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Home , Acrolophitus, Acrolophitus hirtipes, Arethaea, Bootettix argentatus, Cibolacris, Cordillacris crenulata

INSECTA MUNDI, Vol. 15, No.1, March, 2001 35

Preliminary checklist of the orthopteroid (, Mantodea, Phasmatodea,) of

John A. Stidham 301 Pebble Creek Dr., Garland, TX 75040

and

Thomas A. Stidham Department of Integrative Biology, Museum of Paleontology, and Museum of Vertebrate Zoology, University of California, Berkeley, CA 94720,

Abstract: Texas has one of the most diverse orthopteroid assemblages of any state in the United States, reflecting the varied habitats found in the state. Three hundred and eighty-nine species and 78 subspecies of orthopteroid insects (Blattodea, Mantodea, Phasmatodea, and Orthoptera) have published records for the state of Texas. This is the first such comprehensive checklist for Texas and should aid future work on these groups in this area.

Introduction (Flook and Rowell, 1997). The species are listed alphabetically within families or subfamilies, and Overall, the orthopteroid fauna of North Amer­ therefore the alphabetical taxonomic listing does ica is fairly well known. However, details of the not imply any phylogenetic relationships within geographic distribution of many orthopteroid spe­ the subfamily or family. The reference for the cies in the U.S. are not well documented. We have published record is given after the species name compiled the published records of Texas ortho­ and authority. The alpha is still unclear pteroids, as a basis for future study. Most of these for a few species included in this list, and therefore records have been confirmed through collections additional species may be added in the future as the made by the authors throughout Texas over the last number of taxa is clarified. This checklist is meant 20 years. The study of Texas orthopteroids began in to act as an aid to collectors, collection managers, the mid-nineteenth century and continues today field workers, and other researchers working on with 389 species currently recorded in the state. Texas orthopteroids. This places the Texan orthopteroid fauna as one of the most diverse of any state. There are 197 species The Orthopteroidea of Texas of known from Texas, compared to the over 200 species from California (Strohecker et Order Blattodea aI., 1968), though other orthopteroid groups are more diverse in Texas. The large number of species Farrrily Blattelidae is most likely related to the great variety of habitats found in Texas, from desert in the West, to high Blattella germanica (Linnaeus) (Hebard, 1943b) plains in the North, and subtropical climates in the Blattella uaga Hebard (Atkinson et aI., 1991) South. Combined with failure of Pleistocene gla­ Cariblatta lutea lutea (Sa us sure and Zehntner) ciers to reach Texas, the state harbors factors (Atkinson et aI., 1991) necessary for long term orthopteroid diversifica­ Cariblatta lutea minima Hebard (Atkinson et aI., tion, as seen in its speciose fauna. However, the 1991) number of endemic species is low compared to Chorisoneura texensis Saussure and Zehntner (He­ California, since many of the habitats (and hence bard, 1943b) species ranges) in Texas extend into the adjacent Euthlastoblatta abortiua Caudell (Hebard, 1943b) states and into Mexico. Euthlastoblatta gemma (Hebard) (Atkinson et aI., Supraspecific taxonomy follows Otte (1981, 1991) 1984) and Helfer (1987). Tridactylids are placed in Ischnoptera deropeltifonnis (Brunner) (Hebard, following tradition, though recent molec­ 1943b) ular data suggest their placement within 36 Volmue 15, No.1, March, 2001, INSECTA MUNDI

Ischnoptera rufa occidentalis Saussure (Hebard, Order Mantodea 1943b) bolliana (Saussure and Zehntner) (He­ Family Mantidae bard, 1943b) Parcoblatta caudelli Hebard (Hebard, 1943b) Brunnea borealis Scudder (Hebard, 1943b) Parcoblatta desertae (Rehn and Hebard) (Hebard, Litaneutria minor (Scudder) (Hebard, 1943b) 1943b) Oligonicellamexicanus (Saussure) (Hebard, 1943b) Parcoblatta divisa (Saussure and Zehntner) (He­ Oligonicella scudderi (Saussure) (Hebard, 1943b) bard, 1943b) Phyllovates chlorophaea (Blanchard) (Hebard, Parcoblatta fulvescens (Saussure and Zehntner) 1943b) (Hebard, 1943b) Stagmomantis californica Rehn and Hebard (He­ (Brunner) (Hebard, 1943b) bard, 1943b) Parcoblatta pennsylvanica (DeGeer) (Hebard, Stagnwmantis carolina (Johannson) (Hebard, 1943b) 1943b) Parcoblatta virginica (Brunner) (Hebard, 1943b) Stagnwmantis limbata (Hahn) (Hebard, 1943b) Parcoblatta zebra Hebard (Hebard, 1943b) Yersiniops solitarium (Scudder) (Hebard, 1943b) Plectoptera picta Saussure and Zehntner (Atkin- son et aI., 1991) Order Phasmatodea Pseudomops septentrionalis Hebard (Hebard, 1943b) Family Pseudophasmatidae Supella longipalpa (Fabricius) (Hebard, 1943b) Anisomorphaferruginea (Beauvois) (Hebard, 1943b) Family Blattidae Family Heteronemiidae Blatta lateral is (Walker) (Atkinson et aI., 1991) Blatta orientalis Linnaeus (Hebard, 1943b) Diapheromera covillea (Rehn and Hebard) (He- Neostylopyga rhombifolea (Stoll) (Atkinson et aI., bard, 1943b) 1991) Diapheromera femorata (Say) (Hebard, 1943b) Periplaneta americana (Linnaeus) (Hebard, 1943b) Diapheromera persimilis Caudell (Hebard, 1943b) Periplaneta australasiae (Fabricius) (Hebard, DiapherOinera tamaulipennis (Rehn) (Hebard, 1943b) 1943b) Periplaneta brunnea Burmeister (Hebard, 1943b) DiapherOl1lera torquata Hebard (Hebard, 1943b) Periplaneta fuliginosa (Serville) (Hebard, 1943b) Diapheromera velii eucne111is Hebard (Hebard, 1943b) Family Blaberidae Diapheromera velii velii Walsh (Hebard, 1943b) Megaphasma denticrus (StiH) (Hebard, 1943b) Panchlora cubensis Saussure (Hebard, 1943b) Para bacillus coloradus (Scudder) (Hebard, 1943b) Pycnoscelus surinamensis (Linnaeus) (Hebard, Pseudosennyle straminea (Scudder) (Hebard, 1943b) 1943b) Pseudosennyle strigata (Scudder) (Hebard, 1943b) Sennyle mexicana (Saussure) (Hebard, 1943b) Family Polyphagidae Order Orthoptera Arenivaga bolliana (Saussure) (Hebard, 1943b) Arenivaga erratica Rehn (Hebard, 1943b) Suborder Caelifera Arenivaga grata Hebard (Hebard, 1943b) Arenivaga tonlwwa Hebard (Hebard, 1943b) Family Tetrigidae Attaphila fungicola Wheeler (Hebard, 1943b) Compsodes cucullatus Saussure and Zehntner (At- Neotettix femoratus (Scudder) (Rehn and Grant, kinson et aI., 1991) 1961) Compsodes schwarzi (Caudell) (Atkinson et aI., Neotettix nullisinus (Hancock) (Rehn and Grant, 1991) 1961) Eremoblatta subdiaphana (Scudder) (Hebard, Nomotettix cristatus compressus Morse (Rehn and 1943b) Grant, 1961) INSECTA MUNDI, Vol. 15, No.1, March, 2001 37

