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A Preliminary Report of the Fossil Mammals from a New Microvertebrate Locality in the Upper Jurassic Morrison Formation, Grand County, Utah

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GEOLOGY OF THE INTERMOUNTAIN WEST an open-access journal of the Utah Geological Association ISSN 2380-7601 Volume 5 2018 A PRELIMINARY REPORT OF THE FOSSIL MAMMALS FROM A NEW MICROVERTEBRATE LOCALITY IN THE UPPER JURASSIC MORRISON FORMATION, GRAND COUNTY, UTAH Brian M. Davis, Richard L. Cifelli, and Guillermo W. Rougier Theme Issue An Ecosystem We Thought We Knew— The Emerging Complexities of the Morrison Formation SOCIETY OF VERTEBRATE PALEONTOLOGY Annual Meeting, October 26 – 29, 2016 Grand America Hotel Salt Lake City, Utah, USA © 2018 Utah Geological Association. All rights reserved. For permission to copy and distribute, see the following page or visit the UGA website at www.utahgeology.org for information. Email inquiries to [email protected]. GEOLOGY OF THE INTERMOUNTAIN WEST an open-access journal of the Utah Geological Association ISSN 2380-7601 Volume 5 2018 Editors UGA Board Douglas A. Sprinkel Thomas C. Chidsey, Jr. 2018 President Paul Inkenbrandt [email protected] 801.537.3361 Utah Geological Survey Utah Geological Survey 2018 President-Elect Peter Nielsen [email protected] 801.537.3359 801.391.1977 801.537.3364 2018 Program Chair Emily McDermott [email protected] 801.537.3389 [email protected] [email protected] 2018 Treasurer Zach Anderson [email protected] 801.538.4779 2018 Secretary Christopher Kravits [email protected] Bart J. Kowallis Steven Schamel 2018 Past President Bill Loughlin [email protected] 435.649.4005 Brigham Young University GeoX Consulting, Inc. 801.422.2467 801.583-1146 UGA Committees [email protected] [email protected] Education/Scholarship Loren Morton [email protected] 801.536.4262 Environmental Affairs Craig Eaton [email protected] 801.633.9396 Geologic Road Sign Terry Massoth [email protected] 801.541.6258 Historian Paul Anderson [email protected] 801.364.6613 Membership Rick Ford [email protected] 801.626.6942 Public Education Paul Jewell [email protected] 801.581.6636 Matt Affolter [email protected] Publications Roger Bon [email protected] 801.942.0533 Publicity Paul Inkenbrandt [email protected] 801.537.3361 Social/Recreation Roger Bon [email protected] 801.942.0533 Society of Vertebrate Paleontology AAPG House of Delegates Editors 2017–2020 Term Tom Chidsey [email protected] 801.537.3364 Kelli C. Trujillo — University of Wyoming John Foster — Museum of Moab State Mapping Advisory Committe Cary Woodruff — University of Toronto UGA Representative Jason Blake [email protected] 435.658.3423 Octavio Mateus — Universidade Nova de Lisboa Earthquake Safety Committe Production Chair Grant Willis [email protected] 801.537.3355 Cover Design and Desktop Publishing Douglas A. Sprinkel UGA Website www.utahgeology.org Cover Small vertebrate fossils wrapped for transport, Cisco Webmasters Paul Inkenbrandt [email protected] 801.537.3361 Lance Weaver [email protected] 801.403.1636 Mammal Quarry, Upper Jurassic Morrison Forma- tion, Grand County, Utah. UGA Newsletter Newsletter Editor Bob Biek [email protected] 801.537.3356 Become a member of the UGA to help support the work of the Association and receive notices for monthly meetings, annual field conferences, and new publi- cations. Annual membership is $20 and annual student membership is only $5. Visit the UGA website at www.utahgeology.org for information and membership application. This is an open-access article in which the Utah The UGA board is elected annually by a voting process through UGA members. Geological Association permits unrestricted use, However, the UGA is a volunteer-driven organization, and we welcome your distribution, and reproduction of text and figures that voluntary service. If you would like to participate please contact the current are not noted as copyrighted, provided the original president or committee member corresponding with the area in which you would author and source are credited. like to volunteer. Utah Geological Association formed in 1970 from a merger of the Utah Geological Society, founded in 1946, and the Intermountain Association of Geologists, founded in 1949. Affiliated with the American Association of Petroleum Geologists. i GEOLOGY OF THE INTERMOUNTAIN WEST an open-access journal of the Utah Geological Association Volume 5 2018 A Preliminary Report of the Fossil Mammals From a New Microvertebrate Locality in the Upper Jurassic Morrison Formation, Grand County, Utah Brian M. Davis1,2, Richard L. Cifelli2, and Guillermo W. Rougier1 1Department of Anatomical Sciences and Neurobiology, University of Louisville School of Medicine, Louisville, KY, 40202; [email protected], [email protected] 2Department of Vertebrate Paleontology, Sam Noble Museum, University of Oklahoma, 2401 Chautauqua Ave., Norman, OK, 73072; [email protected] ABSTRACT The first Mesozoic mammals in North America were discovered in the Morrison Formation during the closing decades of the 19th century, as by-products of dinosaurs quarried by teams led by O.C. Marsh. These tiny fossils served as foundational specimens for our understanding of Mesozoic mammal evolution. There are now nearly 25 mammal-bearing localities known from the Morrison Formation, distributed across the West- ern Interior from the Black Hills to southern Colorado and west into Utah; the most historically important of these are in Wyoming (e.g., Como Quarry 9). Most Morrison mammals are known by jaws or jaw fragments, and several important Mesozoic groups (e.g., docodonts, dryolestoids, and to a large extent triconodonts and symmetrodonts) were established based on Morrison material, shaping the perception of mammalian diversity on a global scale. Despite heavy sampling of coeval sites elsewhere, the Morrison remains the most systematically diverse (at high taxonomic levels) assemblage of Jurassic mammals in the world. Here, we describe two mammalian specimens and highlight other remains yet to be fully identified from a new microvertebrate locality in the Morrison Formation of eastern Grand County, Utah. The site is positioned low in the Brushy Basin Member and is similar in lithology and stratigraphic level to the famous small vertebrate localities of the Fruita Paleontological Area, located less than 50 km to the northeast. In ad- dition to small archosaurs and squamates, limited excavation to date has yielded at least 20 mammalian spec- imens representing a minimum of six taxa, several of which are new and quite different from typical Morrison taxa. Preservation is generally excellent and includes partially articulated cranial and postcranial elements of small vertebrates. This new site has great potential to contribute new taxa and more complete morphological data than typical Morrison localities, underscoring the importance of continued field work in the Morrison. INTRODUCTION 2004), making it the most diverse Jurassic assemblage in the world and among the most diverse in the Mesozoic. The Upper Jurassic Morrison Formation has yielded Several factors were likely at play in generating this im- one of the most influential mammal assemblages world- pressive record: (1) intense study of the Morrison For- wide. At least 40 taxa are currently described from 20+ mation began in the 1870s (Marsh, 1878; Osborn, 1888; localities (Foster, 2003; Kielan-Jaworowska and others, see summary in Simpson, 1929), and (2) the unit has Citation for this article. Davis, B.M., Cifelli, R.L., and Rougier, G.W., 2018, A preliminary report of the fossil mammals from a new microvertebrate locality in the Upper Jurassic Morrison Formation, Grand County, Utah: Geology of the Intermountain West, v. 5, p. 1–8. © 2018 Utah Geological Association. All rights reserved. For permission to use, copy, or distribute see the preceeding page or the UGA website, www.utahgeology.org, for information. Email inquiries to [email protected]. 1 A Preliminary Report of the Fossil Mammals from a New Microvertebrate Locality in the Upper Jurassic Morrison Formation, Grand County, Utah Davis, B.M., Cifelli, R.L., and Rougier, G.W. a vast latitudinal range, with exposures across much of only from the FPA, highlighting unrecognized diver- the Western Interior. This geographic expanse, coupled sity and rich potential of continued exploration in the with as much as 7 Ma of depositional time (Trujillo and Morrison Formation. Our ongoing work at the CMQ is Kowallis, 2015), suggest that localities in the Morrison certain to increase the already high mammalian diver- possibly represent a wide range of paleoenvironments sity, provide evidence on intraformational geographical of slightly different ages. and temporal faunal variation, and produce further re- Typical Morrison mammal localities (e.g., Como finement of our understanding of the Morrison Jurassic Quarry 9 at Como Bluff, Wyoming) are characterized ecosystem, the standard against which any other Juras- by variations of a standard assemblage of taxa, known sic fauna is to be compared. mainly by jaws and jaw fragments: the mammaliaform Docodon, plagiaulacidan-grade multituberculate mam- INSTITUTIONAL ABBREVIATIONS mals, triconodontids, tinodontid “symmetrodonts,” and various dryolestoids (see Kielan-Jaworowska and oth- DINO = Dinosaur National Monument, Jensen, ers, 2004). Mammal assemblages vary across the extent Utah; OMNH = Oklahoma Museum of Natural Histo- of the Morrison Formation; this has been studied in de- ry, Norman, Oklahoma YPM = Yale Peabody Museum, tail (Foster, 2001, 2003; Foster and others, 2006) using New Haven, Connecticut. taphonomy and the distribution of Docodon, which is abundant
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  • Evidence of Diphyodonty and Heterochrony for Dental

    Evidence of Diphyodonty and Heterochrony for Dental

    第57卷 第1期 古 脊 椎 动 物 学 报 pp. 51–76 2019年1月 VERTEBRATA PALASIATICA figs. 1–9 DOI: 10.19615/j.cnki.1000-3118.180803 Evidence of diphyodonty and heterochrony for dental development in euharamiyidan mammals from Jurassic Yanliao Biota MAO Fang-Yuan1,2 ZHENG Xiao-Ting3,4 WANG Xiao-Li3,4 WANG Yuan-Qing1,2 BI Shun-Dong5 MENG Jin6,1 (1 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences Beijing 100044, China) (2 CAS Center for Excellence in Life and Paleoenvironment Beijing 100044, China [email protected]) (3 Institute of Geology and Paleontology, Linyi University Linyi, Shandong 276005, China) (4 Shandong Tianyu Museum of Nature Pingyi, Shandong 273300, China) (5 Department of Biology, Indiana University of Pennsylvania Indiana 15705, USA) (6 Division of Paleontology, American Museum of Natural History New York 10024, USA) Abstract Evidences for tooth replacement of known euharamiyidans are reported based on eight specimens of four species from the Jurassic Yanliao Biota, Liaoning Province, China. Tooth morphologies, eruptional and wear condition, and tooth germs are directly observed and/or revealed by Micro CT or slab CL scan. The euharamiyidan dentition has definite number of cheek teeth and monophyodont molars that are related to precise occlusion. Incisor germs are found in three specimens of Arboroharamiya but not in Shenshou lui and Xianshou linglong. The incisor germs in the upper jaw, presumably I2, have a large crown with two or three cusps; those in the lower jaw, interpreted as the permanent i2, are positioned dorsal to the root of the erupted incisor, interpreted as di2.