks (Neuroptera: Planipennia) with comments on the evolution of phoresy on spiders in the Mantispidae. Neuroptera International 3: 201-208. Redborg, K. E. 1998. Biology of the Mantispidae. Annual Review of Entomology 43: 175-194. Reeves, W. K., and B. A. Robinson. 1999. Type specimens in the Clemson University Collection. Transactions of the American Entomological Society 125: 459-466. Rehn, J. W. H. 1939. Studies in North American Mantispidae. Transactions of the American Entomological Society 65: 237-264. Rice, M. E. 1987. Morphological variation in Plega dactylota (Neuroptera: Mantispidae) and status of its subspecies. Journal of the Kansas Entomological Socitey 60: 341-344. Say, T. 1824. Order Neuroptera. Pp. 303-310. In W. H. Keating (ed.): Narrative of an expedition to the source of St. Peter's River, Lake Winnepeek, Lake of the Woods, andc. andc. performed in the year 1823, by order of the Hon. J. C. Calhoun, Secretary of War, under the command of Stephen H. Long, Major U.S.T.E., Vol. 2. Carey and Lea, Philadelphia. Say, T. 1825. Mantispa. Plate 25 and unpaginated letterpress [7 pp.]. In American entomology, or descriptions of the insects of North America. Vol. 2. Philadelphia. 1 plate with 7 figures. Schlüter, T. 1978. Zur Systematik und Palökologie harzkonservierter Arthropoda einer Taphozönose aus dem Cenomanium von NW-Frankreich. Berliner geowissenschaftliche Abhandlungen (A) 9: 150 pp. Schneider, W. G. 1843. Monographia generis Rhaphidiae Linnaei. Grassii, Barthii et Socii, Vratislaviae, 96 pp. Schröder, C. 1925. Überordnung: Neuropteroidea Handl. oder Planipennia (Banks) (Netzflügler), pp. 825-839. In Handbuch der Entomologie. Vol. 3. Gustav Fischer, Jena. Scopoli, J. A. 1763. Entomologia Carniolica, exhibens insecta Carnioliae indigena et distributa in ordines, genera, species, varietates, methodo Linneana. Trattner, Vindobonae. 420 pp. Sharp, D. 1901 ["1900"]. Neuroptera [sensu lato]. Zoological Record 37 (Insecta): 337-343. Shelford, R. [W. C.] 1903 ["1902"]. Observations on some mimetic insects and spiders from Borneo and Singapore, with appendices containing descriptions of new species by R. Shelford, Dr. Karl Jordan, C. J. Gahan, the Rev. H. S. Gorham, and Dr. A. Senna. Proc. Zool. Soc. Lond. 1902 (2): 230-284, plate XIX-XXIII [Dating: April 1903 as printed on the wrapper of part II of volume II (comprising pp. 225-483)]. Sherborn, C. D. 1899. Lichtenstein's 'Catalogus rerum naturalium'. The Annals and Magazine of Natural History (7) 3: 272. Ohl: World Catalog of Mantispidae 227

Sherborn, C. D., and B. B. Woodward. 1901. Notes on the dates of publication of the Natural History portions of some French Voyages. - Part I. 'Amérique méridionale'; 'Indes orientales'; 'Pôle Sud' ('Astrolabe' and 'Zélée'); 'La Bonite'; 'La Coquille'; and 'L'Uranie et Physicienne.' Annals and Magazine of Natural History (7) 7: 388-392. Sherborn, C. D., and B. B. Woodward. 1906. On the dates of publication of the natural history portions of the 'Encyclopédie Méthodique'. Annals and Magazine of Natural History (7) 17: 577-582. Stange, L. A. 1965. Systematic and biological notes on the tribe Platymantispini (Neuroptera: Mantispidae) and the description of a new species of Plega from Mexico. Canadian Entomologist 97: 604-612. Stange, L. A. 1967. Catalogo de Neuroptera de Argentina y Uruguay. Acta Zoologica Lilloana 22: 5-87. Stange, L. A., and Y.-H. Wang. 1998. Neuroptera, Megaloptera, Raphidioptera. Guide book to insects in Taiwan. 18. 278 pp. [Text in English and Chinese]. Stitz, H. 1913. Mantispiden der Sammlung des Berliner Museums. Mitteilungen aus dem Zoologischen Museum in Berlin 7: 1-49. Stoll, C. 1787 ["1787-1790"]. Natuurlyke en naar 't leeven naauwkeurig gekleurde Afbeeldingen en Beschryvingen der Spooken, wandelnde Bladen, Zabelspringhanen, Krekels, Trekspringhaanen en Kakkerlakken. In alle vier deelen der Waereld Europa, Asia, Afrika en America huishoudende, by een verzamelt en Beschreeven door Caspar Stoll. Representation exactement colorée d'après nature des Spectres, des Mantes, des Sauterelles, des Grillons, des Criquets et des Blattes. Qui se trouvent dans les quatre parties du monde, l'Europe, l'Asie, l'Afrique et l'Amerique; Rassemblées et décrites par Caspar Stoll. J. C. Sepp, Amsterdam, 20 pp. + pl. I-VI; 56 pp. + pl. Ia-VIa, pl. VII-XI, pl. XIII-XVIII. Mantispidae containing text and plates (probably published in 1787): p. 5, pl. I, fig. 3 ("Le Nain du Cap"), p. 7, pl. II, figs. 6-7 ("Le Nain d'Allemagna", "Le Nain de Suriname"), p. 12, pl. IV, fig. 15 ("Le Nain de Coromandel") [Pagination and plate numbers based on a copy in the ZMB library]. [Dating based on information by Hagen (1862-1863: Bd. 2, 199) and Horn and Schenkling (1929: 1193): Stoll himself published only 4 separately issued parts ("Lieferungen") of the Spectres (pl. I-XVIII) and 1 part of the Locustes (pl. Ia-VIa). They were published between 1787, as printed on the title page, and probably 1790. Houttuyn completed Stoll's work until 1813 (or 1815) after Stoll died in 1795 (cited here as Houttuyn in Stoll, 1813). There is no printed evidence for the exact publication date of the parts of this work. Handschin (1959b: 107) and H. Aspöck (1999: 217) concluded that 1790 is probably the correct date for the Mantispidae containing parts. However, the plates with mantispids are plates I, II, and IV out of 18 plates, which seems to be suggestive of an early date of publication (see also Weidner, 1983, and Boeseman & de Ligny, 2004). We tentatively accept 1787 as the probable date of publication of the mantispid parts. See notes for Houttuyn in Stoll, 1813.] Stork, N. E., and M. J. Samways. 1995. Inventorying and Monitoring. Pp. 453-543. In V. H. Heywood (ed.): Global Biodiversity Assessment. United Nations Environment Programme. Cambridge University Press, Cambridge, 1140 pp. Strand, E. 1942. Miscellanea nomenclatoria zoologica et palaeontologica. X. Folia Zoologica et Hydrobiologica 11: 386-402. Sziráki, G. 1997. Data to the coniopterygid fauna of Yemen, with description of twelve new species (Neuroptera: Coniopterygidae). Acta Zoologica Academiae Scientarum Hungaricae 43: 271-294. 228 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Sziráki, G., and L. Greve. 1996. Some Coniopterygidae (Neuroptera) from a mountain rainforest of Tanzania. Acta Zoologica Academiae Scientarum Hungaricae 42: 81-88. Taylor, C. 1862. An orthopterian defense. Harper's New Monthly Magazine 24: 490-496. Thunberg, C. P. 1784. Dissertatio entomologica novas insectorum species, sistens, cujus partem tertiam, à Carol. P. Thunberg, à et respondens David Lundahl. Edman, Upsaliae. Part 3: Pp. 53-68. Thunberg, C. P. 1815. Hemipterorum maxillosorum genera illustrata. Akademiia nauk SSSR. Memoirs (5) 5: 211-301. Tillyard, R. J. 1925 ["1925-1928"]. Odonata, Neuroptera and Trichoptera from Groote Eylandt, Gulf of Carpentaria. Records of the South Australian Museum 3: 41-44. Tjeder, B. 1963. A new Necyla from Uganda (Neur. Mantispidae). Opuscula Entomologica 28: 121-126. Van der Weele, H. W. 1909. Mecoptera and Planipennia of Insulinde. Notes from the Leyden Museum 31: 1-100. Viette, P. 1958. Mission du Museum dans les îles du Golfe de Guinée. Entomologie: XI. Planipennia Mantispidae. Bulletin de la Société Entomologique de France 63: 216-218 Walker, F. 1853. List [or Catalogue] of the specimens of neuropterous insects in the collection of the British Museum. Part II [= pp. 193-476]. – (Sialides – Nemopterides). British Museum [Natural History], London. Walker, F. 1860. Characters of undescribed Neuroptera in the collection of W. W. Saunders. Transactions of the Entomological Society of London (N.S.) 5: 176-199. Weidner, H. 1972. Die Entomologischen Sammlungen des Zoologischen Instituts und Zoologischen Museums der Universität Hamburg, VIII. Teil Insecta V. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 68: 107-134. Weidner, H. 1983. Schriften von Liebhaber-Entomologen im ausgehenden 18.und beginnenden 19. Jahrhundert im Archiv der entomologischen Sammlungen des Zoologischen Instituts und Zoologischen Museums der Universität Hamburg. Zweiter Teil. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 7 (117): 279-342. Westwood, J. O. 1847 ["1848"]. Order Neuroptera. Pp. 69-70. In The Cabinet of Oriental Entomology being a selection of the rarer and more beautiful species of insects, natives of India and the adjacent islands. The greater portion of which are now, for the first time, described and figured. Smith, London. 88 pp. [Dating: 1848 is printed on the title page, but J. D. Oswald states in the internet Bibliography of Neuropterida that Westwood's original plate was published on 1 December 1847, as indicated on lower left corner of pl. 34, and the associated text was assumed to have been issued concurrently.] Westwood, J. O. 1852. On the genus Mantispa, with descriptions of various new species. Transactions of the Entomological Society of London (N. S.) 1: 252-270. Westwood, J. O. 1867. Description of new Species of Mantispidae in the Oxford and British Museums. Transactions of the Entomological Society of London (3) 5: 501-508. Whalley, P. E. S. 1980. Neuroptera (Insecta) in amber from the Lower Cretaceous of Lebanon. Bulletin of the British Museum of Natural History (Geology) 33: 157-164. Whalley, P. E. S. 1983. Fera venatrix gen. and sp. n. (Mantispidae) from amber in Britain. Neuroptera International 2: 229-233. (published 28 December according to Engel, 2004) Williner, G. J., and N. A. Kormilev. 1958. Notas sobre Mantispidae neotropicales, I (Neuroptera). Revista de la Sociedad Entomologica Argentina 21: 1-18. Ohl: World Catalog of Mantispidae 229

Williner, G. J., and J. C. Mariluis, 1979 ["1978"]. Revision del genero Paramantispa Williner y Kormilev, 1958 (Neuroptera: Mantispidae). Revista de la Sociedad Entomologica Argentina 37:39-46. Willmann, R. 1990. The phylogenetic position of the Rhachiberothidae and the basal sister-group relationships within the Mantispidae (Neuroptera). Systematic Entomology 15: 253-265. Willmann, R. 1994. Die phylogenetische Position ursprünglicher Mantispidae (Insecta, Planipennia) aus dem Mesozoikum und Alt-Tertiär. Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 34: 177-203. Wilson, E. O. 2003. The encyclopedia of life. Trends in Ecology and Evolution 18: 77-80. Yang, C.-k. 1988. Neuroptera: Osmylidae, Dilaridae, Hemerobiidae, Chrysopidae, Mantispidae, Myrmeleontidae, Ascalaphidae, Corydalidae. Pp. 193-216. In Huang, F.-s., Wang, P.-y., Yin, W.-y., Yu, P.-y., Lee, T.-s., Yang, C.-k., and X.-j. Wang (eds.): Insects of Mt. Namjagbarwa region of Xizang. Science Press, Beijing. i-xii + 1-621 pp. [Text in Chinese]. Yang, C.-k. 1992. Neuroptera. Pp. 438-454. In Chen, S. (ed.). Insects of the Hengduan Mountains Region. Vol. 1. Science Press, Beijing, China. xii + 865 pp. [Text in Chinese]. Yang, C.-k. 1999. Neuroptera: Mantispidae. Pp. 132-140, 163-164. In Fauna of Insects. Fujian Province of China. Vol. 3. [Text in Chinese]. Yang, C.-k., and Y.-z. Peng. 1998. A new species of genus Sagittalata from Mt. Funiu (Neuroptera: Mantispidae). Fauna and Taxonomy of Insects in Henan 2: 62-63. Yang, X.-k., and H.-g. Sun (eds.). 1991. Catalogue of the insect type specimens preserved in the insect collections of the Institute of Zoology, Academia Sinica. Agriculture Press, Beijing. 164 pp. [Text in Chinese]. Yazaki, M. 1927. On two new species of Mantispidae from Japan. Science Report, Tohoku Imperial University, Sapporo 2: 361-366. Zimsen, E. 1964. The type material of I. C. Fabricius. Munksgaard, Copenhagen. 656 pp.

Internet Sources

A number of selected electronic sources that are freely available over the internet are listed below. These are sources of particular importance for the production of this catalog, but many more sources have been used for individual data items.

Bibliography of the Neuropterida (by John D. Oswald): http://entowww.tamu.edu/research/neuropterida/neur_bibliography/bibhome.html

Collection depositories of entomologists (in German) http://www.zalf.de/deid/verbleib.phtml

Entomologists of the World (biographies, collection depositories) (in German): http://www.zalf.de/deid/biograph.phtml

Geographic Names Database (worldwide): http://gnpswww.nima.mil/geonames/GNS/index.jsp

Index to the Neuropterida Species of the World (by John D. Oswald): http://entowww.tamu.edu/research/neuropterida/neur_sp_index/ins_search.html

230 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Insect and Spider Collections of the World Web Site (by Al Samuelson and Neal Evenhuis): http://hbs.bishopmuseum.org/codens/codens-r-us.html

Type Specimen Database (CAS) http://www.calacademy.org/research/entomology/types/insecta/mantisp.htm.