Pamtettix aztec us (Saussure) (Rehn and Grant, Menneria bivitatta bivitatta (Serville) (Knutson, 1961) 1940) Pamtettix brevipennis (Hancock) (Rehn and Grant, Mermeria bivitatta maculipennis Bruner (Knut- 1961) son, 1940) Pamtettix cucullatus (Burmeister) (Rehn and Grant, Mermeria picta neomexicana (Thomas) (Otte, 1981) 1961) Menneria picta picta (Walker) (Tinkham, 1948) Paratettix mexicanus (Saussure) (Rehn and Grant, Mermeria texana Bruner (Tinkham, 1948) 1961) Opeia obscum (Thomas) (Otte, 1981) Paratettix rugosus (Scudder) (Rehn and Grant, Orphulella olivacea (Morse) (Gurney, 1940) 1961) Orphulella olivacea halophila Rehn and Hebard Pamtettix toltecus (Saussure) (Rehn and Grant, (Gurney, 1940) 1961) Orphulella pelidna (Burmeister) (Otte, 1981) Tetrix arenosa angusta Hancock (Rehn and Grant, Orphulella speciosa (Scudder) (Otte, 1981) 1961) Pampomala pallida Bruner (Otte, 1981) Tetrix arenosa arenosa Burmeister (Rehn and Pampomala virgata Scudder (Otte, 1981) Grant, 1961) Pampomala wyomingensis (Thomas) (Otte, 1981) Tettigidea armata Morse (Rehn and Grant, 1961) Phlibostroma quadrimaculatum (Thomas) (Otte, Tettigidea latemlis cazieri Rehn and Grant (Rehn 1981) and Grant, 1961) Pseudopomala bmchyptem (Scudder) (Otte, 1981) Tettigidea latemlis latemlis (Say) (Rehn and Grant, Psoloessa delicatula delicatula (Scudder) (Rehn, 1961) 1942) Psoloessa texana pawnee Rehn (Rehn, 1942) Family Psoloessa texana pusilla (Scudder) (Rehn, 1942) Psoloessa texana texana Scudder (Rehn, 1942) Subfamily Rhammatocerus viatorius (Saussure) (Tinkham, 1948) Achurum hilliardi Gurney (Helfer, 1987) Syrbula admimbilis (Uhler) (Otte, 1981) AchurUln Ininimipenne Caudell (Otte, 1981) Syrbula montezuma (Saussure) (Otte, 1981) Achurum sumichmsti (Saussure) (Otte, 1981) hirtipes hirtipes (Say) (Otte, 1981) Subfamily variegatus Bruner (Helfer, 1987) Metaleptea brevicornis (Johannson) (Otte, 1981) Acrolophitus maculipennis (Scudder) (Otte, 1981) Ageneotettix brevipennis (Bruner) (Otte, 1981) Subfamily Oedipodinae Ageneotettix deorum (Scudder) (Otte, 1981) Amphitornis colomdus (Thomas) (Otte, 1981) AncGrl-ia hebardi Rehn (Otte, 1984) Aulocam elliotti (Thomas) (Otte, 1981) Arphia conspersa Scudder (Otte, 1984) Aulocam femomtum (Scudder) (Otte, 1981) Arphia pecos Otte (Otte, 1984) Boopedon auriventris McNeill (Otte, 1981) Arphia pseudonietana (Thomas) (Otte, 1984) Boopedon gmcile Rehn (Otte, 1981) Arphia simplex Scudder (Otte, 1984) Boopedon nubilum (Say) (Otte, 1981) Arphia sulphurea (Fabricius) (Otte, 1984) argentatus Bruner (Otte, 1981) Arphia xanthoptem (Burmeister) (Otte, 1984) Cibolacris parviceps (Walker) (Otte, 1981) Chortophaga austmlior Rehn and Hebard (Otte, Cibolacris samalayucae Tinkham (Otte, 1981) 1984) crenulata (Bruner) (Otte, 1981) Chortophaga viridifasciata (DeGeer) (Otte, 1984) Cordillacris occipitalis (Thomas) (Otte, 1981) Circotettix mbula Rehn and Hebard (Otte, 1984) Dichromorpha elegans (Morse) (Otte, 1981) Conozoa texana (Bruner) (Otte, 1984) Dichromorpha viridis (Scudder) (Otte, 1981) Derotmena haydenii (Thomas) (Otte, 1984) Eritettix abortivus (Bruner) (Otte, 1981) Derotmena laticinctum Scudder (Otte, 1984) (Scudder) (Otte, 1981) Dissosteim carolina (Linnaeus) (Otte, 1984) Heliaula rufa (Scudder) (Otte, 1981) Dissosteim longipennis (Thomas) (Otte, 1984) Ligurotettix planum (Bruner) (Otte, 1981) 38 Volume 15, No.1, March, 2001, INSECTA MUNDI

