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Vol. 3 No.1 April 1999 ISSN 1342-8144 Formerly - Transactions and Proceedings of the Palaeontological Society of Japan Vol. 3 No.1 April 1999 The Palaeontological Society of Japan Co- Editors Kazushige Tanabe and Tomoki Kase Language Editor Marlin Janal (New Yark, USA) Associate Editors Jan Bergstrom (Swedish Museum of Natural History, Stockholm, Sweden), Alan G. Beu (Institute of Geological and Nuclear Sciences, Lower Hutt, New Zealand), Satoru Chiba (Tohoku University, Sendai, Japan), Yoichi Ezaki (Osaka City University, Osaka, Japan), James C.lngle, Jr. (Stanford University, Stanford, USA), Kunio Kaiho (Tohoku University, Sendai, Japan), Susan M. Kidwell (University of Chicago, Chicago, USA), Hiroshi Kitazato (Shizuoka University, Shizuoka, Japan), Naoki Kohno (National Science Museum, Tokyo, Japan), Neil H. Landman (Amemican Museum of Natural History, New York, USA), Haruyoshi Maeda (Kyoto University, Kyoto, Japan), Atsushi Matsuoka (Niigata University, Niigata, Japan), Rihito Morita (Natural History Museum and Institute, Chiba, Japan), Harufumi Nishida (Chuo University, Tokyo, Japan), Kenshiro Ogasawara (University of Tsukuba, Tsukuba, Japan), Tatsuo Oji (University of Tokyo, Tokyo, Japan), Andrew B. Smith (Natural History Museum, London, Great Britain), Roger O.K. Thomas (Franklin and Marshall College, Lancaster, USA), Katsumi Ueno (Fukuoka University, Fukuoka, Japan), Wang Hongzhen (China University of Geosciences, Beijing, China), Yang Seong Young (Kyungpook National University, Taegu, Korea) Officers for 1999- 2000 President : Kei Mori Councillors : Kiyotaka Chinzei, Takashi Hamada, Yoshikazu Hasegawa, Itaru Hayami, Hiromichi Hirano, Noriyuki Ikeya, Junji Itoigawa, Tomoki Kase, Hiroshi Kitazato, Itaru Koizumi, Haruyoshi Maeda, Ryuichi Majima, Makoto Manabe, Hiroshi Noda, Ikuo Obata, Kenshiro Ogasawara, Terufumi Ohno, Tatsuo Oji, Tomowo Ozawa, Yukimitsu Tomida, Tsunemasa Saito, Takeshi Setoguchi, Kazushige Tanabe, Akira Yao Members of Standing Committee: Hiroshi Kitazato (General Affairs), Tatsuo Oji (Laison Officer), Makoto Manabe (Finance), Kazushige Tanabe (Editor in Chief, PRJ, Tomoki Kase (Co- Editor, PRJ, Ryuichi Majima (Planning), Hiromichi Hirano (Membership), Kenshiro Ogasawara (Foreign Affairs), Haruyoshi Maeda (Publicity Officer), Noriyuki Ikeya (Editor, "Fossils"), Yukimitsu Tomida (Editor in Chief, Special Papers), Tamiko Ohana (Representative, Union of Natural History Societies). Secretaries: Masanori Shimamoto, Takao Ubukata (General Affairs), Hajime Taru (Planning), Tokuji Mitsugi (Membership), Shuko Adachi (Foreign Affairs), Kazuyoshi Endo, Yasunari Shigeta (Editors of PRJ, Akira Tsukagoshi (Editor of "Fossils"), Naoki Kohno (Editor of Special Papers) Auditor: Nobuhiro Kotake Notice about photocopying: In order to photocopy any work from this publication, you or your organization must obtain permission from the following organization which has been delegated for copyright for clearance by the copyright owner of this publication. Except in the USA, Japan Academic Association for Copyright Clearance (JAACC), 41 - 6 Akasaka 9- chome, Minato- ku, Tokyo 107- 0052, Japan. Phone: 81 - 3- 3475- 5618, Fax: 81 - 3- 3475- 5619, E- mail: kammori @ msh.biglobe.ac.jp In the USA, Copyright Clearance Center, Inc., 222 Rosewood Drive, Danvers, MA 01923, USA. Phone : (978) 750- 8400, Fax: (978)750- 4744, www.copyright.com Cover: Idealized sketch of Nipponites mirabilis Yabe, a Late Cretaceous (Turonian) nostoceratid ammonite. Various reconstructions of the mode of life of this species have been proposed, because of its curiously meandering shell form (after T. Okamoto, 1988). All communication relating to this journal should be addressed to the PALAEONTOLOGICAL SOCIETY OF JAPAN c/ o Business Center for Academic Societies, Honkomagome 5-16- 9, Bunkyo- ku, Tokyo 113- 8622, Japan Paleontological Research, vol. 3, no.1, pp.1-17, 6 Figs., April 3D, 1999 © by the Palaeontological Society of Japan Permian bivalves from West Spitsbergen, Svalbard Islands, Norway KEIJI NAKAZAWA 28-2, Koyama Shimouchikawara-cho, Kita-ku, Kyoto 603-8132, Japan Received 6 June 1998; Revised manuscript accepted 27 December 1998 Abstract. The bivalve fossils collected by Japanese-Norwegian research groups from the Kapp Starostin Formation in west Spitsbergen are described. They comprise fourteen species belonging to the pterioida and two speCies of the Arcoida. Among them, six species, including two that are indeterminate, are newly described. They are Grammatodon (Cosmetodon)? suzuki;' G. (C.)