Type Specimen Database (MCZ) http://www.calacademy.org/research/entomology/typesdb/families.asp

Mantispidae of Madagascar http://www.calacademy.org/research/entomology/Entomology_Resources/Faunal_Projects/ madagascar_neuroptera/madanote8.htm

Ohl: World Catalog of Mantispidae 231

APPENDIX I. SPECIES DISTRIBUTION

The following table includes only species-group taxa (i.e., species and subspecies) that are available, valid and extant. Species treated in the catalogue as nomina nuda or not unequivocally placed to genus are excluded. Taxa occuring in more than one zoogeographical region are counted for all regions in which they are known to occur. Thus, the total number of species in a genus might be lower than the number of species summed across multiple regions (e.g., Plega: Total species=14; NE=6, NT=10, sum=16. This means that two species occur in both NE and NT). Total numbers in bold are absolute numbers of valid species per genus.

Total AF PA NE NT OR AU Symphrasinae Anchieta Navás 5 5 Plega Navás 14 6 10 Trichoscelia Westwood 13 13 Drepanicinae Ditaxis McLachlan 2 2 Drepanicus Blanchard 5 5 Gerstaeckerella Enderlein 6 6 Theristria Gerstaecker 24 24 Calomantispinae Calomantispa Banks 3 3 Nolima Navás 7 5 2

Mantispinae Asperala Lambkin 2 1 2 Austroclimaciella Handschin 9 4 7 Austromantispa Esben-Petersen 2 1 2 Buyda Navás 2 2 Campanacella Handschin 1 1 Campion Navás 10 10 Cercomantispa Handschin 16 16 Climaciella Enderlein 8 1 8 Dicromantispa Hoffman 5 2 4 Entanoneura Enderlein 7 1 3 3 Euclimacia Enderlein 31 4 27 4 Eumantispa Okamoto 12 4 9 2 Haematomantispa Hoffman 1 1 Leptomantispa Hoffman 3 1 3 Madantispa Fraser 3 3 Manega Navás 1 1 Mantispa Illiger 123 36 11 2 26 42 5 Mimetispa Handschin 1 1 Nampista Navás 1 1 Necyla Navás 15 3 5 8 2 Nivella Navás 1 1 Orientispa Poivre 12 1 12 Paramantispa Williner and Kormilev 3 3 Paulianella Handschin 2 2 Perlamantispa Handschin 10 9 1 Pseudoclimaciella Handschin 21 21 232 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Rectinerva Handschin 1 1 Sagittalata Handschin 10 7 1 2 Spaminta Lambkin 2 2 Stenomantispa Stitz 4 4 Toolida Lambkin 1 1 Tuberonotha Handschin 6 1 6 1 Xaviera Lambkin 1 1 1 Xeromantispa Hoffman 1 1 1 Zeugomantispa Hoffman 3 1 3 Total per geographic region 99 33 19 95 121 67 Total in Mantispidae 410 AF PA NE NT OR AU

APPENDIX II. NUMBER OF SPECIES-GROUP NAMES BY AUTHOR

This table lists the number of published names in the Mantispidae by author based on the present catalog. Author names are sorted alphabetically. "Valid names" and "Synonymized names" are interpreted as treated in this catalog. "Other names" are the remaining available names (i.e., replaced names, emendations) and nomina nuda, which are listed here for the purpose of estimating the total number of species-group names published in the Mantispidae. "Total number of published names" is the sum of the other three categories and includes all names published by a respective author, irrespective of whether these names are available or unavailable.

Author (1) (2) (3) (4) Valid names Synonymized Other names Total names number of published names (1) + (2) + (3) Alayo 2 2 U. Aspöck and H. Aspöck 1 1 Banks 17 10 2 29 Blanchard 1 1 Brauer 2 2 Charpentier 1 1 De Geer 1 1 Enderlein 11 1 12 Erichson 10 4 2 16 Esben-Petersen 13 3 16 Fabricius 1 1 2 Fairmaire 1 1 Fischer von Waldheim 1 1 2 Fraser 1 1 1 3 Gerstaecker 14 5 19 Gmelin 1 1 Guérin-Méneville 6 1 7 Hagen 3 2 5 Handschin 21 2 2 25 Hoffman 5 5 Ohl: World Catalog of Mantispidae 233

Houttuyn in Stoll 1 1 Kolbe 1 1 Kozhanchikov 1 1 2 Kuwayama 2 2 Lambkin 23 23 Lambkin and New 2 2 Latreille 1 1 2 Le Peletier and Audinet-Serville 1 1 Lestage 1 1 Lichtenstein 1 1 Linsley and MacSwain 2 2 Loew 1 1 2 Makarkin 1 1 Matsumura 1 1 2 McLachlan 4 4 Miyake 3 3 Motschulsky 1 1 Nakahara 2 1 3 Navás 135 43 7 185 Newman 1 1 Ohl 4 4 Okamoto 4 2 6 Olivier 1 1 2 Opler 5 5 Pallas 2 2 Parker and Stange 1 1 Paulian 1 1 2 Penny 7 7 Pictet 1 1 Poda 1 1 Poivre 13 1 14 Provancher 1 1 Rambur 1 1 2 Rehn 6 1 7 Say 2 2 Scopoli 1 1 Sharp 1 1 Stitz 29 16 45 Taylor 5 5 Thunberg 1 1 Tillyard 1 1 Tjeder 1 1 Van der Weele 1 1 2 Viette 1 1 Walker 4 1 5 Westwood 25 4 1 30 Williner and Kormilev 1 1 C.-k. Yang 16 16 C.-k. Yang and Peng 1 1 Yazaki 2 2 Total 410 115 36 561

234 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

INDEX

All taxon names used in the catalog are listed alphabetically, followed by the author of the name. The first letter of each supraspecific name is capitalized. Names in boldface are valid names. Other names are in Roman letters, but incorrect original and subsequent spellings are omitted in the Index. The name and author of species-group names is followed by the genus in which the name is now placed, and the author's name is in parentheses if he used a different genus originally. Endings of species-group names have been corrected to agree in gender with the genus to which they are presently referred, even if the names are considered to be invalid and if they are listed in the original combination in the catalog.

acra Whalley †, Paraberotha 211 asiatica Makarkin †, Gerstaeckerella 152 adelungi Navás, Mantispa 178 asiatica Yang, Sagittalata 202 africana (Esben-Petersen), Asperala Lambkin 159 Pseudoclimaciella 199 ata Yang, Sagittalata 202 agapeta (Navás), Mantispa 178 attenuata Lambkin, Theristria 153 alberti (Navás), Pseudoclimaciella 199 aurea (Taylor), Dicromantispa 168 albertisii Navás, Entanoneura 169 auriventris (Guérin-Méneville), Nampista alicante Banks, Mantispa 178 192 amabilis Gerstaecker, Mantispa 178 australasiae (Guérin-Méneville), Campion amapaensis Penny, Climaciella 166 162 ambonensis Ohl, Mantispa 178 austroafrica Poivre, Perlamantispa 197 ambusta (Erichson), Paramantispa 196 Austroclimaciella Handschin 159 amoenula (Gerstaecker), Trichoscelia 149 Austromantispa Esben-Petersen 161 Amycla Rafinesque 177 axillaris Navás, Mantispa 179 anae Penny, Trichoscelia 149 axillata Navás, Mantispa 190 Anchieta Navás 146 azihuna (Stitz), Mantispa 179 anchietai Navás, Gerstaeckerella 152 badia Okamoto, Euclimacia 171 Anisoptera Berthold 146 banksi Enderlein, Trichoscelia 149 Anisoptera Herrich-Schaffer 146 banksi Rehn, Plega 147 Anisoptera Schneider 146 basalis (Navás, 1927), Mantispa 179 Anisopterana Strand 146 basalis (Navás, 1929), Mantispa 178 annulata (Stitz), Xaviera 205 basalis Banks, Theristria 153 annulicornis Gerstaecker, Mantispa 178 basella (Westwood), Trichoscelia 149 antillesensis Hoffman, Leptomantispa 176 basiflava Handschin, Euclimacia 171 aphavexelte U. Aspöck and H. Aspöck, basilei (Navás), Mantispa 179 Mantispa 178 batesella (Westwood), Entanoneura 169 apicalis (Loew), Nampista 192 beardi Penny, Plega 147 apicata Navás, Buyda 161 bella (Kuwayama), Necyla 193 apicipennis (Kolbe), Pseudoclimaciella 199 bella (Westwood), Anchieta 146 arabica Navás, Necyla 193 Bellarminus Navás 157 araucariae Handschin, Eumantispa 174 bequaerti (Navás), Perlamantispa 197 areolaris (Westwood), Entanoneura 170 bicolor (Stitz), Mantispa 179 ariasi Penny, Mantispa 178 bicolor Poivre, Orientispa 195 aruntae Lambkin, Theristria 153 biseriata (Westwood), Ditaxis 151 Ohl: World Catalog of Mantispidae 235

bitschi Poivre, Sagittalata 202 christiana (Charpentier), Perlamantispa 197 boliviana (Navás), Mantispa 179 chrysopinus Brauer, Drepanicus 151 bonhourei (Navás), Mantispa 187 chrysops (Stitz), Campion 163 bouchardi (Navás), Tuberonotha 205 cincticornis (Banks), Dicromantispa 169 braconidiformis Handschin, Rectinerva 202 Climaciella Enderlein 166 brevicollis (Banks), Zeugomantispa 207 cognatella (Westwood), Plega 148 brevicornis (De Geer), Perlamantispa 198 commoni Lambkin, Theristria 154 brevistigma Yang, Mantispa 179 compellens (Walker), Zeugomantispa 206 brianti (Navás), Austroclimaciella 159 completa Banks, Mantispa 180 bruchi (Navás), Dicromantispa 168 condei Poivre, Cercomantispa 164 brunnea (Navás), Perlamantispa 198 confluens Navás, Mantispa 180 brunnea (Say), Climaciella 166 congensis (Navás), Pseudoclimaciella 199 brunneonigra Handschin, Entanoneura 170 coomani (Navás), Mantispa 180 Bucharispa Martynov 192 coorgensis Ohl, Mantispa 180 burgeoni (Navás), Perlamantispa 198 cora Newman, Mantispa 180 burmanella (Westwood), Euclimacia 171 cordieri (Navás), Mantispa 181 burquei (Provancher), Climaciella 166 coronata (Stitz), Pseudoclimaciella 199 Buyda Navás 161 coronata Yang, Orientispa 195 cachani Poivre, Pseudoclimaciella 199 costalis (Erichson), Entanoneura 170 calceata (Navás), Dicromantispa 168 cottami Navás, Euclimacia 171 callosus Lambkin, Campion 162 crenata Navás, Mantispa 181 Calomantispa Banks 157 cruciferus (Navás), Campion 163 Calomantispinae Navás 157 cubana Enderlein, Climaciella 167 Campanacella Handschin 162 dactylota Rehn, Plega 147 Campion Navás 162 debetazi (Navás), Theristria 155 campioni (Navás), Tuberonotha 205 debilis (Gerstaecker), Dicromantispa 168 canadensis Opler, Climaciella 167 decellei Poivre, Cercomantispa 164 capeneri Handschin, Mantispa 179 decepta Banks, Mantispa 181 cardaleae Lambkin, Theristria 154 decolor (Navás), Perlamantispa 197 carnifex Opler, Climaciella 167 decorata (Erichson), Paramantispa 196 castaneipennis Esben-Petersen, Mantispa decumana Hagen, Mantispa 181 179 delamarei Poivre, Sagittalata 202 caucasica Handschin, Mantispa 189 delicata (Navás), Mantispa 181 celebensis Enderlein, Mantispa 179 delicatula (Westwood), Theristria 154 celebica Handschin, Euclimacia 171 deliciosa (Navás), Mantispa 181 centenaria Esben-Petersen, Mantispa 180 denaria (Taylor), Climaciella 166 cercata (Navás), Cercomantispa 166 dentata Lambkin, Theristria 154 Cercomantispa Handschin 164 Dicromantispa Hoffman 168 ceylanica (Stitz), Mantispa 180 dimidiata Matsumura, Mantispa 181 chalybea (Erichson), Climaciella 167 diminuta Matsumura, Mantispa 181 chaos Hoffman in Penny, Leptomantispa dine Rehn, Nolima 157 176 discolor (Westwood), Theristria 154 chiaiensis Ohl, Necyla 193 dispersa (Navás), Mantispa 181 chilensis (Hagen), Gerstaeckerella 152 Ditaxis McLachlan 151 chlorodes (Navás), Mantispa 180 doddi (Navás), Austromantispa 161 chlorotica (Navás), Mantispa 180 dorsalis (Erichson), Perlamantispa 197 chopardi Navás, Entanoneura 169 Drepanicinae Enderlein 151 236 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Drepanicus Blanchard 151 flavinota Handschin, Mantispa 182 Drepaniscus Schröder 151 flavocincta Stitz, Euclimacia 171 duckei Navás, Climaciella 167 flavomaculata (Latreille), Zeugomantispa duckei Penny, Plega 147 206 egella (Westwood), Trichoscelia 149 flavonotata Tjeder, Necyla 193 elisabethae (Navás), Pseudoclimaciella 200 florealis Hagen, Mantispa 182 ellenbergeri (Navás), Mantispa 181 floridana Banks, Mantispa 182 elpidica (Navás), Mantispa 182 Forciada Kozhanchikov 192 enderleini Banks, Mantispa 182 formosana (Okamoto), Necyla 193 enigmatica Grimaldi †, Mantispidiptera fratercula Rehn, Plega 147 210 frontalis (Navás), Mantispa 182 Entanoneura Enderlein 169 fujiana Yang, Orientispa 195 erichsonii (Guérin-Méneville), fuliginosa Loew, Mantispa 183 Pseudoclimaciella 200 fulvicornis (Navás), Pseudoclimaciella 199 erythraea (Brauer), Asperala 159 fumosa Linsley and MacSwain, Plega 147 erythrocephalus Opler, Climaciella 167 fumosella (Westwood), Anchieta 146 Euclimacia Enderlein 170 Fusa Navás 152 Eumantispa Okamoto 174 fusca Stitz, Euclimacia 171 eurydella (Westwood), Anchieta 146 fuscata Navás, Eumantispa 175 eurysticta Gerstaecker, Theristria 154 fuscicolla Yang, Eumantispa 175 exigua Navás, Necyla 193 fuscicornis (Banks), Dicromantispa 169 extrema Navás, Necyla 193 fuscipennis Erichson, Mantispa 183 fasciata (Stitz), Austroclimaciella 160 gayi Blanchard in Gay, Drepanicus 151 fasciatella (Westwood), Plega 147 Gerstaeckerella Enderlein 152 fausta Thunberg, Mantis 208 gerstaeckeri Banks, Euclimacia 171 feae Navás, Entanoneura 170 gigantea Enderlein, Gerstaeckerella 152 feai Navás, Entanoneura 170 gillavryna (Navás), Mantispa 183 felina Gerstaecker, Theristria 154 gilva Lambkin, Theristria 154 femoralis Banks, Mantispa 180 girardi Poivre, Perlamantispa 197 femoralis Navás, Mantispa 182 gounellei (Navás), Dicromantispa 168 fenella (Westwood), Trichoscelia 149 gracilis (Erichson), Dicromantispa 168 fenestralis Navás, Mantispa 182 gracilis Rambur, Mantispa 184 ferdinandi (Navás), Mimetispa 192 gradata (Navás), Mantispa 183 ferrosa (Navás), Tuberonotha 205 grandis (Burmeister), Climaciella 168 ferruginea Stitz, Eumantispa 175 grandis (Erichson), Pseudoclimaciella 200 finoti (Navás, 1915), Cercomantispa 164 grandis (Guérin-Méneville in Duperrey), finoti Navás, 1909, Mantispa 182 Euclimacia 172 flava (Esben-Petersen), Pseudoclimaciella greeni Banks, Mantispa 183 200 griveaudi Poivre, Sagittalata 202 flavacoxa Yang, Orientispa 195 guerinii (Westwood), Euclimacia 172 flaveola (Erichson), Perlamantispa 197 gulosa (Taylor), Zeugomantispa 207 flavescens (Navás), Zeugomantispa 207 guttula Fairmaire in Thompson, Mantispa flavicauda (Navás), Mantispa 182 183 flavicauda Esben-Petersen, Euclimacia 171 habutsuella (Okamoto), Austroclimaciella flavicornis (Navás), Zeugomantispa 207 159 flavicostata Esben-Petersen, Euclimacia haematina (Navás), Mantispa 183 173 Haematomantispa Hoffman 176 Ohl: World Catalog of Mantispidae 237

