The Sarcophagidae (Diptera) of Norwayl
KNUT ROGNES
Rognes, K. 1986. The Sarcophagidae (Diptera) of Norway. Fauna norv. Ser. B, 33,1-26.
Sarcophagid material from Norway in the museum collections of the Universities in Bergen, Oslo and Tromsø and in author's own collection has been revised and records of the 49 spe- ciespresenflyknown fromNorwaypresented. 5 species,vrz. Amobiadistorta Gllen, 1926), Blaesoxipha rossica Villeneuve,1912, Helicophagella noverca (Rondani, 1869), Heterony- chia boettcheriana (Rodendorf, 1937) and H. ostensackeni (Rodendorf, 1937), arc recorded from Scandinavia for the first lime. Phrosinella septentrionalis Rodendorf, 1970 is establis- hed as a new junior synonym of Tachina sannioZettprstedt 1838 (now rn Phrosinella). Sar- cophaga tuberosa verticina Ringdahl, I 945 is tentatively established as a new junior syno- nym of Parasarcophaga (Liosarcophaga) pleskeiRodendorf, 1937. The male terminalia of Macronychia agrestis Gall6n, 1820\, M. polyodon (Meigen, 1824), M. stigineryis(Zetter- stedt, I 838) and Parasarcophaga pleskeiRodendorf, 1937 are illustrated. The zoogeography of the Norwegian species is discussed.
K. Rognes, Stavanger l,ærerhøgskole, Postboks 2521 Ullandhaue, N-4001 Stavanger, Nor- way.
INTRODUCTION given elsewhere. The genus has for the sake of convenience been included separately at the end The Sarcophagidae (rather unappropriately cal- of the present paper. led fleshflies) is a large and cosmopolitan fly fa- Strøm, (1 78 1784), Zetterstedt (1 1844, mily whose members usually can be recognised l, 838, 1845, 1855), Siebke (1877), (1889), by silvery grey and black colours, striped tho- Schøyen Storm (1891, 1896, 1907), Strand (1900), Biden- rax, striped, banded, tessellate or spotted abdo- kap (l 901), Ringdahl (1944a, 1944b, 1952, men, and a strongly setose body. In the world I 95 4) Rognes ( I 98 published fauna about 2300 species are known (Hennig and 2) have records of Sarcophagidae from Norway, and a total 197 3). They are very small to very large callpte- of 29 species (Helicobosca palpalis excluded) have rate flies with strong setae in a row on the hy- previously been recorded. The present popleuron (meron). All members are larvipa- revision lists 49 species from Norway, which 5 are rous. In the Scandinavian fauna they can be dis- of first records from Scandinavia. In comparison tinguished from the Tachinidae by an undevelo- it may be noted Ltrndbeck (1927) published ped postscutellum, from most Rhinophoridae by that 45 species from Denmark, Hackman (1980) 57 a posteriorly broad and truncate lower squama, from Finland, Ringdahl (195» 65 from Sweden and from most Calliphoridae by their non-me- and Pont 0975) 55 from the British Isles. tallic colour and the position of the outer ph seta on or medial to a line through the prst seta paral- MATERIALAND METHODS lel with the long æris of the fly. However, some Nearly all available Norwegian material of Sar- non-metallic calliphorids are similar to sarco- cophagidae, i.e. about 700 specimens, partly in phagids even in these respects. This applies to my own collection (380 specimens), partly in the some species of the genus Pollenia Robineau- museum collections of the Universities in Ber- Desvoidy, recognisable e.g. by presence of long gen (mainly O. Bidenkap, A. Fjellberg, L. Greve, waty golden yellow hairs in addition to the nor- A. l-øken, N. Knaben, T.R. Nielsen leg.), Oslo mal black thoracal vestiture, and to the genus (mainly J. Knaben and J.H.S. Siebke leg.), and Helicobosca Bezzi, recognisable by yellow palpi Tromsø (mainly T. Soot-Ryen lee.) form the and very strong setae in the lower half of the pa- basis the present study. In addition a few rafiacialia. My reasons for treating Helicobosca of small private collections (B. Sagvolden, J.H. Si- its a calliphorid rather than a sarcophagid will be monsen) have been examined. I have not seen the Trondheim collection (mainly V. Storm leg.) (1944a), l) Paper no. 4 in a series based on investigations fi- revised by Ringdahl nor foreign collec- lanced by a grant («vikarstipend») from Stavanger tions, except for occasional specimens. Lærerhøgskole 1982- I 983. The Norwegian species have been assigned to
Fauna norv. Ser. B, 33: l-26. Oslo 1986. 4 subfamilies, i.e. Miltogrammatinae (Miltog- 1980), and the sources used for identification ramminae), Macronychiinae, Paramacronychii- (see above). A few other papers have also been nae (Agriinae) and Sarcophaginae, in accor- consulted (Collart 1954, 1958, Saccå & Rivo- dance with the definitions (mainly based on the secchi 1954, Stackelberg 1962). In addition structure of the male postabdomen) of these or more detailed information on the North Euro- equivalent groups given by Rodendorf (1937, pean distribution is given, extracted from Pont 1967), Zumpt 0952, 1956) and Kurahashi (197 5) for the British Isles, Lundbeck (1927) for (1970, 1972, 1975). A short diagnosis for each Denmark, Tiensuu (1939, 1964), Junnikkala subfamily based mainly on Norwegian material ( I 960), Brander ( I 964), Hedstrom ( 1964), Heds- is given below. trcim & Nuorteva (1971), Hanski & Nuorteva (197 (197 The main works used for identification of spe- 5), Hanski 6), Hanski & Kuusela (l 977 , cies are those of Bottcher (1913 - l9l4), Stein 1980), Pyornilå & Pyornilå (1979), Hackman (1924), Lundbeck (1927), Rodendorf (1930, (1980) and Kuusela & Hanski (198 2)for Finland, 1935, 1937, 1970, l97la, 1975a, 1982), Seguy and Ringdahl (1937, 1945b, 1951, 1952, 1958) (1941), Ringdahl (1945b), Emden (1954), Kano, and Persson (1983) for Sweden. Field & Shinonaga (1967), Mihalyi 097 9c) and Verves (1982a). Numerous other papers have also been consulted, especially those of Aldrich (1916), Allen (1926), Draber-Monko 0964-- SYSTEMATIC LIST 1981), Gregor & Povolny (1961), Grunin (1964), Kurahashi (1970-1975), I-ehrer (1959- l98l). S u bfamily Miltogram m atinae Iæonide & llonide (197 3 - 1982), Mihålyi Diagnosis. Flies of small size. Arista bare. Gena (197 5 197 9b), Parker (l9l 1923), Patton in profile much narrower than half eye height. - 'Wainwright 6- (1934), Patton & (1935 - 1937), Po- Prosternum, propleuron and basisternum bare. volnf (1961 - I 977), Povolny & Slamedkovå Hind coxae bare behind. Posterior spiracle of (195 9 - 197 9), Povolnf & Stan6k ( 1969), Roden- thorax operculate with unequal lappets or rhi- dorf 0925-197 8), Rodendorf & Grunin (1938), nophorid-like with two subequal fringes. Male Rodendorf &, Verves 0977a- 1980), Shewell postabdomen with a T6, a compound T7 + 8, (197 l), Strukan (1967), Venturi 0947 - 1960), and an epandrium. Spiracle VI in membrane, Verves (197 3 - 1983) and Wainwright spiracle VII in T7 + 8. Processus longus absent (1928 - 1940). Within the Sarcophaginae (ex- or present. Epiphallus absent or present. cept Blaesoxipha Loew) no attempt has been The species in this subfamily prefer dry habi- made at naming female specimens. Even though tats and are associated with fossorial hymenop- Kulikova (1982) has given a key to 15 species terans, the larvae being parasites, predators or based on the structure of the female postabdo- guests in their nests. men, many females are still unknown or un- The nomenclature follows Rodendorf (1970). described. Almost all males which do not have One new combination and one new synonymy visible genitalia have been dissected for exami- are established. nation of these parts. The terminalia have been stored in glycerol in microvials to avoid distor- Genus Amobia Robineau-Desvoidy, I 830 tion or compression. Drawings have been made Pachyophthalmøs Brauer & Bergenstamm, 1889 from such preparations. The faunistic data have been presented in ac- l. Amobia distorta (Allen) cordance with the system of ØkJand (1981). The Pachyophthalmus distortus Allen, 1926 distribution in terms of the European Inverte- Material: AK, Frogn: Sønderstøa Degerud. EIS brate Survey (EIS) system for Norway (for refe- 28. rences, see Rognes 1980) is also given. For the Data of capture: l6 June. benefit of the reader a map (Fig. 26) is included Note. The single male specimen has been misi- showing the numbers assigned to each square of dentified by Ringdahl (1944b) as signata Mei- the EIS system. gen. However, the dissected genitalia leave no For each species I have given some notes on doubt as to its identity. Two females in the Oslo its biology. I have also indicated its world distri- collection (EIS 28, 47) and 3 females in my col- bution in broad terms. This information has lection (EIS 17 , 28) belong either to distorta og been extracted from the available catalogues for signata. different parts of the world (Downes 1965, So- Biology. Bred from nests of Trypoxylon politum vza l,opes 1969, Souza Lopes et al 19V7, Dear Say (Sphecidae) (Allen 1926) and Odynerus cras-
