研究報告
(ᖪ 22: 193-214 (2002ٿέ៉ Formosan Entomol. 22: 193-214 (2002)
Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan
Hsien-Tzung Shih! Department of Applied Zoology, Taiwan Agricultural Research Institute, 189 Chung-Cheng Rd., Wufeng, Taichung 413, Taiwan, R.O.C. Jeng-Tze Yang*! Department of Entomology, National Chung Hsing University, 250 Kuo Kung Rd., Taichung 402, Taiwan, R.O.C.
ABSTRACT
The family Aphrophoridae consists of over 900 species in the world. Seventy-three species have been recorded from Taiwan, with 50 species (68.5%) considered endemic. The checklist revises the data of geographical distribution, and provides information on type deposition and type localities.
Key words: Homoptera, Cercopoidea, Aphrophoridae, Taiwan, checklist.
Introduction ptyelus, Toroptyelus, and Jembroides, were synonymized with related genera or The family Aphrophoridae was reinstated to the original ones by Liang erected by Amyot and Serville in 1843 (1998, 1999). based on the type genus Aphrophora After a literature review, the Germar 1821 (Amyot and Serville, 1843). checklist contains 1 subfamily, 6 tribes, Aphrophoridae contains 2 subfamilies, 8 17 genera, and 73 Taiwanese aphrophorid tribes, and over 900 species in the world species. Fifty of the species in the present at present, of which 62.2% of the species checklist are endemic to Taiwan. In this are from the Palearctic and Oriental paper, we have tried to organize the Regions. information to make it much more Seventy-eight species of Aphropho- complete and clear to facilitate future ridae were recorded as occurring in studies of this group. Taiwan before 1943 (Metcalf, 1962), among which, 40 species and 9 Materials and Methods monotypical genera were published by Matsumura (1903, 1907, 1934, 1938, 1940, The type depositions are listed below. 1942a, b), i.e., Awaphora, Macrofukia, The abbreviation of each museum Mimoptyelus, Toroptyelus, Jembroides, collection was checked by the authors Megagukia, Jembrophora, Nokophora and from the Website of the Bishop Museum, Kotophora. Matsumura’s monotypical http://www.bishopmuseum.org/research/na genera, Awaphora, Macrofukia, Mimo- tsci/ento/codens-inst.html.
*Correspondence address e-mail:[email protected]! Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan 193 BMNH – The Natural History Museum, was cited by authors from Amyot and London, UK [formerly the British Serville (1843), Distant (1908), Matsu- Museum (Natural History)]. mura (1940, 1942a), Metcalf (1962), DEI – Deutschen Entomologischen Metcalf and Horton (1934), and Stål Institutes, Eberswalde, Germany. (1863, 1866). The checklist of all EIHU – Laboratory of Systematic scientific names is arranged in Entomology, Faculty of Agriculture, alphabetical sequence by specific epithet. Hokkaido University, Sapporo, New distributional data are recorded Japan. from the TARI (Taiwan Agricultural ICRI – Research Institute of Entomology, Research Institute) collection and are Zhongshan (Sun Yat-Sen) University, indicated in boldface. Guangzhou, Guangdong, China. NCSU – National Carolina State Family APHROPHORIDAE ڰUniversity Insect Collection, Raleigh, Amyot and Serville, 1843 ы NC, USA. ᖩࡊ SMTD – Staatliches Museum für Tierkunde, Dresden, Germany. Subfamily APHROPHORINAE Licent, ᖩֲࡊڰUSNM – National Museum of Natural 1912 ы History, Washington, D.C, USA Aphrophorinae Licent, 1912, Thèses [formerly the United States présentées à la Faculté des Sciences National Museum]. de Nancy pour obtenir le grade de The distribution records, synonymes docteur es-sciences naturelles 1912: of genus and species are extracted mainly 134. from the original publications of Chou and Liang (1987), Chou et al. (1992), Tribe APHROPHORINI Lallemand, ᖩڰDistant (1908), Jacobi (1904, 1921), Kato 1924 ы (1933a, b), Komatsu (1997), Kwon and Aphrophorini Lallemand, 1924, Paris Mus. Lee (1979), Lee et al. (1976), Liang (1992, d’Hist. Nat. Bull. 1924: 296. (Metcalf, 1997, 1999, 2000, 2001), Matsumura 1962) (1903, 1907, 1934, 1938, 1940, 1942a, b), ڰMelichar (1903), Metcalf (1962), Metcalf Genus Aphrophora Germar, 1821 ы and Horton (1934), Nast (1972), ᖩᛳ Ossiannilsson (1981), Schmidt (1920), Aphrophora Germar, 1821, Mag. Entomol. Schumacher (1915a, b), Shih et al. (2000), 3: 48. Shih and Yang (2002), and Walker (1851, Awaphora Matsumura, 1940, J. Fac. 1858). Metcalf (1962), probably based on Agric. Hokkaido Imp. Univ. 45(2): 59 the articles of Kato (1933a, b), listed (Type species: Awaphora kikuchii Japan and Taiwan as the distributional Matsumura, 1940) [synonymised by area of 18 Taiwanese Aphrophoridae Liang, 1998: 241]. species. However, Kato (1933a, b) only Macrofukia Matsumura, 1940, J. Fac. listed Taiwan as the distributional area Agric. Hokkaido Imp. Univ. 45(2): 58 of Taiwanese Aphrophoridae. We, (Type species: Macrofukia habonis therefore, emended these 18 species as Matsumura, 1940) [synonymised by endemic to Taiwan in this checklist, and Liang, 1998: 241]. mark each species name with an Miphora Matsumura, 1940, J. Fac. Agric. asterisk Ŏ in the text. Hokkaido Imp. Univ. 45(2): 61 (Type The taxonomic system of this study species: Miphora arisanella Matsu- is based mainly on that of Metcalf (1962). mura, 1940) [synonymised by Liang, Information of type species for each genus 1998: 241].
ௐˬഇס˟˩˟ᖪௐٿέ៉ 194 Nagafukia Matsumura, 1940, J. Fac. 1904) [synonymised by Komatsu, Agric. Hokkaido Imp. Univ. 45(2): 57 1997: 82]. (Type species: Nagafukia mushana Nagophora Matsumura, 1942a, Ins. Matsumura, 1940) [synonymised by Matsumurana 16: 62 (Type species: Liang, 1998: 241]. Nagophora hirayamai Matsumura, Todophora Matsumura, 1940, J. Fac. 1942) [synonymised by Liang, 1998: Agric. Hokkaido Imp. Univ. 45(2): 56 241]. (Type species: Aphrophora abieti Obiphora Matsumura, 1942a, Ins. Matsumura, 1904) [synonymised by Matsumurana 16: 62 (Type species: Komatsu, 1997: 82]. Aphrophora intermedia Uhler, 1896) Yezophora Matsumura, 1940, J. Fac. [synonymised by Nast, 1972: 168; Agric. Hokkaido Imp. Univ. 45(2): 61 synonymy confirmed by Liang, 1998: (Type species: Yezophora kurilensis 242]. Matsumura, 1940) [synonymised by Ogaphora Matsumura, 1942a, Ins. Kwon and Lee, 1979: 15; synonymy Matsumurana 16: 65 (Type species: confirmed by Liang, 1998: 241]. Aphrophora bizonalis Matsumura, Atuphora Matsumura, 1942a, Ins. 1907) [synonymised by Liang, 1998: Matsumurana 16: 48 (Type species: 242]. Aphrophora stictica Matsumura, 1903) Omalophora Matsumura, 1942a, Ins. [synonymised by Nast, 1972: 168; Matsumurana 16: 66 (Type species: synonymy confirmed by Liang, 1998: Cicada spumaria salicis De Geer, 241]. 1773) [synonymised by Nast, 1972: Boniphora Matsumura, 1942a, Ins. 168; synonymy confirmed by Liang, Matsumurana 16: 48 (Type species: 1998: 242]. Aphrophora bicolor Matsumura, 1907) Petaphora Matsumura, 1942a, Ins. [synonymised by Liang, 1998: 241]. Matsumurana 16: 69 (Type species: Dophora Matsumura, 1942a, Ins. Aphrophora maritime Matsumura, Matsumurana 16: 49 (Type species: 1903) [synonymised by Kwon and Aphrophora vitis Matsumura, 1904) Lee, 1979: 15; synonymy confirmed [synonymised by Komatsu, 1997: 82]. by Liang, 1998: 242]. Europhora Matsumura, 1942a, Ins. Sabphora Matsumura, 1942a, Ins. Matsumurana 16: 51 (Type species: Matsumurana 16: 72 (Type species: Aphrophora alpina Melichar, 1900) Aphrophora takagii Matsumura, 1940) [synonymised by Nast, 1972: 168; [synonymised by Liang, 1998: 242]. synonymy confirmed by Liang, 1998: Sagophora Matsumura, 1942a, Ins. 241]. Matsumurana 16: 74 (Type species: Formophora Matsumura, 1942a, Ins. Aphrophora tsuruana Matsumura, Matsumurana 16: 52 (Type species: 1907) [synonymised by Liang, 1998: Aphrophora arisana Matsumura, 242]. 1940) [synonymised by Liang, 1998: Seiphora Matsumura, 1942a, Ins. Matsu- 241]. murana 16: 76 (Type species: Hosophora Matsumura, 1942a, Ins. Seiphora seitonis Matsumura, 1942) Matsumurana 16: 53 (Type species: [synonymised by Liang, 1998: 242]. Hosophora nomurella Matsumura, Takaphora Matsumura, 1942a, Ins. 1942) [synonymised by Liang, 1998: Matsumurana 16: 84 (Type species: 241-242]. Aphrophora brevis Matsumura, 1904) Jophora Matsumura, 1942a, Ins. [synonymised by Komastsu, 1997a: Matsumurana 16: 58 (Type species: 82]. Aphrophora compacta Matsumura, Tamaphora Matsumura, 1942a, Ins.
Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan 195 Matsumurana 16: 85 (Type species: murana 16: 52 [reinstated Tamaphora domonsis Matsumura, combination by Liang, 1998: 245]. 1942) [synonymised by Liang, 1998: Type deposition: EIHU (Komatsu and 242]. Hayashi, 1998). Tilophora Matsumura, 1942a, Ins. Matsu- Type locality: TAIWAN: Mt. Ari [Ali- murana 16: 88 (Type species: shan], Chiayi Co. (Matsumura, 1940). Aphrophora flavipes Uhler, 1896) Distribution: Taiwan. [synonymised by Nast, 1972: 168; synonymy confirmed by Liang, 1998: 2. Aphrophora arisanella (Matsumura, ᖩڰπᗝы (1940 .[242 Tobiphora Matsumura, 1942a, Ins. Miphora arisanella Matsumura, 1940, J. Matsumurana 16: 88 (Type species: Fac. Agric. Hokkaido Imp. Univ. Aphrophora rugosa Matsumura, 1903) 45(2): 61. [synonymised by Kwon and Lee, 1979: Formophora karenkonis Matsumura, 16]. 1942a, Ins. Matsumurana 16: 52 Tonkaephora Matsumura, 1942a, Ins. [synonymised by Liang, 1998: 245]. Matsumurana 16: 89 (Type species: Miphora shirakii Matsumura, 1942a, Ins. Tonkaephora oshodenella Matsumura, Matsumurana 16: 61 [synonymised 1942) [synonymised by Liang, 1998: by Liang, 1998: 245]. 243]. Aphrophora arisanella (Matsumura, 1940) Trigophora Matsumura, 1942a, Ins. [transferred by Liang, 1998: 245] Matsumurana 16: 91 (Type species: Type deposition: EIHU (Komatsu and Cercopis alni Fallén, 1805) Hayashi, 1998). [synonymised by China, 1951: 279]. Type locality: TAIWAN: Mt. Ari [Ali- Tukaphora Matsumura, 1942a, Ins. shan], Chiayi Co. (Matsumura, 1940). Matsumurana 16: 94 (Type species: Distribution: Taiwan. Tukaphora sinalca Matsumura, 1942) [synonymised by Liang, 1998: 243]. 3. Aphrophora flavicosta Kato, 1933 ŏ ᖩڰYamaphora Matsumura, 1942a, Ins. ਂเቡы Matsumurana 16: 95 (Type species: Aphrophora flavicosta Kato, 1933b, Yamaphora yatugadakeana Matsu- Entomol. World 1: 230. mura, 1942) [synonymised by Type deposition: Unknown. Komastsu, 1997a: 82]. Type locality: TAIWAN: Karenko Yaphora Matsumura, 1942a, Ins. Matsu- [Hualien], Hualien Co. (Kato, 1933b). murana 16: 96 (Type species: Distribution: Taiwan. Aphrophora yatsugadakensis Matsu- mura, 1934) [synonymised by 4. Aphrophora habonis (Matsumura, ᖩڰыਂܜKomastsu, 1997a: 82]. 1940) ൺᗝ Macrofukia habonis Matsumura, 1940, J. Type species: Aphrophora alni (Fallén, Fac. Agric. Hokkaido Imp. Univ. 1805) 45(2): 59. Aphrophora habonis (Matsumura, 1940) 1. Aphrophora arisana Matsumura, [transferred by Liang, 1998: 246]. ᖩ Type deposition: EIHU (Komatsu andڰᗕโᕇы 1940 Aphrophora arisana Matsumura, 1940, J. Hayashi, 1998). Fac. Agric. Hokkaido Imp. Univ. Type locality: TAIWAN: Musha [Wushe], 45(2): 63. Nantou Co. (Matsumura, 1940; Formophora arisana (Matsumura, 1940); Komatsu and Hayashi, 1998). Matsumura, 1942a, Ins. Matsu- Distribution: Taiwan.
ௐˬഇס˟˩˟ᖪௐٿέ៉ 196 5. Aphrophora horishana Matsumura, Obiphora intermedia (Uhler, 1896); .ᖩ Matsumura, 1942a, Insڰᗝૄโы 1940 Aphrophora horishana Matsumura, 1940, Matsumurana 16: 63 [reinstated J. Fac. Agric. Hokkaido Imp. Univ. combination by Liang, 1998: 246]. 45(2): 63. Type deposition: USNM (Uhler, 1896). Type deposition: EIHU (Komatsu and Type locality: JAPAN: Mt. Ibuki (Liang, Hayashi, 1998; Komatsu and 2000). Hayashi, 1998). Distribution: China, Korea, Japan Type locality: TAIWAN: Horisha [Puli], (Hokkaido, Kyushu, Honshu, Nantou Co. (Matsumura, 1940). Shikoku), Taiwan. Distribution: Taiwan. 8. Aphrophora karenkoensis Kato, 1933ŏ ᖩڰᖩыڰAphrophora horizontalis Kato, 1933 ˬ֎เˬ֎เы .6 ,ᖩ Aphrophora karenkoensis Kato, 1933bڰᖩыڰᗕโᗕโы Aphrophora horizontalis Kato, 1933b, Entomol. World 1: 229. Entomol. World 1: 228. Type deposition: Unknown. Obiphora mushana Matsumura, 1942a, Type locality: TAIWAN: Karenko Ins. Matsumurana 16: 63 [Hualien], Hualien Co. (Kato, 1933b). [synonymised by Liang, 1998: 246]. Distribution: Taiwan. Obiphora rectella Matsumura, 1942a, Ins. Matsumurana 16: 64 [synonymised 9. Aphrophora kikuchii (Matsumura, ᖩڰby Liang, 1998: 246]. 1940) ᗕϨы Type deposition: Unknown. Awaphora kikuchii Matsumura, 1940, J. Type locality: TAIWAN: Karenko Fac. Agric. Hokkaido Imp. Univ. [Hualien], Hualien Co. (Kato, 1933b). 45(2): 60. Distribution: China (Henan (new Aphrophora kikuchii (Matsumura, 1940) record), Anhui, Zhejiang, Hubei, [transferred by Liang, 1998: 247]. Jiangxi, Fujian, Guangdong, Guangxi, Type deposition: EIHU (Komatsu and Sichuan, Guizhou, Yunnan), Russia Hayashi, 1998). (Maritime Territory, Kurile Island), Type locality: TAIWAN: Kyuhabon, Korea Peninsula (Sakhalin, Quelpart Nantou Co. (Matsumura, 1940). Islands), Japan (Hokkaido, Honshu, Distribution: China, Taiwan. Kyushu, Shikoku), Taiwan. Remarks: We have examined one male 10. Aphrophora memorabilis Walker, ᖩڰᖩыڰspecimen of this species from the 1858 Ϩโ৳Ϩโ৳ы Insect Collection of Anhui Aphrophora memorabilis Walker, 1858, Agricultural Univ., Anhui Prov., Suppl. List Homopterous Insetcs in China, which was collected by Dr. BMNH: 186. Ping Cai from Gikon Mt., Hsinyang Aphrophora notabilis Walker, 1858, Suppl. Co., Henan Prov., China on July 9, List Homopterous Insetcs in BMNH: 1997. 186 [synonymised by Metcalf, 1962: 466]. 7. Aphrophora intermedia Uhler, 1896 Cercopis memorabilis (Walker, 1858); ᖩ Lallemand, 1912, Gen. Insect 143: 61ڰᖩыڰϨϨы Aphrophora intermedia Uhler, 1896, US [reinstated combination by Metcalf, Natl. Mus. Proc. 19: 288. 1962: 466]. Cercopis intermedia (Uhler, 1896); Type deposition: BMNH (Walker, 1858). Lallemand, 1912, Gen. Insect 143: 60 Type locality: CHINA (Walker, 1858). (Metcalf, 1962). Distribution: China (Anhui, Hubei,
Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan 197 Hunan, Shaanxi, Jiangxi, Jiangsu, Aphrophora taiwana Kato, 1933b, Sichuan, Yunnan, Guizhou, Zhejiang, Entomol. World 1: 231. Guangdong, Guangxi, Fujian), Miphora taiwana (Kato, 1933b); Matsu- Taiwan. mura, 1942a, Ins. Matsumurana 16: 61 [reinstated combination by Liang, 11. Aphrophora mushana (Matsumura, 1998: 251]. ᖩ Type deposition: Unknown (Komatsuڰቡਂы (1940 Nagafukia mushana Matsumura, 1940, J. and Hayashi, 1998). Fac. Agric. Hokkaido Imp. Univ. Type locality: TAIWAN: Karenko [Hua- 45(2): 58. lien], Hualien Co. (Kato, 1933b). Aphrophora mushana (Matsumura, 1940) Distribution: Taiwan. [transferred by Liang, 1998: 248]. Type deposition: EIHU (Komatsu and 15. Aphrophora tsuruana Matsumura, ᖩڰHayashi, 1998). 1907 ነਂโᕇы Type locality: TAIWAN: Musha [Wushe], Aphrophora tsuruana Matsumura, 1907, Nantou Co. (Matsumura, 1940). Annot. Zool. Jpn. 6: 110. Distribution: Taiwan. Cercopis tsuruana (Matsumura, 1907); Lallemand, 1912, Gen. Insect 143: 63 12. Aphrophora nagasawae Matsumura, [reinstated combination by Liang, .[ᖩ 1998: 251-252ڰᖩыڰᑢᑢϨϨϨы 1907 Aphrophora nagasawae Matsumura, 1907, Aphrophora kanoniella Matsumura, 1940, Annot. Zool. Jpn. 6: 108. J. Fac. Agric. Hokkaido Imp. Univ. Cercopis nagasawae (Matsumura, 1907); 45(2): 536 [synonymised by Liang, Lallemand, 1912, Gen. Insect 143: 61 1998: 251]. [reinstated combination by Metcalf, Formophora kohozana Matsumura, 1942a, 1962: 467]. Ins. Matsumurana 16: 52 [synony- Type deposition: EIHU (Komatsu and mised by Liang, 1998: 251]. Hayashi, 1998). Formophora kononiella (Matsumura, 1940) Type locality: TAIWAN: Taihoku [Taipei] [transferred by Matsumura, 1940: 52, (Matsumura, 1907). 54]. Distribution: Taiwan. Sagophora tsuruana (Matsumura, 1907) [transferred by Matsumura, 1942b: 13. Aphrophora nomurella (Matsumura, 75]. ᖩ Type deposition: EIHU (Komatsu andڰπᗝবਂы (1942 Hosophora nomurella Matsumura, 1942a, Hayashi, 1998). Ins. Matsumurana 16: 54. Type locality: TAIWAN: Taikokan Hosophora rarasana Matsumura, 1942a, (Matsumura, 1907; Komatsu and Ins. Matsumurana 16: 54 [synony- Hayashi, 1998). mised by Liang, 1998: 248]. Distribution: China (Hubei, Jiangxi, Aphrophora nomurella (Matsumura, 1942) Fujian, Guangdong, Guangxi, [transferred by Liang, 1998: 248]. Sichuan, Guizhou), Taiwan. Type deposition: EIHU (Komatsu and Hayashi, 1998). Genus Aphropsis Metcalf and ᖩᛳڰType locality: TAIWAN: Arisan [Ali- Horton, 1934 ਖ shan], Chiayi Co. (Matsumura, 1942a). Distribution: Taiwan. Aphropsis Metcalf and Horton, 1934, Lingnan Sci. J. 13: 409. 14. Aphrophora taiwana Kato, 1933ŏ ᄂ Type species: Aphropsis gigantean ᖩ Metcalf and Horton, 1934ڰᖩыڰᑢϨ៉ᑢϨы៉
ௐˬഇס˟˩˟ᖪௐٿέ៉ 198 Distribution: Taiwan. 16. Aphropsis maxima Jacobi, 1921 ̂ ,ᖩ 20. Jembrana daitoensis Matsumuraڰਖ ᖩڰਖڡᖩڰਖڡAphropsis maxima Jacobi, 1921, Arch. f. 1940 ነነ Natuegesch. 87: 28. Jembrana daitoensis Matsumura, 1940, J. Type deposition: SMTD (Emmrich, 1974). Fac. Agric. Hokkaido Imp. Univ. Type locality: TAIWAN: Hoozan [Feng- 45(2): 50. shan], Kaohsiung Co. (Jacobi, Type deposition: EIHU (Komatsu and 1921). Hayashi, 1998). Distribution: Taiwan. Type locality: TAIWAN: Mt. Toran [Tatung], Taichung Co. (Matsumura, Genus Jembrana Distant, 1908 1940). .ᖩᛳ Distribution: Taiwanڰਖڡ
Jembrana Distant, 1908, Rhynchota IV- 21. Jembrana flavomaculata Kato, ᖩڰਖڡᖩڰਖڡHomoptera. The fauna of British 1933ŏ เเ India including Ceylon and Burma: Jembrana flavomaculata Kato, 1933b, 104. Entomol. World 1: 223. Type species: Jembrana buprestioides Type deposition: Unknown. Distant, 1908 Type locality: TAIWAN: Mt. Niitaka, Nantou Co. (Kato, 1933b). 17. Jembrana arisanensis Chou and Distribution: Taiwan. ᖩڰਖڡఀܜ Liang, 1987 Jembrana arisanensis Chou and Liang, 22. Jembrana formosana Kato, 1933ŏ ᑢ ᖩڰਖڡᖩڰਖڡEntomotaxonomia 9(4): 283- ነነ :1987 288. Jembrana formosana Kato, 1933b, Type deposition: ICRI (Hua, 1989). Entomol. World 1: 226. Type locality: TAIWAN: Alishan, Type deposition: Unknown. Chiayi Co. (Chou and Liang, 1987). Type locality: TAIWAN: Suiha (Kato, Distribution: Taiwan. 1933b). Distribution: Taiwan. 18. Jembrana albinotata Kato, 1933ŏ Ϩ ,ᖩ 23. Jembrana kankonis Matsumuraڰਖڡᖩڰਖڡ৳৳ ᖩڰਖڡᖩڰਖڡJembrana albinotata Kato, 1933b, 1942 ఀბπఀბπ Entomol. World 1: 224. Jembrana kankonis Matsumura, 1942a, Type deposition: Unknown. Ins. Matsumurana 16: 55. Type locality: TAIWAN: Musha [Wushe], Type deposition: EIHU (Komatsu and Nantou Co. (Kato, 1933b). Hayashi, 1998). Distribution: Taiwan. Type locality: TAIWAN: Kwanko (Matsumura, 1942a). 19. Jembrana centralis Matsumura, Distribution: Taiwan. ᖩڰਖڡᖩڰਖڡព̚ਖព̚ਖ 1940 Jembrana centralis Matsumura, 1940, J. 24. Jembrana kanoniella Matsumura, ᖩڰਖڡᖩڰਖڡFac. Agric. Hokkaido Imp. Univ. 1940 ᐝ̚โᐝ̚โ 45(2): 51. Jembrana kanoniella Matsumura, 1940, J. Type deposition: EIHU (Komatsu and Fac. Agric. Hokkaido Imp. Univ. Hayashi, 1998). 45(2): 51. Type locality: TAIWAN: Mt. Ari [Ali- Type deposition: EIHU (Komatsu and shan], Chiayi Co. (Matsumura, 1940). Hayashi, 1998).
Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan 199 Type locality: TAIWAN: Mt. Ari [Ali- Matsumura, 1942 shan], Chiayi Co. (Matsumura, 1940). Distribution: Taiwan. 27. Jembrophora sawadai Matsumura, ᖩڰఀ̂ᐙ̚ਖۡ 1942 25. Jembrana punctipennis Kato, 1933ŏ Jembrophora sawadai Matsumura, 1942a, .ᖩ Ins. Matsumurana 16: 57ڰਖڡᖩڰਖڡბเਂბเਂ Jembrana punctipennis Kato, 1933b, Type deposition: EIHU (Komatsu and Entomol. World 1: 225. Hayashi, 1998). Type deposition: Unknown. Type locality: TAIWAN: Hattsukan Type locality: TAIWAN: Musha [Wushe], [Patungkuan], Nantou Co. (Matsu- Nantou Co. (Kato, 1933b). mura, 1942a). Distribution: Taiwan. Distribution: Taiwan.
Genus Jembra Metcalf and Genus Megafukia Matsumura, ᖩᛳڰᖩᛳ 1940 ̂ыڰHorton, 1934 ˬਖ
Jembra Metcalf and Horton, 1934, Megafukia Matsumura, 1940, J. Fac. Lingnan Sci. J. 13: 408. Agric. Hokkaido Imp. Univ. 45(2): 59. Jembroides Matsumura, 1942a, Ins. Type species: Megafukia gigas (Kato, Matsumurana 16: 55 (Type species: 1933) Jembroides inouyei Matsumura, 1942) [synonymised by Liang, 1999: 339]. 28. Megafukia gigas (Kato, 1933) ŏ ᄂ៉ ᖩڰType species: Jembra nigronervosa ̂ы (Lallemand, 1924) Aphrophora gigas Kato, 1933b, Entomol. Remarks: Metcalf and Horton (1934) World 1: 227. erected the genus Jembra, which Megafukia gigas (Kato, 1933) [transferred only contains the single species J. by Matsumura, 1940: 59]. pallida Metcalf and Horton. But, Type deposition: Unknown. Liang (1999) noted that this species Type locality: TAIWAN: Shinchiku [Hsin- is the synonym of J. nigronervosa chu], Hsinchu Co. (Kato, 1933b). (Lallemand, 1924). Distribution: Taiwan.