? sp. ind., Streblochondria win­ snesi, Vorkutopecten svalbardensis, Deltopecten sp. ind., and Palaeolima nakamurai. The fauna belongs to the Boreal bioprovince, but a single species, Cassianoides sexcostatus (Stuckenberg) has also been reported from the Central Rocky Mountains of the United States. The bivalve fauna suggests an Artinskian-Kungurian age for the Kapp Starostin Formation. This is somewhat earlier than the age deduced from brachiopods and bryozoans, but it is not decisive because the materials are poor. The Kapp Starostin Formation is conformably overlain by the Otoceras-bearing, earliest Triassic Vardebukta Formation, so a time-gap corresponding at least to the Dorashamian and Dzhulfian (= Tatarian) is inferred between the two formations. Key words: Kapp Starostin Formation, Permian bivalves, Spitsbergen Introduction (Aequipecten)? keyserlingiformis Licharew, Pecten (Aequi­ pecten) keyserlingi Stuckenberg, Pecten (Pseudamusium) cf. The study materials were collected by Nakamura et al. in ufaensis Tschernyschew, Pecten (Pseudamusium) ex aft. 1984 and 1990, and by Nakazawa et al. in 1986 from the sericeus Verneuil, Parallelodon sp. ind., genus [=Paral­ Permian Kapp Starostin Formation of west Spitsbergen, in lelodon?] et sp. ind., Clidophorus? sp. ind., Pecten wilc­ the Svalbard Islands, Norway. Brachiopods and bryozoans zeki Toula, and Leda sp. indo are most abundant among the macrofossils. Many species Among the species described by Toula, Aviculopecten of brachiopods have been described by various authors, bouei was referred to Aviculopecten (Deltopecten) mutabilis notably, Frebold (1937) and Gobbet (1963). Bivalves are by Licharew (1927). Pecten cf. ellipticus was identified as found rather rarely and the descriptive studies are few. Pecten (Pseudamusium) ex aft. sericeus Verneuil, and Pecten Toula (1873,1875a, b) first described the following species, (Aviculopecten) cf. dissimilis was compared to Aviculopecten collected from "Carboniferous-Permian" strata on S¢rkapp netschajewi Licharew and reported as A viculopecten ? sp. Island oft the southerntip of Spitsbergen, and Axel Island in indo by Frebold (1937). Aviculopecten kokscharofi reported Bellsund and on Hornsund, on the west coast of Spitsber­ by Toula (1873) is probably identical with Aviculopecten cf. gen: hiemalis illustrated by Licharew (1927, especially pI. 5, fig. 20), Pecten (Aviculopecten) bouei Verneuil, Pecten and is referred to Etheripecten cf. mutabi/is Licharew in the (Aviculopecten) kokscharofi Verneuil, Pecten (Aviculopecten) present paper. cf. ellipticus Phillips, Pecten (Aviculopecten) cf. dissimilis Among the species reported by Frebold, Aequipecten Fleming, Pecten (Aviculopecten) wilczeki Toula, Gervillia cf. keyserlingi was considered to belong to the genus Mor­ antiqua Munster, Aviculopecten draschei Toula and Gervillia rispecten Muromtseva and Guskov by Muromtseva (1984, p. sp. 75). Morrispecten is, however, a junior synonym of Undo­ Frebold (1937) described the following species, mostly from pecten Waterhouse, 1982 (Newell and Boyd, 1995). Pecten the upper part of the Kapp Starostin Formation in the (Aequipecten)? keyserlingiformis and Aviculopecten Festningen section, located on the southern coast of Spits­ (Deltopecten) cf. mutabilis and A. (D.) cf. hiemalis of Frebold bergen at the entrance to Isfjorden and from S¢rkapp are referred to Etheripecten keyserlingiformis and E. cf. Island: mutabilis ?, respectively. Pseudamusium cf. ufaensis and Aviculopecten (Deltopecten) cf. mutabilis Licharew and A. Parallelodon sp. indo are identified as Streblochondria win­ (D.) cf. hiemalis Salter, Aviculopecten? sp. indet., Pecten snesi sp. nov. and Grammatodon (Cosmetodon) ? suzukii sp. - 2 Keiji Nakazawa nov., respectively, in the present paper. Sixteen species The Hovtinden Member, about 225 m thick, consists of including four new species and two indeterminable ones are seven units, 6 to 12, each composed mainly of spicularite, described. All the materials are kept at the Department of alternation of spicularite and spicularitic shale, bioclastic Geology and Mineralogy, Faculty of Science, Kyoto Univer­ limestone, spicularite, bioclastic limestone (partly silicified), sity. alternation of siltstone and paper shales, and quartzose sandy shale or siltstone, respectively, in ascending order. Stratigraphy Bivalve fossils have been obtained from Units 5, 7, 8, 9 and 11. The fossils were collected at four localities in west Spits­ (2) Reinodden section (Figure 2B) bergen, namely, the Festningen section along the southern At Reinodden the formation reaches more than 300 m in coast of the entrance to Isfjorden, the Skansbukta section at thickness. It is classified into eight units, A to H, in ascend­ Billfjorden, and the Reinodden and Ahlstrandodden sections ing order (Nakazawa et a/., 1990). Unit A, less than 5 m along the southern coast near the entrance of Van Keulenf­ thick, is the Vrpringen Member consisting of fossiliferous jorden, in Bellsund (Figure 1). Faunas from the first and bioclastic
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