2 sicornis (Panzer) (Eumenidae) (Draber-Monko hetsli. EIS 4,7,24,28,32,33, 49, 55,62, 147. te64). Date of earliest and latest capture; 5 June, I Distribution. Palaearctic and Nearctic Regions. Sept. Not on the British Isles. Not previously recorded Biology. Bred from a Pompilus nest (Pompilidae) from Scandinavia. by Lundbeck (1927). Also reported as a clepto- parasite (feeding on the provisions of the host Genus Metopia Meigen, 1803 nest) of Sphex rufocinctus Brulle and Gorytes la- ticinctus (tæpeletier) (Sphecidae) (tomholdt 2. Metopia argyrocephala (Meigen) te7 s). Musca leucocephala Rossi, 1790, pr€occ.; Ta- Distribution. Palaearctic (cf. map in Verves china argyrocephala Meigen , 1824. 198 2b) and Nearctic Regions. British Isles. Den- Material: AK, Oslo: Alunsj øen, Bygdøy, Tøyen; mark (also specimens in my own collection), Frogn: Sønderstøa Degerud; Asker: Brønnøya. Finland (north to Ostrobottnia borealis and Re- ON, Vang: Øylo. BØ, Øvre Eiker: Burud, Ulle- gio kuusamoensis) and Sweden (north to Torne land. BV, Rollag: ?loc. TEY, Drangedal: Tørnes. Lappmark). TEI, Tokke: Dalen. AAI, Åmli, Bjorevja. RY, Sandnes: Melsheia, Myrland; Stavang€r: Kross- berg; Klepp: Øksnevad; Rennesøy: Vikevåg. 4. Metopia rondaniana Venturi HOY, Askøy: Herdla. HOI, Kvinnherad: Rosen- Metopia rondaniana Venturi, 1952 dal. SFI, l,eikaogef: Fosse. NSI, Beiarn: KvåI. Material. HES, Elverum: Grundset. EIS 55. EIS 7, 10, 14, 16,19,27,29, 31,35, 39,50, 52, Date of captur€: Unknown. 126. Biology. Unknown. Date of earliest and latest capture: I June, I I Distribution. Palaearctic Region east to Altai Aug. Mts. Not on the British Isles. Denmark (speci- Note. Only males have been included. Females mens in my own collection) and Finland. are not separable from those of rondaniana or tshernovae (among others) also occuring in Nor- way. 5. Metopia tshernovae Rodendorf Biology. Seguy (1941) and Venturi (19 52) cite a Metopia tshernovae Rodendorf, I 955 number of Apidae, Sphecidae and Vespidae spe- Material: AK, Oslo: Oslo. BØ, Ringerike: Høne- cies as hosts for the larvae. Lomholdt (197 S, foss. HEN, Åmot: ?loc. EIS 28, 36, 64? !97 6) reports it as parasitoid of crabro peltarius Dates of captur€: Unknown. (Schreb er), Sphex rufocinctus Brulle, Ammophila Biology: Unknown. pubescens Curtis and Oxybelus argentatus Cur- Distribution. Europe, Y:azakhstan. Not on the tis. However, as argyrocephala only relatively British Isles. Denmark (specimen in my own recently has been distinguished from species collection), Finland. such as tshernovae and rondaniana these re- cords may not be reliable. Genus Miltogramma Meigen, 1803 Distribution. Palaearctic (cf. map in Verves 1982b), Nearctic, Neotropical and Oriental (Ver- ves 197 9d). Regions. British Isles. Denmark 6. Mittogramma punctataMeigen (also specimens in may own collection), Finland Miltogramma punctata Meig€tr, 1824 (north to Ostrobottnia borealis) and Sweden Material: VAY, Lyngdal. Tofteland. EIS 4. (north to Torne Lappmark). Date of captur€: 20 Aug. Biology. Bred from nests of Ammophila hirsuta 3. Metopia campestr:is (Fallen) ScoD. (= Podalonia viatica) (Sphecidae) and Cot- Tachina campestris Fallen, 1820; Tachina ama- letes davesianus Smith, C. fodiensLatt., C. inex- bilis Meigen , 1824, teste Lundbeck 1927 : 224. pectatus Nosk. , C. succinctus L. and Halictus Material. AK, Oslo: Ljan, Tøyen. HES, Elve- sexnotatulus Nyl. (Apidae) (Tiensuu 1939, 56- rurl: Grundset. OS, Gausdal: Espedal (780 m). guy l94l , Draber-Monko 1969). ON, Nord-Fror: Moen. TEI, Vinje: Haukeliseter Distribution. Palaearctic Region. British Isles. (980 m). VAI, Kvinesdal: Gjemlestad. RY, Sta- Denmark, Finland (north to Tavastia australis) vanger, Krossberg; Strand: Prekestolhytta. RI, and Sweden (north to Gætrikland). Forsand: Røssdalen, Songesand. HOI, Eidfiord: Hjølmodalen (350 m), Sysenvatndammen (980 Genus Miltogrammidium Rodendorf, I 930 m). SFI, Balestrand: Brekke. TRI, Målselv: Fri- Pseudomiltogramma Rodendorf, I 930 7 . Miltogrammidium brevipilum (Yrlleneuve) 1020; Tachina pilitarsrs Zetterstedt, 1844: l02l; Miltogramma brevipilum Villeneuve, l9l I Phrosinella septentrionalis; Stackelberg 1962, Material: HES, Kongsvinger: Vinger. EIS 38? nomen nudum; Phrosinella septentrionalis Ro- Date of captur€: June (cf. Siebke 1877: 90). dendorf, 1970: 643, N.SYN. Note. The specimen stood in the Oslo collection Note on taronomy. I have examined the a ho- under Miltogramma oestracea Fall. In 1944 it lotype of Tachina sannio captured in Sweden at was sent to O. Ringdahl, who identified it ils «Åsele d. 28. Jul. I 8 3 2>> (in Museum of Zoology , «Miltogramma ? testaceifrons Ros.» according to Lund). I find it to be conspecific with the a pa- attached label. The specimen must be the one ratype of Phrosinella septentrionalis from from Kongsvinger mentioned in Siebke (1877: «Norv. Dowre» (in Naturhistorisches Museum, 90). The frons measures 0.32x and the face Vienna) (also examined). Furthermore, I have 0.23x head width and it runs to brevipilum rn examined the d holotype of Tachina pilitarsrs Venturi's key (1960). It fits Rodendorfs (1930: which was captured in Norway at «Oestre Næs 47) description well. Yærdaltæ 10 Jul. 1840» (in Museum of Zoo- Biology. Unknown. logy, Lund). Zetterstedt (1844: 102» suggested Distribution. Europe, Turkestan (in Kazakh that sannio and pilitarsrs were conspecific, and SSSR). Not on the British Isles. Sweden (Oland, Ringdahl (1934, 1937, 1945a) agreed with this, Gotland). though misidentifying both €N Phrosinella na- suta (Meigen). The abdominal pattern of the Genus Oebalia Robineau-Desvoidy, I 863 male and female, and the very characteristic fore Ptychoneura Brauer & Bergenstamm, 1889 tarsus of the male are shown in Figs. I -3. The true P. nasuta which apparently does not occur 8. Oebalia cylindrica (Fall6n) in Scandinavia (see below), is smaller, has a dif- ,Tachina cylindrica Fallen, I 8 10 ferent abdominal pattern (cf. Rodendorf l97la: Material: AK, Oslo: Tøyen. EIS 28. figs. 106, 109), and unmodified fore tarsi in the Date of captur€: I I June. male sex. I have seen I d I g from southern Biology . L.awae live in the nests of Crabro ambi- Yugoslavia (n.f-. Coe) (cf. Mihålyi 198 0), 2 g A guus Dahlb., C. capitosus Schuck, C. cinxius from southern France (C.f. Wainwright) and Dahlb. , C. tibialis Fabr. , Trypoxylon attenuatum I g from Spain (O.W. Richards) of nasuta (all in Smith (Sphecidae) (Tiensuu lg3g, Rodendorf British Museum (Natural History), London). Ro- 197 5a). The Larvae nourish themselves from pa- dendorf ( I 970 , 197 la, 197 lb) knew his sep ten- ralysed chironomid midges (Rodendorf 1975a). trionalis only from the female sex. Verves Distribution. Europe. British Isles. Denmark, (1980f) described the male of septentrionalis on Finland (north to Ostrobottnia borealis) and Sweden (north to Lycksele t appmark).
9. Oebalia minuta (Fallen) Tachina minuta Fallen, I 8 l0; Tachina ruJitars,s Meigen, 1824 (teste Ringdahl 1937: 38) Material: AK, Oslo: Tøyen. VE, Tjøme: ?loc. EIS 19, 29. Date of earliest and latest capture: 26 June, 7 Aug. Biology. Lårvae live in the nests of Crabro cin- xius Dahlb., Rhopalum clavipes (L.) and R. co- arctatum (Dahlb.) (Sphecidae) (Rodendorf 197 5a, Lomholdt 1,97 6). Distribution. Europe, Japan British Isles. Den- mark and Sweden (north to Ångermanland).
Genus Phrosinella Robineau-Desvoidy, I 863 Euhilarella Townsend, l91 5 ; Pedimya Roden- dorf, 1924 Figs. 1 -3 . Phrosinella sannio (Zetterstedt, 1838). I - Male abdominal pattern (Sande), 2 - Female ab- I 0. Phrosinella sannio (Zetterstedt), N.COMB. dominal pattern (Lauve), 3 Fore tarsus of male, Tachina sannio Zetterstedt, 1838: 636 1844: mediat view, (Sandå).
4 the basis of 2 d d from Taymyr in the Asiatic Biology. Bred from nests of Halictus gracilis part of the USSR. It is the same EN Zetterstedt's Mor. , H. subauraløs Rossi (Apidae) and Bembex pilitarsis and conspecific with Scandinavian ma- integra Panz., Crabro scutellaløs Sch€v., Oxybe- terial. Most interestingly, the Palaearctic sannio lus uniglumis L., Philanthus apivorus Latr., P. and all the Nearctic Phrosinella species listed by triangulum Fabr. , Sphex albisectuslnp., Tachy- Downes (1965) have modified fore tarsi in the tes unicolor Panz., Thyreopus peltarius Schreb. male (cf. Allen 1926, Reinhard 196l). Verves (Sphecidae) (Lundbeck 1927, Tiensuu 1939, 56- (1980f) therefore united them in a separate sub- guy l94l , Draber-Monko 1978). genus Euhilarella Townsend, as opposed to Distribution. Palaearctic Region. British Isles. Phrosinella (s.str.) containing the remaining Pa- Denmark, Finland (north to Ostrobottnia borea- laearctic species (including nasuta) which have lis and Regio kuusamoensis), Sweden (north to unmodified male tarsi. Norrbotten). Material: TEI, Kviteseid: lauve (E of Skredtve- it), Nordbø; Tokke: Lårdal. AAI, Åmli' Sandå. I 2. Senotainia imberbis (Zetterstedt) ON, Dovre: ?loc. (paratype a of septentriona- Tachina imberbis Zetterstedt, I 83 8; Ptychoneura /rs). NTI, Verdal: Østre Nes (holotype d of pili- crabronum Kramer, 1920 tarsii. EIS 10, 17 , 7l?, 98. Material: AK, Oslo: Tøyen; Bærum:Østerås. OS, Date of earliest and latest capture: 28 June, l0 Lunner: Grua. HOI, Ullensv€rog: Lofthus. EIS July. 28, 32, 36. Habitat: Roadsides, gravel pits, sandy ground. Date of earliest and latest capture: 8 June, 27 Biology. Unknown. July. Distribution. Iæningrad region and Taymyr Biology. Bred from nests of «Coelocrabo cinxius N.O. in USSR. All records from Finland (Tien- Dahlb. et capitosus Schuck.» (Seguy l94l). suu 1939, Hackman 1980) of nasuta (north to Distribution. Europe. Not on the British Isles. Ostrobottnia borealis and Regio kuusamoensis) Finland (north to Regio kuusamoensis), Sweden probably belong to the present species. I have (north to Torne lappmark). seen 3 d d from Sortavala (Karelia ladogensis) (now USSR) (Tiensuu leg.) and 3 d d I g from Paanajårvi (Regio kuusamoensis) (now USSR) Su bfamily Macronych iinae (R. Frey leg.) (a11 Museum of Zoology, Helsinki) reported as nasuta by Tiensuu (1939), which all Diagnosis. Flies of moderate to large size. Arista belong to sannio. All records from Sweden bare. Gena in profile about half of eye height. (north to Lule Lappmark) also probably belong Head shorter at vibrissae than at lunula, in pro- to the present species (cf. Ringdahl 195».I have file view. Prosternum, propleuron and basister- seen 6A d 17Q a fromSmåland,Norrbotten num bare. Hind coxae bare behind. Posterior and Lule Lappmark (Ringdahl leg.) (all Museum spiracle non-operculate, similar to the one in of Zoology, Lund) identified as nasuta by Ring- Rhinophoridae, with hairs along circular rim dahl which all belong to sannio. One of the fem- very short ventrally, developed as longer fringes ales (