hagenella (Westwood), Plega 148 kononenkoi Makarkin, Mantispa 190 hamiltonella (Westwood), Campanacella kroombitensis Lambkin and New, Campion 162 163 harmandi (Navás), Eumantispa 175 laciniata Stitz, Climaciella 166 haugi Navás, Mantispa 183 lacolombierei (Navás), Austroclimaciella hauseri Poivre, Mantispa 190 159 hemichroa (Navás), Asperala 159 latifascia McLachlan, Trichoscelia 149 hemichroa (Navás), Perlamantispa 197 latifrons Enderlein, Mantispa 184 henrotayi Nel †, Climaciella 167 leonina Navás, Gerstaeckerella 153 henryi Grimaldi †, Mantispidiptera 211 leopoldi (Lestage), Austroclimaciella 160 hilaris (Navás), Sagittalata 202 leopoldi Navás, Necyla 194 hillieri Navás, Theristria 154 Leptomantispa Hoffman 176 honesta (Navás), Cercomantispa 165 lilliputiana (Olivier), Zeugomantispa 206 horstaspoecki Ohl, Euclimacia 172 limbata (Gerstaecker), Entanoneura 169 hypogastrica Navás, Eumantispa 175 lineaticollis Enderlein, Mantispa 184 hypophoea (Navás), Perlamantispa 197 lineatifrons (Enderlein), Perlamantispa 198 icterica Pictet, Mantispa 183 lineolata Westwood, Mantispa 184 ilsae (Stitz), Stenomantispa 204 lipanica Rehn, Plega 147 imbecilla (Gerstaecker), Austromantispa loanga (Navás), Pseudoclimaciella 200 161 lobata Navás, Mantispa 185 immaculata (Stitz), Mantispa 183 lombokensis Handschin, Eumantispa 175 imperfecta Lambkin, Theristria 155 longyana Yang, Orientispa 195 implexa Navás, Gerstaeckerella 152 loveni (Navás), Mantispa 185 impressus (Navás), Campion 163 luddemanni (Navás), Manega 208 indica Westwood, Mantispa 184 luederwaldti Enderlein, Mantispa 185 infensa Navás, Nolima 158 lugubris (Navás), Sagittalata 203 infrequens Lambkin, Toolida 204 lurida Walker, Mantispa 185 insignis (Navás), Tuberonotha 205 lutea (Stitz), Mantispa 185 instabilis Opler, Climaciella 167 luzonensis Navás, Mantispa 185 interrupta (Say), Dicromantispa 168 luzonica (van der Weele), Austroclimaciella iridella (Westwood), Trichoscelia 149 160 iridipennis Guérin-Méneville, Mantispa luzonica Navás, Mantispa 185 184 machadoi (Handschin and Markl), irrorata (Erichson), Gerstaeckerella 153 Pseudoclimaciella 200 Isla Navás 162 maculata (Stitz), Austroclimaciella 160 ivoiriensis Poivre, Pseudoclimaciella 200 maculata Banks, Calomantispa 157 jacobsoni Handschin, Euclimacia 172 maculosa Lambkin, Theristria 155 januaria (Navás), Mantispa 184 Madantispa Fraser 177 japonica McLachlan, Mantispa 184 magna (Miyake), Tuberonotha 205 javanica Westwood, Mantispa 184 maindroni Navás, Mantispa 185 jocosa (Gerstaecker), Trichoscelia 149 major (Stitz), Necyla 193 jocosa Navás, Entanoneura 170 major (Stitz), Zeugomantispa 207 jucunda (Navás), Sagittalata 203 manca (Gerstaecker), Xaviera 205 jucunda Navás, Necyla 193 mandarina Navás, Mantispa 185 kantsi Rehn, Nolima 158 Manega Navás 208 keiseri Handschin, Cercomantispa 164 mantispa (Scopoli), Mantispa 190 kibumbana (Navás), Perlamantispa 197 Mantispa Illiger 177 238 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Mantispidiptera Grimaldi † 210 nanyukina (Navás), Mantispa 187 Mantispilla Enderlein 177 nassonovi (Navás), Trichoscelia 149 Mantispinae Leach 159 natalensis Navás, Necyla 194 margaritacea (Fischer von Waldheim), natalica (Navás), Cercomantispa 164 Zeugomantispa 207 navasi Handschin, Mantispa 187 mariae (Navás), Entanoneura 170 nawae (Miyake), Eumantispa 175 marshalli (Navás), Mantispa 186 ndjallai Poivre, Cercomantispa 164 maynei (Navás), Mantispa 186 necopina (Navás), Paulianella 196 meadewaldina (Navás), Mantispa 186 Necyla Navás 193 medialis Banks, Theristria 155 negusa Navás, Mantispa 187 melanocera (Navás), Mantispa 186 nelsoni Navás, Euclimacia 172 melitomae Linsley and MacSwain, Plega Neoclimaciella Handschin 196 148 neotropica Navás, Mantispa 187 meridiei Lambkin, Ditaxis 151 neptunica Navás, Mantispa 187 Mesomantispa Makarkin † 210 newmani (Banks), Mantispa 187 Mesomantispinae Makarkin † 210 nigra (Stitz), Necyla 194 metallica Esben-Petersen, Euclimacia 172 nigra Handschin, Euclimacia 172 militaris (Navás), Mantispa 186 nigra Lambkin, Theristria 155 mimetica (Sharp), Mimetispa 192 nigrata Banks, Calomantispa 157 Mimetispa Handschin 192 nigricornis (Enderlein), Mantispa 188 minjerribae Lambkin, Spaminta 203 nigricornis (Navás), Cercomantispa 165 minor (Stitz), Necyla 194 nigricornis (Stitz), Dicromantispa 168 minor Banks, Theristria 155 nigricoxa Yang, Orientispa 195 minuta (Fabricius), Zeugomantispa 206 Nivella Navás 208 minuta (Kozhanchikov), Mantispa 189 nobilis Navás, Anchieta 146 minuta Handschin, Madantispa 177 Nobrega Navás 166 mista (Navás), Dicromantispa 168 nodosa (Westwood), Euclimacia 172 miyakei (Okamoto), Austroclimaciella 160 Nolima Navás 157 moesta Hagen, Mantispa 186 notha (Erichson), Anchieta 146 Molinella Navás 151 nubeculosa (Navás), Haematomantispa 176 moluccensis Banks, Mantispa 186 nubila (Stitz), Mantispa 188 moluccensis Handschin, Eumantispa 175 nuchalis (Gerstaecker), Euclimacia 172 morosa (Gerstaecker), Euclimacia 172 nuda (Stitz), Dicromantispa 168 moucheti (Navás), Mantispa 186 nymphe Hoffman in Penny, Leptomantispa mouldsorum Lambkin, Theristria 155 177 moulti (Navás), Drepanicus 151 obscura Handschin, Cercomantispa 165 moulti Navás, Mantispa 186 obscurata (Navás), Mantispa 188 mozambica (Westwood), Cercomantispa obtusa Hoffman in Penny, Climaciella 167 164 occidentalis (Paulian), Pseudoclimaciella muhni (Navás), Paramantispa 196 200 mutata Motschulsky, Mantispa 187 occidentis (Banks), Climaciella 166 myrapetrella (Westwood), Trichoscelia 150 oligocenica Nel †, Prosagittalata 209 nabota Olivier, Mantis 208 olsufievi (Navás), Paulianella 196 Nampista Navás 192 ophryuta Yang, Orientispa 195 nana (Navás), Mantispa 188 orientalis (Esben-Petersen), Necyla 194 nana Erichson, Mantispa 187 Orientispa Poivre 194 nana Lichtenstein, Mantis 208 ornata (Stitz), Pseudoclimaciella 199 Ohl: World Catalog of Mantispidae 239

pagana (Fabricius), Mantispa 190 punctata (Stitz), Zeugomantispa 207 pallescens (Navás), Zeugomantispa 207 pupa Navás, Necyla 194 pallescens Stitz, Mantispa 188 pusilla (Pallas), Perlamantispa 198 pallida Lambkin, Theristria 155 pusilla Yang, Orientispa 195 paolina (Navás), Cercomantispa 165 pygmaea (Stitz), Mantispa 189 papuana (van der Weele), Xaviera 206 quadrituberculata (Westwood), Paraberotha Whalley † 211 Austroclimaciella 160 paraensis Penny, Plega 148 radialis (Navás), Mantispa 189 paraguayana Ohl, Mantispa 188 radiata (Navás), Mantispa 189 Paramantispa Williner and Kormilev 196 ragazziana (Navás), Pseudoclimaciella 200 partheniella (Westwood), Anchieta 147 Rectinerva Handschin 202 partita Enderlein, Euclimacia 173 regia (Navás), Tuberonotha 205 parvula Penny, Mantispa 188 regina Esben-Petersen, Euclimacia 173 pasteuri Navás, Mantispa 188 reinhardi (Stitz), Stenomantispa 204 Paulianella Handschin 196 relicta (Cockerell) †, Vectispa 210 pauliani Handschin, Madantispa 177 relicta (Kozhanchikov), Nampista 192 pavida (Gerstaecker), Spaminta 203 remipes (Gerstaecker), Trichoscelia 150 pehlkei Enderlein, Mantispa 188 Retinoberotha Schlüter † 211 perla (Pallas), Perlamantispa 197 rhombica Navás, Euclimacia 173 Perlamantispa Handschin 197 riedeliana (Fischer von Waldheim), perparva (Esben-Petersen), Cercomantispa Gerstaeckerella 153 165 rieki Lambkin, Theristria 155 personata (Stitz), Climaciella 167 rimata (Navás), Mantispa 189 phaeonota Navás, Mantispa 188 romani (Esben-Petersen), Anchieta 146 phthisica (Gerstaecker), Buyda 161 royi Poivre, Perlamantispa 198 picea (Esben-Petersen), Cercomantispa 165 rubella Navás, Nivella 208 picta Navás, Entanoneura 169 rubellus Navás, Campion 163 picta Stitz, Calomantispa 157 rubescens Stitz, Climaciella 166 pictiventris (Gerstaecker), Campion 162 rubicunda (Navás), Campion 163 pinal Rehn, Nolima 158 rubida (Stitz), Pseudoclimaciella 200 pirioni (Navás), Gerstaeckerella 152 rubricata (Navás), Zeugomantispa 207 platycephala (Stitz), Campion 163 rufa Esben-Petersen, Euclimacia 173 Platymantispa Rehn 146 rufescens Latreille, Mantispa 208 Plega Navás 147 ruficauda Enderlein, Euclimacia 173 plicicollis Handschin, Mantispa 189 rufipes (Tillyard), Spaminta 203 porosa Hoffman in Penny, Climaciella 167 rufocincta Handschin, Euclimacia 173 praeliator Navás, Nolima 158 rugicollis (Navás), Eumantispa 176 prasinus Esben-Petersen, Drepanicus 152 rugicollis (Stitz), Dicromantispa 168 prolixa (Erichson), Paramantispa 196 sacra Navás, Necyla 194 Promantispa Jarzembowski † 209 Sagittalata Handschin 202 Promantispa Panfilov † 209 salana (Navás), Mantispa 189 Prosagittalata Nel † 209 salonii Navás, Gerstaeckerella 152 Pseudoclimaciella Handschin 199 sanguinea (Navás), Pseudoclimaciella 201 pugnax (Navás), Nolima 158 sankitana (Navás), Perlamantispa 198 pulchella (Banks), Leptomantispa 177 santareni (Navás), Trichoscelia 150 pulla (Navás), Cercomantispa 165 sarta (Stitz), Pseudoclimaciella 201 pullula (Banks), Austromantispa 161 sasakii (Miyake), Eumantispa 175 240 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