26. Jembra inouyei (Matsumura, 1942) Tribe CLOVIINI Schmidt, 1920 ᖩڰJembroides inouyei Matsumura, 1942a, ᙣᐝ Ins. Matsumurana 16: 56. Jembra inouyei (Matsumura, 1942) Cloviini Schmidt, 1920a, Stettin Entomol. [transferred by Liang, 1999: 345]. Ztg. 81: 48. Type deposition: EIHU (Komatsu and ᖩڰHayashi, 1998). Genus Clovia Stål, 1866 ᙣᐝ Type locality: TAIWAN: Arisan [Ali- ᛳ shan], Chiayi Co. (Matsumura, 1942a). Distribution: Taiwan. Clovia Stål, 1866, Hemiptera Africana 4: 75. Genus Jembrophora Matsumura, Betaclovia Matsumura, 1940, J. Fac. ᖩᛳ Agric. Hokkaido Imp. Univ. 45(2): 38ڰᐙ̚ਖ̂ 1942 (Type species: Betaclovia longicephala Jembrophora Matsumura, 1942a, Ins. (Kato, 1933)) [synonymised by Liang, Matsumurana 16: 57. 1998: 243]. Type species: Jembrophora sawadai Type species: Clovia bigoti (Signoret,
ௐˬഇס˟˩˟ᖪௐٿέ៉ 200 1858) 31. Clovia lineatocollis (de Motschulsky, ᖩڰᖩᙣᐝڰโᓂቢโᓂቢᙣᐝ (1859 29. Clovia bipunctata Kirby, 1891 ˟ᕇᙣ Aphrophora lineatocollis de Motschulsky, .(ᖩ 1859: 110 (Metcalf, 1962ڰᐝ Clovia bipunctata Kirby, 1891, Linn. Soc. Clovia perstrigata Kirby, 1891a, Linn. Soc. London J. Zool. 24: 163 (Metcalf, London J. Zool. 24: 162. 1962). Clovia lineatocollis (de Motschulsky); Type deposition: Unknown. Melichar, 1903, Homopteren- Fauna Type locality: CEYLON [SRILANKA] von Ceylon 1903: 135. (Metcalf, 1962). Type deposition: Unknown. Distribution: India (Pondicherry), Bengal, Type locality: CEYLON [Sri Lanka] Sri Lanka, China (Fujian), Japan, (Metcalf, 1962). Taiwan, Java. Distribution: India, Sri Lanka, Taiwan.
30. Clovia conifera (Walker, 1851) ᐄԛ 32. Clovia longicephala Kato, 1933ŏ ܜ ᖩڰᖩ ᙣᐝڰϰਨᙣᐝ Ptyelus conifer Walker, 1851, List Clovia longicephala Kato, 1933b, Entomol. Homopterous Insetcs in BMNH 3: World 1: 220 [reinstated combination 711. by Liang, 1998: 243]. Ptyelus simulans Walker, 1851, List Betaclovia longicephala (Kato) [trans- Homopterous Insetcs in BMNH 3: ferred by Matsumura, 1940: 76]. 717 [synonymised by Metcalf, 1962: Type deposition: Unknown. 100]. Type locality: TAIWAN: Musha [Wushe], Ptyelus frenulatus Stål, 1854, Akad. Nantou Co. (Kato, 1933b). Forhandl. 11: 250 [synonymised by Distribution: Taiwan. Metcalf, 1962: 100]. Clovia frenulata (Stål, 1854) [reinstated 33. Clovia multilineata (Stål, 1865) ቢ ᖩڰcombination by Stål, 1866: 75]. ᙣᐝ Clovia conifer (Walker, 1851) [transferred Ptyelus multilineata Stål, 1865, Ofversigt by Atkinson, 1885: 19]. af Svenska Vetensk. Akad. Forhandl. Type deposition: BMNH (Walker, 1851). 22: 154 (Metcalf, 1962). Type locality: East Indies (Walker, Clovia multilineata (Stål, 1865) [trans- 1851). ferred by Stål, 1866: 76]. Distribution: India, Bangladesh [Bengal], Type deposition: Unknown. Sikkim, Burma, Thailand, Laos, Type locality: CHINA (Metcalf, 1962). Cambodia, Vietnam, China (Xizang, Distribution: China (Guangdong, Sichuan, Tsinghai, Gansu, Yunnan, Guizhou, Yunnan), Taiwan, Oriental region. Guangxi, Guangdong, Fujian), Japan, Taiwan, Philippines, Malay Peninsula, 34. Clovia peracuta Jacobi, 1921 ᄂ៉ ᖩڰᖩᙣᐝڰMalaysia (Malacca, Pahang (new ቢቢᙣᐝ record), Perak (new record), Clovia peracuta Jacobi, 1921, Arch. f. Sarawak), Singapore, Indonesia. Natuegesch. 87: 13. Remarks: We have examined 4 male and Type deposition: SMTD (Emmrich, 11 female specimens of this species 1974). from TARI. Those specimens were Type locality: TAIWAN (Jacobi, 1921). collected by Drs. C.W. & L.B. O’Brien Distribution: Taiwan. from Pahang (1Ɛ, 3Ə) and Perak (3 Ɛ, 8Ə), Malaysia on 7-30 July 1992. 35. Clovia puncta (Walker, 1851) ˘ᕇᙣ ᖩڰᐝ
Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan 201 Ptyelus punctum Walker, 1851, List 37. Nagaclovia formosana Matsumura, ᖩڰᙣᐝܜ Homopterous Insetcs in BMNH 3: 1940 718, 719. Nagaclovia formosana Matsumura, 1940, Ptyelus bipunctipennis Stål, 1854, J. Fac. Agric. Hokkaido Imp. Univ. Ofversigt af Svenska Vetensk. Akad. 45(2): 48. Forhandl. 11: 250 [synonymised by Nagaclovia piceipectus Matsumura, 1940, Metcalf, 1962: 118]. J. Fac. Agric. Hokkaido Imp. Univ. Clovia punctum (Walker, 1851); Metcalf, 45(2): 49 [synonymised by Liang, 1962: 118, 119. 1998: 252]. Type deposition: BMNH. Type deposition: EIHU (Komatsu and Type locality: North Bengle (Walker, Hayashi, 1998). 1851). Type locality: TAIWAN: Kotosho [Orchid Distribution: India (Madras, Bombay Is.], Taitung Co. (Matsumura, 1940). [Mumbai], Uttar Pradesh, Mysore, Distribution: Taiwan (Eastern Taiwan; Bihar), Sikkim, Vietnam (Tonkin, Orchid Island; Green Island (new Tonlon), Bangladesh, China (Sichuan, record)), China, Japan (Okinawa). Hubei, Jiangsu, Yunnan, Guizhou, Remarks: The present authors collected Hunan, Jiangxi, Zhejiang, Guangxi, this species from the plant Guangdong, Hong Kong, Hainan, Miscanthus floridulus (Labill.) Warb. Fujian), Japan (Okinawa, Ryuku), ex Schum. & Laut. (Poaceae) from Taiwan (Kinmen, Orchid Island), 1998 to 2002 in Taiwan and its Java. adjacent islands, such as Orchid Island and Green Island. Its 36. Clovia quadrangularis Metcalf and host-plant range is very narrow by .ᖩ the authors’ collection recordsڰHorton, 1934 ͞ᙣᐝ Clovia quadragularis Metcalf and Horton, 1934, Lingnan Sci. J. 13: 422. Tribe LEPYRONIINI Lallemand ᖩڰClovia rectangularis Lallemand, 1942, and Synave, 1952 Notes Entomol. Chinoise 9: 69 [synonymised by Metcalf, 1962: 120]. Lepyroniini Lallemand and Synave, 1952, Type deposition: ICRI (formerly called Brussels Inst. Roy. Sci. Nat. Bull. 28: Lingnan Univ.) (Hua, 1989) and 17. NCSU (pers. comm.). Type locality: CHINA: Hainan (Metcalf Genus Ariptyelus Matsumura, ᖩᛳڰand Horton, 1934). 1940 Ύᗝਖ Distribution: China (Sichuan, Hubei, Anhui, Hunan, Jiangxi, Zhejiang, Ariptyelus Matsumura, 1940, J. Fac. Yunnan, Guizhou, Guangxi, Hainan, Agric. Hokkaido Imp. Univ. 45(2): 50. Guangdong, Fujian), Taiwan Toroptyelus Matsumura, 1942b, Ins. (Kinmen). Matsumurana 16: 90 (Type species: Toroptyelus arisanus Matsumura, Genus Nagaclovia Matsumura, 1942) [synonymised by Liang, 1998: .[ᖩᛳ 243ڰᙣᐝܜ 1940 Type species: Ariptyelus auropilosus Nagaclovia Matsumura, 1940, J. Fac. (Matsumura, 1907). Agric. Hokkaido Imp. Univ. 45(2): 48. Type species: Nagaclovia formosana 38. Ariptyelus arisanus (Matsumura, ᖩڰMatsumura, 1940 1942) Ύᗝነਖ Toroptyelus arisanus Matsumura, 1942b,