6 Denmark, Finland and Sweden (north to Lule Lappmark).
S u b family Param acronych iinae Diagnosis. Flies of moderate to large size. Arista long pubescent or bare. Prosternum, propleuron and basisternum bare (but propleuron hairy in Dexagria Rodendorf, 1978). Hind coxae bare behind. Posterior spiracle of thorax operculate, posterior lappet larger than anterior one. Male postabdomen with T6,T7 and T8 all fused into a single large sclerite in front of the epandrium, carrying at least a transverse row of strong setae near fore margin. Behind this row a trace of a suture indicating posterior border of the T6 part. spiracle vI and vII within the sclerite. surstyli sometimes fused with the epandrium. Processus longus absent or very strongly reduced. Epi- phallus absent (Norwegian material) or present. The larvae are associated with other live or dead insects. Members of some genera are known to cause myiasis in man (wohtfartia Bra- uer &, Bergenstamm, Angiometopa Brauer & Bergenstamm). The nomenclature follows Verves (1982d. Note that the genus Helicobosca Bezzi is placed in this subfamily by most authors (cf. verves 1980c, I 982il, but that I consider it as belonging in the Calliphoridae. My reasons for this will be given elsewhere. Genus Agria Robineau-Desvoidy, I 830 Figs. 15 Macronychia striginervis (zetterstedt, -20. (see 1838). l5 - scutellar and abdominal pattern, inclu- Pseudosarcophaga Kramer, 1908 Shewell, ding bases of marginal setae, 16 Epandrium, sur- te7 t) styli posterior - and cerci, view, 17 Epandrium, 16. Agria mamillata (Pandelle) surstyli and cerci, left lateral view, 18 ST 5, ven- - Sarcophila mamillata Pandelle, I 896. tral view, 19 male postabdomen (T6, T7, T8 and - Material: AK, Oslo: Sognsvatn, Sørkedalen; ST 6) (epandrium removed), posterolateral view ,, 20 Bæ- phallosome, epiphallus, hypandrium, pregonite, rurll: Steinshøgda. TEI, Kviteseid: Kviteseid postgonite, phallapodeme and ejaculatory sclerite, gamle kirke. EIS 17 , 28, 36. left lateral view, (Stangenes). Date of earliest and latest capture: 2l June, 26 June. Biology. Bred from various Hyponomeuta spe- I 5 . Macrony chia striginervis (Zetterstedt) cies (Lundbeck 1927, Tiensuu 1939, Seguy Xysta striginervis Zetterstedt, I 8 3 8; Mittog- 1941 , Junnikkala 1960, Pyornilå & Pyornilå ramma ungulares Pandelle, 1895. 1979) and Lasiocampa pini (Seguy l94l) (both Material. HEN, Tynset: Tylldalen. TEI, Tokke: Læpidoptera). The fly u. . . oviposits on the skin Dalen. VAY, Kristiansand: Stangenes. EIS 2, of the last-instar lawae of Hyponomeuta species. 16, 7 2. The hatchinglawae penetrates the skin and lives Date of earliest and latest capture: 28 June, 4 as an endoparasite of its host. After destroying July. the first host, the parasite larva attacks a second Note. The male from Stangenes has been dissec- one and so on until it is ready for pupation. The ted. The scutellum and abdomen, including ter- parasite hibernates as a pupa in the ground.» minalia are shown in Figs. l5 -20. (Junnikkala 1960: 34- 35). Biology. L,anrae in the nests of Clytochrysus ca- Distribution. Palaearctic Region. British Isles. vifrons Thoms. (Sphecidae) (Emden 1954). Denmark, Finland (north to Ostrobottnia borea- Distribution. Palaearctic Region. British Isles. lis) and Sweden (Skåne, Vdrmland). Genus Brachicoma Rondani, 1856. singly it was attracted to meat bait by Gregor & Brachycoma, emend. Povolny (1961). Distribution. Palaearctic and Nearctic Regions. 17 . Brachicoma devia (Fallen) British Isles. Denmark, Finland and Sweden Tachina devia Fallen, 1820; Tachina vilisZetter- (north to Torne lappmark). stedt, 1844; Brachicoma devia borealis; Ring- dahl 1944b. Material: AK, Oslo: Tøyen. HEN, Åmot: ?loc.; Su bfamily Sarcoph aginae Tynset: Tylldalen. OS, Sør-Froo: Harpefoss. ON, Diagnosis. Flies of very small to very large size. Vågå: Vågåmo. BØ, Kongsber§: Hvittingfoss; Arista with long hairs in basal half. Prosternum Hurum: Filtvet; Flesberg: Hvila. BV, Rollag: and basisternum hairy. Propleuron bare. Hind Rollag stasjon. VE, Tjøme: Tjøme. TEI, Bø: W- coxae hairy behind. Posterior spiracle of thorax reb; Seljord: Ulver€s; Kviteseid: Kviteseid, operculate, with large posterior lappet. Coxople- Skredi; Fyresdal: between Aslestad and Slystøyl ural streak absent (cf. Downes 1955). Male post- (650 m); Tokke: Langes æ 650 m). VAY, Lindes- abdomen with T6 reduced to narrow transverse I1€s: Ytre Jørgenstad. VAI, Kvinesdal: Gjemles- strips carrytng long hairs or completely absent. tad. RY, Stavan§€r: Byhaugen, Krossb€rg, Sta- T7 + 8 (first genital segment) without indications vanger, Tjensvoll; Randab€r§: near Hålands- of sutures and never with transverse row of se- vatn. RI, Forsand: lærang. HOY, Bergen: Lakes- tae near fore margin, but often with marginal våg, Storetveit, Åstveit (Golfbanen). I.[I, Stein- row. Spiracle VI in membrane, spiracle VII in §er: Gulbergaunet. EN, Porsanger (FI, Karasjok T7 + 8. Surstylus small. Processus longus very ?), Vuorje. EIS 1,4,7,16,17,18, 19,27,28, 30, strongly reduced (see also Patton 1934, Patton & 39, 62, 64?,71,72, 101, 166. Wainwright 1935a, 1935b, 1936a, 1936b, 1937 , Additional material (not seen). «Tromsø» (TRY, as «spiracle VIII» or «proximal segment of ninth Tromsø: Tromsø, EIS 162) (Ringdahl 1954). coxite», and Zumpt & Heinz (1950), as «lOth Date of earliest and latest capture: 24 April, 6 sternum»). Epiphallus absent. Aug. Some species have larvae which are sapro- Note. The specimen from Vuorje («Vuorjegai- phagous, developing in excrement, small-size sa») has'been identified by O. Ringdahl as <
8 Distribution. Palaearctic and Afrotropical Re- 2l . Sarcotachinella sinuata (Meigen) gions. Not on the British Isles. Not reported Sarcophaga sinuata Meigen, I 8 iG; Sarcophaga from Denmark by Lundbeck (1927), but i ha,oe arvorum Metgen, 1826. collected 22 q in that a country on a locality Material: Ø, Halden: reåf Halden. Ry, Sta- where it was very numerous on dry sun-expo- vanger: Sunde (indoors on window). ptS 7 20. qe_q ground (East Jutland, , Mols Bjerge 12 July Dates of captur€: The specimen from sunde w€§ 197 9). Æso known from Sweden (north to Hål- captured 20 June. singland). Note. Zetterstedt (1845: 1289) reported this spe- cies in Norway «ad Næs Verdaliæ» (NTI, V-er- dal: Nes) (cf. Schøyen 1889: 7), but the record 19 . Blaesoxipha gladiatrix (pandelle) was retracted by zetterstedt ( I 8 5 9: 3267). I have pandelle, not sqrcophaga gladiatrix l g96 ; Blaesox- seen this material. ipha laticornis: Rodendorf 1937 (only males); Biology. Parasite of grasshoppers (Aldrich 1916, Wainwright 1940 Rodendorf & verves 1978). Lundbeck reared it Material: AK, oslo: Kværner; Bærurn: Østerås, from a pupa found in the ground. Attracted to (Rodendorf Øverland. BØ, Øvre Eiker: Ulleland. TEI, sau- faeces 1959). A single specimen cap- herad: Nordagutu. MRy, Gjemnes: Osmarka tured on food markets in Budapest (Aradi & Mi- (Raumyr). EIS 18, 28, 84. hrilyi 197 l). Distribution. Date of earliest and latest capture: 23 June, 5 Palaearctic and Nearctic Regions. Aug. British Isles. Denmark, Finland (north to Karelia Note. The males and females agree with Iionide borealis) and sweden (north to Gdllivare in & llonide's (1979, 1982) conCept of gladiatrix. Torne Lappmark). B_1ology. Parasitic in grasshoppeis, moitty of the c horthippus bicolor - mollis - biiuttulus grorp (t,- Genus Helicophagella Enderlein, l92B onide & L6onide 1979). C. bicolor and C.btgut_ Bellieria: authors, not Robineau-Desvoidy, teste tulus occur in Norway (Knaben 1943). §ouza Lopes (1955) and Souza Lopes et al Distribution. Incompletely known becaus e gla- Qe77). 