Monographs of the Western North American Naturalist

Volume 1 Article 3

11-25-2002 Annotated checklist of the millipeds of California (Arthropoda: Diplopoda) Rowland M. Shelley [email protected]

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Recommended Citation Shelley, Rowland M. (2002) "Annotated checklist of the millipeds of California (Arthropoda: Diplopoda)," Monographs of the Western North American Naturalist: Vol. 1 , Article 3. Available at: https://scholarsarchive.byu.edu/mwnan/vol1/iss1/3

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ANNOTATED CHECKLIST OF THE MILLIPEDS OF CALIFORNIA (ARTHROPODA: DIPLOPODA)

Rowland M. Shelley1

ABSTRACT.—The milliped fauna of California consists of 11 orders, 24 families, 83 genera, and 226 species and sub- species. A complete listing of these taxa and intergrades is provided, with published and new records from the state and type localities. Cylindroiulus caeruleocinctus (Wood), C. truncorum (Silvestri), Ophyiulus pilosus (Newport) (all Julida: Julidae), and Opiona fisheri Gardner and Shelley (Chordeumatida: Caseyidae) are newly recorded from California. The new combination Atopetholus wheeleri (Chamberlin) is proposed.

Key words: milliped, California.

Nearly 40 years have elapsed since Buckett any other state in the continental United States. (1964) published a listing of California Diplo- Four diplopods—Cylindroiulus caeruleocinctus poda. This work preceded 2 periods of activity (Wood), C. truncorum (Silvestri), Ophyiulus on the state’s fauna, the first from 1964 through pilosus (Newport), and Opiona fisheri Gardner 1975 by Buckett and M.R. Gardner, and the and Shelley—are newly recorded from Cali- second from 1989 through 1997 by myself. fornia. These species plus Brachyiulus lusi- Knowledge of California diplopods is thus sig- tanus Verhoeff are European julids that have nificantly advanced over the condition in 1964, been imported into urban environments; like- to the point that a new checklist is in order. wise, the Asian species Oxidus gracilis (C.L. Buckett (1964) documented a fauna of 9 orders, Koch) and the Australian/New Zealand dip- 23 families, 80 genera, and 178 species and lopod Akamptogonus novarae (Humbert and subspecies, 126 of which (70.1% of the total) DeSaussure) have been introduced into Cali- were known only from the type locality; the fornia cities and towns. Generic and/or specific present contribution documents a fauna of 11 names cannot be provided for certain repre- orders, 24 families, 83 genera, and 226 species sentatives of the Siphonophoridae (Siphono- and subspecies, 78 of which (34.5% of the phorida), Atopetholus (Spirobolida: Atopetholi- total) are known only from the type locality. dae), Caseya (Chordeumatida: Caseyidae), Rhis- While considerable work has been done on cosomides (Chordeumatida: Rhiscosomididae), the California fauna, much remains, and this and Striaria (Chordeumatida: Striariidae). For contribution is intended to stimulate future Paeromopus angusticeps (Wood) (Julida: Paero- research. mopodidae), and Motyxia sequoia (Chamber- With the publication of this checklist, Cali- lin), Xystocheir dissecta (Wood), and X. stolon- fornia becomes the 4th state with a modern ifera Shelley (all Polydesmida: Xystodesmi- milliped listing, the others being Michigan dae), I present accounts of intergrades, which (Snider 1991, an update of an old list by John- are prominent in sections of the state. I also son [1954]), North Carolina (Shelley 2000a), propose the new combination Atopetholus and Florida (Shelley 2001). An earlier publica- wheeleri (Chamberlin). The format for the tion on Ohio (Williams and Hefner 1928) is now accounts provides a citation of the type local- obsolete. In addition, the faunas of eastern ity (given as a state or country when not in and western Canada have been documented California), occurrence in California including (Shelley 1988, 1990), and there are now 2 con- published and new records, pertinent refer- tinental checklists, by Chamberlin and Hoffman ences, and miscellaneous remarks where appro- (1958) and Hoffman (1999), from which faunal priate. For new records, locality details are information can be gleaned for California or provided when there are 6 or fewer new county

1Research Laboratory, North Carolina State Museum of Natural Sciences, 4301 Reedy Creek Rd., Raleigh, NC 27607. E-mail: [email protected]