Encoptolophus sordidus costalis (Scudder) (Knut­ Subfamily son, 1940) Encoptolophus subgracilis texensis Bruner (Knut- Aeoloplides elegans (Scudder) (Tinkham, 1948) son, 1940) Aeoloplides turnbulli bruneri (Caudell) (Wallace, Eximacris superbUln (Hebard) (Otte, 1984) 1955) Hadrotettix magnifica (Rehn) (Otte, 1984) Agroecotettix nwdestus aristus Hebard (Hebard, Hadrotettix trifasciatus (Say) (Otte, 1984) 1943a) Heliastus benjamini Caudell (Otte, 1984) Agroecotettix modestus crypsidomus Hebard (He­ Heliastus subroseus Caudell (Otte, 1984) bard, 1943a) Heliastus sumichrasti (Saussure) (Otte, 1984) Campylacantha lamprotata Rehn and Hebard (Rehn Hippiscus ocelote (Saussure) (Otte, 1984) and Hebard, 1909) Hippopedon capito (StaI) (Otte, 1984) Campylacantha olivacea olivacea (Scudder) (Knut­ Leprus intermedius Saussure (Otte, 1984) son, 1940) Leprus wheeleri (Thomas) (Otte, 1984) Campylacantha olivacea vivax (Scudder) (Helfer, Mestobregnw plattei corrugata (Scudder) (Otte, 1987) 1984) Chloropus cactocaetes Hebard (Helfer, 1987) Mestobregma terricolor Rehn (Otte, 1984) Clematodes larrae Scudder (Tinkham, 1948) Metator pardalinus (Saussure) (Otte, 1984) DactylotUln bicolor pictUln (Thomas) (Knutson, Pardalophora haldemanii (Scudder) (Otte, 1984) 1940) Pardalophora phoenicoptera (Burmeister) (Otte, Dactylotum bicolor variegatum (Scudder) (Helfer, 1984) 1987) Pardalophora saussurei (Scudder) (Otte, 1984) Dendrotettix quercus Packard (Helfer, 1987) Platylactista aztecus (Saussure) (Tinkham, 1948) Hesperotettix speciosus (Scudder) (Knutson, 1940) Psinidia amplicornis Caudell (Otte, 1984) Hesperotettix viridis pratensis Scudder (Knutson, fenestralis (Serville) (Otte, 1984) 1940) Psinidia fenestral is frater Rehn (Rehn, 1919) Hesperotettix viridis viridis (Thomas) (Knutson, Spharagemon bolli Scudder (Otte, 1984) 1940) Spharagemon bolli inornatum Morse (Tinkham, Hypochlora alba (Dodge) (Knutson, 1940) 1948) alexanderi Hilliard (Hilliard, 2001) Spharagemon campestris (McNeill) (Otte, 1984) Melanoplus angularis Little (Knutson, 1940) Spharagenwn collare (Scudder) (Otte, 1984) Melanoplus angustipennis angustipennis (Dodge) Spharagemon cristatum (Scudder) (Otte, 1984) (Helfer, 1987) Spharagenwn equale (Say) (Otte, 1984) Melanoplus angustipennis impiger Scudder (Knut- Spharagemon mannorata picta (Scudder) (Otte, son, 1940) 1984) Melanoplus aridus (Scudder) (Spooner, 1989) Trachyrhachis aspera Scudder (Otte, 1984) Melanoplus ariz07we Scudder (Helfer, 1987) Trachyrhachis coronata Scudder (Otte, 1984) Melanoplus bispinosus Scudder (Knutson, 1940) Trachyrhachis kiowa fuscifrons (StaI) (Otte, 1984) Melanoplus bivittatus (Say) (Knutson, 1940) Trachyrhachis kiowa kiowa (Thomas) (Otte, 1984) Melanoplus bowditchi Scudder (Rehn and Hebard, Trimerotropis citrina Scudder (Otte, 1984) 1909) Trimerotropis cyaneipennis Bruner (Otte, 1984) Melanoplus confusus Scudder (Knutson, 1940) Trimerotropis latifasciata Scudder (Otte, 1984) Melanoplus dakini Hilliard (Hilliard, 2001) Trinwrotropis melanoptera McNeill (Otte, 1984) Melanoplus desultorius Rehn (Spooner, 1989) Trimerotropis pallidipennis (Burmeister) (Otte, Melanoplus differentialis nigricans Cockerell (Rob- 1984) erts, 1942) Trimerotropis pistrinaria Saussure (Otte, 1984) Melanoplus discolor (Scudder) (Knutson, 1940) Trimerotropis salina McNeill (Otte, 1984) Melanoplus eumera Hebard (Tinkham, 1948) Trimerotropis saxatilis McNeill (Isely, 1935) Melanoplus femur-rubrum (DeGeer) (Knutson, Trimerotropis schaefferi Caudell (Otte, 1984) 1940) Trimerotropis strenua McNeill (Otte, 1984) Melanoplus flabellatus Scudder (Knutson, 1940) Tropidolophus fonnosus (Say) (Otte, 1984) Melanoplus flavidus Scudder (Knutson, 1940) Xanthippus corallipes pantherinus (Scudder) Melanoplus foedus fluviatilis Brunner (Knutson, (Helfer, 1987) 1940) INSECTA MUNDI, Vol. 15, No.1, March, 2001 39