ௐˬഇס˟˩˟ᖪௐٿέ៉ 202 Ins. Matsumurana 16: 91. 40. Lepyronia okadae (Matsumura, 1903) ᖩڰAriptyelus arisanus (Matsumura, 1942) [transferred by Liang, 1998: 252]. Euclovia okadae Matsumura, 1903, Type deposition: EIHU (Komatsu and Sapporo Col. Agric. J. 2: 25. Hayashi, 1998). Lepyronia bifasciata Liu, 1942, Lingnan Type locality: TAIWAN: Arisan [Ali- Sci. J. 20: 245 [synonymised by Nast, shan], Chiayi Co. (Matsumura, 1972: 160]. 1942b; Komatsu and Haya- Lepyronia okadae (Matsumura, 1903) shi, 1998). [transferred by Nast, 1972: 161]. Distribution: Taiwan. Type deposition: EIHU (Komatsu and Hayashi, 1998). 39. Ariptyelus auropilosus (Matsumura, Type locality: JAPAN: Schizuoka (Matsu- ,ᖩ mura, 1903; Komatsu and HayashiڰΎᗝਖͨܛ (1907 Aphrophora auropilosus Matsumura, 1998). 1907, Annot. Zool. Jpn. 6: 113. Distribution: China (Anhui, Fujian, Cercopis auropilosus (Matsumura, 1907) Guangdong, Guizhou, Hebei, Henan, [transferred by Lallemand, 1912: 60]. Hubei, Jianxi, Jiangsu, Jiling, Peuceptyelus auropilosus (Matsumura, Sinkiang, Shandong, Munchuria, 1907) [transferred by Schumacher, Sichunan, Yunnan), Korea Peninsula, 1915a: 88]. Japan (Honshu, Kyushu), Taiwan Ariptyelus auropilosus (Matsumura, 1907) (Liang, 1992). [transferred by Matsumura, 1940: 50]. Genus Nokophora Matsumura, ᖩᛳڰಚыܜ Ariptyelus kikuchii Matsumura, 1942a, 1940 Ins. Matsumurana 16: 47 [synony- mised by Liang, 1998: 252]. Nokophora Matsumura, 1940, J. Fac. Ariptyelus shokanus Matsumura, 1942a, Agric. Hokkaido Imp. Univ. 45(2): 56. Ins. Matsumurana 16: 47 [synony- Type species: Nokophora nokoensis mised by Liang, 1998: 252]. Matsumura, 1940 Type deposition: EIHU (Komatsu and Hayashi, 1998). 41. Nokophora nokoensis Matsumura, ᖩڰಚыܜ Type locality: TAIWAN: Koshun [Hen- 1940 chun], Pingtung Co. (Matsumura, Nokophora nokoensis Matsumura, 1940, J. 1907; Komatsu and Hayashi, 1998). Fac. Agric. Hokkaido Imp. Univ. Distribution: China (Yunnan), Japan, 45(2): 56. Taiwan. Type deposition: EIHU (Komatsu and Hayashi, 1998). Genus Lepyronia Amyot and Type locality: TAIWAN: Noko [Nengkao], .(ᖩᛳ Nantou Co. (Matsumura, 1940ڰServille, 1843 Distribution: Taiwan. Lepyronia Amyot and Serville, 1843, Hemipteres. 1843: 567. Genus Peuceptyelus Sahlberg, ᖩᛳڰEuclovia Matsumura, 1903, Sapporo Col. 1871 Ӊ Agric. J. 2: 24 (Type species: Euclovia okadae Matsumura, 1903) Peuceptyelus Sahlberg, 1871, Not. Fenn. [synonymised by Nast, 1972: 160]. (n.s.) 9(12): 84 (Metcalf, 1962). Type species: Lepyronia coleoptrata Jembropsis Matsumura, 1940, J. Fac. (Linnaeus, 1758) Agric. Hokkaido Imp. Univ. 45(2): 52 (Type species: Jembropsis roku-
Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan 203 rinzana Matsumura, 1940) [synony- Peuceptyelus dimidiatus Matsumura, mised by Liang, 1998: 243]. 1904) [synonymised by Liang, 1998: Kitaptyelus Matsumura, 1940, J. Fac. 244]. Agric. Hokkaido Imp. Univ. 45(2): 36 Oiptyelus Matsumura, 1942a, Ins. (Type species: Peuceptyelus miyabei Matsumurana 16: 66 (Type species: Matsumura, 1911) [synonymised by Peuceptyelus oiwakeanus Matsumura, Liang, 1998: 243]. 1934) [synonymised by Liang, 1998: Ainoptyelus Matsumura, 1942a, Ins. 244]. Matsumurana 16: 46 (Type species: Sappoptyelus Matsumura, 1942b, Ins. Peuceptyelus nigroscutellatus Matsumurana 16: 75 (Type species: Matsumura, 1904) [synonymised by Sappoptyelus jozanus Matsumura, Nast, 1972: 162; synonymy confirmed 1942) [synonymised by Liang, 1998: by Liang, 1998: 244]. 244]. Eguptyelus Matsumura, 1942a, Ins. Type species: Peuceptyelus coriaceus Matsumurana 16: 50 (Type species: (Fallen, 1826) Jembropsis exacavatus Matsumura, 1940) [synonymised by Liang, 1998: 42. Peuceptyelus coriaceus (Fallen, 1826) ᖩڰᐘӉ .[244 Futaptyelus Matsumura, 1942a, Ins. Cercopis coriacea Fallen, 1826, Hemiptera Matsumurana 16: 50 (Type species: Sveciae 2: 14 (Metcalf, 1962). Futaptyelus hiraozanus Matsumura, Aphrophora carixia Walker, 1851, List 1942) [synonymised by Liang, 1998: Homopterous Insetcs in BMNH 3: 244]. 701 [synonymised by Metcalf, 1962: Iwaptyelus Matsumura, 1942a, Ins. 209]. Matsumurana 16: 50 (Type species: Peuceptyelus coriacea (Fallen, 1826) Peuceptyelus ogasawarai Matsumura, [transferred by Sahlberg, 1871: 272] 1940) [synonymised by Liang, 1998: (Metcalf, 1962). 244]. Type deposition: Unknown. Kageptyelus Matsumura, 1942a, Ins. Type locality: SWEDEN (Metcalf, 1962). Matsumurana 16: 58 (Type species: Distribution: Sweden, Norway, Denmark, Kageptyelus kageyamai Matsumura, Finland, Latvia, Estonia, Germany, 1942) [synonymised by Liang, 1998: Hungary, Austria, Poland, Central 244]. Europe, Altai Mountains, Ural Koreptyelus Matsumura, 1942a, Ins. Mountains, Belorussia, Kurile Matsumurana 16: 59 (Type species: Islands, Siberia, China, Taiwan. Kitaptyelus ikumai Matsumura, 1940) [synonymised by Liang, 1998: 244]. 43. Peuceptyelus excavatus (Matsumura, ᖩڰMaptyelus Matsumura, 1942a, Ins. 1940) ਂቡነ୧Ӊ Matsumurana 16: 59 (Type species: Jembropsis excavatus Matsumura, 1940. J. Maptyelus depressus Matsumura, Fac. Agric. Hokkaido Imp. Univ. 1942) [synonymised by Liang, 1998: 45(2): 54. 244]. Eguptyelus excavatus (Matsumura, 1940) Mimoptyelus Matsumura, 1942a, Ins. [transferred by Matsumura, 1942a: Matsumurana 16: 61 (Type species: 50]. Peuceptyelus takaosanus Matsumura, Peuceptelus excavatus (Matsumura, 1940) 1934) [synonymised by Liang, 1998: [transferred by Liang, 1998: 244]. 244]. Type deposition: EIHU (Komatsu and Nikkoptyelus Matsumura, 1942b, Ins. Hayashi, 1998). Matsumurana 16: 102 (Type species: Type locality: TAIWAN: Numanohira
ௐˬഇס˟˩˟ᖪௐٿέ៉ 204 [Chaopin], Alishan, Chiayi Co. 1940. J. Fac. Agric. Hokkaido Imp. (Matsumura, 1940). Univ. 45(2): 52. Distribution: Taiwan. Peuceptelus rokurinzana (Matsumura) [transferred by Liang, 1998: 254]. 44. Peuceptyelus kanmonis (Matsumura, Type deposition: EIHU (Komatsu and .(ᖩ Hayashi, 1998ڰᗕነؾӉਂ (1940 Jembropsis kanmonis Matsumura, 1940. Type locality: TAIWAN: Mt. Rokurin J. Fac. Agric. Hokkaido Imp. Univ. [Lulinshan], Chiayi Co. (Matsumura, 45(2): 53. 1940). Peuceptelus kanmonis (Matsumura, 1940) Distribution: Taiwan. [transferred by Liang, 1998: 253]. Type deposition: EIHU (Komatsu and 48. Peuceptyelus sigillifer (Walker, 1851) ᖩڰHayashi, 1998). ѳӉ Type locality: TAIWAN: Kanmon [Kuang- Aphrophora sigillifer Walker, 1851, List man], Nantou Co. (Matsumura, 1940; Homopterous Insetcs in BMNH 3: Komatsu and Hayashi, 1998). 700. Distribution: Taiwan. Aphrophora maculipes Stål, 1870, Ofversight at Svenska Vetensk. Akad. 45. Peuceptyelus nigriceps (Matsumura, Forhandl.27: 723. [synonymised by [ᖩ Metcalf, 1962: 214ڰαࢵӉ (1940 Jembropsis nigriceps Matsumura, 1940. Aphrophora facialis Kirby, 1891, Linn. Hokkaido Imp. Univ. Fac. Agric. J. Soc. London J. Zool. 24: 161 45 (2): 54. [synonymised by Metcalf, 1962: 214]. Peuceptelus nigriceps (Matsumura) [trans- Aphrophora deserta Melichar, 1903, ferred by Liang, 1998: 254]. Homopteren- Fauna von Ceylon 1903: Type deposition: EIHU (Komatsu and 131 [synonymised by Metcalf, 1962: Hayashi, 1998). 214]. Type locality: TAIWAN: Kanmon [Kuan- Cercopis deserta (Melichar, 1903); gman], Nantou Co. (Matsumura, Lallemand, 1912, Gen. Insect 143: 60 1940). (Metcalf, 1962). Distribution: Taiwan. Cercopis fascialis (Kirby, 1891); Lalle- mand, 1912, Gen. Insect 143: 60 46. Peuceptyelus nitobei (Matsumura, (Metcalf, 1962). -ᖩ Cercopis maculipes (Stål, 1870); LalleڰϨࢵӉ (1940 Jembropsis nitobei Matsumura, 1940. mand, 1912, Gen. Insect 143: 60 Hokkaido Imp. Univ. Fac. Agric. J. (Metcalf, 1962). 45 (2): 53. Peuceptyelus sigilifer (Walker, 1851) Peuceptelus nitobei (Matsumura) [trans- [transferred by Jacobi, 1921: 25]. ferred by Liang, 1998: 254]. Type deposition: BMNH (Walker, Type deposition: EIHU (Komatsu and 1851). Hayashi, 1998). Type locality: Northern India (Walker, Type locality: TAIWAN: Kanmon [Kuang- 1851). man], Nantou Co. (Matsumura, Distribution: India (Mumbai, Madaras, 1940). Uttar Pradesh), Sri Lanka, Burma, Distribution: Taiwan. Bengal, Sikkim, Vietnam (Cochin China, Tonkin), China (Anhui, 47. Peuceptyelus rokurinzana (Matsu- Zhejiang, Hubei, Jiangxi, Hunan, ,ᖩ Fujian, Guangdong, Guangxiڰmura, 1940) เሸӉ Jembropsis rokurinzana Matsumura, Sichuan, Guizhou, Yunnan), Japan,
Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan 205 Taiwan, Philippines, Indonesia Type locality: TAIWAN: Karenko [Hua- (Sumatra, Java, Siberut, Lombok, lien], Hualien Co. (Kato, 133b). Coorg, Flores). Distribution: Taiwan.