satsumensis (Yazaki), Austroclimaciella strigipes (Westwood), Spaminta 204 160 stuermeri Schlüter †, Retinoberotha 211 sauteri Navás, Euclimacia 171 styriaca (Poda), Mantispa 190 sayi (Banks), Dicromantispa 169 subcostalis Navás, Mantispa 190 scabricollis McLachlan, Mantispa 189 subflava (Nakahara), Austroclimaciella 160 scabrosa (Banks), Xeromantispa 206 subfusca (Nakahara), Austroclimaciella 160 schajovskoyi Williner and Kormilev, sumatrana (Stitz), Necyla 194 Drepanicus 152 superba Lambkin, Euclimacia 173 schoutedeni (Navás), Mantispa 189 suzukii Okamoto, 1910, Eumantispa 175 scutellaris (Westwood), Spaminta 204 Symphrasinae Navás 146 semeriai Poivre, Sagittalata 203 Symphrasis Hagen 149 semifurva Yang, Orientispa 195 synapsis Hoffman in Penny, Dicromantispa semihyalina (Le Peletier de Saint-Fargeau 169 and Audinet-Serville), Climaciella 167 synoeca Opler, Climaciella 167 sequella (Westwood), Trichoscelia 150 taeniata Stitz, Eumantispa 176 seyrigi Fraser, Madantispa 177 tagalensis Banks, Euclimacia 173 shirozui (Nakahara), Orientispa 195 taina (Alayo), Mantispa 190 sibirica Gmelin, Mantis 209 taiwanensis Kuwayama, Eumantispa 176 sibirica Makarkin †, Mesomantispa 210 tanegashimensis (Yazaki), Austroclimaciella signata (Hagen), Plega 148 160 similata (Navás), Perlamantispa 198 taroom Lambkin and New, Theristria 156 similis Handschin, Entanoneura 170 taylori (Navás), Austroclimaciella 160 similis Panfilov in Dolin et al. †, tenella Erichson, Mantispa 190 Promantispa 209 tenera (Navás), Mantispa 190 simplex (Navás), Cercomantispa 166 tenuistriga Gerstaecker, Campion 163 simplex Stitz, Mantispa 190 tessmanni (Stitz), Mantispa 191 simplicia (Fraser), Cercomantispa 166 thaumasta (Navás), Trichoscelia 149 simulatrix (McLachlan), Mimetispa 192 Theristria Gerstaecker 153 sinica Yang, Entanoneura 170 Theristriella Esben-Petersen 153 smithersi Lambkin, Theristria 155 thomensis (Viette), Pseudoclimaciella 201 Spaminta Lambkin 203 tibetana Yang, Eumantispa 176 speciosa Navás, Nampista 192 tillyardi (Esben-Petersen), Campion 162 spectabilis Banks, Calomantispa 157 tillyardi Handschin, Theristria 156 spilonota Banks, Mantispa 190 timmerhansi (Navás), Pseudoclimaciella spiniferus Lambkin, Campion 163 201 Stenispa Baly 174 tincta (Navás), Perlamantispa 198 Stenispa Navás 174 tinctus (Navás), Climaciella 168 Stenomantispa Stitz 204 tobari (Navás), Trichoscelia 150 stenoptera Gerstaecker, Mantispa 190 tonkinensis Navás, Mantispa 191 stenostigma (Navás), Cercomantispa 165 Toolida Lambkin 204 stigma (Esben-Petersen), Theristria 156 torquata Navás, Euclimacia 174 stigmalis Banks, Theristria 156 torquilla Hagen, Mantispa 184 stigmata (Stitz), Zeugomantispa 207 transversa (Stitz), Mantispa 191 stitzi Handschin, Pseudoclimaciella 201 trevori Lambkin, Austromantispa 161 storeyi Lambkin, Theristria 156 triangularis Handschin, Euclimacia 174 strenua (Gerstaecker), Tuberonotha 205 Trichoscelia Westwood 148 striatella (Navás), Cercomantispa 165 trichostigma (Navás), Zeugomantispa 207 Ohl: World Catalog of Mantispidae 241

trifasciata (Stitz), Trichoscelia 150 vespiformis Okamoto, Euclimacia 174 trilineata (Navás), Dicromantispa 168 Veura Navás 153 trilineata Navás, Necyla 194 victor Navás, Nolima 158 tristella (Navás), Cercomantispa 165 victorii (Guérin-Méneville), Perlamantispa tristis (Navás), Cercomantispa 165 198 trivenata (Stitz), Pseudoclimaciella 201 virescens (Rambur), Zeugomantispa 207 tropica (Westwood), Pseudoclimaciella 201 viridata (Navás), Zeugomantispa 207 Tuberonotha Handschin 204 viridis (Stitz), Zeugomantispa 207 uhleri (Banks), Dicromantispa 169 viridis (Walker), Zeugomantispa 207 umbra (Paulian), Pseudoclimaciella 201 viridula (Erichson), Zeugomantispa 207 umbripennis (Navás), Mantispa 187 viridula (Houttuyn in Stoll), Zeugomantispa umbripennis (Navás), Pseudoclimaciella 206 201 vittatus (Guérin-Méneville), Campion 164 umbripennis Walker, Mantispa 191 vulpes (Stitz), Cercomantispa 166 unicolor Stitz, Climaciella 166 wagneri (Navás), Paramantispa 196 uniformis (Navás), Zeugomantispa 206 weelei Handschin, Austroclimaciella 161 uptoni Lambkin, Theristria 156 Whalfera Engel † 210 usambarica (Fraser), Mantispa 191 woodhousei Navás, Euclimacia 174 varia (Erichson), Climaciella 166 Xaviera Lambkin 205 varia (Walker), Trichoscelia 150 Xeromantispa Hoffman 206 variegata Navás, Plega 148 xuthoraca Yang, Orientispa 195 variolosa Navás, Mantispa 191 yuata Yang and Peng, Sagittalata 203 vassei (Navás), Perlamantispa 198 yucatanae Parker and Stange, Plega 148 Vectispa Lambkin † 209 zayasi (Alayo), Mantispa 191 venatrix (Whalley) †, Whalfera 210 Zeugomantispa Hoffman 206 venulosa (Navás), Mantispa 191 zikani Navás, Plega 148 venusta Lambkin, Calomantispa 157 zonalis Navás, Euclimacia 174 verendus (Navás), Campion 162 zonaria Navás, Mantispa 191 verruculata (Navás), Mantispa 191 zonata Navás, Mantispa 192 verticalis (Banks), Spaminta 203

242 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

ANNOTATIONS

1 Some of the Mantispidae types in the ANIC are labelled as MCZ types, including a MCZ type number. These specimens have been transferred from the MCZ to the ANIC by H. E. Evans in the 1970s, and no type specimens of these species are retained in the MCZ (P. D. Perkins, pers. comm. to MO, 2004). 2 Whalley (1980) studied Lebanese amber inclusions from the collection of Professor Aftim Acra, Beirut, Lebanon, which were temporarily stored in the BMNH when examind by Whalley. 3 The types of Mantispidae described by Enderlein (1910) were originally deposited in the Natural History Museum in Stettin (= Szezcyn), Poland. The parts of the Stettin collections that survived World War II are now housed in the MZPW (Horn et al., 1990). Presence of the Enderlein types in the MZPW, except for the type of M. pehlkei, which is likely to have been destroyed in World War II, was recently confirmed by W. Tomaszewska (pers. comm. to MO, Jan. 2004). 4 The entire holometabolous collection, including catalogs, of the Hamburg museum (ZMUH) was destroyed during an air raid on 30 July 1943, except for some material on loan at that time and some specimens in alcohol. According to Weidner (1972), there is no indication that any Neuroptera types survived World War II. Thus, all type material of Mantispidae recorded from the ZMUH must be regarded as lost. 5 The species epithet of Plega paraense was explicitly based on the Brazilian province Pará, the type locality. Since Plega is a feminin noun, the proper ending is –ensis, and the name needs a mandatory change to paraensis under ICZN Art. 34.2 and 31.2. 6 Monserrat (1985: 241) listed a specimen of Symphrasis signata in the MZBS as a dubious type from Navás' personal collection. However, Hagen (1877: 210) based the original description of this species explicitly on a single female from Fort Tejon, California, which is housed in the MCZ. 7 Penny (1982a: 438) considered the possibility that P. zikani might be a darkly pigmented form of Trichoscelia varia, without having seen the holotype of P. zikani. Penny and de Costa (1983) placed P. zikani in Trichoscelia as a distinct species. However, P. zikani has a subbasal spine on the forefemur and a curving forewing second radial cell, which is a diagnostic combination of characters for Plega. The flagellomeres of the holotype of P. zikani are almost as long as wide, which places it in the P. melitomae species group, where it probably represents a distinct species. Unfortunately, because the holotype is a female (according to the original description; tip of abdomen including ovipositor now lacking) the exact identification of this taxon requires further study. 8 Penny (1982a: 429) stated that the "holotype" of M. egella was present in the BMNH. However, Westwood (1867: 502) based the description of M. egella explicitly on two males, without designating a holotype. Penny's holotype citation is not a valid lectotype designation because it does not identify a single specimen from the syntypical series to be the name-bearing type (ICZN Art. 74.5). Paul A. Brown (pers. comm. to MO, March 2004) confirmed the presence of two syntypes of M. egella in the BMNH.

Ohl: World Catalog of Mantispidae 243

9 The holotype of Anisoptera jocosa Gerstaecker is supposed to be in the Greifswald collection (EMAU), but it could not be located by J.-C. Kornmilch in 2001 (pers. comm. to MO). Since there is no indication that the specimen is on loan, it must be tentatively regarded as lost. 10 Penny (1982a: 422) placed S. trifasciata in Plega, but the lack of a subbasal spine on the forefemur of the holotype clearly places it in Trichoscelia. 11The type material of species described by F. M. Brauer can be expected to be housed in the NHMW (Horn et al., 1990). Ulrike Aspöck (pers. comm. to MO, Nov. 2003) informed one of us (MO) that there is a single specimen of D. chrysopinus in the NHMW, which, however, does not bear an explicit type label. The status of this specimen needs to be clarified. 12 Williner and Kormilev (1958: 16) listed D. chrysopinus as a synonym of D. gayi. In the original description of D. chrysopinus, Brauer (1867a: 284) also noted that it might be the male of D. gayi. We follow Penny (1982b: 211) in treating D. chrysopinus as a valid species. 13 The repository of the types of Drepanicus gayi Blanchard in Gay has not been confirmed, but other neuropteran materials collected by Claude Gay and described by Charles Emile Blanchard in Gay's "Historia Fisica y Politica de Chile" are known to be present in the MNHN (Horn et al., 1990). 14 The holotype of D. prasinus bears a label reading "Nd. Cal." (= "Nieder-Kalifornien"), which is German for Baja California. The type locality of D. prasinus as "Greenville, California" (as cited by Esben-Petersen (1912: 272) in the original description) is in all probability incorrect (this species is not known to occur in America north of Mexico) and probably resulted from a mislabelling or misinterpreted locality label. Lambkin (1986a: 18) stated: "Lionel Stange (pers. comm. to MO) has informed me that the Californian D. prasinus was actually based on a Chilean specimen whose label had been mixed with that of a Californian snake fly". We were unable to confirm this statement, and Lionel Stange has informed one of us (MO) that it was probably an opinion arrived at without having seen the actual specimen (pers. comm. to MO Sept 2003). However, the species is quite obviously southern South American in origin, though its precise type locality remains unclear. 15 The MCZ currently has one syntype specimen of M. chilensis, but Hagen explicitly mentioned two syntypes: "[sent to me by] Dohrn, … [from] Dr. Caesar in Bremen". The whereabouts of the second syntype is unknown to us. 16 Navás did not explicitly mention the depository of the type(s) of Fusa pirioni, but the introduction to the publication in which it was described suggests that material collected by Anastasio Pirión might be in either Pirión's or Navás' personal collection, which is probably now in MZBS. The type of another species described in this work was deposited in Navás personal collection. 17 Raphidia riedeliana was synonymized with Gerstaeckerella irrorata (as Mantispa irrorata) by Hagen (1861: 322), but was subsequently considered to be a valid species (Hagen, 1866: 375, 428). The possible synonymy of R. riedeliana with G. irrorata implies affinities to Gerstaeckerella. Additionally, Hagen (1866: 375) stated: "The genus Mantispa includes heterogeneous material, which will subsequently divided in different genera. Particularly different are M. riedeliana and chilensis" (my translation). Fischer von Waldheim's (1834)