90 2002] CALIFORNIA MILLIPEDS 91 records; counties alone are presented when Alameda, Contra Costa, Humboldt, Los Angeles, there are more than 6 new counties. Details Marin, Mendocino, Napa, Nevada, Orange, about junior synonyms should be sought in Plumas, Riverside, San Diego, San Joaquin, San either Hoffman (1999) or the reference given Luis Obispo, San Mateo, Santa Clara, Shasta, for the relevant taxa. Acronyms of sources of Solano, Sonoma, Tehama, Trinity, and Ventura preserved material are as follows: cos. (CDFA, FSCA, LACMNH, NMNH, UCB, UCD, UCR, WAS). AMNH – American Museum of Natural History, REFERENCE.—Pierce 1940. New York, NY REMARKS.—For convenience I list all sam- BMNH – The Natural History Museum, London, ples of California Polyxenus that I have seen England under P. anacapensis. Some may refer to other BYU – Monte L. Bean Life Science Museum, species. Provo, UT CAS – California Academy of Sciences, San Subclass Chilognatha Francisco CDFA – California Department of Food and Infraclass Pentazonia Agriculture, Sacramento Order Glomerida FMNH – Field Museum of Natural History, Family Glomeridae Chicago, IL Genus Glomeroides Chamberlin, 1922 FSCA – Florida State Collection of Arthropods, Gainesville Glomeroides primus (Silvestri, 1929) LACMNH – Los Angeles County Museum of TYPE LOCALITY.—Mill Valley, Marin County. Natural History, Los Angeles, CA OCCURRENCE IN CALIFORNIA.—Published MCZ – Museum of Comparative Zoology, records: Marin and Monterey counties. New Harvard University, Cambridge, MA records: Contra Costa Co., Redwood Park NCSM – North Carolina State Museum of (FSCA, NMNH) and Morega Redwoods (CAS). Natural Sciences, Raleigh NMNH – National Museum of Natural History, Santa Cruz Co., Ben Lomond (FSCA) and Santa Smithsonian Institution, Washington, DC Cruz City Cave (FSCA). TMM – Texas Memorial Museum, Austin REFERENCE.—Shear 1986. UAZ – Department of Entomology, University of Arizona, Tucson Infraclass Helminthomorpha UCB – Entomology Department, University of Subterclass Colobognatha California, Berkeley Order Polyzoniida UCD – Entomology Department, University of Family Polyzoniidae California, Davis Subfamily Buzoniinae UCR – Entomology Department, University of Tribe Buzoniini California, Riverside Genus Buzonium Cook and UID – Entomology Department, University of Loomis, 1928 Idaho, Moscow WAS – Private collection of W.A. Shear, Buzonium crassipes Cook and Loomis, 1928 Hampden-Sydney, VA WSU – Entomology Department, Washington TYPE LOCALITY.—Sunnyside Mine near State University, Pullman Seneca, Plumas County. WTAMU – Department of Life, Earth, and OCCURRENCE IN CALIFORNIA.—Published Environmental Sciences, West Texas A&M records: Calaveras, Del Norte, El Dorado, University, Canyon, TX Humboldt, Mendocino, Nevada, Placer, Siski- you, Sonoma, and Trinity counties. New records: Subclass Penicillata None. Order Polyxenida REFERENCE.—Shelley 1997a. Family Polyxenidae Genus Polyxenus Latreille Tribe Bdellozoniini Genus Bdellozonium Cook and Polyxenus anacapensis Pierce, 1940 Loomis, 1928 TYPE LOCALITY.—Middle Anacapa Island, Santa Barbara County. Bdellozonium cerviculatum Cook and Loomis, 1928 OCCURRENCE IN CALIFORNIA.—Published record: Santa Barbara County. New records: TYPE LOCALITY.—Belden, Plumas County. 92 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

OCCURRENCE IN CALIFORNIA.—Published species should be regarded as valid until the records: Del Norte, El Dorado, Fresno, Hum- type specimen, in the Zoologisches Museum boldt, Mariposa, Monterey, Placer, Shasta, der Humboldt-Universität, Berlin, Germany, Tuolumne, and Tulare counties. New records: could be examined. He suggested that it might None. be a senior name to B. producta Loomis. REFERENCE.—Shelley 1997a. Brachycybe picta Gardner, 1975 Bdellozonium quicki (Chamberlin, 1954) TYPE LOCALITY.—1 mi (1.6 km) N Mendo- TYPE LOCALITY.—Chowchilla Mountain, 0.75 cino, Mendocino County. mi (1.2 km) N Signal Peak, Mariposa County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published records: Marin and Mendocino counties. New record: Mariposa County (known only from records. Marin Co., Sam P. Taylor State Park the type locality). (UCD), Mill Valley (BMNH), and Lagunetas REFERENCE.—Shelley 1997a. Canyon (FSCA). Mendocino Co., 5 mi (8 km) E Anchor Bay (AMNH). Genus Stenozonium Shelley, 1997 REFERENCE.—Gardner 1975. Stenozonium exile Shelley, 1997 Brachycybe producta Loomis, 1936