4,gfA only recently (tæonide & I_eonide l§7 9, 198» has been kept apart'from Blaesoxipha 22. Helicophagella agnata (Rondani) grylloctona [,oew. Known with certainty from Sarcophaga agnata Rondani, 1860 France (including Corsica) and the Britisir Isles. Material: RI, Forsand: Songesand. SFI, Bale- strand: Bøyum. EIS 7, 59. Additional material (not seen). «Aurland I ex.» (SFI, Aurland: Aurland, EIS 5l ) (Rinedahl 20. Blaesoxipha rossicø Villeneuve 1954). The specimen could not be founO in Blaesoxipha rossica Villeneuve, l9l2 Ringdahl's collection in Lund (R. Danielsson in Material: MRY, Gjemnes' osmarka (Raumyr). litt. 2 Sept. 1983). EIS 84. Fiology. Bred from Helix aspersa Mtiller, 1774 Date of capture : 14 July. (Emden 1954). This snail ii not known from Note. The bvipositor is very well visible and the Norway (cf. Kerney & Cameron 197 9). specimen fits the description of Rodendorf Distribution. Europe. British Isles. Denmark, (193 7) and Emden (1954) very well. Finland and Sweden (Skåne, Småland). Biology. Lawae parasitic in grasshoppers (Ro- dendorf 1932, 1937, Seguy l94l , L"eonide I967, 23. Helicophagella crassimargo (Pandelle) I-eonide & I-eonide l97l ). Of the hosts mentio- Sarcophaga crassimargo Pandelle, I 896 led by Iionide & Iæonide ( l97l), Chorthippus Material: AK, Oslo: Tøyen; Eidsvoll: Eidsvoll. bicolor and C biguttulus occur in Norway (cf. HES, Elverum: Grundset. OS, Sør-Fron: Harpe- Knaben 1943). foss. ON, Nord-Froo: Vinstra. VAI, Kvinesdal, Distribution. Palaearctic Region. British Isles. Gjemlestad. NNØ, Sørfold. Røsvik. EIS 4, 28, Not previously reported from scandinavia. It 37,55,62, l3l. frdy, however, have been confused with B. eryt- Date of earliest and latest capture: 8 July, 20 hrura (Meigen) which has been recorded from Aug. Denmark, Finland and Sweden. Biology. Keilin (1919) bred a female of suppo- se.dly this species from Helicella virgata (da Cos- Genus Sarcotachinella Townsend, lglT ta), which is unknown from Norway (cf. Ker- Arhopocnemis Enderlein, l92B ney & Cameron 197 9). A specimen was captu- red on food-markets in Budapest (Aradi & Mihå- single, rather northern locality in Scandinavia lyi l9t l). suggests a disjunct boreo-alpine or boreo-mon- Distribution. Palaearctic Region east to Central tane distribution in Europ€, even though it has Asia. British Isles. Denmark, Finland (north to been taken in Belgium (Collart 1958) and near Ostrobottnia kajanensis) and Sweden (north to the Polish baltic coast (Draber-Monko 197 3a). Norrbotten).
(Meigen) 24. Helicophagella melanura 26. Helicophagella rosellei (Bottcher) Sarcophaga melanura Meig€n, 1826; Myiophora Sarcophaga rosellei Bcittcher, l9l2 striata Robineau-Desvoidy, I 8 30 Material: AK, Oslo: Bygdøy; Frogn: Drøbak; Material: AK, Oslo: Bekkelaget, ?loc.; Bærum: Bærum: Nordli NW of Østerås. OS, Ringebu: Nordli NW of Østerås. HEN, Rendalen: Solbak- Elstad; Nordre Land: Dokka. VE, ?loc. VAY, ken near Storsjøen. AAY, Grimstad: Ryvingen. Flekkefiord: Lauvnes. NSY, Meløy: Storglom- (Kvisla). VAY, Mandal: Mandal RY, Klepp: Bo- vatn (above tree-line at about 520 m). NSI, Salt- resanden, Orresanden; Randaberg: Sandestran- dal: Rognan. EIS 4, 19?, 28, 45, 63, 126?, 127. den. SFI, Årdal: Svalheim i Utladalen. EIS 2, 6, Date of earliest and latest capture: 3l May, 30 7,28,51,64. July. Date of earliest and latest capture: I June, 27 Biology. Unknown. Aug. Distribution. Europe. Altai Mts in Central Asia. Biology. Breeds in human and animal faeces, British Isles. Northern Sweden (HarjeAalen, (Roden- animal carcasses, privies and garbage Jdmtland, Ångermanland, Torne Lappmark). A dorf 1959, Mihalyi 1965, Ishijima 1967, Kano, disjunct boreo-alpine or boreo-montane distri- Field & Shinonaga 1967, Sj.chevskaya 1970, bution in Europe. Rodendorf & Verves 197 8), sometimes parasitic (Keilin on various molluscs l92l , Seguy l94l), Genus Thyrsocnema Enderlein, lg28 insects (Seguy l94l), birds (Emden 1954) and mammals (Seguy l94l , Emden 1954, Nielsen, Nielsen & Walhovd 1978). Adults visit faeces, meat (including dead snails) and fruits for nou- 27 . Thyrsocnema incisilobata (Pandelle) rishment (Kirchberg 1954, Gregor &, Povolny Sarcophaga striata Meigen, 1826, preocc. in 196l , Aradi & Mihalyi l97l ). Synanthropic spe- Sarcophaga by Musca striata Fabricius, 1794 cies of hygienic importance. (= Ravinia pernix Harris); Sarcophaga incisilo- Distribution. Palaearctic, Nearctic and Oriental bata Pandelle, 189 6; Sarcophaga privigna Pan- Regions. British Isles. Denmark, Finland (north delle, 1896, preocc. to Ostrobottnia media) and Sweden (north to Material: AK, Oslo: Bekkelaget, HovedØyz, Torne Lappmark). Oslo; Frogn: Sønderstøa Degerud; Bærum: Øverland. OS, Sør-Fror: Harpefoss; Øyer: Øyer. 25. Helicophagella noverca (Rondani) ON (OS?), «Fron>> (Nord-Fron or Sør-Fron): Sarcophaga novercø Rondani, 1860 ?loc. BØ, Lier: Gjellebekk; Røyken: ?loc. BV, Å1, Material: NSY, Meløy: Storglomvatn (above Torpo. VE, Brunlafl€s: Mølen. TEI, Sauherad: tree-line at about 520 m). EIS 126? Nordagutu; Kviteseid: I.auve, Skredi; Tokke: Note. In dorsal view first genital segment lårdal. VAY, Flekkefiord: Hidra (Dragøy, Kirk- (TZ + 8) of examined male specimen with large hamn, Kråkedal). RY, Sandnes: Sviland. RI, circular dusted spot surrounded by narrow band Forsand: Songesand. HOI, Granvin: Skjervet. of black gloss, hind margin dusted, the marginal EIS 4, 7, I l, 17, 18, 28, 4l , 43, 54, 62. setae hairlike and long. Terminalia dissected. Date of earliest and latest capture: 8 June, I I Biology. Bred from a dead Helix sp. (Schmitz Aug. l9l7) and from 'Helix pomatia L. (Enderlein Biology. Larvae coprophagous, also reported to 1928, Seguy l94l). Eberhard (1955) bred it parasitise a grasshopper (Seguy l94l). Bred from horse meat baits and kept it easily in cul- from a snail (Keilin 1919, Rodendorf 1967). ture on meat. No larval development occured on Adults attracted to faeces, meat and dead snails faeces. A single specimen captured on the food- (Kirchberg 1954, Gregor & Povolny l96l). markets in Budapest (Aradi & Mihålyi 197 l). Distribution. Palaearctic Region east to Central Distribution. Europe (mainly in mountainous Asia. British Isles. Denmark, Finland (north to districts), Caucasus. Not on the British Isles. Not Karelia borealis) and Sweden (north to Jåmt- previously reported from Scandinavia. The land). l0 28. Thyrsocnema kentejarea Rodendorf Date of earliest and latest capture: 30 May, I I Thyrsocnema kentejanø Rodendorf, 1937; Thyr- Aug. socnema kentejana lapponica Tiensuu, 1939 Biology. Occasional parasite «de l'escargot» (Se- Material: NSI, Grane' Majavatn (3 l0 m). EIS guy l94l), of Helix aspersa (Rodendorf 1967, lll. Draber-Monko 1978). Date of captur€: 22 July. Distribution. Europe, Caucasus. British Isles. Biology. Unknown. Denmark and Sweden (Skåne, Gotland). Distribution. North China, Mongolia, USSR (South Siberia, Indigirka Basin, Sikhote-Alin), Genus Discachaeta Enderlein, 1928 Central Europe (Romania). Not on the British Isles. Finland (tapponia inarensis) and North (Meigen) Sweden (south to Hårjedalen). A disjunct boreo- 32. Discachaeta pumila alpine distribution in Europe. Sarcophaga pumila Meigen, 1826 Material: AK, Oslo: HovedØyz, Tåsen; Bærum: Genus Bellieromima Rodendorf, 1937 Nordli NW of Østerås, Østerås. BØ, Røyken: Bellieromima Rodendorf, 1937, as subgenus of Slemmestad. EIS 28. Thyrsocnema. Date of earliest and latest capture: 15 July, 16 Aug. 29 . Bellieromima subulata (Pandelle) Biology. Unknown. Sarcophaga subulata Pandelle, 1896 Sarcop- Distribution. Europe. North Africa. British Isles. haga laciniata Pandelle, 1896. Denmark, Finland (north to Tavastia borealis) (north Material: AK, Oslo: Bygdøy. EIS 28. and Sweden to Uppland). Date of capture: 25 June. Biology. Unknown. Genus Heteronychia Brauer & Bergenstamm, Distribution. Europe, Caucasus. British Isles. 1889 (misidentification) Denmark, Finland (Nylandia) and Sweden Pierretiø: Rodendorf 1937 (Skåne, Ångermanland). 