Melanoplus foedus foedus Scudder (Hebard, 1936) Schistocerca albolineata (Thomas) (Hubbell, 1960) Melanoplus foedus iselyi Hebard. (Knutson, 1940) Schistocerca alutacea (Harris) (Hubbell, 1960) Melanoplus gladstoni Scudder (Helfer, 1987) Schistocerca americana (Drury) (Knutson, 1940) Melanoplus glaucipes (Scudder) (Knutson, 1940) Schistocerca dwnnifica (Saussure) (Knutson, 1940) Melanoplus herbaceous Bruner (Rehn and Hebard, Schistocerca emarginata (Scudder) (Knutson, 1940) 1909) Schistocerca nitens (Thunberg) (Helfer, 1987) Melanoplus inconspicnus Caudell (Hebard, 1917a) Schistocerca obscnra (Fabricius) (Helfer, 1987) Melanoplus lleeleri (Thomas) (Knutson, 1940) Schistocerca rubiginosa (Harris) (Helfer, 1987) Melanoplus lahinus (Scudder) (Knutson, 1940) Schistocerca shoshone (Thomas) (Hubbell, 1960) Melanoplus mastigiphallus Strohecker (Stroheck- er, 1941) Subfamily Leptysminae Melanoplus mexican us (Saussure) (Helfer, 1987) Melanoplus occidentalis (Thomas) (Helfer, 1987) Leptysma marginicollis (Serville) (Knutson, 1940) Melanoplus pachardii Scudder (Knutson, 1940) vitreipennis (Marschall) (Hilliard, 1982) Melanoplus plebejus (Stal) (Helfer, 1987) Melanoplus ponderosus (Scudder) (Knutson, 1940) Subfamily Melanoplus ponderosus viola (Thomas) (Hebard, 1941) Brachystola magna (Girard) (Rehn and Grant, 1961) Melanoplus punctulatus arboreus Scudder (Rehn, Phrynotettix robustus occultus Rehn and Grant 1946) (Rehn and Grant, 1961) Melanoplus regal is (Dodge) (Helfer, 1987) Phrynotettix robustus robustus (Bruner) (Rehn and Melanoplus msticns obovatipennis Blatcheley (Cop­ Grant, 1961) pock, 1962) Phrynotettix tschivavensis tschivavensis (Halde­ Melanoplus sanguinipes defectus Scudder (Helfer, man) (Rehn and Grant, 1961) 1987) Phrynotettix tschivavensis varuculatus Glover Melanoplus sanguinipes vulturnis Gurney and (Rehn and Grant, 1961) Brooks (Gurney and Brooks, 1959) Romalea guttatus (Stoll) (Rehn and Grant, 1961) Melanoplus scudderi latus Morse (Knutson, 1940) Taeniopoda equues (Burmeister) (Rehn and Grant, Melanoplus scudderi texensis Hart (Knutson, 1940) 1961) Melanoplus tenninalis Scudder (Scudder, 1897) Melanoplus texanus (Scudder) (Knutson, 1940) Suborder Ensifera Melanoplus thomasi Scudder (Helfer, 1987) Melanoplus tubercnlatus Morse (Helfer, 1987) Family Gryllacrididae Melanoplus wameri Little (Knutson, 1940) Netrosonw nigropleura Scudder (Helfer, 1987) Paraidenwna fratercnla Hebard (Hebard, 1918) Subfamily Rhaphidophorinae Paraidemona latifurcnla Hebard (Hebard, 1918) Paraidenwna mimica (Scudder) (Hebard, 1918) Ammobaenetes phrixocnemoides (Caudell) (Helfer, Paraidemona nuttingi Yin and Smith (Yin and 1987) Smith, 1989) Ceuthophilus carlsbadensis Caudell (Hubbell, 1936) Paraidemona olsoni Yin and Smith (Yin and Smith, Ceuthophilus conicaudus Hubbell (Hubbell, 1936) 1989) Ceuthophilus cunicularis Hubbell (Hubbell, 1936) Paraidemona punctata (Stal) (Hebard, 1918) Ceuthophilus divergens Scudder (Hubbell, 1936) Paratylotropidea bmnneri Scudder (Knutson, 1940) Ceuthophilus isletae Hubbell (Hubbell, 1936) atlantica Scudder (Knutson, 1940) Ceuthophilus nodulosus Brunner (Hubbell, 1936) Paroxya clavuliger (Serville) (Helfer, 1987) Ceuthophilus pallidus Thomas (Hubbell, 1936) Phaedrotettix accola (Scudder) (Hebard, 1917b) Ceuthophilus polingi Hubbell (Hubbell, 1936) Phaedrotettix dwnicola (Scudder) (Hebard, 1917b) Ceuthophilus rehni Hubbell (Hubbell, 1936) Phaedrotettix palmeri (Scudder) (Hebard, 1917b) Ceuthophilus secret us Scudder (Hubbell, 1936) Phaulotettix compressus Scudder (Helfer, 1987) Ceuthophilus wnbratilis Hubbell (Hubbell, 1936) Phaulotettix eurycercns Hebard (Helfer, 1987) Ceuthophilus utahensis Thomas (Hubbell, 1936) Philoeleon nigrovittata (Stal) (Tinkham, 1948) Ceuthophilus variegatus Scudder (Hubbell, 1936) Photaliotes nebrascensis (Thomas) (Knutson, 1940) Daihinia brevipes Haldeman (Helfer, 1987) 40 Volume 15, No.1, March, 2001, INSECTA MUNDI

Daihiniella bellicosus (Hubbell) (Hubbell, 1936) Subfamily Pseudophyllinae Daihiniodes hausteriferum (Rehn) (Hubbell, 1936) Udeopsylla robusta (Haldeman) (Helfer, 1987) Paracyrtophyllus excels us (Rehn and Hebard) (He­ bard, 1941) Subfamily Stenopelmatinae Paracyrtophyllus robustus Caudell (Hebard, 1941) Pterophylla camellifolia camellifolia (Fabricius) Stenopelllwtus fuscus Haldeman (Hebard, 1916a) (Hebard, 1941) Pterophylta camellifolia intennedia (Caudell) (He­ Family bard, 1941) Pterophylla furcata furcata (Ca udell) (Hebard, 1941) Subfamily Pterophylla furcata laletica Hebard (Hebard, 1941)

Bucrates malivolens (Scudder) (Helfer, 1987) Subfamily Conocephalus brevipennis (Scudder) (Rehn and Hebard, 1915a) Amblycorypha floridana Rehn and Hebard (Rehn Conocephalus fasciatus (DeGeer) (Rehn and He­ and Hebard, 1914c) bard, 1915a) Alllblycorypha hausteca (Saussure) (Rehn and He­ Conocephalus hygrophilus Rehn and Hebard (Rehn bard, 1914c) and Hebard, 1915a) Amblycorypha insolita Rehn and Hebard (Rehn Conocephalus resacensis Rehn and Hebard (Rehn and Hebard, 1914c) and Hebard, 1915a) Amblycorypha oblongifolia (DeGeer) (Rehn and Conocephalus saltans (Scudder) (Rehn and He­ Hebard, 1914c) bard, 1915a) Amblycorypha rotundifolia parvipennis StaI (Rehn Conocephalus spartinae (H. Fox) (Rehn and He­ and Hebard, 1914c) bard, 1915a) AmblycoryphauhleriStaI(RehnandHebard,1914c) Conocephalus strictus (Scudder) (Rehn and He­ Arethaea ambulator Hebard (Rehn and Hebard, bard, 1915a) 1914a) exiliscanorus (Davis) (Helfer, Arethaea arachnopyga Rehn and Hebard. (Rehn 1987) and Hebard, 1914a) Neoconocephalus nebrascensis (Brunner) (Walker, Arethaea constricta comanche Hebard (Hebard, 1998) 1936b) Neoconocephalus robustus crepitans (Scudder) Arethaea constricta constricta Brunner (Hebard, (Tinkham, 1944) 1936b) Neoconocephalus triops (Linnaeus) (Helfer, 1987) Arethaea gracilipes cerciata Hebard. (Rehn and Neoconocephalus velox Rehn and Hebard (Walker, Hebard, 1914a) 1998) Arethaea gracilipes gracilipes (Thomas) (Rehn and agile (DeGeer) (Morris and Walker, Hebard, 1914a) 1976) Arethaea grallator (Scudder) (Rehn and Hebard, Orchelillwm bullatum Rehn and Hebard (Rehn 1914a) and Hebard, 1915b) Arethaea phantasm a Rehn and Hebard (Rehn and Orchelimum calcaratum Rehn and Hebard (Rehn Hebard, 1914a) and Hebard, 1915b) Arethaea semialata Rehn and Hebard (Rehn and Orchelimum concinnum concinnum Scudder (Rehn Hebard, 1914a) and Hebard, 1915b) Dichopetala brevihastata Morse (Rehn and He­ Orchelimum concinnum delicatulum Bruner (Tho­ bard, 1914b) mas and Alexander, 1962) Dichopetala castanea Rehn and Hebard (Rehn and Orchelimum nigripes Scudder (Rehn and Hebard, Hebard, 1914b) 1915b) Dichopetala catinata Rehn and Hebard (Rehn and Orchelimum vulgare Harris (Rehn and Hebard, Hebard, 1914b) 1915b) Dichopetala emarginata Brunner (Rehn and He­ Pyrgocorypha uncuinata (Harris) (Helfer, 1987) bard, 1914b) INSECTA MUNDI, Vol. 15, No.1, March, 2001 41

Dichopetalagladiator Rehn and Hebard (Rehn and Family Tridactylidae Hebard, 1914b) Dichopetala oreoeca Rehn and Hebard (Rehn and Tridactylus apicalis Say (Helfer, 1987) Hebard, 1914b) Tridactylus minutis Scudder (Helfer, 1987) Dichopetala pollicifera Rehn and Hebard (Rehn and Hebard, 1914b) Family Myrmecophilidae Dichopetala seeuersi Strohecker (Strohecker, 1941) Insara elegans (Scudder) (Tinkham, 1944) Mynnecophila nebrascensis Lugger (Helfer, 1987) Microcentrum latifrons Spooner (Spooner, 1989) Microcentrum minus Strohecker (Strohecker, 1952) Family Microcentrum rhombifolium (Saussure) (Helfer, 1987) Montezumina modesta (Brunner) (Walker, 1998) Acheta domesticus (Linnaeus) (Blatchley, 1920) Scudderia curuicauda-laticauda Brunner (Rehn Allonemobius fasciatus (DeGeer) (Hebard, 1913) and Hebard, 1914) Allonemobius fasciatus socius (Scudder) (Hebard, Scudderia furcata Brunner (Rehn and Hebard, 1913) 1914) Allonemobius grise us funeral is (Hart) (Hebard, Scudderia furcata furcifera (Scudder) (Rehn and 1913) Hebard, 1914) Allonemobius sparsalus (Fulton) (Walker, 1998) Scudderia mexicana (Saussure) (Rehn and He­ Anaxipha exigua (Say) (Helfer, 1987) bard, 1914) Anurogryllus muticus (DeGeer) (Helfer, 1987) Scudderia texensis Saussure and Pictet (Rehn and Cycloptiliwn antillarum (Redtenbacher) (Hebard, Hebard, 1914) 1916b) Cycloptilium comprehendens copmprehendens He­ Subfamily Decticinae bard (Hebard, 1916b) Cycloptilium comprehendens fortior Hebard (He­ bard, 1916b) Anabrus simplex Haldeman (Caudell, 1907) Cycloptilium kelainopum Love and Walker (Love Capnobotes fuliginosus (Thomas) (Tinkham, 1944) and Walker, 1979) Eremopedes couilleae Hebard (Tinkham, 1948) Cycloptilium pigrum Love and Walker (Love and Eremopedes scudderi Cockerell (Tinkham, 1948) Walker, 1979) Neobarrettia spinosus (Caudell) (Helfer, 1987) Cycloptilium slossoni Scudder (Love and Walker, Neobarrettia uictoriae (Caudell) (Helfer, 1987) 1979) Pediodectes daedalus (Rehn and Hebard) (Rehn Cycloptilium squamosum Scudder (Love and Walk­ and Hebard, 1920) er' 1979) Pediodectes grand is (Rehn) (Tinkham, 1944) Cycloptilium trigonipalpum Rehn and Hebard (Love Pediodectes haldemanii (Girard) (Tinkham, 1944) and Walker, 1979) Pediodectes nigromarginata (Caudell) (Rentz and Cyrtoxipha columbiana Caudell (Walker, 1969) Birchim, 1968) carolinus (Scudder) (Hebard, 1913) Pediodectes pratti (Caudell) (Rentz and Birchim, Eunemobius carolinus neomexicanus (Scudder) 1968) (Hebard, 1913) Pediodectes steuensoni (Thomas) (Tinkham, 1944) Falcicula hebardi Rehn (Blatchley, 1920) Pediodectes tinkhami Hebard (Rentz and Birchim, Gryllodes sigillatus Walker (Alexander and Walker, 1968) 1962) firma Scudder (Alexander, 1968) Family Gryllotalpidae Gryllus fultoni Alexander (Alexander, 1968) Gryllus pennsyluanicus Burmeister (Alexander, Gryllotalpa cultriger Uhler (Frank et aI., 1995) 1968) Neocurtilla hexadactyla (Perty) (Helfer, 1987) Grylluspersonatus Uhler (Rehn and Hebard, 1915c) Scapteriscus borelli Giglio-Tos (Frank et aI., 1995) Gryllus rubens (Scudder) (Walker, 1998) Scapteriscus uicinus Scudder (Frank et aI., 1995) Gryllus texensis Cade and Otte (Cade and Otte, 2000) Hapithus agitator Uhler (Helfer, 1987) 42 Volume 15, No.1, March, 2001, INSECTA MUNDI

Hoplosphyrwn boreale (Scudder) (Helfer, 1987) Flook, P.K. and C.H.F. Rowell. 1997. The phy­ Miogryllus lineatus (Scudder) (Hebard, 1915) logeny of the Caelifera (Insecta, Orthoptera) as Miogryllus uerticalis (Serville) (Hebard, 1915) deduced from mtrRNA gene sequences. Mol. Neonel1wbius cubensis monnonius (Scudder) (He- Phylog. Evol. 8: 89-103. bard, 1913) Frank, J.H., T.R. Fasulo, and D.E. Short. 1995. Neoxabea bipunctata (DeGeer) (Walker, 1963) McricketKnowledgebasehttp://gnv.lfas.ufl.edu/ Neoxabea formosa (F. Walker) (Walker, 1967) -entIlmcricket/. Oecanthus argentinus Saussure (Walker, 1962) Gurney, A.B. 1940. A revision of the grasshoppers Oecanthus califomicus Saussure (Walker, 1963) ofthe genus Orphulella from America North of Oecanthus celerinictus T.J. Walker (Walker, 1962) Mexico. Entomologica Am .. 20: 85-157. Oecanthus exclamationis Davis (Walker, 1963) Gurney, A.B. and A.R. Brooks. 1959. Grasshop­ Oecanthus fultoni T.J. Walker (Walker, 1963) pers of the Mexicanus group, genus Melano­ Oecanthus leptogramnws T.J. Walker (Walker, plus (Orthoptera: Acrididae). Proc. U.s. Natl. 1963) Mus. 110: 1-93. Oecanthus niueus (DeGeer) (Walker, 1963) Hebard, M. 1913. A revision of the species of the Oecanthus pini Beutenmuller (Walker, 1962) genus Nemobius (Orthoptera: Gryllidae) found Oecanthus quadripunctatus Beutenmuller (Walk- in North America North of the Isthmus of er, 1962) Panama. Proc. Acad. Nat. Sci. Phila. 65: 394- Orocharis saltator Uhler (Walker, 1969) 492. Phyllopalpus pulchellus (Uhler) (Walker, 1998) Hebard, M. 1915. The American species of the Trigonidomimus belfragei Caudell (Caudell, 1912) genus Miogryllus. J. New York Entomol. Soc. 23: 101-12l. References Hebard, M. 1916a. A study of the species of the genus Stenopelmatus found in the United States. Alexander, R.D. 1968. Life cycle origins, specia­ J. New York Entomol. Soc. 24: 70-86. tion, and related phenomena in crickets. Quart. Hebard, M. 1916b. The Mogoplistinae (Ortho­ Rev. BioI. 43: 1-4l. ptera, Gryllidae) of the United States. Trans. Alexander, R.D. and T.J. Walker. 1962. Two Am. Entomol. Soc. 42: 135-160. introd uced field crickets new to Eastern United Hebard, M. 1917a. New genera and species of the States (Orthoptera: Gryllidae). Ann. Entomol. Melanopli found within the United States and Soc. Am. 55: 90-94. Canada (Orthoptera: Acrididae). Parts X to Atkinson, T.A., P.G. Koehler, and R.S. Patter­ XIV. Trans. Am. Entomol. Soc. 43:147-163. son. 1991. Catalog and atlas of the Hebard, M. 1917b. Notes on Mexican Melanopli (Dictyoptera) of North America North of Mexi­ (Orthoptera: Acrididae). Proc. Acad. Nat. Sci. co. Misc. Publ. Entomol. Soc. ArneI'. 78: 85 pp. Phila. 69: 251-275. Blatchley, W.S. 1920. Orthoptera of Northeastern Hebard, M. 1918. New genera and species of America. Nature Publ. Co., Indianapolis, Indi­ Melanopli found within the United States (01'­ ana. thoptera: Acrididae). Trans. Am. Entomol. Soc. Cade, W.H. and D. Otte. 2000. Gryllus texensis n. 44: 141-169. sp.: a widely studied field (Orthoptera; Hebard, M. 1936a. New genera and species of the Gryllidae) from the Southern United States. Melanopli found within the U.S. and Canada, Trans. ArneI'. Entomol. Soc. 126: 117-123. Part VIII. Trans. Am. Entomol. Soc. 62: 182- Caudell, AX 1907. The Decticinae (a group of 190. Orthoptera) of North America. Proc. U.S. Natl. Hebard, M. 1936b. VI. A revision of the genus Mus. 32: 285-410. Arethaea (Tettigoniidae, Phaneropterinae). Caudell, A.N. 1912. A new genus and species of Trans. Am. Entomol. Soc. 62: 231-256. Gryllidae from Texas. Proc. Entomol. Soc. Wash. Hebard, M. 1941. The group Pterophyllae as found 14: 187-189. in the United States (Tettigoniidae: Pseudo­ Coppock, S., Jr. 1962. The grasshoppers of Okla­ phyllinae). Trans. Am. Entomol. Soc. 67: 197- homa (Orthoptera: Acrididae) Oklahoma State 219. Univ. Agric. Exp. Sta. Report P-399: 143 pp. Hebard, M. 1943a. New genera and species of Melanoplifound within the U.s., PartlV. Trans. Am. Entomol. Soc. 68: 49-66. INSECTA MUNDI, Vol. 15, No.1, March, 2001 43

Hebard, M. 1943b. The Dermaptera and Ortho­ Rehn, J.A.G. 1942. On the locust genus Psoloessa pterous families Blattidae, Mantidae, and Phas­ (Orthoptera: Acrididae: Acridinae). Trans. Am. midae of Texas. Trans. Am. Entomol. Soc. 48: Entomol. Soc. 68: 167-237. 239-313. Rehn, J.A.G. 1946. On the Punctulatus species­ Helfer, J.R. 1987. How to know the grasshoppers, group of the genus Melanoplus (Orthoptera, crickets, cockroaches and their allies. Dover Acrididae, Cyrtacanthacridinae). Proc. Acad. Publications, Inc., New York. Nat. Sci. 98: 241-269. Hilliard, J.R., Jr. 1982. Endophytic oviposition by Rehn, J.A.G. and H.J. Grant Jr. 1961. A mono­ LeptysJrw marginicollis marginicollis and Ste­ graph of the Orthoptera of North America (North nacris vitreipennis (Orthoptera: Acrididae: Lep­ of Mexico) , Vol. 1. Monographs of the Acad. Nat. tysminae) with life history notes. Trans. Am. Sci. Phila., 12: 257 pp. Entomol. Soc. 108: 153-180. Rehn, J.A.G. and M. Hebard. 1909. An ortho­ Hilliard, J.R., Jr. 2001. Two new pterological reconnaisance of the Southwest­ species in the Texanus group of the genus ern United States. Part II: New Mexico and Melanoplus (Orthoptera: Acrididae: Melano­ Western Texas. Proc. Acad. Nat. Sci. Phila. 61: plinae) with biological notes on the group. Trans. 111-175. Am. Entomol. Soc. 127: 31-68. Rehn, J.x.G. and M. Hebard. 1914a. A revision of Hubbell, T.H. 1936. A monographic revision ofthe the orthopterous group Insarae (Tettigoniidae, genus Ceuthophilus (Orthoptera, Gryllacrid­ Phaneropterinae). Trans. Am. Entomol. Soc. idae, Rhaphidophorinae). Univ. Fla. Pub. BioI. 40: 37-184. Sci. Series 2: 1-551. Rehn, J.A.G. and M. Hebard. 1914b. A study of Hubbell, T.H. 1960. The sibling species of the the species of the genus Dichopetala (Ortho­ Alutacea group of the bird-locust genus Schis­ ptera: Tettigoniidae). Proc. Acad. Nat. Sci. Phi­ tocerca (Orthoptera, Acrididae, Cyrtacan­ lao 66: 64-160. thacridinae). Misc. Publ. Mus. Zool., Univ. Mich. Rehn, J.A.G.and M. Hebard. 1914c. Studies in 116: 1-91. American Tettigoniidae (Orthoptera) I and II. Isely, F.B. 1935. Acridianresearches within North­ Trans. Am. Entomol. Soc. 40: 271-344. eastern Texas (Orthoptera). Entomol. News 46: Rehn, J.A.G. and M. Hebard. 1915a. A synopsis 37-75. of the species of the genus Conocephalus Knutson, H. 1940. A key to the Acrididae (Ortho­ (Xiphidiwn of authors) found in North America ptera) of Northeastern Texas with ecological North of Mexico. Trans. Am. Entomol. Soc. 41: notes. Field and Laboratory 8: 33-58. 155-224. Love, R.E. and T.J. Walker. 1979. Systematics Rehn, J.A.G. and M. Hebard. 1915b. A synopsis and acoustic behavior of scaly crickets (Ortho­ of the species of the genus Orchelintwn. Trans. ptera: Gryllidae: Mogoplistinae) of Eastern Am. Entomol. Soc. 41: 11-83. United States. Trans. Am. Entomol. Soc. 105: 1- Rehn, J.A.G. and M. Hebard. 1915c. The genus 66. Gryllus as found in America. Proc. Acad. Nat. Morris, G.K. and T.J. Walker. 1976. Calling Sci. Phila. 77: 293-322. songs of Orchelimwn meadow katydids (Tet­ Rehn, J.A.G. and M. Hebard. 1920. Description tigoniidae), 1. mechanism, terminology, and of new genera and species of North American geographic distribution. Can. Entomol. 108: Decticinae (Orthoptera: Tettigoniidae). Trans. 785-800. Am. Entomol. Soc. 46: 225-265. Otte, D. 1981. The North American grasshoppers, Rentz, D.C. and J.D. Birchim. 1968. Revisionary Volume I Acrididae, Gom p hocerinae, and Acrid­ studies in the Nearctic Decticinae. Mem. Pacif­ inae. Harvard Univ. Press, Cambridge, Massa­ ic Coast Entomol. Soc. 3: 1-173. chusetts. Roberts, H.R. 1942. Two subspecies of Melano­ Otte, D. 1984. The North American grasshoppers, plus differentialis and related new subspecies Volume II Oedipodinae. Harvard Univ. Press, from Mexico with discussion of their variation. Cambridge, Massachusetts. Trans. Amer. Entomol. Soc. 68: 151-166. Rehn, J.A.G. 1919. Descriptions of new and criti­ Scudder, S.H. 1897. Revision of the orthopteran cal notes upon previously known forms of North group Melanopli (Acrididae), with special refer­ American Oedipodinae (Orthoptera: Acrididae). ence to North American forms. Proc. U.S. Natl. Trans. Am. Entomol. Soc. 45: 229-255. Mus. 20: 293-294. 44 Volume 15, No.1, March, 2001, INSECTA MUNDI