244 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

original description and the illustration also imply placement of R. riedeliana in Gerstaeckerella. 18 In the original description of Theristria basalis, Banks (1939: 475) explicitly mentioned 10 syntypes in the MCZ. Lambkin (1986a: 56) listed one presumed "holotype" and four additional "paratypes" in the ANIC. Since there are no syntypes left in the MCZ (P.D. Perkins, pers. com. 2004), the location of the remaining 5 syntypes remains unknown. Lambkin's "holotype" citation is deemed here to constitute a valid lectotype designation, because Lambkin clearly indicated that Banks based his description on more than one specimen, and because Lambkin explicitly selected one particular specimen of the type series to be the name-bearing type (ICZN Art. 74.5). 19 Lambkin (1986a: 62) cited the "holotype" of M. delicatula to be in the OUM. He also correctly pointed out that Westwood (1852) based the original description of this species explicitly on multiple specimens. Thus, as Westwood did not explicitly designate a primary type, the OUM specimen cannot be a holotype. Lambkin also mentioned another specimen in the OUM, which is labelled "delicatula Westw. var.". Lambkin was informed by the OUM that this specimen "has never stood with the type specimen and [that] there is no reason to suppose it is a syntype". However, Westwood explicitly mentioned specimens of the newly described M. delicatula that vary in some respects. It seems quite possible that this specimen might have been labelled accordingly as a variety, and, from available information, it cannot be rigorously concluded that it was not part of the original syntype series. There is also a specimen in the MCZ that is labelled as a type of M. delicatula, probably by Hagen. This specimen may be another syntype. In any case, Lambkin's "holotype" citation is deemed here to be a valid lectotype designation under ICZN Art. 74.5, because he clearly indicated that Westwood based his description on more than one specimen, and because he explicitly selected a particular specimen (the only OUM specimen of M. delicatula with a type label) of the type series to be the name-bearing type. 20 The holotype of Theristria hillieri Navás is a female according to the original description, but a male according to Lambkin (1986a: 70). 21 Lambkin (1986b: 5) listed holotypes and paratypes of C. spectabilis. However, the original description of C. spectabilis is explicitly based on multiple syntypes, and none was selected as a primary type by Banks. Lambkin's "holotype" citation is deemed here to constitute a valid lectotype designation under ICZN Art. 74.5, because he clearly indicated that Banks based his description on more than one specimen, and because he explicitly selected a particular specimen of the type series to be the name-bearing type. One paralectotype is in the BMNH (P. A. Brown, pers. comm, March 2004). 22 The syntypes of Mantispa erythraea are considered here to be destroyed (see annotation under ZMUH), but Esben-Petersen (1917: 218) examined them in the Hamburg museum before their loss and provided a photograph of one of them. 23 Handschin (1961:286) gave a list of six species that, at Handschin's time, have been considered to belong in Climaciella. At the end of Handschin's text that followed this list, he concluded that "these species […] exclusively occur in the eastern hemisphere and should, thus, be named Austroclimaciella" (Handschin, 1961: 287). We interprete this statement as an implicite new

Ohl: World Catalog of Mantispidae 245

generic placement of C. brianti in Austroclimaciella, although Handschin has never proposed this combination explicitly. 24 Climaciella habutsuella was synonymized with C. quadrituberculata by Kuwayama (1962: 379), but was listed as a valid species by New (2003: 153). 25 Yang and Sun (1991: 68) mentioned a holotype of A. lacolombierei in the IZAS, but the holotype status of this specimen, which was originally deposited in the "Mus. Heude, Shanghai" (Navás, 1931b), needs confirmation. 26 The original spelling of this species, "Mantispa 4-tuberculata", is an incorrect original spelling and has been corrected to M. quadrituberculata, a mandatory change under ICZN Art. 32.5.2.6. 27 Climaciella subflava Nakahara is considered here to be an unavailable name, because it was published before 1931 and was not accompanied by a description, a definition, or an indication (ICZN Art. 12.1.). It is a possible a misspelling of Climaciella subfusca Nakahara. 28 The holotype of B. apicata bears no information about its geographic origin, but Navás presumed that it was from central Africa because of its similarity to Pseudoclimaciella tropica and other African Mantispidae. However, in the catalog of Navás types in the MZBS, Monserrat (1985: 241) gave Costa Rica as type locality. Hoffman (1992) confirmed the strictly New World distribution of Buyda. 29 Oswald and Penny (1991) mentioned that they had been unable to critically determine the priority of the synonymous names Campion and Isla. They listed Campion as the senior synonym following Lambkin (1986b: 10). New evidence demonstrates conclusively that Campion, published in Navás 1914a, predates Isla, published in Navás 1914d. The serial issue containing Navás 1914a – Boletín de la Sociedad Aragonesa de Ciencias Naturales, vol. 13, no. 3 – bears the imprint date "Marzo 1914." The copy of this serial issue in the library of the National Museum of Natural History (Washington, D.C., USA) bears a Smithsonian Institution date of receipt stamp of 27 April 1914, demonstrating its physical existence by at least that date. Navás (1914d) concludes with the text "Zaragoza, 30 de Abril de 1914". Whether this date is considered to be the date on which Navás 1914d was completed, submitted or published, it could not have been published any earlier than this date, which postdates the known existence date of Navás 1914a 30 The holotype (or syntypes) of Mantispa australasiae could also not be located by Lambkin (1986b: 15). Most of Guérin-Menéville's material is known to be housed in the MNHN, but Horn et al. (1990) and Evenhuis (1997) listed additional museum collections where Guérin- Menéville materials are known to be deposited. The sex of the type material was not indicated in the original description but is considered to be of the male sex according to New (1996: 29). We could not find any printed information whether the type material of M. australasiae has been examined after the original description. 31 Esben-Petersen based the description of M. tillyardi on two specimens, a male (Western Australia, WAM) and a female (Victoria, ZMUC), without assigning either of them a particular type status. Lambkin (1986b: 16) incorrectly listed the male as the "holotype" and the female as the "paratype". Lambkin's "holotype" citation is deemed here to constitute a valid lectotype designation under ICZN Art. 74.5, because he clearly indicated that Esben-Petersen based his

246 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

description on more than one specimen, and because he explicitly selected a particular specimen (the male from the WAM) of the type series to be the name-bearing type. 32 Lambkin (1986b: 54) stated that Mantispa chrysops is either Campion callosus or C. impressus, which implies placement in Campion. The type has lost its taxonomically distinctive color and lacks the abdomen, so its definitive placement cannot be established on these bases. We follow Lambkin in treating this species as a nomen dubium in Campion. 33 Navás (1914a: 62) explicitly based the description of Mantispa crucifera on multiple specimens of unknown sex, and did not explicitly indicate a primary type. These specimens were listed as the "holotype" and three "paratypes" (one female, two males) by Lambkin (1986b: 22). Lambkin's "holotype" citatoin is deemed here to constitute a valid lectotype designation under ICZN Art. 74.5, because he clearly indicated that Navás based his description on more than one specimen, and because he explicitly selected a particular specimen of the type series to be the name-bearing type. Lambkin (1986b: 22) considered the female paralectotype of M. crucifera to be misidentified and included it as a paratype in the type series of Campion callosus Lambkin. 34 The taxonomic identity of C. vittata cannot be established with certainty, because the current depository of its type, or types, is unknown. Published references refer this taxon to various species now placed in Campion. Lambkin (1986b: 54) pointed out that "it is almost certainly" one of five species of Campion and treated it as a nomen dubium, which we follow here. Lambkin also critically discussed the potential type depository and the taxonomic placement. 35 The holotype of Cercomantispa decellei was originally in Poivre's private collection, which is now in the MNHN. 36 Westwood gave "Mus. D. Miers" as the type depository of M. mozambica. The private collection of Miers was completely transferred to the OUM in 1880, but the type of M. mozambica could not be located in Oxford recently (J. Hogan, pers. comm. to MO, Febr. 2004). Fraser (1955: 21) listed specimens from Ruanda as M. usambarica and attributed the authorship to Westwood. To the best of my knowledge, no such species was ever published by Westwood, and it is possible that M. usambarica is a gross misspelling of M. mozambica. However, since M. usambarica is a species name significantly different from M. mozambica, it is listed here as a nomen nudum in Mantispa. 37 In the original description of Necyla paolina, Navás stated that the the material upon which it was based was in the "Mus. Observat. Phytopatolog. Chiavari" (= Museo degli Studi Fitopatologici di Chiavari). This museum is now an archaeological collection (Barbara Agabiti, pers. comm. to MO, Febr. 2004). The former Paoli collection was divided into three parts, which were transferred to the natural history museums in Genova, Milano, and Firenze. The African collection of Paoli is apparently in Genova (MCSN), where the type(s) of N. paolina should be sought. 38 Cercomantispa perparva var. obscura Handschin 1963 ["1962"] is an unavailable name because it was published explicitly as a variety after 1960 (ICZN Art. 15.2). 39 The holotype of Mantispilla vulpes was originally deposited in the ZMB, but only the pinned original labels are currently present. In the card file of the ZMB Neuroptera collection there is a

Ohl: World Catalog of Mantispidae 247

note with the following information written by R. Schulze, a former ZMB technician (translated from German): "according to a note by Dr. Winkler on the loan form: specimen destroyed, on 31 May 1959 received back from a loan to Prof. Handschin, Basel. R. Schulze, 1981". Thus, the holotype of M. vulpes must be regarded as destroyed. 40 Opler (1981) described five color "morphs" of C. brunnea: "instabilis", "canadensis", "carnifex", "synoeca", and "erythrocephalus". These names are unavailable as infrasubspecific "morphs" (ICZN Arts. 1.3.4 and 45.6.2). The Climaciella brunnea "instabilis morph" was subsequently used as "C. brunnea var. instabilis" by Boyden (1983: 508). This usage is unavailable as an infrasubspecific "variety" (ICZN Art. 15.2). 41 Westwood (1852: 253) was the first to discuss the confusing nomenclature of Climaciella semihyalina and gave a list of synonyms known to him. He included "Mantispa grandis Burmeister" in this list, which he apparently interpreted as a valid species different from the homonymous M. grandis Guérin-Méneville and M. grandis Erichson. The original publication of Burmeister (1839: 967) clearly shows that Burmeister referred to M. grandis Guérin- Méneville and that it was not his intent to introduce a new name. Consequently, M. grandis is not available with Burmeister as author. Subsequent uses of "grandis Burmeister" by Westwood (1852: 253) and Penny (1982a: 453) as synonyms of Mantispa semihyalina and Climaciella semihyalina, respectively, are not sufficient to make this name available. 42 The holotype of Mantispilla debilis nuda Stitz, 1912, has been examined by one of us (MO) and shows a color pattern that is typical for the genus Dicromantispa and the species D. debilis, respectively (Hoffman in Penny, 2002: 259). Thus, M. debilis nuda is considered synonymous with D. debilis. 43 The original description of Mantispa gracilis Erichson (1839) does not explicitly record the number of specimens he studied. However, the description gives only one definitive length ("5 lin[es]."), instead of a length range, which might imply that the type series consisted of only a single specimen. This conclusion is supported by the accession number 208 on the label of the ZMB specimen, which refers to the historical ZMB accession catalog, where only one specimen (marked with an asterisk to indicate its type status) is listed. There is another specimen in the MCZ labelled as the holotype of M. gracilis, which probably has no type status at all. 44 The holotypes of Mantispilla debilis nigricornis Stitz and Mantispilla debilis rugicollis Stitz have been examined by one of us (MO). The distinctive forewing coloration (Sc distinctly lighter than costal crossveins; Hoffman in Penny, 2002: 259) is identical with that in Dicromantispa gracilis (Erichson). Since no further differences could be found, M. debilis nigricornis and M. debilis rugicollis is considered synonymous with D. gracilis. 45 For synonymy of Mantispilla debilis rugicollis Stitz with Dicromantispa debilis see annotation at Mantispilla debilis nigricornis Stitz. 46 Available information on the depository of the type of Mantispa bruchi is contradictory. Navás (1915d:135) indicates that it is housed in the MLPA. Williner and Kormilev (1958: 7) mentioned a specimen in the MACN that they considered to be "the type". Penny (1982a: 439) examined a putative holotype at the MNHN.

248 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

47 Navás (1930c: 24) explicitly changed the spelling of "Entatoneura [sic] albertisii" to "E. albertisi". Although demonstrably intentional under ICZN Art. 33.2, the altered spelling "albertisi" is an unavailable incorrect subsequent spelling under ICZN Art. 33.4. 48 In the original description of Entanoneura similis, Handschin explicitly mentioned two specimens, neither of which was designated as the holotype in the text. However, a holotype was designated in the legend to Plate 1, Fig. 2. 49 Navás (1930c: 24) explicitly changed the original spelling "Entatoneura [sic, for Entanoneura] feae" to "E. feai", probably because he considered "E. feae", which is based on the name of a man (Leonardo Fea), to be incorrectly latinized. The corrected name Entanoneura feai is a demonstrably intentional change of Entanoneura feae under ICZN Art. 33.2 and is separately available as an unjustified emendation (incorrect latinization is not justifiable grounds for an emendation, ICZN Art. 32.5.1). 50 One of us (MO) examined about 20 specimens of Euclimacia badia from Taiwan, most of which were identified by Navás, Stitz, and Esben-Petersen. The overall coloration is reddish- brown, with a varying amount of black, ranging from almost completely reddish to an almost completely black thoracic and abdominal venter. These two extremes are linked by a range of intermediates, and the almost complete range from all reddish to largely black can be found on Taiwan. The holotype of Euclimacia sauteri, also from Taiwan, apparently represents the almost reddish phenon, and, as a consequence, E. sauteri is synonymized here with E. badia. The holotype of E. fusca Stitz from Taiwan also resembles E. badia and might be a color variety of E. badia (see notes on E. fusca below). 51 The holotype of Euclimacia fusca Stitz is quite similar to E. badia Okamoto, and Stange and Wang (1998: 94) suspected that the former might be a color variant of the latter (although the specimen they illustrated as E. fusca (Fig. 42) is very unlikely to be this species). Euclimacia badia shows a broad range of intraspecific variation in overall body color (see notes on E. badia above). However, the holotype of E. fusca is characterized by an unusual combination of color characters, which is apparently not in the phenotypical range of E. badia that one us (MO) could observe. For example, the antennae are uniformly orange in E. fusca but largely dark brown to black with a orange tip in E. badia. Additionally, the thorax of E. fusca is almost completely dark brown to black, markedly contrasting to the reddish-orange pronotum (in E. badia, the pronotum, mesonotum and metanotum are reddish, with some irregular black markings, particular on the metanotum). The status of E. fusca needs to be studied in more detail. 52 The whereabouts of the type material of Mantispidae described by Guérin-Méneville in Duperrey, 1831 [1830] is unknown to us. A general account of the current depositories of Guérin-Méneville's material is given by Horn et al. (1990). 53 Mantispa guerinii Westwood was obviously intended to be a replacement name of M. grandis Guérin-Méneville, because it as considered to be a junior primary homonym of M. grandis Erichson. (Westwood also cited M. grandis Burmeister, which is not available under the ICZN.) Since M. grandis Guérin-Méneville, 1831 [1830], is a senior homonym of M. grandis Erichson, 1839, Westwood's replacement is invalid. M. grandis Erichson was replaced by Mantispa erichsonii Guérin-Méneville (now in Pseudoclimaciella).