33. Heteronychia boettcheriareø (Rodendorf) Genus Pierretla Robineau-Desvoidy, I 863 Pierretrø (s.str.) boettcheriarea Rodendorf, 1937 Thyrsoctt€lna: Rodendorf 1937 , in part. Material: TEI, Tokke: LårdaL EIS 17. Date of captur€: 4 July. 30. Pierretia clathrata (Meigen) Biology. Unknown. Sarcophaga clathrata Meigen, 1826; Sarcop- Distribution. Europe, Caucasus. Not on the Bri- haga hinei Aldrich, 1916 tish Isles. Not previously known from Scandina- Material: HEN, Åmot: ?loc. VE, Hof: Thorrud. via. Seguy 0941) reports it north to South Swe- TEI, Kviteseid: Skredi; Fyresdal: between Asles- den, but his source appears to be Lundbeck's tad and Slystøyl (650 m); Tokke: Dalen. VAY, (1927) information concerning «offuscata», lis- Flekkefiord, Nuland. HOI, Granvin: Eide. EIS 4, ted as a synonym by Seguy. Lundbeck's offu- 16, 17 , 28, 4l , 64? scata has the postabdomen black and is not the Date of earliest and latest capture: 24 June, 28 present species (probably obscurala Rodendorf). July. Biology. Bred from egg-sacks of «Epeira cornu- ta» (= Araneus cornutus Clerck, 17 57 of l"ocket 34. Heteronychia haemorrhoa (Meigen) & Millidge 1953) (lvtit< 1890, Lundbeck 1927) Sarcophaga haemoruhoa Meigen, 1826 and Clubiona sp. (Grunin 1964), and from grass- Material: AK, Oslo: Bekkelaget, Grefsen; Frogn: hoppers (Seguy I 941). Drøbak, Sønderstøa Degerud; Bærum: Høvik, Distribution. Palaearctic and Nearctic Regions. Nordli NW of Østerås, Østerås. VE, ?loc. TEI, British Isles. Denmark, Finland (north to lappo- Tokke: LårdaL VAY, Mandal: Mandal. EIS 2, nia enontekiensis) and Sweden (north to Torne 17 , 19?, 28. Lappmark). Date of earliest and latest capture: l0 June, 27 Aug. 31. Pierretia soror (Rondani) Biology. Bred from Cepaea hortensrs (L.) (Vtit< Sarcophaga soror Rondani, 1860 I 890, Schmitz 1917, Keilin I 9l 9). Material: AK, Oslo: Tøyen; Bærum: Nordli NW Distribution. Europe. British Isles. Denmark, of Østerås, Østerås. BØ, Røyken: Slemmestad; Finland (Alandia) and Sweden (north to Upp- Hurum: Filtvet. EIS 28. land). ll 35. Heteronychia obscurata (Rodendorf) Denmark, Finland (north to Lapponia inarensis) Pierretra (s.str .) obscurata Rodendorf, 1937 and Sweden (north to Torne Lappmark). Repor- Material: AK, Bærum: Øverland. EIS 28. ted from above tree-line in Jåmtland (Sweden) Date of capture: 23 June. (Ringdahl l95l). Note. Ringdahl (1944b) recorded a male from AK, Bærum: Høvik l6 Aug. 1935 (EIS 2$. The specimen was not in the museum collections ex- 38. Heteronychia vicina (Macquart) amined. Sarcophaga vicina Macqu&rt, 183 5; Sarcophaga Biology. Unknown. ebrachiata Pandelle, I 896 Distribution. Palaearctic and Oriental Regions. Material: AK, Bærum: Nordby gård. HEN, British Isles. Denmark, Finland (Regio Aboen- Åmot: ?loc. OS, Gausdal: Aulestad; Nordre sis) and Sweden (north to Hårjedalen). [,and: 3 km N of Dokka. ON, Vang: Øylo. VAY, Flekkefiord: Rasvåg. VAI, Kvinesdal: Gjemles- 36. Heteronychia ostensacke,rzi (Rodendorf) tad. RY, Karmøy: Kopervik. RI, Forsand: to- Pierretra s.str . osten-sackeni Rodendorf, 1937 wards Prekestolen. STI, Oppdal: Kongsvoll Material: AK, Oslo: Bekkelaget. EIS 28. (Blesbekken). NSY, Bodø: Bodø. EIS 4, 7 , 13, Date of captur€: 27 Aug. 28, 45, 52, 54, 64?, 79, 130. Note. The two specimens (Oslo collection nos. Date of earliest and latest capture: 18 May, I I 77 64 ( d ), 77 63 ( q )) were associated in the ori- Aug. ginal drawer and stood under the name «,Sør- Note. Ringdahl's male specimen (1944b, &s cophila latifrons Fall.» Both sexes carry identical <») from TRY, Karlsøy: Måkeskjær labels in Siebke's handwriting «Bæklag (EIS 170) could not be found in the Tromsø or 27 .8.46>>. The vein 17 has a single seta on the up- other Norwegian collections. The identification per wing surface in both specimens. Each mea- is queried by Ringdahl himself so it should pro- sures 4 mm. bably not be accepted. The recorded from ON, Biology. Unknown. Dovre: Hjerkinn (EIS 7l?) (Ringdahl I 954) is ac- Distribution. Czechoslovakia, Poland (Pienin), cepted. West Germany (Heidelberg, terra typica) (Dra- Biology. Unknown. ber-Monko 1978). Not previously known from Distribution. Mountainous regions in Central Scandinavia. The few records available suggest Europe, Caucasus. British Isles. North Sweden a boreo-alpine or boreo-lnontane distribution in (Torne tappmark south to Hårjedalen). Recor- Europe. ded from above the tree-line in Jåmtland (Swe- den) (Ringdahl l95l). A disjunct boreo-alpine 37. Heteronychia vagares (Meigen) distribution in Europe. Sarcophaga vagans Meigen, 1826; Sarcophaga frenata Pandelle, 1896; Sarcophaga cruenta Genus Parasarcophaga Johnson & Tiegs, l92l Pandelle, 1896 Material: Ø, Halden: fl€&r Halden. AK, Oslo: Oslo, Tåsen; Bærum: Nordby gård. HEN, 39. Parasarcophaga aratrix (Pandelle) Åmot: ?loc. OS, Lillehamfii€r: Maihaugen in Lil- Sarcophaga aratrix Pandelle, 1896 lehammer, near Gausa river by Flåkåli bru W of Material. AK, Oslo: Sognsvatn. OS, Sør-Fror: E Fåberg; Sør-Froo: E of Hundorp. ON, Nord- of Hundorp. AAY, Hisøy: Hisøy. VAY, Kristi- Fron: 3 km E of Vinstra.BØ, Ringerike, Løvlia. ansand: Vrånes. HOY, Fitjar: ?Fitjar. EIS 2, 6, VE, ?loc. TEI, Kviteseid: Heggtveit, Skredtveit; 22, 28, 63. Tokke: Lårdal. HOY, Bergen: Fantoft; Sam- Date of earliest and latest capture: 2l June, 22 nanger' Ådland. SFI, Aurland: Fretheim. NSI, July. Saltdal: Saltfiell. TRY, Tromsø: Tromsøya. EIS Biology. Bred from Lymantria monacha L. (tæ- 17,20,28,30,36, 40, 41, 54, 62,63,64?, 127?, pidoptera) and Prionus coriarius L. (Coleoptera) 162. (Lundbeck 1927 , Seguy l94l ), from horse meat Date of earliest and latest capture: I I June, I baits (Eberhardt 1955), and from a bird bait (Ku- Aug. usela & Hanski 1982). Adults attracted to faeces, Note. The female recorded by Rognes (1982) meat and fruits (Gregor & Povolnf l96l). from Dovre is not accepted here. Distribution. Palaearctic Region east to Kamc- Biology. Bred from snails of the family Succinei- hatka. British Isles. Denmark, Finland (north to dae (Verves 197 O. Ostrobottnia borealis) and Sweden (Skåne, Små- Distribution. Palaearctic Region. British Isles. land). t2 40. Parasarcophaga emdeni Rodendorf Sarcophaga teretirosfrus; Lundbeck 1927, Para- sa r c o p ha § a (L io sa r c o p h a ga) t e r e t i roslris ; Roden - dorf 1937, not Pandelle (misidentifications); Pa- rasarcophaga emdeni Rodendorf, 1959, nomen nudum; Parasarcophaga emdeni Rodendorf, 1969,1970 Note. Previously some confusion existed as to the correct use of the name teretiroslris Pandelle, 1896. Emden (1954) and Rodendorf 0969, 1970) applied it to the species described as kroe- beri Rodendorf, 1937 . Rodendorf 0969, 1970) introduced the name emdenifor the species misi- dentified as «teretirostris» in his 1937 work. See Povolny & Slamedkovå 097 9) for a discussion of this problem. Note that these authors miscite the date of emdeni, Rodendorfs paper of 1969 having apparently been overlooked. Note also that Nordic records of «teretirostris» (Tiensuu 1939, Hedstrom 1964, Brander 1964, Ringdahl 1952) probably all refer to emdenf. The true te- retirostns is apparently absent from the Nordic countries but occurs in Western Europe from the British Isles southwards to Spain, and also in E. Germany and Poland (Povolnlf & Slame- dkovå 197 9). Hackman (1980) appears to regard the names as synonyffis, though provides both names with a question-mark. Material. TEI, Tokke: I-ardal. EIS 17. Date of capture: I July. Figs. 2l -24. Parasarcophaga pleskei (Rodendorf, Biology. Adults attracted to faeces (Rodendorf 193T. 2l Cerci, posterior vrew, 22 Cerci and pregonite- postgo- 1959). Collected on the food-markets in Buda- surstyli, left lateral view, 23 - and view, 24 phallosome, distal hall pest (Aradi & Mihalyi l97l). Breeds in dead He- nite, ieft lateral - left iateral view (both styti shown), (Åmot). licella candicares (Veryes & Kuz'movich 1979). This snail does not occur in Norway (Kerney & Cameron 1979, as H. obvia). been identified and published by Ringdahl Distribution. Palaearctic Region (including (1944b) as Sarcophaga tuberosa Pand. The spe- North Africa) east to Altai Mts. Not on the Bri- cimen from Åmot has been dissected by me tish Isles. Denmark, Finland (north to Ostrobott- (Figs. 2l -24). Note that in most specimens I nia borealis) and Sweden (Skåne, Oland, Got- have examined (not the one figured) there is a land). slight bulge distally on the lower edge of the cerci its seen in profile as shown in Rodendorfs 41. Parasarcophaga pleskei Rodendorf (193T fig. 328. I have examined 4 A d (Mu- Parasarcophaga Øiosarcophaga) pleskei Roden- seum of Zoology, Lund) from Torne Lappmark dorf, 1937; Sarcophaga tuberosa Pandelle ssp. (labelled «Abisko» and «Jebrenj. ») (Sweden) verticina Ringdahl, 1945a, TENTATIVE (Ringdahl leg.) identified as tuberosa Pand. (ac- N.SYN .; Sarcophaga tuberosa Yar. harpax; Ald- cording to labels) by Ringdahl and placed in his rich 1916 (excluding right part of his fig. 79); collection as tuberosa spp . verticina. The genita- Sarcophaga harpax; Roback 1954, Downes lia which are in an extended position are closely I 965. similar to those of the Norwegian specimens. Material: HEN, Åmot: ?loc. TRI, Balsfiord: Recently I have collected 2 A d on Newfound- Aspenes. EIS 64?, 154. land (Canada) which also are conspecific with Dates of capture: The specimen from Aspenes the Norwegian material. They fit Aldrich's ( has been captured 1 I July. I 9 I 6) and later American authors' concept of Notes. The male specimen from Aspenes, which «harpax», and G.E. Shewell, Ottawa, has identi- has the genitalia extended and fully visible, has fied my specimens as such. Note that this con-