Spooner, J.D. 1989. Microcentrum latifrons, a lidae: Oecanthinae). 1. The Genus Neoxabea new species of angular-winged katydid from and the Niueus and Varicornis Groups of the Southwestern United States (Orthoptera, Tet­ Genus Oecanthus. Ann. Entomol. Soc. Am. 55: tigoniidae, Phaneropterinae). Trans. Am. En­ 303-322. tomol. Soc. 114: 311-317. Walker, T.J.1963. The taxonomy and calling songs Strohecker, H.F. 1941. Two new species of Or tho­ of the United States tree crickets (Orthoptera: ptera from Texas. Ann. Entomol. Soc. Am. 34: Gryllidae: Oecanthinae). II. The Nigricornis 539-542. Group of the Genus Oecanthus. Ann. Entomol. Strohecker, H.F. 1952. Descriptions of new spe­ Soc. Am. 56: 772-789. cies and notes on North American Orthoptera. Walker, T.J. 1967. Revision of the Oecanthinae Am. Midland Naturalist 48: 683-688. (Gryllidae: Orthoptera) of America South of the Strohecker, H.F., W.W. Middlekauff, and D.C. United States. Ann. Entomol. Soc. Am. 60: 784- Rentz. 1968. The grasshoppers of California 796. (Orthoptera: ). Bull. California In­ Walker, T.J. 1969. Systematics and acoustic be­ sect Survey 10: 177pp. havior of United States crickets of the genus Thomas, E.S. and R.D. Alexander. 1962. Sys­ Orocharis (Orthoptera: Gryllidae). Ann. Ento­ tematic and behavioral studies on the meadow mol. Soc. Am. 62: 752-762. grasshoppers of the OrchelimUln concinnum Walker, T.J. 1969. Systematics and acoustic be­ group (Orthoptera: Tettigoniidae). Occ. Papers havior of United States crickets of the genus Mus. Zool., Univ. Mich. 626: 1-30. Cyrtoxipha (Orthoptera: Gryllidae). Ann. En­ Tinkham, E.R. 1944. Biological, taxonomic and tomol. Soc. Am. 62: 945-952. faunistic studies on the shield-back katydids of Walker, T.J. 1998. Handbook of katydids and the North American deserts. Am. Midland Nat­ crickets of America North of Mexico. http:// uralist 31: 257-328. csssrvr .entnem. ufl.ed u/ -walker/handbook/ Tinkham, E.R. 1948. Faunistic and ecological hdbkhmpg.htm. studies on the Orthoptera of the Big Bend Wallace, H.S. 1955. Revision of the genus Aeol­ Region of Trans-Pecos Texas, with especial oplides (Orthoptera, Acrididae). Ann. Entomol. reference to the orthopteran zones and faunae Soc. Am. 48: 453-480. of Midwestern North America. Am. Midland Yin, X. and R.L. Smith. 1989. Three new grass­ Naturalist 40: 521-663. hoppers from the Western United States (01'­ Walker, T.J.1962. The taxonomy and calling songs thoptera: Acrididae). Pan-Pacific Entomol. 65: ofU nited States tree crickets (Orthoptera: Gryl- 166-171.