Ohl: World Catalog of Mantispidae 249

54 We have not examined the type of M. nodosa but the original description and illustration clearly show a character combination (short, humped prothorax; partly darkened wings; large body size) that is typical of Euclimacia. Additionally, M. nodosa was mentioned as a species "closely allied " to Euclimacia rufocincta by Shelford (1903 [1902]: 236, pl. 19, fig. 27; as Mantispa sp.). Thus, M. nodosa most likely belongs in Euclimacia. 55 Euclimacia regina was described based on a holotype of unstated geographic origin (Esben- Petersen, 1917: 9). It bears a label with the collector's name, Carl Aurivillius. Esben-Petersen noted that Aurivillius collected in the Sunda Islands (see also Horn et al., 1990), and assumed that the type of E. regina came from those islands. 56 Handschin (1961: 270) considered E. rufocincta to be identical to the mantispid illustrated by Shelford (1903 ["1902"]: 236, pl. 19, fig. 27) as an unidentified Mantispa together with its putative Batesian mimicry model, the vespid Polistes sagittarius Saussure. Despite the fact that E. rufocincta and the mantispid illustrated by Shelford are rather similar with respect to overall body and wing coloration, Handschin's interpretation, however, is probably incorrect, because E. rufocincta has terga IV-V red, whereas terga II-III are red in the mantispid illustrated by Shelford. 57 Stenispa Navás, 1914i was originally described as a monotypic genus to contain Eumantispa hypogastrica Navás, 1914i only, and no other species have been placed in the genus subsequently. Navás (1941i) separated Stenispa from Eumantispa based on two characters. According to him, Stenispa has four radial forewing cells only, whereas Eumantispa has six. Additionally, the forecoxae in Stenispa are markedly divided, but are indistinctly divided in Eumantispa according to Navás. Eumantispa is indeed known to have "supernumery radial crossveins" (New, 2003), resulting in relatively many radial cells, but there is a considerable amount of intraspecific and even intraindividual variation involved. One of us (MO) has studied long series of Eumantispa from India, New Guinea, and other southeast Asian countries, and most individuals have different numbers of radial cells on the right and the left forewing. Additionally, the number of radial cells varies between four and six among conspecific individuals. As a consequence, four radial cells is not an appropriate character to differentiate Stenispa from Eumantispa. The degree of subdivision of the forecoxae is difficult to evaluate objectively and should be studied in more species of Eumantispa. Okamoto (1911; the German tranbslation of Okamoto, 1910) stated that "the subdivision of the forecoxae is usually indistinct" (translation by MO), which implies that he has already seen specimens of Eumantispa with a more distinct subdivision, when he described the genus. In summary, Stenispa is not properly defined as a distinct genus different from Eumantispa and, thus, it is here considered to be a junior subjective synonym of Eumantispa. 58 Handschin (1961: 297) synonymized Eumantispa hypogastrica Navás with E. ferruginea Stitz. His synonymization is based on a comparison of the holotype of E. ferruginea with a specimen in the NHRS, which was identified as E. hypogastrica by Esben-Petersen. However, the original illustration of the wings of E. hypogastrica in Navá's original description clearly show obvious differences to the holotype of E. ferruginea. For example, the hindwing vein Cu-a is markedly curved in E. ferruginea, which is quite unusual in Eumantispa and resembles

250 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

Campanacella, whereas Cu-a is straight in E. hypogastrica. In summary, it is highly unlikely that E. hypogastrica and E. ferruginea are conspecific. 59 Two paratypes of the Leptomantispa antillesensis of Hoffman in Reeves and Robinson (1999) were listed in a catalog of type specimens in the Clemson University Arthropod collection. This name is unavailable in this publication because it is not accompanied by a description or definition (ICZN Art. 13.1). Reeves and Robinson's attribution of the name to Hoffman makes reference to the unpublished and unavailable dissertation name Leptomantispa antillensis of Hoffman (1992: 170). 60 Mantispa is obviously a collective, non-monophyletic genus with weakly defined delimitations. Future attempts to achieve a more objective concept of this genus will possibly alter the corresponding distribution range. 61 The new name, ambonensis, derives from the Indonesian island Ambon, the collecting locality of the type(s) of Mantispilla basalis Navás, 1929c. 62 The new name, coorgensis, refers to the former Indian state Coorg, which is now part of the state Karnataka. Coorg was indicated as the collecting locality of the syntype series of Mantispa femoralis Banks, 1933. 63 Mantispa decumana Hagen is considered here to be an unavailable name, because it was published before 1931 and was not accompanied by a description, definition, or indication (ICZN Art. 12.1.). 64 Mantispa florealis Hagen is considered here to be an unavailable name, because it was published before 1931 and was not accompanied by a description, definition, or indication (ICZN Art. 12.1.). It is listed by Hagen with "Mus[eo] Berol[inensis]" (= ZMB) and the locality Sri Lanka (as "Ceylon"), probably refering to a specimen in the ZMB that was so labeled. 65 Hagen (1859: 406) explicitly stated that he literally repeated the description sent to him by Loew and that he did not examin any material. In the original description, "Patria: Nubia, Rueppel. Mus. Senkenbergianum [= SMFD]" is given. W. Nässig (pers. comm. to MO, March 2004) informed one of us (MO) that there is a single mantispid in the SMFD, which bears a label with the information "Nubia" and "Rueppel" but no explicit type label. The status of this specimen needs to be clarified. 66 Poivre (1982c: 669) designated a "neotype" for "Perlamantispa icterica", which was demonstrated to be an unnecessary designation by U. Aspöck and H. Aspöck (1994: 108), because the holotype still exists in the MCZ. Consequently, Poivre's neotype is set aside under ICZN Art. 75.8. Additionally, the "neotype" was misidentified by Poivre and was corrected to Mantispa aphavexelte by U. Aspöck and H. Aspöck (1994: 108). U. Aspöck and H. Aspöck reexamined the holotype of M. icterica and confirmed that it was conspecific with M. perla. Because it is still an open question whether the population of M. perla occupying the Iberian Pensinsula is distinct at subsepcific rank (as Mantispa perla icterica) (H. Aspöck et al., 2001), M. icterica is listed here as a valid species name. 67 Mantispa torquilla Hagen (1858b) is considered here to be an unavailable name, because it was published before 1931 and was not accompanied by a description, definition, or indication

Ohl: World Catalog of Mantispidae 251

(ICZN Art. 12.1.). Hagen listed it from the "Mus[eo] Berol[inensis]" (= ZMB), probably refering to a specimen in the ZMB that was so labeled. 68 According to the original description, the holotype of Mantispa japonica was deposited "in [Percy C.] Wormald's coll.", which has since been transferred to the NMWC (Horn et al., 1990: 428). One specimen labelled as a type of M. japonica is currently present in the BMNH (P. A. Brown, pers. comm. to MO, March 2004). 69 The type of Mantispa javanica was stated by Westwood (1852) to be in the museum of the East India House. These collections were mostly transferred to the BMNH after the East India Company was dissolved in 1858. A type specimen of M. javanica is in the BMNH (P. A. Brown, pers. comm. to MO March 2004). 70 The confusing taxonomic history of Mantispa lobata Navás was discussed by U. Aspöck and H. Aspöck (1994). Most published records of M. lobata actually refer to different Palaearctic species (H. Aspöck et al., 2001), so most published distributional information for it is not reliable. The taxonomic identity and geographic distribution of M. lobata need further study. 71 Navás mentioned "Br. O. Afr." (= British East Africa) as the type locality of Mantispa loveni, a name formerly applied to either Kenya, or collectively to the former British dependencies of Tanzania, Kenya, and Uganda. Navás also mentioned that the large specimen of the syntype series bears a locality label with the information "Elgon, Lindblom, 1700 meter, maj". Elgon is an extinct vulcano on the border between Kenya and Uganda. 72 Ulrike Aspöck and Horst Aspöck (1999: 106, 110) pointed out that Mantispa mandarina was generally misconceived in the literature, and they corrected most published records to M. aphavexelte (e.g., H. Aspöck et al., 1980: 184, and several publications cited in H. Aspöck et al., 2001: 204). The true distribution of M. mandarina is unknown and needs careful re- examination based on its separation from M. aphavexelte. The type locality of M. mandarina, in northern China, is currently the only reliable record. 73 Mantispa maynei was listed as a synonym of M. verruculata by Handschin (1959a: 200). However, New and Haddow (1973: 162) pointed out differences in the pterostigmal color to differentiate M. verruculata, M. maynei, and M. nanyukina, and we follow them in treating M. maynei as a valid species. 74 Penny at al. (1997: 74) gave the MCZ as the depository of the type of Mantispa moesta, but it could not be located there (P. D. Perkins, pers. comm. to MO). In the original description, Hagen mentioned "Saussure" behind the type locality. Since most of Saussure's collection is now in the MHNG (Horn et al., 1990), the type of M. moesta might also be there. 75 Mantispa mutata is considered here to be an unavailable name, because it was published before 1931 and was not accompanied by a description, definition, or indication (ICZN Art. 12.1.). 76 If Mantis nana Lichtenstein, currently treated as a synonym of Mantispa nabota (Olivier) (a nomen dubium in Mantispidae), proves to be a true Mantispa, then Mantispa nana Erichson will become a junior secondary homonym and will need to be replaced by the junior synonym, M. bonhourei Navás.

252 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

77 The new name, paraguayana, derives from the country of the collecting locality of the type(s) of Mantispilla nana Navás, 1912b. 78 The type of Mantispa pehlkei was originally deposited in the MZPW, but could not be located there recently (W. Tomaszewska, pers. comm. to MO, Jan. 2004). 79 Although Navás (1933d: 215) originally indicated that Mantispa phaeonota was from Kenya, he later corrected that statement to indicate Madagascar instead (Navás, 1935 ["1934"]: 66). Handschin (1963 ["1962"]) then listed Kenya again along with Madagascar. To our knowledge however, apart from the incorrect citation in the original description, M. phaeonota has yet to be validly recorded from outside Madagascar. 80 Handschin (1959:190 [footnote]) states: "Mantispilla minuta ist schon 1775 von FABRICIUS für eine Form aus Peru gebraucht worden. Minuta Kozhantshikov ist also als Synonym einzuziehen. Falls die Form sich als gut erweist, schlage ich vor, sie als caucasica zu bezeichenen" [English translation: "Mantispilla minuta has already been used by Fabricius 1775 for a species from Peru. Minuta Kozhanchikov is therefore a synonym (= homonym). If the species (of Kozhanchikov) proves to be good, I propose to name it caucasica"]. There appears to be at least two ways to interpret Handschin's statement. One is that Handschin was attempting to point out a putative primary homonymy between minuta Fabricius and minuta Kozhanchikov. However, Handschin's apparent attribution of the combination Mantispilla minuta to Fabricius (1775) is incorrect. Fabricius' (1775:278) minuta was originated described in the genus Mantis (the genus Mantispilla was not published until 1910), so there is no primary homonymy between these names. Under this interpretation, adopted by H. Aspöck et al. (2001: 203 [footnote]), Handschin's "caucasica" seems best interpreted as 'Mantispilla caucasica', an unnecessary replacement name for Mantispilla minuta Kozhanchikov. A second interpretation of Handschin's statement considers the larger context of the text in which his statement was made. The quoted statement is printed as a footnote asterisked to the name "Mantispilla minuta Kozhantshikov 1949", which is contained in a list of Mantispa and Mantispilla species from Asia and southeastern Europe. This list, a similar list of African Mantispa, Mantispilla and Necyla species, and the surrounding text are part of a discussion headed "Die Gruppe der Mantispa styriaca Poda." This text focuses on taxonomic difficulties within the styriaca group of the genus Mantispa, and specifically notes Handschin's belief that some of the species belonging to this group have been decribed under different names – sometimes in different genera, particularly Mantispa and Mantispilla. Seen in this context, Handschin's statement can be interpreted as noting the apparent secondary homonomy between Mantispa minuta (Fabricius, 1775) [Mantis] and 'Mantispa minuta (Kozhanchikov, 1949) [Mantispilla]' – a combination not explicitly used by Handschin, but plausibly implicit in his text. Under this interpretation, Handschin's "caucasica" seems best interpreted as 'Mantispa caucasica', an apparently necessary replacement name for the junior secondary homonym 'Mantispa minuta (Kozhanchikov)' of Handschin's era. We have not been able to identify any evidence that will permit an unambiguous choice among these two interpretations. This ambiguous result with regard to the possible implicit generic combinations for "caucasica", together with the fact that Handschin did not explicitly combine "caucasica" with any genus-group name, leads us to the conclusion that this name fails ICZN

Ohl: World Catalog of Mantispidae 253

Art. 11.9.3 ("A species-group name must be published in unambiguous combination with a generic name (either explicit, or implicit by context)"), and, therefore, must be considered unavailable. 81 Evenhuis (1997: 617) stated that the "whereabouts of [the] personal collection of insects [of Poda is] unknown; probably destroyed or scattered and lost". 82 The authorship of Raphidia mantispa is frequently attributed in the literature to Linnaeus (1767), but Linnaeus cearly referenced Scopoli (1763), who first introduced this name in the species-group. 83 The type of Mantis pagana was originally in the collection of "D. Mallet" (Fabricius, 1775: 278). The depository of this type was not given by Zimsen (1964: 614). Evenhuis (1997: 245) listed a number of additional collections as possible sources for material described in Fabricius' 1775 monograph. These should be checked for the type of M. pagana. 84 The holotype of Mantispilla tessmanni is destroyed except for two wings mounted on small cards pinned with the original labels. In the card file of the ZMB Neuroptera collection there is a note with the following information written by R. Schulze, a former ZMB technician (translated from German): "According to a note by Dr. Winkler on the loan form: specimen destroyed, on 31 May 1959 received back from a loan to Prof. Handschin, Basel. R. Schulze, 1981". 85 Fraser (1955: 21) attributed authorship of Mantispilla usambarica to Westwood, but, to the best of our knowledge, no such species was ever published by Westwood. It is possible that M. usambarica is a grossly incorrect subsequent spelling of M. mozambica Westwood. However, since this has not been conclusively demonstrated, we treat it separately here. It is an unavailable name, having been published after 1930 and not accompanied by a description or definition (ICZN Art. 13.1.). 86 In the original description of Mantispa variolosa, Navás (1914d: 649) gave the following information concerning the geographic origin of its type(s): "Patria, Oceanía: "Graham's town, Albany Museum, J. Linskey" (Mus. de Londres)". We interpret this statement to mean that the specimen(s) were collected in "Oceania"; that it(they) were formerly contained in the Albany Museum of Grahamstown, South Africa, and that it/they belonged to the BMNH when examined by Navás. An "Oceania" type locality is consistent with the title of Navás' paper: "Neurópteros de Oceanía". "Oceania" is a collective name for the lands of the Pacific Ocean, especially including Micronesia, Melanesia and Polynesia, but sometimes also including Australia, New Zealand and the Malay Archipelago. Navás' concept of "Oceania" was probably very broad, as most of the mantispid species described in this publication are from Australia. Esben-Petersen (1923:599) listed M. variolosa as a questionable record from Grahamstown, New Zealand. But, this interpretation is probably incorrect, as it conflicts with the label reference to the Albany Museum, which is in South Africa. Mantispa variolosa was not included in Lambkin's revision of Australian Mantispidae (1986a, b), and its precise type locality remains unknown. 87 The type data for Mantispa venulosa are given in the original description is "América central: "Caldera, 1200 ft., Champion" (Mus. de Londres)". There are a number of populated places in Central America with the name "Caldera" in e.g., Mexico, Panama, Columbia, so the type

254 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

locality of M. venulosa remains unclear. The cited elevation and collector may ultimately be useful in narrowing the number of possible sites. 88 In the original description, the type locality of Mantispa verruculata is given as "Afrique Orientale Anglaise: Landjoro, dans le Pori". The type material was collected during an expedition to East Africa by Ch. Alluaud and R. Jeannel in 1911-1912, who travelled mostly in Kenya and Tanzania. The localities indicated by Navás have not yet been relocated in any country of the former British East Africa (Kenya, Tanzania, Uganda). 89 Mantispa verruculata is widely distributed in East Africa south to South Africa. A suspect northern record from Tunisia (Navás, 1930d: 136) requires confirmation. 90 McLachlan (1900: 127) stated that the holotype of M. simulatrix was only temporarily being held in the Oxford collection and that it would "ultimately be deposited in the Sarawak Museum". It is currently deposited in the OUM (J. Hogan, pers. comm. to MO June 2004). 91 Mantispa mimetica Sharp was introduced in the 1900 volume of the Zoologial Record and is obviously identical to M. simulatrix McLachlan based on the bibliographical information given by Sharp. Mantispa mimetica may be a manuscript name that was submitted by McLachlan (the compiler of the Neuroptera section of the Zoological Record until 1885) to Sharp (editor of the 1900 volume), but was later changed to M. simulatrix. The identity of M. mimetica and M. simulatrix was first proposed by van der Weele (1909: 96). Mantispa mimetica is an unavailable name, having been first published before 1931 and not accompanied by a description, definition, or indication (ICZN Art. 12.1.). 92 Euclimacia ferdinandi was listed as a valid species in Euclimacia by New (2003: 154). However, Handschin (1961: 276) compared Navás' original description with Mantispa simulatrix in detail, and he could demonstrate that the two are identical. The unique coloration of this species along with some structural characters prompted Handschin to described a new genus, Mimetispa. 93 The genus-group name Bucharispa is apparently unavailable, having been apparently first published by Martynov (1936: 437), which contains an illustration labelled "Bucharispa" and the Russian equivalent of "new genus, new species". No species were attributed to the genus by Martynov and, apparently, no description or diagnosis was provided. Because Bucharispa was published after 1930, the presence of an illustration cannot serve as an indication to make the name avaliable. Kozhantshikov (1949: 355) listed Bucharispa as a synonym (of Forciada) and attributed it, still without included species, to "Bianchi, Martynov, in litt.", actions which are also not sufficient to make Bucharispa available. 94 See annotation for Mantispa australasiae. 95 Navás (1913a:280-281) contains two original spellings of this genus group name: "Nicyla", in the header of its generic treatment, and in the header of the treatment of its type species; and "Necyla", in the caption to figure 8, illustrating the forewing of its type species. Under ICZN Art. 32.5.1, the correct original spelling is "Necyla", because Navás explicitly stated the etymology of the name on p. 280 as: "Etym. Anagramma vocis Ceylan." (English translation: Etym[ology]. An anagram of Ceylan.). See also Navás 1914k:119, where Navás explicitly notes

Ohl: World Catalog of Mantispidae 255

that Nicyla was an unintended spelling, reiterates the etymology of the name as based on the word "Ceylan", and notes that Necyla was the intended spelling. 96 The type of Mantispa formosana bella was originally contained in Kuwayama's collection, which was partially transferred to the ITLJ according to Horn et al. (1990). 97 Necyla formosana chiaiensis was originally described as a variety of Mantispa formosana (as Mantispa formosana bella), which was transferred to Necyla by Stange and Wang (1997). Based on this transfer, chiaiensis is herewith tentatively placed in Necyla as well, but its final generic placement and taxonomic status require further research. The new name, chiaiensis, derives from Chia-I, the current spelling of the type locality Kagi on Taiwan. 98 Mantispilla formosana var. major Stitz, 1913, is a primary homonym of Mantispilla punctata var. major Stitz, 1913, published in the same paper. Since neither name is considered valid, neither name is replaced here. However, if either of these names becomes valid, it will need to be replaced by a junior synonym or a new name. 99 Necyla formosana minor was originally described as a variety of Mantipa formosana, which was transferred to Necyla by Stange and Wang (1997). Based on this transfer, minor is herewith tentatively placed in Necyla as well, but its final generic placement and taxonomic status require further research. 100 Mantispa orientalis Esben-Petersen and Mantispilla nigra Stitz were both published in 1913. Esben-Petersen (1917) synonymized them and assigned the former priority over the latter, but without giving his rationales for his decision. On the title page of Stitz (1913) the date on which the paper was send out is giving as October 1913 ("Eingesandt im Oktober 1913"). Esben- Petersen (1913) was officially send on 1 September 1913 and has an accession date by the ZMB library dated 22 September 1913. Thus, Esben-Petersen (1913) was apparently published prior to Stitz (1913). 101 Navás stated (1927: 89) that he based the description of Necyla pupa on a single specimen ("Un ejemplar en alcohol capturado por el Dr. Guido Paoli"), but he did not explicitly state its depository. The Paoli collection was transferred to the MCSN (see annotation at N. paolina), and Navás also mentioned the MCSN in the introduction of the paper containing the description of N. pupa. Thus, unless the specimen was retained in Navás' collection (?),the type of M. pupa might be found in the MCSN. 102 Necyla formosana sumatrana was originally described as a variety of Mantispa formosana, which was transferred to Necyla by Stange and Wang (1997). Based on this transfer, sumatrana is herewith tentatively placed in Necyla as well, but its final generic placement and taxonomic status require further research. 103 Neoclimaciella Handschin is a nomen nudum, having been proposed after 1930 and not having its type species fixed in its original publication (ICZN Art. 13.3.). No species have ever been included in this genus. 104 The historical accession catalog of the ZMB lists four syntypes of Mantispa decorata. The original entry was subsequently corrected to three specimens, with the annotation "one specimen probably in the museum Budapest in 1964". Gyorgy Sziraki from the HNHM (pers.

256 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

comm. to MO, Aug. 2001) confirmed that among the two specimens of M. decorata in the HNHM, one bears the same labels as the three syntype specimens in the ZMB and probably represents a fourth syntype. 105 Stange (1967: 22) explicitly indicated that he had personally examined the type of M. muhni, which is deposited in the SMBA. Hoffman (1992) pointed out that the type of M. muhni must be considered as lost because it was deposited in the destroyed collections of the ZMUH. Navás (1930e) explicitly stated that in this particular publication specimens from the ZMUH are marked by "M.H.". This marker is absent from the original description of M. muhni, which states "col. Mühn" instead. Thus, Hoffman's statement appears to be in error, and the type material of M. muhni probably resides in the SMBA, though this has not been confirmed. 106 Navás (1936a ["1935"]) named Climaciella olsufievi in honour of the collector of the holotype, Mr. Olsoufieff, hence reducing the '-ou-' in the collectors' name to '-u-' and the '-ff' to '-v-'. Handschin (1960b, 1961, 1963 ["1962"]) altered the name to P. olsoufievi. However, Navás repeatedly used both the species name olsufievi and the collector's name Olsoufieff in the original work, so there is no clear evidence that the taxon name spelling as olsufievi was the result of an inadvertent error that would require correction under ICZN Art. 32.5.1. Consequently, olsufievi is herewith accepted as the correct original spelling. 107 Perlamantispa was synonymized with Mantispa by H. Aspöck et al. (1980: 182) because (translated from German) "several genera separated from Mantispa [, such as Perlamantispa,] can not be convincingly differentiated". Consequently, the type, and only European, species of Perlamantispa, P. perla, was treated as a member of Mantispa in subsequent publications by these authors (e.g., H. Aspöck, 1999; H. Aspöck et al., 2001; U. Aspöck and H. Aspöck, 1994). However, Perlamantispa is considered to be a valid genus in many other publications, and particularly by Oswald and Penny (1991: 48) in the catalog of genus-group names in Neuropterida. We follow these authors and tentatively accept Perlamantispa as valid in this catalog. The final decision requires a taxonomic revision of the species currently placed in Perlamantispa and is deferred to future research. 108 The holotype of Perlamantispa austroafrica is mentioned with the information "collection Pictet", but the description is published in a work entitled as a review of the mantispids in the MHNG. 109 Poivre (1982a ["1981"]: 194) stated that the holotype of Perlamantispa girardi "retourné à l'I.F.A.N., Dakar", which might imply that the type was still in Poivre's collection in 1981. Since Poivre's collection is now in the MNHN, both the IFAN and the MNHN should be considered when locating the holotype. 110 The type locality and type depository of Mantis perla and its possible synonyms and subspecies were discussed in detail by U. Aspöck and H. Aspöck (1994). Horst Aspöck et al. (2001: 204) considered the type of M. perla to be no longer in existence. 111 The historical accession catalog of the ZMB lists six specimens under accession number 195. The entry is in Erichson's handwriting: "Mantispa perla Pall[as]. Er[ichson].* / Ross[ia]. meridionalis [= southern Russia]". Erichson (1839) obviosuly accepted these specimens as the syntypes of Mantispa christiana, when he synonymized it with M. perla. In 1981, a handwritten note was added to the accession catalog that only four specimens could be found at that time.

Ohl: World Catalog of Mantispidae 257

The four specimens bear identical label sets: "195 / S. Rußland Redow". U. Aspöck & H. Aspöck (1994:101) discussed the putative type status of these specimens in detail (they listed three specimens only and did not label which specimens they studied). Charpentier (1825:93) explicitly mentioned the ZMB as the depository and "Russia europaea" as the origin of the material he studied. The four specimens correspond perferctly with Charpentier's short description of M. christiana and they are the only candidates now in the ZMB that are plausible as Charpentier's types. As U. Aspöck & H. Aspöck (1994) emphasized however, they lack any direct indication of type status. Hagen (1859:408) mentioned that he possessed "a type of Charpentier's M. christiana from Rumelien, informed about from Kunze to Winthem" (translated from German). This specimen is now deposited in the MCZ. "Rumelien" [= Rumelia or Roumelia] was a poorly defined European division of the Ottoman Empire. However, it generally included (in current geopolitical terms) Albania, Macedonia, Thrace (European Turkey) and the southern parts of Bulgaria. "Rumelien" was not part of European Russia, and, thus, the specimen mentioned by Hagen (1859) is probably not a syntype of M. christiana. 112 Mantispa flaveola Erichson was synonymized with Mantispa minuta (Fabricius) by Penny (1982a: 459), but the holotype of M. flaveola does not belong to any known New World genus. This fact is mentioned on labels by R.G. Beard pinned under the holotype and by Hoffman (1992) [both unpublished sources], who both suggested that M. flaveola belongs in Perlamantispa and is similar to P. perla. One of us (MO) has compared the holotype of M. flaveola to P. perla, and to most species currently placed in Perlamantispa. He found perfect agreement between M. flaveola and P. perla, which are herewith synonymize. The original, green locality label on the holotype of M. flaveola reads "Para / Sieber". Sieber was an insect collector paid by J. C. von Hoffmannsegg and who is known to have collected in Brazil in 1808/1809 (see also Ohl, 2002, for the history of Hoffmannsegg's collections). Sieber's material was incorporated into the ZMB collections in 1810 (Horn et al., 1990: 365). The green label color was also used to indicate a New World locality. Thus, the label obviously refers to Brazil, and the specimen was clearly mislabeled at some point in the past. This must have happened before 1839, when Erichson described M. flaveola. 113 Ulrike Aspöck & H. Aspöck (1994:110) pointed out that the type or the types of Mantispa victorii could not be located in the MNHN, but Hagen (1859: 408) listed one type of this species as being in his collection ("one type of Guèrin's M. Victorii from Armenia, by Motschulsky" (translated from German). A specimen exists in the MCZ that is labelled as a type of M. victorii. It bears exactly the same label data as indicated by Hagen (1859). Additionally, Hagen (1858a: 128) also pointed out that Guérin-Menéville described M. victorii upon a note sent by Motschulsky, whose name is also on the label. There is a slight discrepancy in the label locality data. In the original description, "Caucasus" is given by Guérin-Menéville, whereas "Armenien" is on the label of the putative type. Since the Caucasus mountains extend into Armenia, Guérin-Menéville has probably simply used a broader geographic term. So we conclude that the MCZ specimen labelled as M. victorii is likely a syntype of this species. 114 Mantispa perla var. brunnea is in all probability an unusually dark, southern Italian (Puglia region) color phenon of Perlamantispa perla. Based on the known variation and geographical distribution of P. perla (see H. Aspöck et al. 1980, H. Aspöck et al. 2001, U. Aspöck 1996, and U. Aspöck and H. Aspöck 1994), and personal examination of much material of this species, it