49. Peuceptyelus takaosanus Matsumura, 52. Philaenus flavovittatus Kato, 1933 ŏ ᖩڰܜᖩ เ୧ڰਂӉ 1934 Peuceptelus takaosanus Matsumura, 1934, Philaenus flavovittatus Kato, 1933b, Ins. Matsumurana 9: 78 [reinstated Entomol. World 1: 222. combination by Liang, 1998:254]. Type deposition: Unknown. Mimoptyelus takaosanus (Matsumura, Type locality: TAIWAN: Karenko [Hua- 1934) [transferred by Matsumura, lien], Hualien Co. (Kato, 133b). 1942: 61]. Distribution: Taiwan. Type deposition: EIHU (Komatsu and Hayashi, 1998). 53. Philaenus minutus Kato, 1933 ŏ ̈ ᖩڰܜ Type locality: TAIWAN: Mt. Takao [Kaohsiung], Kaohsiung Co. (Matsu- Philaenus minitus Kato, 1933b, Entomol. mura, 1934). World 1: 222. Distribution: Japan (Honshu), Taiwan. Type deposition: Unknown. Type locality: TAIWAN: Hori [Puli], Tribe PHILAENINI Metcalf, Nantou Co. (Kato, 133b). .ᖩ Distribution: Taiwanڰܜ 1955
Philaenini Metcalf, 1955, J. Wash. Acad. 54. Philaenus mushanus Matsumura, ᖩڰܜSci. 45: 264. 1940 ̣୧ᗝ Philaenus mushanus Matsumura, 1940. .Hokkaido Imp. Univ. Fac. Agric. J ڰܜ Genus Philaenus Stal, 1864 ᖩᛳ 45 (2): 44. Type deposition: EIHU (Komatsu and Philaenus Stal, 1864, Stettin Entomol. Hayashi, 1998). Ztg. 25: 66 (Metcalf, 1962). Type locality: TAIWAN: Musha [Wushe], Type species: Cicada spumarius (Lin- Nantou Co. (Matsumura, 1940). naeus, 1758) Distribution: Taiwan.
50. Philaenus arisanus Matsumura, 55. Philaenus nigripectus (Matsumura, ᖩڰܜᖩ 1903) โڰܜ֧̋ܠ 1940 Philaenus arisanus Matsumura, 1940, J. Ptyelus nigripectus Matsumura, 1903, Fac. Agric. Hokkaido Imp. Univ. Sapporo Col. Agric. J. 2: 51. 45(2): 43. Philaenus nigripectus (Matsumura, 1903) Type deposition: EIHU (Komatsu and Mstsumura, 1940. Hokkaido Imp. Hayashi, 1998). Univ. Fac. Agric. J. 45 (2): 76. Type locality: TAIWAN: Mt. Ari [Ali- Type deposition: Unknown. shan], Chiayi Co. (Matsumura, 1940). Type locality: JAPAN: Sapporo (Matsu- Distribution: Taiwan. mura, 1903). Distribution: Japan (Hokkaido, Shikoku), 51. Philaenus castaneus Kato, 1933ŏ ፘ Taiwan. ᖩڰܜ Philaenus castaneus Kato, 1933b, 56. Philaenus pallidus (Melichar, 1903) ᖩڰܜEntomol. World 1: 234. ୶Ғ Type deposition: Unknown. Ptyelus pallidus Melichar, 1903, Homo-
ௐˬഇס˟˩˟ᖪௐٿέ៉ 206 pteren- Fauna von Ceylon 1903: 135. 58. Philagra fusiformis (Walker, 1858) ᖩڰPhilaenus pallidus (Melichar, 1903) ᔧԛ෪ [transferred by Matsumura, 1940: Cyrene fusiformis Walker, 1858, Suppl. 48]. List Homopterous Insetcs in BMNH Type deposition: Museum in Budapest, 1858: 47. Hungary (Melichar, 1903). Philagra fusiformis (Walker, 1858) Type locality: CEYLON [Sri Lanka] [transferred by Distant, 1908: 108]. (Melichar, 1903). Tonga fusiformis (Walker, 1858) [trans- Distribution: India, Sri Lanka, China, ferred by Schmidt, 1910: 174] Japan, Taiwan, Malaya. (Metcalf, 1962). Type deposition: BMNH (Walker, 1858). Tribe PHILAGRINI Schmidt, Type locality: Unknown. ᖩ Distribution: India (Assam, Uttanڰ෪ 1920 Pradesh), Sikkim, Burma, Bang- Philagrini Schmidt, 1920a, Stettin Ento- ladesh, Tailand, Vietnam (Tonkin), mol. Ztg. 81: 51. China (Guangdong, Hong Kong), Taiwan. ᖩڰGenus Philagra Stal, 1863 ෪ ᛳ 58-1. Philagra fusiformis var. numerosa ᖩڰLallemand, 1912 ൺᗝᔧԛ෪ Philagra Stal, 1863, Entomol. Soc. Philagra fusiformis numerosa Lallemand, London Trans. (3)1: 593 [replaced 1912, Gen. Insect 143: 72 (Metcalf, name of Chalepus Walker, 1851 by 1962). Distant in 1908]. Philagra numerosa Kato, 1933a, Three Chalepus Walker, 1851, List Homopterous Colour Illustrated Insects of Japan 4: Insetcs in BMNH 3: 731. pl. 43. Philagrina Lallemand, 1946, Soc. Type deposition: DEI (Gaedike, 1971). Entomol. De Belg. Bul. Et Ann. 82: Type locality: INDIA (Metcalf, 1962). 193 (Type species: Philagrina Distribution: India, Taiwan, Japan. espadon Lallemand, 1946) [synony- mised by Liang, 2001: 444]. 59. Philagra insularis Jacobi, 1921 বវ ᖩڰType species: Philagra douglasi Stål, ᔧԛ෪ 1863 Philagra insularis Jacobi, 1921, Arch. f. Remarks: Metcalf and Horton (1934) Natuegesch. 87: 30. noted the name Chalepus was Type deposition: SMTD (Emmrich, preoccupied by Thunberg (1805). The 1974). genus Chalepus of the family Type locality: TAIWAN (Jacobi, 1921). Chrysomelidae (Insecta: Coleoptera) Distribution: Taiwan. was erected by Thunberg in 1805. 60. Philagra kanoi Matsumura, 1940 ᑢ ᖩڰϨ৳෪֧̋ܠ ֧̋ܠ Philagra arisana Kato, 1933ŏ .57 .ᖩ Philagra kanoi Matsumura, 1940. J. FacڰϨ৳෪ Philagra arisana Kato, 1933b, Entomol. Agric. Hokkaido Imp. Univ. 45(2): 65. World 1: 236. Type deposition: EIHU (Komatsu and Type deposition: Unknown. Hayashi, 1998). Type locality: TAIWAN: Arisan [Ali- Type locality: TAIWAN: Arisan [Ali- shan], Chiayi Co. (Kato, 1933). shan], Chiayi Co. (Matsumura, 1940). Distribution: Taiwan. Distribution: Taiwan.
Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan 207 61. Philagra kuskusuana Matsumura, .ᖩ Ptyelinae Fowler, 1897, Biol. Centrali-Amڰᗝ෪ܜᖩڰᗝ෪ܜՅͩڗՅͩڗ 1942 Philagra kuskusuana Matsumura, 1942b, 2: 174 (Metcalf, 1962). Ins. Matsumurana, 16: 72. Ptyelini Schmidt, 1920, Arch. f. Type deposition: EIHU (Komatsu and Naturgesch. 85: 115. Hayashi, 1998). Type locality: TAIWAN: Kusukusu Genus Kotophora Matsumura, ᖩᛳڰKaoshih], Pingtung Co. (Matsumura, 1940 ᑢൺᐝਖ] 1942b). Distribution: Taiwan. Kotophora Matsumura, 1940. J. Fac. Agric. Hokkaido Imp. Univ. 45(2): 47. 62. Philagra longirostris Schumacher, Type species: Kotophora botelensis (Mat- (ᖩ sumura, 1938ڰᖩ෪ڰᗝᔧԛᔧԛᔧԛ෪ܜᗝܜ 1915ŏ Philagra fusformis longirostris Schu- macher, 1915b, Suppl. Entomol. 4: 65. Kotophora botelensis (Matsumura, ᖩڰᑢൺᐝਖ (1938 .114 Philagra koshunensis Kato, 1933a, Three Aphrophora botelensis Matsumura, 1938, Colour Illustrated Insects of Japan 4: Ins. Matsumurana 12:149. Pl. 43. Kotophora botelensis (Matsumura, 1938) Type deposition: DEI (Gaedike, 1971). [transferred by Matsumura, 1940: Type locality: TAIWAN: Kankau, 75]. Koshun [Hengchun], Pingtung Co. Type deposition: EIHU (Komatsu and (Schu- macher, 1915b). Hayashi, 1998). Distribution: Taiwan. Type locality: TAIWAN: Kotosho [Orchid Island], Taitung Co. (Matsumura, 63. Philagra montana Kato, 1933 ŏ ͉π 1938). .(ᖩ Distribution: Taiwan (Orchid Islandڰ෪̋ Philagra montana Kato, 1933b, Entomol. Remarks: Matsumura (1940) noted the World 1: 235. species is very allied to Poophilus Type deposition: Unknown. costalis (Walker, 1851). Type locality: TAIWAN: Mt. Taihei [Taipingshan], Ilan Co. (Kato, Genus Mesoptyelus Matsumura, ᖩᛳڰ1933b). 1904 ̚ਖ Distribution: Taiwan. Mesoptyelus Matsumura, 1904. Annot. 64. Philagra subrecta Jacobi, 1921 เਂ Zool. Jpn. 5: 48. ᖩ Type species: Mesoptyelus nigrifronsڰ෪ Philagra subrecta Jacobi, 1921, Arch. f. Matsumura, 1904 Natuegesch. 87: 30. Type deposition: SMTD (Emmrich, 66. Mesoptyelus arisanus Matsumura, ᖩڰ̚ਖ˘֧̋ܠ 1940 .(1974 Type locality: TAIWAN (Jacobi, 1921). Mesoptyelus arisanus Matsumura, 1940. J. Distribution: China (Zhejiang, Jiangxi, Fac. Agric. Hokkaido Imp. Univ. Fujian, Guangdong, Hainan, 45(2): 45. Sichuan, Yunnan, Xizang [Tibet]), Type deposition: EIHU (Komatsu and Taiwan, Vietnam. Hayashi, 1998). Type locality: TAIWAN: Arisan [Ali- Tribe PTYELINI Fowler, 1897 shan], Chiayi Co. (Matsumura, 1940; .(ᖩ Komatsu and Hayashi, 1998ڰਖ
ௐˬഇס˟˩˟ᖪௐٿέ៉ 208 Distribution: Taiwan. 23]. Ptyelus natalensis Stål, 1855, Ofversigt af 67. Mesoptyelus auropilosus Kato, 1933ŏ Svenska Vetensk. Akad. Forhandl. 12: :ᖩ 97 [synonymised by Metcalf, 1962ڰᆵ̚ਖ Mesoptyelus auropilosus Kato, 1933b, 23]. Entomol. World 1: 234. Ptyelus rotundatus Fairmair and Signoret, Type deposition: Unknown. 1858: 332. (Metcalf, 1962). Type locality: TAIWAN: Arisan [Ali- Ptyelus dolosus Walker, 1858, Suppl. List shan], Chiayi Co. (Kato, 1933b). Homopterous Insetcs in BMNH 1858: Distribution: Taiwan. 189 [synonymised by Metcalf, 1962: 23]. 68. Mesoptyelus fascialis Kato, 1933ŏ ˘ Poophilus natalensis Stål, 1866, -ᖩ Hemiptera Africana 4: 74 [synonyڰ̚ਖ Mesoptyelus fascialis Kato, 1933b, mised by Metcalf, 1962: 23]. Entomol. World 1: 233. Poophilus costalis (Walker, 1851) Type deposition: Unknown. [transferred by Atkinson, 1885: 114]. Type locality: TAIWAN: Musha [Wushe], Poophilus actuosus Distant, 1892, Homo- Nantou Co. (Kato, 1933b). ptera. A naturalist in the Transvaal Distribution: Russian Maritime Territoy, 1892: 248 [synonymised by Metcalf, Taiwan, China. 1962: 24]. Gallicana flava Metcalf and Horton, 1934, 69. Mesoptyelus karenkonis Matsumura, Lingnan Sci. J. 13: 420 [synony- .[ᖩ mised by Metcalf, 1962: 26ڰ̚ਖˬ 1940 Mesoptyelus karenkonis Matsumura, 1940. Type deposition: BMNH (Walker, 1851). J. Fac. Agric. Hokkaido Imp. Univ. Type locality: Northern India (Walker, 45(2): 46. 1851). Type deposition: EIHU (Komatsu and Distribution: Western Africa (Dahomey Hayashi, 1998). [Benin], Guinea, Ivory Coast), Type locality: TAIWAN: Karenko Central Africa (Angola, Cameroon, [Hualien], Hualien Co. (Matsumura, Central African Republic, Congo, 1940). Gabon), Southern Africa (Nyasaland Distribution: Taiwan. [Malawi], South Africa (Natal, Transvaal)), Eastern Africa (Somali- Genus Poophilus Stål, 1866 ൺᐝ land, Tanganyika [Tanzania], ,ᖩᛳ Uganda), Russia, Iran, Afghanistanڰਖ India (Assam, Bihar, Mumbai, Poophilus Stål, 1866, Hemiptera Africana Darjeeling, Madras, Uttar Pradesh), 4: 72. Bangladesh, Sri Lanka, Nepal, China Type species: Poophilus actuosus (Stål, (Xinjiang, Fujian, Hainan, Guang- 1855) dong). Japan (Okinawa), Taiwan (Kinmen), Philippines, Malay Penin- 70. Poophilus costalis (Walker, 1851) sula, Singapore. ᖩڰළൺᐝਖ Ptyelus costalis Walker, 1851, List 71. Poophilus grisescens (Schaum, 1853) ᖩڰHomopterous Insetcs in BMNH 3: ѷൺᐝਖ 707. Aphrophora grisescens Schaum, 1853, Ber. Ptyelus concolor Walker, 1851, List Akad. Berlin 1853: 359 (Metcalf, Homopterous Insetcs in BMNH 3: 1962). 715 [synonymised by Metcalf, 1962: Poophilus grisecens (Schaum, 1853)
Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan 209 [transferred by Jacobi, 1904: 776]. Type deposition: Unknown. 73. Ptyelus vittatus Kato, 1933ŏ ᓂ୧৳ ᖩڰType locality: MOZAMBIQUE (Metcalf, ਖ 1962). Ptyelus vittatus Kato, 1933b, Entomol. Distribution: Central Africa (Cameroon, World 1: 221. Congo), Eastern Africa (Tanganyika, Type deposition: Unknown. Uganda), Western Africa (Dahomey), Type locality: TAIWAN: Karenko Southern Africa (Mozambique, Natal, [Hualien], Hualien Co. (Kato, 1933b). South Africa), India, Taiwan, Distribution: Taiwan. Malaysia. Acknowledgments Genus Ptyelus Le Peletier and ᖩᛳ The authors wish to express theirڰServille, 1825 ਖ cordial thanks to Drs. Lois B. O’Brien Ptyelus Le Peletier and Serville, 1825, and Charles W. O’Brien (Florida A&M Histoire Naturelle. Entomologie, ou Univ., USA) for the Oriental Cercopoidea histoire Naturelle des Crustaces, des specimens they kindly sent us, Dr. Lois B. Arachnides et des Insectes 10: 608 O’Brien for her critical reading and (Metcalf, 1962). valuable comments in improving the Ptyela Germar, 1833, Rev. Entomol. contents of the paper, Prof. Dr. P. Cai for Silbermann 1: 179 [synonymised by loaning us their collection, and Dr. Bob Metcalf, 1962: 33]. Blinn for the information of Metcalf’s Plinthacrus Spinola, 1850, Soc. Ital. types in the NCSU. We also thank to Dr. Modena Mem. 25: 56 [synonymised Reinhard Gaedike (DEI), Dr. Michael Ohl, by Metcalf, 1962: 34]. Dr. Ursula Göllner-Scheiding, Ms. Isodes Ampleus Licent, Thèses présentées à la Dorant and Mr. Peter Schönefeld Faculté des Sciences de Nancy pour (Museum für Naturkunde der Humboldt obtenir le grade de docteur Universität, Berlin, Germany) for their es-sciences naturelles 1912: 28 kind assistance during the first author [synonymised by Metcalf, 1962: 38]. visit to their museums during the PPP Type species: Ptyelus flavescens projects (8702PPP16), PPP (0900045437) (Fabricius, 1794). and thanks to Dr. Prof. A. F. Emel’yanov (Zoological Institute, Russian Academy of 72. Ptyelus tamahonis Matsumura, 1940 Sciences, Russian Federation) for ᖩ bibliographic assistance. This work wasڰܜ֧̋ܠᑢ Ptyelus tamahonis Matsumura, 1940, J. financially supported mostly by NSC Fac. Agric. Hokkaido Imp. Univ. 89-2313-B-005-102 and partialy by the 45(2): 74. Council of Agriculture, Executive Yuan, Type deposition: EIHU (Komatsu and Taiwan, Republic of China (91AS-2.4.3- Hayashi, 1998). CI-C1). Type locality: TAIWAN: Tomaho (near Mt. Niitaka), Nantou Co. (Mat- References sumura, 1940). Distribution: Taiwan. Amyot, C. J. B., and J. G. A. Serville. Remarks: Matsumura (1940) noted the 1843. Deuxieme partie. Homopteres. species is very allied to Philaenus Homoptera Latr. Histoire Naturelle arisanus Matsumura, but it lacks a des Insectes. Hemipteres 1843: transverse furrow along each side of 1-676. the vertical ridge. Atkinson, E. T. 1885. Notes on Indian