TYPE LOCALITY.—Arcata Community Forest TYPE LOCALITY.—Baja California Norte. near Arcata, Humboldt County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published records: Amador, Calaveras, El Dorado, Lake, record: Humboldt County. New records: None. Marin, Mariposa, Napa, Nevada, Placer, Sacra- REFERENCE.—Shelley 1997a. mento, Shasta, Solano, Sonoma, and Tuolumne counties. New records: Butte Co., Forest Ranch Family Hirudisomatidae (NCSM) and 2.5 mi (4.0 km) SW Cherokee Genus Octoglena Wood, 1864 (WSU). REFERENCES.—Gardner 1975, Chamberlin Wood, 1864 Octoglena bivirgata and Hoffman 1958. TYPE LOCALITY.—2 mi (3.2 km) N Ft. Ross, Murray, 1877 Sonoma County (neotype locality). Brachycybe rosea OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—3 mi (4.8 km) SW Placer- records: Colusa, Del Norte, Humboldt, Lake, ville, El Dorado County (neotype locality). Marin, Mendocino, Napa, San Mateo, Santa OCCURRENCE IN CALIFORNIA.—Published Clara, Santa Cruz, and Sonoma counties. New records: El Dorado, Nevada, Plumas, and records: None. Shasta counties. New records: Butte Co., 4 mi REFERENCE.—Shelley 1995a. (6.4 km) SW Stirling City (LACMNH), Big Bear Mountain near Pulga (NMNH), and Shelley, 1995 Octoglena sierra Forest Ranch (NCSM). Lassen Co., Monton TYPE LOCALITY.—4 mi (6.4 km) W Newcas- (FSCA). Mendocino Co., 4.1 mi (6.6 km) SW tle, Placer County. Leggett (AMNH). Placer Co., 5 mi (8 km) N OCCURRENCE IN CALIFORNIA.—Published Placerville (NCSM). Sierra Co., Sierra City records: Placer County. New records: None. (AMNH). Tehama Co., Deer Creek (NMNH), REFERENCE.—Shelley 1995a. S of Lassen Peak (NMNH), and Dales, 10 mi (16 km) E Red Bluff (NCSM). Yuba Co., 5 mi Superorder Typhlogena (8 km) N Smartville (UCB) and 1 mi (1.6 km) Order Platydesmida W Strawberry Valley Ranger Station (UCB). Family Andrognathidae REFERENCE.—Gardner 1975. Genus Brachycybe Wood, 1864 Genus Gosodesmus Brachycybe californica (Karsch, 1880) Chamberlin, 1922 TYPE LOCALITY.—California without further Chamberlin, 1922 specification. Gosodesmus claremontus OCCURRENCE IN CALIFORNIA.—Unknown. TYPE LOCALITY.—Claremont, Los Angeles REMARKS.—Hoffman (1999) thought this County. 2002] CALIFORNIA MILLIPEDS 93

OCCURRENCE IN CALIFORNIA.—Published finding more specimens. Re-collection of I. records: Amador, Kern, Los Angeles, Marin, plenipes is much to be desired. Mariposa, Mendocino, Monterey, Placer, San Mateo, Santa Barbara, Santa Clara, Santa Cruz, Family Siphonophoridae and Tulare counties. New records: Alameda, Undetermined genus and species Del Norte, El Dorado, Fresno, Madera, Orange, San Bernardino, San Diego, San Joaquin, OCCURRENCE IN CALIFORNIA.—Published Shasta, and Stanislaus cos. (AMNH, CAS, records: Los Angeles and Orange counties. New FSCA, LACMNH, NCSM, UAZ, UCB, UCD, records: None. UCR). REFERENCE.—Shelley 1996a. REFERENCE.—Gardner 1975. Subterclass Eugnatha Genus Ischnocybe Cook and Superorder Juliformia Loomis, 1928 Order Julida Family Julidae Ischnocybe plicata Cook and Loomis 1928 Genus Brachyiulus Berlese, 1884 TYPE LOCALITY.—14 mi (22.4 km) up North Brachyiulus lusitanus Verhoeff, 1898 Fork of Feather River from Belden, Plumas County. TYPE LOCALITY.—Portugal. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published records: Plumas County. New records: Hum- record: Contra Costa County. New records: boldt Co., Highway of Giants, 2 mi (3.2 km) N Kern Co., 15 mi (24 km) S Shirley Meadows Mendocino County line (AMNH). Sonoma Co., (LACMNH). Los Angeles Co., Los Angeles locality not specified (BMNH). (LACMNH), Pasadena (CAS), Claremont and REFERENCE.—Gardner 1975. Pomona (NMNH). Riverside Co., Riverside (UCR). Santa Barbara Co., Santa Barbara Genus Mitocybe Cook and (NMNH). Santa Clara Co., Palo Alto (NMNH). Loomis, 1928 Tulare Co., E of Visalia (WTAMU). Yolo Co., Davis (UCD). Yuba Co., 5 mi (8 km) N Smart- Cook and Loomis, 1928 Mitocybe auriportae ville (UCB). TYPE LOCALITY.—Mount Tamalpais, Marin REFERENCE.—Reeves 2000. County. OCCURRENCE IN CALIFORNIA.—Published Genus Cylindroiulus record: Marin County (known only from the Verhoeff, 1893 type locality). Cylindroiulus caeruleocinctus (Wood, 1864) REFERENCE.—Gardner 1975. TYPE LOCALITY.—Pennsylvania. Order Siphonophorida OCCURRENCE IN CALIFORNIA.—Published Family Siphonorhinidae records: None. New records: Los Angeles Co., Genus Illacme Cook and Long Beach, San Pedro, and Torrance (LAC- Loomis, 1928 MNH). Orange Co., La Habra (LACMNH). REFERENCES.—None. Illacme plenipes Cook and Loomis, 1928 REMARKS.—Cylindroiulus caeruleocinctus TYPE LOCALITY.—Near the divide between is a new species for California. Salinas and San Juan Bautista, San Benito (Curtis, 1845) County. Cylindroiulus latestriatus OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—England. record: San Benito County (known only from OCCURRENCE IN CALIFORNIA.—Published the type locality). records: Los Angeles and Santa Barbara coun- REFERENCE.—Shelley 1996a. ties. New records: Los Angeles Co., Artesia, REMARKS.—Illacme plenipes is the milliped Long Beach, Pasadena, and Torrance (CAS, and the animal with the most legs; a female FMNH, LACMNH). Orange Co., Anaheim, paratype has 750. I have searched in the vicin- Newport Beach, and Santa Ana (LACMNH). ity of the type locality several times without Sacramento Co., Sacramento (NMNH). San 94 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

Bernardino Co., Victorville (LACMNH). San OCCURRENCE IN CALIFORNIA.—Published Francisco Co., San Francisco (CAS). San Joa- record: Inyo County. New record: Inyo Co., quin Co., Manteca (CAS). Santa Barbara Co., Nelson Range, Grapevine Canyon, (FSCA). Los Alamos (LACMNH). REFERENCE.—Shelley and Bauer 1997a. REFERENCE.—Chamberlin 1914. Californiulus chamberlini (Brolemann, 1922) REMARKS.—Buckett (1964) said that this species occurs from Los Angeles as far north TYPE LOCALITY.—Mt. Shasta, Siskiyou as San Francisco Bay. County. OCCURRENCE IN CALIFORNIA.—Published (Silvestri, 1896) Cylindroiulus truncorum records: Shasta and Siskiyou counties. New TYPE LOCALITY.—Tunisia. records: None. OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Shelley 1994a. records: None. New records: Alameda Co., Californiulus dorsovittatus Verhoeff, 1938 Berkeley (UCB). Butte Co., Chico (NCSM). Los Angeles Co., Pasadena (CAS). Sacramento TYPE LOCALITY.—Mt. Harkness, Lassen Vol- Co., North Highlands, Dry Creek (UCD). San canic National Park, Plumas County. Bernardino Co., Mt. Baldy Village (BYU). San OCCURRENCE IN CALIFORNIA.—Published Mateo Co., Juniperro Serra County Park (CAS). records: Lassen, Plumas, Shasta, