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Pollination Biology Movie

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Movie: Sexual Encounters of the Floral Kind

Lab Quiz today.

Guest Lecture Wednesday: Evo-devo of floral organs.

Guest Lecture next Monday: Conservation and management of rare in Washington.

MHG Project due next Wednesday. Phylogeny of (or Tricolpates) Eudicots (or Tricolpates)

“Basal eudicots”

(Soltis et al. 2011) Synapomorphies for Asterids

Iridoid compounds

www.scielo.br/img/fbpe/jbchs/v12n2/a04fig01.gif

Unitegmic ovules

http://www.nature.com/nature/journal/v450/n7173/images/nature06393-i2.0.jpg Tenuinucellate ovules

http://www.biologie.uni-hamburg.de/b-online/library/webb/BOT410/Angiosperm/LeucaenaOvule300NucellusLab.jpg Phylogeny of Asterids

Asterids

Cornales

Ericales

Lamiids: Campanulids: (APG 2003, Judd & Olmstead 2004, Soltis et al. 2005) N. hemisphere woody families with opposite

Clayton J. Antieau Ben Legler

Richard Olmstead http://static.guim.co.uk/sys-images/Guardian/Pix/pictures/2008/04/02/horsechestnut460x276.jpg Cornaceae (Dogwood family)

Cornus kousa Kousa dogwood

Textbook DVD KRR Cornaceae (Dogwood family)

13 genera, 130 species (, Davidia, Nyssa)

Habit:

Leaves:

http://www.naturekind.org/imgs/robbin/r/Cornaceae_Cornus_canadensis_29113.html

Ben Legler 2004 Cornaceae (Dogwood family)

13 genera, 130 species (Cornus, Davidia, Nyssa)

Habit:

Leaves:

Textbook DVD WSJ Ben Legler 2004 Cornaceae (Dogwood family)

Cornus unalaschkensis

Western bunchberry

Note this exceptional species that is a “subshrub” and not obviously woody.

Ben Legler 2004 Cornaceae (Dogwood family)

Inflorescence:

Flowers:

Ben Legler 2003

Sex of plant:

Rod Gilbert 2005 Cornaceae (Dogwood family)

Flower symmetry:

Perianth parts:

# of :

G. D. Carr

# of pistils/flower:

# of carpels/pistil:

Ovary position:

Textbook DVD KRR & DLN Cornaceae (Dogwood family)

Fruit type:

Textbook DVD JRA

Tim Hagan

Ben Legler 2005 Phylogeny of Asterids

Asterids

Cornales Cornaceae

Ericales Ericaceae Polemoniaceae

Lamiids: Campanulids: Gentianales Apiales Apiaceae Apocynaceae Rubiaceae Dipsacales Lamiales Caprifoliaceae Asterales Lamiaceae Asteraceae Scrophulariaceae Campanulaceae Solanales Solanaceae Boraginaceae (APG 2003, Judd & Olmstead 2004, Soltis et al. 2005) Ericaceae (Heather family)

http://georgian.wunderground.com/data/wximagenew/f/Feather3/5431.jpg Textbook DVD KRR & DLN macrophyllum Pacific Rhododendron (WA State )

Ryan Miller Ericaceae (Heather family)

Vaccinium contains ~450 species, including , , huckleberry, etc.

Textbook DVD WSJ corymbosum northern highbush blueberry

Textbook DVD KRR & DLN

Textbook DVD KRR & DLN Ryan Miller Ericaceae (Heather family)

Ben Legler 2004 124 genera, 4100 species (, , , , Rhododendron, Vaccinium)

Habit:

Leaves:

Tim Hagan 2004 Ericaceae (Heather family)

http://www.apsnet.org/education/illustratedglossary/PhotosI-M/mycorrhiza.jpg

Textbook DVD WSJ Ericaceae are strongly associated with mycorrhizal fungi and often uniflora mycotrophic. Indian-pipe

Ericaceae include a mycoparasitic Mycotrophic = nutritionally dependent on fungi underground, which, in turn are (holoparasitic) group: the dependent on other green plants through Monotropoids. mycorrhizal connections. (Kron 1996) Ericaceae (Heather family)

Inflorescence:

Flowers:

http://www.nestlerode.org/Cremation_Urn/urn_finished.jpg

Sex of plant: G. D. Carr 2006

Yaowu Yuan Yaowu Yuan Ericaceae (Heather family)

Flower symmetry:

Perianth parts:

Textbook DVD KRR & DLN http://www.rampantscotland.com/colour/supplement060527.htm

http://www.nestlerode.org/Cremation_Urn/urn_finished.jpg Ericaceae (Heather family)

# of stamens:

Yaowu Yuan Ben Legler

Textbook DVD KMN Ericaceae (Heather family)

# of pistils/flower:

# of carpels/pistil:

Ovary position:

Clayton J. Antieau Textbook DVD KRR & DLN Ericaceae (Heather family)

Fruit type:

Ben Legler 2005

Textbook DVD KRR Textbook DVD KRR & DLN Animal Pollination Syndromes (continued)

“Buzz” pollination by bees:

1) no nectar reward (pollen sole reward) 2) connivent anthers (anthers held together) at center of flower 3) anthers opening by terminal pores; pollen shakes out when the bee buzzes the flower

Yaowu Yuan

Ryan Miller Polemoniaceae ( family)

Textbook DVD KRR & DLN

Yaowu Yuan Phlox sp.

Ryan Miller Polemoniaceae (Phlox family)

Ipomopsis aggregata scarlet

Yaowu Yuan grandiflora large-flower mountain-trumpet

Textbook DVD KRR & DLN

Ryan Miller Yaowu Yuan Polemoniaceae (Phlox family)

18 genera, 380 species (Collomia, Gilia, Phlox, )

Habit:

Ben Legler 2005

Leaves:

Ben Legler 2004 Ben Legler 2004 Polemoniaceae (Phlox family)

Inflorescence:

Flowers:

G. D. Carr 2008 Sex of plant:

Ben Legler 2003 Polemoniaceae (Phlox family)

Flower symmetry:

Perianth parts:

Ryan Miller

G. D. Carr 2009 Polemoniaceae (Phlox family)

# of stamens:

# of pistils/flower:

# of carpels/pistil:

Ovary position:

Ben Legler Textbook DVD KRR & DLN Polemoniaceae (Phlox family)

Fruit type:

http://www.wnmu.edu/academic/nspages2/gilaflora/i_aggregata3.jpg