ௐˬഇס˟˩˟ᖪௐٿέ៉ 210 Rhynchota, No. 2. J. Asiatic Soc. Colour Illustrated Insects of Japan Bengal 54: 5-23. 4: 1-9 (in Japanese). China, W. E. 1951. A note on Kato, M. 1933b. Notes on Japanese Matsumura’s recent paper on the Homoptera, with descriptions of one Palaearctic Cercopoidea (Homo- new genus and some new species. ptera). Entomol. Mon. Mag. 87: 279. Entomol. World 1: 220-237 (in Chou, I., and A. P. Liang. 1987. Two Japanese). new species of Jembrana distant Komatsu, T. 1997. A revision of the from China (Homoptera: Aphro- froghopper genus Aphrophora phoridae). Entomotaxonomia 9: 283- Germar (Homoptera, Cercopoidea, 288 (in Chinese). Aphrophoridae) from Japan. Parts 1. Chou, I., F. Yuan, and A. P. Liang. 1992. Jpn. J. Entomol. 65: 81-96. Homoptera: Aphrophoridae. pp. 234- Kwon, Y. J., and C. E. Lee. 1979. 242. In: S. X. Chen, ed. Insects of Morphological and phylogenetic the Hengduan Mountains region. studies on the male genitalia of the Vol. 1. Science Press, Beijing (in Korean Cercopoidea (Homoptera: Chinese). Auchenorrhyncha). Nature and Life Distant, W. L. 1908. Rhynchota IV. 9: 1-31. (Homoptera and Appendix). pp. Lallemand, V. 1942. Notes sur quelques 79-156. In: The fauna of British espèces recueillies par le R. Piel India including Ceylon and Burma. (Musée Heude, Shanghai) et le R. P. Taylor and Francis, London. de Cooman (Hoa Binh, Tonkin). Emmrich, R. 1974. Das Typen-Material Notes Entomol. Chinoise 9: 69-77. der Zikaden (Homoptera, Auchenor- Lallemand, V. 1946. Quatrième note sur rhyncha) des Staatlichen Museums les cercopides. Soc. Entomol. Belg. für Tierkunde Dresden III. Tiel: Bul. Ann. 82: 189-197. Cercopoidea. Entom. Abh. Mus. Lallemand, V., and H. Synave. 1952. Tierk. Dresden 40: 105-126. Contribution a l’ètude des Gaedike, H. 1971. Katalog der in den Hémiptères. I. - Note sur quelques Sammlungen des ehemaligen Deuts- Homoptères de l’Institut royal des chen Entomologischen Institutes sciences naturelles de Belgique. aufbewahrten Typen-VI. Beitr. Ento- Brussels Inst. Roy. Sci. Nat. Bull. mol. 21: 315-340. 28: 1-23. Germar, E. F. 1821. Bemerkungen über Lee, C. E., S. M. Lee, and Y. J. Kwon. einige Gattungen der Cicadarien. 1976. An insect list of the Mag. Entomol. 4: 1-106. Auchenorrhyncha preserved in the Hua, L. Z. 1989. List of insect specimens National Science Museum of Korea. of Zhongshan (Sun Yatsen) Univer- Nature & Life (Kyungpook J. Biol. sity. Guangzhou, China, 158 pp. (in Sci.), 6: 51-64. Chinese). Liang, A. P. 1992. Homoptera: Cerco- Jacobi, A. 1904. Homopteren aus poidea. pp. 126-129. In: F. S. Huang, Nordost-Afrika, gesammelt von Oscar ed. Insects of Wuling Mountains Neumann. Zool. Jahrb. Abt. F. area, southern China. Science Press, System. 19: 761-782. Beijing (in Chinese). Jacobi, A. 1921. Kritische Bemer- Liang, A. P. 1997. Homoptera: kungen uber die Cercopidae Cercopoidea: Aphrophoridae and (Rhynchota Homoptera). Arch. f. Cercopidae. pp. 310-315. In: X. K. Naturgesch. 87: 1-65. Yang, ed. Insects of the Three Kato, M. 1933a. Homoptera. Three Gorges Reservoir Area of Yangtze
Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan 211 River. Chongoing Publishing House, 12: 147-153. Chongqing, China. Matsumura, S. 1940. New species and Liang, A. P. 1998. Oriental and eastern genera of Cercopidae in Japan, Korea Palaearctic aphrophorid fauna and Formosa, with a list of the (Homoptera: Aphrophoridae): tax- known species. Hokkaido Imp. Univ. onomic changes and nomenclatural Fac. Agric. J. 45: 35-82. notes. Oriental Ins. 32: 239-257. Matsumura, S. 1942a. New species and Liang, A. P. 1999. On the spittlebug new genera of Palaearctic Super- Genus Jembra Metcalf and Horton family Cercopoidea with a tabular with description of one new species key to the classification (I). Ins. (Homoptera: Cercopoidea: Aphro- Matsumurana, 16: 44-70. phoridae). Oriental Ins. 33: 337-348. Matsumura, S. 1942b. New species and Liang, A. P. 2000. Lectotype desig- new genera of Palaearctic nations and taxonomic notes on P. Superfamily Cercopoidea with a R. Uhler’s Japanese Cercopoidea tabular key to the classification (II). (Homoptera) in the US National Ins. Matsumurana, 16: 71-106. Museum of Natural History. J. New Melichar, L. 1903. Homopteren- Fauna York Entomol. Soc. 108: 268-272. von Ceylon 1903: I-IV, 1-248. Liang, A. P. 2001. Generic assignment Metcalf, Z. P. 1955. New names in the and synonymy of some Indo-West Homoptera. J. Wash. Acad. Sci. 45: Pacific Aphrophoridae (Hemiptera: 262-267. Cercopoidea) in the Natural History Metcalf, Z. P. 1962. General catalogue of Museum, London. Fla. Entomol. 84: the Homoptera Fasc. VII, Cercopoidea. 442-445. Part 3. Aphrophoridae. 600 pp. Licent, P. E. 1912. Recherches d’ana- Waverly Press, Baltimore, MD. tomie et de physiologie comparées Metcalf, Z. P., and G. Horton. 1934. The sur le tube digestif des Homoptères Cercopoidea (Homoptera) of China. supérieurs. Thèses présentées à la Lingnan Sci. J. 13: 367-429. Faculté des Sciences de Nancy pour Nast, J. 1972. Palaearctic Auchenorr- obtenir le grade de docteur hyncha (Homoptera) an annotated es-sciences naturelles 1912: 7-161. checklist. 550 pp. Warszawa. Matsumura, S. 1903. Monographie der Ossiannilsson. F. 1981. The Auchenorr- Cercopiden Japans. Sapporo Col. hyncha (Homoptera) of Fennoscandia Agric. Jour 2: 15-52. and Denmark. Part 2. The Families Matsumura, S. 1904. Additamenta zur Cicadidae, Cercopidae, Membracidae, Monographie der Cercopiden Japans, and Cicadellidae (excl. Deltoce- mit der Beschreibung einer neuen phalinae). Fauna Entomol. Scand. Cicada-Art. Annot. Zool. Jpn. 5: 7(Pt. 2): 223-593. 31-54. Shih, H. T., J. T. Yang, and M. M. Yang. Matsumura, S. 1907. Die Cicadinen 2000. Cercopoidea (Homoptera: Au- Japans. [Tokyo] Annot. Zool. Jpn. 6: chenorrhyncha) of Orchid Island. 83-116. 2000 Symposium of The Biological Matsumura, S. 1934. Insects collected at Society of China and Society of the foot of Mt. Yatsugadake and its Streams: 63 (in Chinese). environment. Ins. Matsumurana 9: Shih, H. T., and J. T. Yang. 2002. Three 60-80. New Records Aphrophoridae (Homo- Matsumura, S. 1938. Homopterous insects ptera) of Kinmen Islands. Polish J. collected by Mr. Tadao Kano at Entomol. 71: 91-100. Kotosho, Formosa. Ins. Matsumurana. Stål, C. 1854. Nya Hemiptera. Ofversight
ௐˬഇס˟˩˟ᖪௐٿέ៉ 212 at Svenska Vetensk. Akad. Forhandl. Schumacher, F. 1915b. Homoptera in H. 11: 231-255. Sauter’s Formosa – Ausbeut. Suppl. Stål, C. 1863. Hemipteroum exoticorum Entomol. 4: 108-142. generum et specierum nonnullarum Uhler, P. R. 1896. Summary of the novarum descriptions. Entomol. Soc. Hemiptera of Japan presented to the London Trans. 1: 571-603. United States National Museum by Stål, C. 1866. Hemiptera Homoptera Latr. Professor Mitzukuri. US Natl. Mus. Hemiptera Africana 4: 1-276. Proc. 19: 255-297. Schmidt, E. 1920a. Neue Zikaden von den Walker, F. 1851. List of the specimens of Philippinen, Sumatra und Java. homopterous insects in the collection (Rhynchota-Homoptera). Stettin of the British Museum 3: 637-907. Entomol. Ztg. 81: 43 -56. Walker, F. 1858. Supplement. List of the Schumacher, F. 1915a. Der genen- specimens of homopterous insects in wartige stand unserer Kenntnis von the collection of the British Museum der Homopteren – Fauna der Insel 1858: 1-307. Fomosa unter besonderer Beruck- sichtigung von Sauter’s schem Received June 24, 2002 Material. Berlin Zool. Mus. Mitt. 8: Accepted July 23, 2002 73-134.
Checklist of Aphrophoridae (Homoptera: Cercopoidea) from Taiwan 213 !ᖩᓁࡊğڰᖩࡊЩᐂĞТਂϫĈڰᄂ៉யы
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