13 cept of «harpax» is quite different from that of baits Gncluding liver, horse meat, dead snails) Rodendorf (1937), Gregor & Povolny (1961), (Kirchberg 1954, Eberhardt 1955, Gregor & Po- Kano, Field & Shinonaga 0967) and Mihålyi volny 196l , Mihålyi 1965, Hanski l976,Hanski 097 9c). & Kuusela 1977 , 1980). ((. . . one of the most Biology. Unknown. common sarcophagid flies in Japan. . . Larvae Distribution. Caucasus, Central Asia, China breed in garbage, dead animals, animal excre- (Rodendorf 1937 , Fan 1965, Verves 197 8d). ment and occasionally in lavatories. Some cases Sweden (Torne Lappmark, Hdrjedalen) (alpine of intestinal myiasis . . . have been reported in and subalpine zones). Also in North America Japan.» (Kano, Field & Shininaga 1967). (according to own specimens from Newfound- Distribution. Palaearctic and Oriental Regions. land, see above). British Isles. Finland (Nylandia, Tavastia austra- lis), Sweden (I have examined a male from Upp- 42. Parasarcophaga portschinskyi Rodendorf land, Sandhamn 2l July 1929, Ringdahl leg., Parasarcophaga (Liosarcophaga) portschinskyi misidentified as tuberosa by Ringdahl, see above Rodendorf, 1937. under P. portschinskyi) (not recorded by Ring- Material: TEI, Tokke: lårdal. EIS 17. dahl 1952), but not reported from Denmark. Date of captur€: 4 July. Biology. Larvae coprophagous (Rodendorf Genus Robineauella Enderlein, 1928 1959). Adults attracted to meat baits (Gregor & Povolny 196l). Also collected on food-markets 44. Robineauella scoparia (Pandelle) in Budapest (Aradi & Mihålyi I 97 l). «The larvae Sarcophaga caerulescens Zetterstedt, I 8 3 8, teste are necrophagous and may be facultative preda- Ringdahl 1944a, 1945a, 1945b, 1952; Sarcop- tors of butterfly larvae.» (Rodendorf & Verves haga scoparia Pandelld, 189 6; Sarcophaga sco- 197 8).» . . . can be reared with decomposing paria nearctica Parker, 1916. animal matter in the laboratory. Someti- Notes. I have retained Pandelld's name pending mes. . . parasites of lepidopterous larvae.» a modern revision of all Zetterstedt's types. I (Zhang 198». have also examined 2 d d of nearctica from Distribution. Palaearctic Region east to China. Newfoundland (Canada). They have the epan- Not on the British Isles. In the Nordic countries drium and partly the T7 + 8 red but otherwise previously listed only from Finland (Hackman the terminalia appear identical with those of the 1980). I have collected a male in Denmark (East Norwegian specimens. Jutland, Mols Bjerge 12 July 1979), and in the Material: Ø, Marker: Flagghytta. AK, Oslo: Bergen collection is a male from Sweden (Lule Alunsj øen. HES, Stange: Stange. HEN, Åmot: Lappmark, Messaure I -8 July 1974, K. Mtiller ?loc. BØ,Øvre Eiker: Burud, Ulleland. TEI, Kvi- l.g.). Of l0 males in coll. Ringdahl (Lund) posi- teseid: Skredi; Tokke: Dalen, Lårdal. AAY, Hi- tioned under the name <
15 land), Finland (Nylandia, Karelia australis), Dale and Sel, Hodnefiell; Karmøy: Kopervik. Sweden (according to specimen in British Mu- HOY, Bergen: Espegrend, Fana kirke, Fjøs- seum (Natural History), London, from Skåne, anger, Grønnestølen, Hop, Myravatn (Tveiter- Rovarekulan 2 Aug. 1980, A.C. Pont leg., ås), between Rådal and Stend, Storetveit, Åst- which I have dissected and identified as the pre- veit; Fitjar:? Fitjar; Samn€rri§€r: Ådtand; Askøy: sent species). Danish and Swedish collections Herdla. SFI, læikar§€r: Njøs. MRI, Rauma: [ær- are in need of revision. The specimen from Lår- heim. NTI, Steinkjer: Gulbergaunet. NSY, gård is the northernmost known (cf. Tiensuu Bodø: Bodø. EIS 2,7, l3 , 14, 17 ,22?, 28, 30 ,39, 1939 and map in Verves 1980d). 40,50,63,77, l0l,130. Date of earliest and latest capture: I I May, 29 47 . Sarcophaga schulzi Mueller Sept. Sarcophaga carnaria varietas schulzi Mueller Biology. Adults attracted to faeces and dead sna- 1922; Saicophaga carnaria: Patton 1934: 581' ils, apparently for feeding only (Kirchberg fig. i; Sariophåsa subvicina vulgaris Roden] tes4). dorf, 1937; Sarcophaga carnaria f. vulgaris: Distribution. Palaearctic Region east to Ural and Emden 1954: I16, fig. 39 G; Sarcophaga carna- Caucasus. Vladivostok. In the USSR north to ria: Kirchberg 1954: I 10, Abb. 1; Sarcophaga Leningrad region (Verves 1981b). British Isles. dolosa I-ehrer, 1967 Denmark, Finland (north to Tavastia australis) Material: Ø, Fredrikstad: Øra. AK, Oslo: Byg- and Sweden (Skåne, Dalarne). The record from døy, Oslo; Bærum: Østerås. HES, Elverum: Bodø appears to be the northernmost known. ,S. Grundset. HEN, Stor-Elvdal: Koppang. BØ, subvicina is the most common sarcophagid in Modum: ?loc. AAY, Hisøy: Hisøy. VAY, Kristi- the coastal areas of Western Norway. ansand: Hamre. RI, Suldal: Sandsa. HOY, Ber- §er: Stend. SFI, I-æikarg€r: Fylkesgarden (near Nybø). EIS 2. 6, 15, 20,28,30, 50, 55, 64. Genus Ravinia Robineau-Desvoidy, I 863 Date of earliest and latest capture: 2l May, I Aug. 49. Ravinia pernix (Harris) Biology. Larvae parasitic in earthworms (Kirc- Musca pernix Harris, 1780; Musca striataFabri- hberg 1954, Grunin 1964). Adults attracted to cius, 1794, preocc.; Sarcophaga haematodes faeces and meat (including dead snails) (Kirc- Meigen, 1826; R avinia striata: Rodendorf, hberg 1954, Gregor & Povolny l96l). 1937, 1970 Distribution. Palaearctic Region east to Altai Material: Ø, Halden: n€&r Halden. AK, Oslo: Tø- Mts. British Isles. In the USSR north to the Kola yen; Frogn; Sønderstøa Degerud; Bærum: Peninsula (Verves l98lb). Finland (north to Ta- Nordli NW of Østerås. HES, Elverum: Grund- vastia borealis and Savonia borealis), Sweden set. ON, Vågå: Vågåmo; Sel: Lårgård; Vang: (according.to specimen from Uppland, I-edsun- Nystuen. BØ, Ringerike: Hønefoss; Flesberg: det W of Oregrund 15 16 July 1980, A. C. Hvila. VAY, Mandal: Mandal. EIS 2,20,27 ,28, Pont leg., in British Museum (Natural History), 36, 52, 55, 7 l. London,which I have dissected and identified as Date of earliest and latest capture: 5 July, 9 Aug. the present species). Probably mixed with car- Biology. Breeds in faeces, also human (Kirc- naria and lehmanni rn Danish and Swedish col- hberg 1954, Mihtilyi 1965, Sfchevskaya 1970). letions. Occasional producer of intestinal myiasis in man (Rodendorf & Verves 197 8). Bred from snails, 48. Sarcophaga subvicina Rodendorf grasshoppers and a beetle (Seguy l94l , Roden- Sarcophaga vicina Villeneuve, 1899, preocc.; dorf & Verves 197 8). Adults attracted to faeces, Sarcophaga subvicina subvicina Rodendorf meat and fruits (Eberhardt 1955, Gregor & Po- tg37 . volny I 96 I ). Captured on the food-markets in Material: AK, Oslo: Bygdøy; Bærum: Høvik Budapest (Aradi & Mihålyi 1971). «. . . breeds in Nordby gård, Ostøya; Asker: Brønnøya. OS', human faeces and dead animals in nature and is Ringebu: Elstad. BØ: Drammen: Drammen. easily reared from such media in the laboratory» TEI, Tokke: LårdaL VAY, Mandal: Mandal. (Kano, Field & Shinonaga 1967). RY, Sandnes: Austrått, Bogafiell, Melsheia, Distribution. Palaearctic and Oriental Regions. Stølsvik; Stav&rS€r: Forus, Godalen, Gosen, British Isles. In the USSR north to the Arctic Krossb€rg, Lille Stokkavatn, Madlalia, Sunde, Circle near the Kola Peninsula (Verves 1981b). Tjensvoll, Ullandhaug; Klepp: Øksnevad; Denmark, Finland (north to Lapponia kemensis) Strand: I km E of Jøssang; Rennesøy: between and Sweden (north to Lule Lappmark). l6 Notesonotherspecies Siebke (1877: 90) who cites two localities, i.e. Below are given some annotåtions concerning Kongsvinger' and sarpsborg' The specimen from sarpsborg could not be found in the collec- species rec6rded from Norway by Ringdahl iisif,b"t;hi"htdonotacceptasbetongirgto tion' The specimen from Kongsvinger is a fem' ale see un- the Norwegian fauna on the basis of presenfly Miltogrammidium brevipilum' above der name (no' 7)' available evidence. However, they may ultima- that tely be captured in this country.
ZOOGEOGRAPHIC DISCUSSION Hilarella hilarella (Zetterstedt, I 844) Recorded from Norway by Ringdahl (1952, The highest number of sarcophagid species are 147, No. 325). The record is probably based on known from the southern or sout-eastern parts Siebke (1877:90, as Miltogramma hilarella) who of Norway (Fig. 25), which have the highest reports to have taken a female «Ad Christianiam summer temperatures (Abrahamsen et al 197 7 , in horto botanico. . . 14 August 1850». No cor- Gjessing & Ouren 1983). This is in accordance responding specimen could be traced in the Oslo with the distribution of sarcophagids in Finland museum, neither is there any evidence in the and Sweden (Tiensuu 1939, Ringdahl 1952). museum correspondence that Ringdahl ever re- The Norwegian species can be divided into ceived any specimen under this name for identi- four groups according to their presently known fication. As Siebke's collection was rearranged distribution in Norway. However, as the num- by W.M. Schøyen after its reception by the mu- ber of specimens captured in Norway is very seum (cf. Natvig 1943), the specimen actually low, the presented picture is likely to undergo before Siebke may be impossible to locate. Re- considerable changes in the future. (l) Widely cord not accepted. distributed species: 2. Metopia argyrocephala, 3. M. campestris, 17. Brachicoma devia, 23. Heli' Phrosinella nasuta (Meigen, 1824) cophagella crassimargo, 26. H. rosellei, 37 . He- Recorded from Norway by Ringdahl (1952, teronychia vagans, 38 . H. vicina, 4l . Parasar- 147, No. 323). Record probably based on holo- cophaga pleskei, 45. Sarcophaga carnaria and type of Tachina pilitarsis Zetterstedt, 1844 from 48. ,S. subvicina. (2) Species known only from Norway (cf. Siebke 1877:81) erroneously syno- Northern Norway: 25 . Helicophagella noverca nymised by Ringdahl with nasuta Meigen. See and 28. Thyrocnema kentejana. (3) Species above under 9. Phrosinella sannio. Record not known only from Western Norway: 20. Blae- accepted. soxipha rossica antl 22. Helicophagella agnata. (a) Species known only from Southern Norwå!: Angiometopa ruralis (Fall6n, I 8 I 7) All species not mentioned above under ( I ), (2) or Recorded from Norway by Ringdahl (1952, (3). The majority of species in this fourth group 147, No. 309). The recorded is probably based is known exclusively from the southern or on Siebke ( I 87 7 : 96, as Sarcophaga ruralis) who south-eastern parts of Norw&Y, but about one reports to have captured a female «ad Aaset in third are known also from Western Norway. par. Aamodt Østerdaliae 27 Juli 1870. . .)). In As far as their world distribution is concer- the Oslo collection a single specimen (No. 7749) ned, the Norwegian species appear to belong to stood under the name ,Sa rcophaga ruralis Fall. three distributional types: (l) Species occuring It was labelled (l) «Aamot» Q) «Siebke» and only in Fennoscandia and northern parts of the must be the one referred to by Siebke (1.c.). It is a USSR. Only one species, the miltogrammine female Billaea trianguhfera (Zetterstedt) (Tachi- Phrosinella sannio belongs here. Its closest relati- nidae). Record not accepted. ves, judged by the apomorphic fore tarsus in the males, live in North America. (2) Species with Brachicoma devia borealis Ringdahl, 1932 disjunct northern and southern distributional Recorded from Norway by Ringdahl (1944b, areas in Eurasia (boreo-alpine or boreo-montane 195». I have seen the specimen on which this distribution). Six species belong here, vrz. 25. record was based. See above under 17 . Brachi- Helicophagella noverca, 26. H. rosellei, 28. coma devia. Thyrsocnema kentejana, 36. Heteronychia os- tensackeni Q), 38. H . vicina and 4l . Parasarcop - Miltogramma oestraceum (Fallen, 18 20). haga pleskei. (3) Species with a more or less con- Recorded from Norway by Ringdahl (1952: tinuous southern distribution in Europe or Eu- 147, No. 3l». The record is probably based on rasia, reaching north to the lowlands of
t7 Fig. 26.The numbered squares of the EIS system for Fig. 25. Number of sarcophagid species recorded Norway. frdm each square of the EIS system for Norway.
palpalrs (Robineau- Southern Norway to a greater or lesser extent, Note on Helicobosca 1830) sometimes high into the mountains or farther Desvoidy, The 42 remaining north, often iynanthropic. Bezzi, 19-06 as this group. No doubt several I consider the genus Helicobosca species belong to (see abcve, Intro- groups are included within this belonging in ttri Catliphoridae ,bog.ographiial For conveni- concept uro.rth.ir, (cf. e.g. Rodendorf 1959), OuctiJ" , Paramacronychiinae). the known Norwegian material is but an analysis is not possible on the basis of the ence, dataon material at hand. presented below
18 Material. VE, ?loc. AK, Bærum: Steinshøgda, Aldrich, J.M. 1916. Sarcophaga and allies in North Ostøya. TEI, Tokke: Dalen. NSI, Saltdal: Rog- America.The Thomas Say Foundation of the En- nan. EIS 16, 19?, 28, 127. tomological Society of America, LaFayette, Indi- Date of earliest and latest capture: 19 June, 29 ana, 302 pp, l6 pls. July. Allen, H.W. 1926. North American species of two- winged flies belonging the tribe Miltogram- Note. The specimens from VE, labelled < Many thanks are due to L. Greve Jensen, Mu- (Dipt.) (Fortsetzung.). D. ent. Zeitschr. l9l 3 , seum of Zoology, Bergen, J.E. Raastad, Mu- l-16. seum of Zoology, Oslo, and A. Fjellberg, Bottcher, G. 1913b. Die månnlichen Begattungs- Tromsø Museum, for having made the collec- werkzeuge bei dem Genus Sarcophaga Meig. und ihre Bedeutung ftir die Abgrenzung tions of calypterate Diptera in their care avai- der Arten. (Dipt.) (Fortsetzung.). D. ent. Zeitschr. l9l 3 lable to me; to R. Danielsson, Lund, R. Contre- ll5-130. , ras-Lichtenberg, Vienna, B. Lindeberg, Helsinki, Bcittcher, G. l9l 3c. Die mannlichen Begattungs- and A.C. Pont, London, for loan of material; to werkzeuge bei dem Genus Sarcophaga Meig. und A.C. Pont, L. Greve Jensen and F. Midtgaard for ihre Bedeutung ftir die Abgrenzung der Arten. (Fortsetzung). help with literature; to G.E. Shewell, Ottawa, (Dipt.) D. ent. Zeitschr. l9l 3 , for having checked my identification of some 239 -254. Newfoundland Sarcophagidae and for useful in- Bottcher, G. 1913d. Die mdnnlichen Begattungs- formation; to J.-C. L6onide, Marseille, for ha- werkzeuge bei dem Genus Sarcophaga Meig. und ving checked ihre Bedeutung fiir die Abgrenzung der Arten. the identity of the Blaesoxipha ma- (Dipt.) (Fortsetzung). terial; D. ent. Zeitschr. l9l3 , to Torbj ørn Rognes for help with the pre- 351-378. paration of the maps; to S. Andersen, Copenha- Bcittcher, G. 1914. Sarcophaga lunigera nov. spec. B€tr, T. Jonassen, F. Midtgaard, T.R. Nielsen, (Dipt.). D. ent. Zeitschr. 1914, 434-436. A.-J. Nilsen and S. Svendsen for gift of material; Brander, T. 1964. Lounais-Håmeen kdrpdsistå, Bra- and to B. Sagvolden and J.H. Simonsen for let- chycera (Diptera). Lounais-Hcimeen Luonto 16, ting me see their collections. The grant from Sta- 5l -69. vanger Lærerhøgskole 1982- 1983 is gratefully Collart, A. 1954. Quelques Sarcophaginæ d'Italie acknowledged. (Diptera, Calliphoridae). Bull. Ann.,Soc. ent. Belg. 90,260-263. Collart, A. 1958. Notes sur la faune des Hautes-Fag- REFERENCES nes en Belgiques. Inst. r. Sci. nat. Belg. 34 (3 l), I -5. Abrahamsen, J., Dahl , E., Jacobsen, N.K., Kalliola, Dear, J. 1980. 91. Family Sarcophagidae. In: R., Påhlsson, L. & Wilborg, L. 1977. Naturgeog- Crosskey, R.W. (ed.), Catalogue of the Diptera of rafisk regionindelning av Norden. Nordiske Ut- the Afrotropical Region, British Museum (Natural redninger 1977 no. 34, Stockholm, ISBN History), [,ondon, pp. 801 - 8l 8. 9l-7052-339-8, 130 pp. Downes, W.L. 1955. 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K faune ription of a new Nearctic species (Diptera, Sarcop- Sarcophagidae (Diptera) Mongolskoi Narodnoi hagidae). Can. Ent. 103, ll79- 1 191. Respubliki. I. Sarcophaginae i Sarcotachininae Siebke, J.H.S. 1877 . Enumeratio Insectorum Norvegi- ( On the fauna of Sarcophagidae (Diptera) of the corum Fasciculum IV. Catalogum Dipterorum Mongolian People's Republic. I. Sarcophaginae Continentem. A.W. Brøgger, Christianra ( = Oslo), and Sarcotachininae ) . Nasekomye Mongolii 5, 255 pp. 7t6-730. Soot-Ryen, T. 1943. A review of the literature on Rodendorf, B.B. & Verves, Yu. G. 1977b. New spe- Norwegian Diptera until the year 1940. Tromsø cies of the genus Blaesoxipha Low (Diptera, Sar- Mus. Aarsh. 65 (1942) 6), l-46. cophagidae) from Transcaucasia and the Kopet- Souza Lopes, H.de 1955. Considera coes sobre os 96- dag. Ent. B"u. Wash. 56, 134- 136. (Russian ori- neros de Sarcophagidae (Diptera) propostos por ginal rn Ent. obozr. not seen.) Robineau-Desvoidy em 1830 e 1863 . Arquivos do Rodendorf, B.B. & Verves, Yu. G. 1978. Sarcophagi- Museu Nacional 42, 265-272. nae (Diptera, Sarcophagidae) from Mongolia. Souza Lopes, H. de 1969. 103. Family Sarcophagi- Ann. hist.-nat. Mus. nat. hung. 70, 241 -258. dae. - In: Papavero, N. (ed.), A. catalogue of the Rodendorf, B.B. & Verves, Yu.G. 1979a. K faune Diptera of the Americas South of the United States. Sarcophagidae (Diptera) Mongolskoi Narodnoi Departamento de Zoologia, Secretaria da Agricul- Respubliki. II. Novye dannye o Sarcophaginae tura, Sao Paulo, pp. I - 88. ( On the fauna of Sarcophagidae (Diptera) of the Souza Lopes, H. de, Kano, R., Shinonaga, S. & Kura- Mongolian People's Republic. II. New data on hashi, H. I 977 . Family Sarcophagidae. - In: Del- Sarcophaginae ) . Nasekomye Mongolii 6, finado, M.D. & Hardy, D.E. (eds.), .4 Catalog of 475-497. the Diptera of the Oriental Region. Vol. IIL Subor- Rodendorf, B.B. & Verves, Yu.G. 1979b. NoWe der Cyclorrhapha (excluding Division Aschiza). dvukrylye podsem. Sarcophaginae (Diptera, Sar- The University Press of Hawaii, Honolulu, pp. cophagidae) tz Palearktiki ( New Palaearctic Sar- 557 -583. cophaginae (Diptera, Sarcophagidae) ) . Ent. Stackelbgrg, A.A. 1962. Materialy po faune dvukr:i- obozr. 58, 190- 199. (English translation in Ent. likh læningradskoi oblasti. VI. Diptera, Calyq,t- Rev, Wash. 58, ll4-121. rata, chast' I < List of Diptera of the Leningrad Rodendorf, B.B. & Verves, Yu.G. 1980. K faune Sar- Region, VI. Diptera Calyptrata, Part I > Trudy cophagidae (Diptera) Mongolskoi Narodnoi Re- 24 Zoologicheskogo Instituta Akademii Nauk,SS,SR Verves, Yu.G. 197 3. Novi vidi sarkofaeid (Diptera, 31,319-399. Sarcophagidae) z Ukraini. Dopovidi Akademii stein, P. 1924. Die verbreitetsten Tachiniden Mittele- Nauk Ukrains'koi Å,SR , Seriya B, Geologiya, Geo- uropas nach ihre Gattungen und Arten. Arch. Na- Jizika, Khimiya ta Biologiya, Kiiv 1973 , (A) (6), turg. 90 I -27 I . 946 -948. Stop-Bowitz, C. 1969. A contribution to our know- Verves, Yu.G. 1974. K faune sarkofagid (Diptera, ledge of the systematics and zoogeography of Sarcophagidae) SSSR. I. Triba Sarcophagini (Sar- Norwegian earthworms (Annelida Oligochaeta: cophaginae) (po materialam koltektsii Instituta Lumbricidae). Nytt. Mag. Zool. 17, 169-280. Zoologii A N USSR). Vestnik zoologii 1974, Storm, V. I 891 . Diptera brachycera, quæ observavit 30-37. determinavitque. K. norske Vidensk. Selsk. Skr. Verves, Yu.G. 197 5a. Norfe vidy mukh roda Hete- Iggt,2g3-313. ronychia B.B. (Diptera, Sarcophagidae) < New Storm, V. 1896. Dipterologiske undersøgelser. K. species of the genus Heteronychia B.B. (Diptera, norske Vidensk. Selsk. Skr. 1895,225-241. Sarcophagidae)). Ent. obozr. 54, 665-667. Storm, V. 1907. Supplerende iagttagelser over In- (Enelish translation in Ent . Rev. Wqsh. 5 4 , secta Diptera ved Trondhjem. K. norske Vidensk. 130-131.) Selsk. Skr. 1907 (5), l-ll. Verves, Yu.G. 197 5b. K faune sarkofagid (Diptera, Strand, E. 1900. Bidrag til Hatlingdals og Lyngørs in- Sarcophagidae) SSSR. II. Podsemeistva Sarcopha- sektfauna. Nyt Mag. Naturv. 37, 46-72. ginae (Triba Agriini), Macronichiinae i Miltog- Strøm, H. 1781 . Udkast til en Beskrivelse over Har- rammatinae (po materialam kollektsii Instituta danger i Bergens Stift i Norge samlet og sammens- Zoologli A N USSR). Vestnik zoologii 1975, krevet af Sal. Hr. Marcus Schnabels efterladte Pa- 73-77. pirer ved H.S. Kiøbenhavn. (Not seen. Cited Verves, Yu.G. 1976. K izucheniyu sarkofagid (Dip- from Soot-Ryen 1943 > tera, Sarcophagidae) parazitov nazemnikh Strøm, H. 17 84. Physisk-Oeconomisk Beskrivelse bryukhonogik mollyuskov. Vestnik zoologii I 976, over Eger-Præstegjeld i Aggerhus-Stift i Norge; 28. tilligemed et geographisk Kort over samme, al Verves, Yu.G. 1977a. Novii vid rody Heteronychia Hans Strøm, Prof. Theol. og Sognepræst til Eger. B.B. zTyan'-Shanya (Sarcophagidae) ( New spe- Kiøbenhavn. ( Not seen. Cited from Soot-Ryen cies of the genus Heteronychia B.B. from the Tien t943 > Shan (Sarcophagidae) ) . Dopovidi Akademii Nauk Strukan, D. 1967. Remarques sur deux Sarcophagi- Ukrains'koi R,SR , Seriya B, Geologichni, Khi- nae peu connus de Yougoslavie (Diptera, Sarcop- michni ta Biologichni Nauki, Kiiv 1977, hagidæ). Cahiers des Naturalistes, Bult. N.P., n.s. 266-267. 23, 45-47. Verves, Yu.G. 1977b. Novi vidi sarkofagid (Diptatz; Sychevskaya, V.I. 1966. O sinantropnykh mukhakh Sarcophagidae) z Kavkazy i Seredn'oi Azii ( New Pamira. Zool. Zh. 45,390- 399. species of sarcophagid flies (Diptera: Sarcophagi- Sychevskaya, V.I. 1970. Zonal'noe raspredelenie dae) from the Caucasus and Middle Asia ) . Dopo- koprofil'nykh i skhizofil'nfkh mukh (Diptera) v vidi Akademii Nauk Ukrains'koi RSR, Seriya B, srednei Asii (, zonal distribution of coprophilus Geologichni, Khimichni ta Biologichni Nauki, Kiiv and schizophilous flies (Diptera) in middle I 977, 35 I - 354. Asia ) . Ent. obozr.49,8l9-S3t. (Engtish trans- Verves, Yu.G. 1978a. Vertikal'no-zonal'nii rozpodil lation in Ent. Rev. Wash. 49, 498-505). ' sarkofagid Krimy (Diptera, Sarcophagidae). Vis- Thompson, W.-R. 1921. Contributions å la connais- nik Kiivs'kogo Universitety, Biologiya 20, 93 -96. sance des formes larvaires des Sarcophagides Verves, Yu.G. 1978b. Noqfe i maloizvestnye Sarcop- (Dipt.). II. Ptychoneura rufitarsrs Meig . Bull. Soc. hagidae (Diptera) Sibiri i Dal'nego Vostoka ent. Fr. 192 I , 219 -222. ( New and little-known species of Sarcophagidae Tiensuu, L. 1939. Die Sarcophagiden (Dipt.) Finn- from Siberia and Far East > . Taksonomiya i eko- lands. Ann. ent. fenn. 5, 255-266. logiya chlenistonogikh Sibiri (Novye i maloizvest- Tiensuu, L. 1964. Huomion arvoisia kårprisloytdjå nye vidy fauny Sibiri), Novosibirsk I 9 78 , Lounais-Hameestå ( Bemerkenswerte Fliegen- 184-187. funde aus Stidwest-Håme ) . Lounais-Hcimeen Verves, Yu.G. 1978c. Sarkofagidi (Diptera, Sarcop- Luonto 15, 76-77. hagidae) z baseinu Indigirki ( Sarcophagid flies Venturi, F. 1947. Notulae dipterologicae. I. Miltog- (Diptera, Sarcophagidae) from the Indigirk a ba- rammini e Metopiini (Dipt. Sarcophagidae) sin ) . Dopovidi Akademii Nauk Ukrains'koi RSR, dell'Italia centrale. Redia 32, I l9- 139. Seriya B, Geologichni, Khimichni ta Biologichni Venturi, F. 1952. Notulae dipterologicae. V. Revi- Nauki, Kiiv 1978, I 80 - 182. sione sistematica del genere Metopia Meigen (Dip- Verves, Yu.G. 1978d. K faune Sarcophaginae (Sar- tera, Sarcophagidae) in Italia. Boll. entom. Bo- cophag idae, Diptera) Kavkaz a. Nauchnye Doklady logna. 19, 147 Vysshei Shkoly, Biologicheskie I 9 78 -170. Nauki , Venturi, F. 1960. Sistematica e geonemia dei Sarcofa- 36-45. gidi (excl. Sarcophaga Meig. s.l.) itatiani (Diptera). Verves, Yu.G. 1979a. Palearkticheskie vidyi podroda Frustula entomologica 2 (7), l-124. Spatulapica Fan roda Heteronychia (Diptera, Sar- 25 cophagidae) ( Palearctic species of the subgenus miya Nauk SSSR, Zoologicheskii Institut, læning- Spatulapica Fan of the genus Heteronychia (Dip- rad. tera, Sarcophagidae) ) . Zool. Zh. 58, 860-870. Verves, Yu.G. 1982a. 64h. Sarcophaginae. [n: Verves, Yu.G. 1979b. Description of Paramacrony- Lindner, E. (ed.), Die Fliegen der palaearktischen chia hackmarci sp.n. (Diptera, Sarcophagidae, Pa- Region, Lieferung 327, pp. 235-296. ramacronychiinae). Ann. ent. fenn. 45, 3l-32. Verves, Yu.G. 1982b. I