258 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

seems extremely unlikely that there exits in Italy a geographically distinct population that merits subspecific rank. Based on these observations, var. brunnea is herewith synonymized with the nominate species, P. perla. Furthermore, Mantipsa perla var. brunnea Navás, 1906 is a junior primary homonym of Mantispa brunnea Say in Keating, 1824 (now in Climaciella), and would require a replacement name if considered valid. 115 The whereabouts of the collections and types of Peter Simon Pallas were discussed in detail by Evenhuis (1997: 592), and efforts should be made to locate the type(s) of M. pusilla in the collections mentioned by him. 116 The surviving types and collections of Carl De Geer are housed in the NHRS according to information in Evenhuis (1997: 192). Horst Aspöck (1999: 216) expressed doubts about the identity of Perlamantispa brevicornis and listed it as an unplaced Mantispidae (as "Mantidae g.sp.", which is a lapsus, H. Aspöck, pers. comm. to MO, Dec. 2003). 117 The original publication of Pseudoclimaciella cachani contains at least three different spellings of its species-group name [pages on which each spelling occur are representative, but should not be considered exhaustive]: (1) cahani, p. 3 [a prominent spelling in the species description heading]; (2) cachani, pp. 6, 7, 8, 9, 10, 19, 21, 23; (3) chachani, p. 8. Poive, p. 3, states explicitly that "Cette espèce est dédiée au Professeur Cachan de l'Institut agronomique de Nancy, ...". Acting as first revisors (ICZN Art. 24.2.3), and in accordance with Poivre's apparent intention (based on his explicit dedication and on the frequency of its occurrence in the text), we select "cachani" to be the correct original spelling. 118 The status and sex of the type of Pseudoclimaciella timmerhansi were determined by Handschin (1960b: 217). 119 Paulian (1957: 76) listed three species treated by Navás (1936a ["1935"]:102-105) in Climaciella as C. olsufievi and C. necopina (both now in Paulianella) and C."umbra" Although the latter name does not occur Navás' text, it is clear from Paulian's bibliographic citation and Navás' text that "umbra" is simply a gross misspelling of C. umbripennis, which is the third (and only other) mantispid name mentioned by Navás. Accordingly, Paulian's "umbra" is considered here to be an incorrect subsequent spelling of umbripennis. 120 New (1996: 27) excluded Mantispa scutellaris from the catalog of Australian Neuroptera, because it "is almost certainly not Australian, and the type specimen has no locality label". 121 New (1996: 27) pointed out that Mantispa strigipes was included in the Australian fauna on the basis of a hardly legible locality label only. He concluded that "no subsequent specimens have been found in Australia and it seems doubtful that this is an Australian species". 122 Stenomantispa was originally proposed as a subgenus of Mantispa. It was raised to generic rank by Handschin (1961: 297), listed as a subgenus of Mantispa by Oswald and Penny (1991: 34), and listed as a genus by New (1986: 129, 1998: 179). The ranking used by New (1998) is followed here. The type species of Stenomantispa is Mantispa (Stenomantispa) ilsae, by monotypy. A second species, Mantispa reinhardi Stitz, was only doubtfully included by Stitz (1913), and is therefore not available for type selection under ICZN Art. 72.4.1, although its placement in Stenomantispa was confirmed by Handschin (1961: 299).

Ohl: World Catalog of Mantispidae 259

123 Penny (1982a: 456) mentioned a female as the holotype of Mantispa compellens in the BMNH, but Walker (1860) did not designate a holotype. He differentiated two "variations A and B", which is suggestive of at least two original syntype specimens. One of them, the "holotype"of Penny (1982a), is a female. P. A. Brown (pers. comm. to MO, March 2004) confirmed that only one specimen in the BMNH is currently labelled as a type. The whereabouts of the remaining syntype(s) is unknown to us. 124 The type(s) of Necyla uniformis was retained by Navás in his personal collection and could not be found in the MZBS (Monserrat, 1985). However, the original description is sufficiently detailled to allow a definitive placement in Zeugomantispa ("Prothorax […] minute rugosus et granulous", Navás, 1927c: 61) and confirms the identity with Z. compellens ("Meso- et metanotum scutellis flavis. […] Femora anteriora basi et macula grandi ad medium interne et externe fuscis", p. 61) 125 Mantis lilliputiana Olivier and Mantispa flavomaculata Latreille were independently described to formalize the mantispid listed as "Le Nain de Suriname" by Stoll (1787 [1787- 1790]: plate II, fig. 7), and, thus, are synonyms. This was already stated by Erichson (1839: 173), although he incorrectly assigned M. flavomaculata priority over M. lilliputiana. Houttuyn in Stoll (1813: 77) also proposed a name for this species, Mantis viridula. In the original description of M. lilliputiana, Olivier mentioned "Du Cabinet de L. F. Holthuisen", which is the same collection from which Stoll's material also originated. It is unclear if Olivier ever personally studied the material in the Holthuisen collection, or if he just refered to this collection as Stoll's source. Based on the original description of M. flavomaculata, it seems unlikely that Latreille had examined Stoll's original material, nor any other material. Horst Aspöck (1999: 216) interpreted M. lilliputiana as "Mantispidae g.sp.", but it seems likely that both M. lilliputiana and M. flavomaculata are junior subjective synonyms of Zeugomantispa minuta. The Holtuisen collection, parts of which were illustrated by Stoll, was auctioned in Hamburg, Germany, on 16 May 1797 (auction postponed from 3 February 1796). The collection inventory was published as an auction catalog (Lichtenstein, 1796). Species are listed in the catalog with references to their respective illustrations in Stoll (1787 [1787-1790]). The species illustrated by Stoll on plate II, fig. 7 is listed as "Mantis minuta Fabricius" (now in Zeugomantispa) (Lichtenstein, 1796: 81), which is herewith accepted as the basis for synonymizing M. lilliputiana, M. flavomaculata, and M. viridula with M. minuta, although this was not Lichtenstein's explicit intention. See also the annotation for M. nabota and its synonyms. 126 The whereabouts of the type of Raphidia margaritacea was unknown to Penny (1982a: 459). According to Horn et al. (1990), most of the type material of G. Fischer von Waldheim is deposited in the ZMUM. A smaller portion is housed in the ZIL, but Raphidia margaritacea was not mentioned by Krivokhatsky (1995: 11) from this collection. 127 Mantispilla punctata Stitz and Mantispa viridis Stitz were synonymized with Zeugomantispa minuta (as Mantispa minuta) by Penny and da Costa (1983: 683). They did not distinguish between Z. minuta and Z. virescens, which were differentiated (as Z. minuta and Z. viridula) by Hoffman (2002: 273) largely based on one forewing character. Hence, the synonymy of M. punctata and M. viridis with Z. virescens is newly proposed here.

260 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

One of us (MO) examined the types of Mantispa viridula Erichson, Mantispa virescens Rambur, Mantispilla punctata Stitz, Mantispilla punctata major Stitz, Mantispilla stigmata Stitz, and Mantispa viridis Stitz. No structural differences could be found, and the morphology corresponds to the diagnostics of Zeugomantispa virescens as given by Hoffman in Penny (2002; as Z. viridula). Thus, Z. viridula, M. virescens, M. punctata, M. punctata major, M. stigmata, and M. viridis are considered to be synonymous, with Z. viridula (Erichson) as the oldest name. However, Z. viridula (Erichson) is a junior secondary homonym of Z. viridula (Houttuyn in Stoll) [= Z. minuta (Fabricius)] and, thus, is not available. Instead, the oldest available name is Z. virescens (Rambur). 128 One syntype of Mantispa viridula Erichson was considered a misidentification by Stitz (1913: 20) and was subsequently attributed to Mantispa punctata var. major Stitz, 1913: 20 (as its holotype). 129 The type locality of Mantispa virescens was unknown to Rambur (1842: 433: "patrie inconnue"), but is South America according to Esben-Petersen (1917: 14). 130 After we realized that Z. viridula (Erichson) is a junior secondary homonym of Z. viridula (Houttuyn in Stoll) [= Z. minuta (Fabricius)], effort has been made to replace Erichson's viridula by the next available name. Hoffman (1992) in his unpublished doctoral thesis proposed the synonymy of Z. viridula (Erichson) and Mantispa virescens Rambur, so that Rambur's name would be the oldest available name for that species. Although we decided generally not to use the results proposed by Hoffman (see Miscellaneous Annotations above), we decided to make an exception here. One of us (MO) re-examined the holotype of M. virescens, which confirms the synonymy of M. virescens with Z. viridula (Erichson). Thus, Rambur's name is the oldest available name for the species previously known as Z. viridula (Erichson). We are thankful to Kevin Hoffman to give us the permission to use his results in this case (pers. comm. to MO, June 2004). The single type specimen of Mantispa virescens in the ISNB is in bad condition and lost most of his color. The head, the prothorax, and the forelegs are glued on a small card pinned below the specimen. Mots of the left fore and the right hind leg are lacking. The terminalia are dissected and preserved in glycerol in a microvial on a separate pin near the specimen. 131 The holotype of Mantispa punctata var. major was originally part of the syntype series of Mantispa viridula (now in Zeugomantispa) (see Stitz, 1913: 20). 132 Mantispilla punctata var. major Stitz, 1913, is a primary homonym of Mantispilla formosana var. major Stitz, 1913. See annotation for this species. 133 Mantispa viridis Stitz is a junior primary homonym of Mantispa viridis Walker, 1853 (now in Zeugomantispa). 134 Lambkin (1986b: 54) lists Manega luddemanni and Nivella rubella as nomina dubia, their types presumed destroyed in the bombing of the Hamburg Museum (see annotation for the ZMUH). Thus, the taxonomic identities of the monotypic genera Manega and Nivella cannot be established with certainty without neotype designations for their type species. Following Lambkin, both genera are also treated here as nomina dubia. 135 See annotation for Manega.

Ohl: World Catalog of Mantispidae 261

136 Thunberg (1815: 293) placed Mantis fausta in Mantispa, but this species name was apparently not mentioned in subsequent publications. Although most of Thunberg's type material is deposited in the Uppsala collection (UZIU), the type specimen of M. fausta could not be located there (H. Mejnol, pers. comm. to MO, 2003). However, because the original labels are still present in the UZIU, the type specimen must be regarded as lost. The placement of M. fausta in the Mantispidae and its true taxonomic affinities remain uncertain. 137 Stoll (1787 [1787-1790]: 12; pl. IV, fig. 15) illustrated and briefly described a mantispid from the Holthuisen collection as "Le Nain de Coromandel" without assigning it a binominal name. Mantis nabota Olivier, Mantis nana Lichtenstein, and Mantispa rufescens Latreille were all independently described in order to make Stoll's species formally available. Based on their original descriptions, it seems unlikely that Olivier, Lichtenstein, or Latreille ever examined Stoll's original,or any other, material of this species. Erichson (1839: 173) established the synonymy of M. rufescens and M. nabota, incorrectly giving the former priority over the latter. Since the original description of Mantis nana also exclusively referenced Stoll's illustration and description, it is here treated as another synonym of M. nabota, which is the senior species- group name. Horst Aspöck (1999: 216) interpreted M. nabota as "Mantispidae g.sp.", and we agree with him that its identity cannot be confirmed at the moment. Thus, it is listed here as a nomen dubium. Charpentier (1844: 309) stated that Mantis nana Lichtenstein is the female of the mantid Mantis monacha Fabricius (now Miomantis monacha; see Ehrmann, 2003: 229), but Charpentier obviously confused M. nana Lichtenstein and M. nana Stoll, a mantid (= Miomantis fenestrata (Fabricius). Stoll's illustration clearly shows that M. nana Lichtenstein is a mantispid. See also the annotations for M. lilliputiana and M. flavomaculata. 138 Mantispa sibirica was synonymized with M. perla by Charpentier (1844: 311) and listed accordingly as a synonym by Hagen (1859: 408, 1866: 425). However, it was not included in the synonymical lists of M. perla by H. Aspöck et al. (1980), U. Aspöck and H. Aspöck (1994), and H. Aspöck et al. (2001). Horst Aspöck (1999: Tab.1) listed M. sibirica as "Mantispa sp.", hence implying doubts about its taxonomic identity. We agree with H. Aspöck, and consequently, M. sibirica is listed here as a nomen dubium. The whereabouts of a personal insect collection held by Gmelin, if any existed at all, are unknown (see Evenhuis, 1997: 309, for details). Mantispa sibirica was described with reference to an unnamend mantispid described by Lepechin (1774). 139 Panfilov (1980) placed the monotypic genus Promantispa in a family of its own, Promantispidae. Lambkin (1986a) showed that P. similis belongs in Mantispidae, but with unknown generic or subfamily affinities within the family. 140 Vectispa relicta was tentatively placed in the Drepanicinae by Lambkin (1986a: 24). 141Willmann (1994) re-examined the type of Whalfera venatrix (as Fera venatrix) and critically discussed its affinities with the Symphrasinae. Ulrike Aspöck & Mansell (1994) and Engel (2004) considered it a questionable member of the Rhachiberothidae. Whalfera venatrix is here listed as a possible mantispid with unknown affinities, because of the ambiguous evidence for a definite position in Neuroptera.

262 Contrib. Ent. Internat., Vol. 5, No. 3, 2004

142 Retinoberotha stuermeri Schlüter, 1978, was considered to be a basal mantispid by Willmann (1994: 194). This interpretation was based on the assumption that raptorial forelegs were a synapomorphy of the Rhachiberothidae + Mantispidae (Willmann, 1990). This view was rejected by U. Aspöck and Mansell (1994), who favoured a sister-group relationship between the Rhachiberothidae and Berothidae. Retinoberogha stuermeri may be related either to Mantispidae, Rhachiberothidae, or one of their stem groups. 143 In the original description of Retinoberotha stuermeri, Schlüter stated that the holotype is in the Palaeontological Institute of the Freie Universität, Berlin. However, the specimen has been kept in Schlüter's personal collection since its description (Schlüter, pers. comm. to MO). It is currently on loan to one of us (MO) and will ultimately be deposited in the ZMB.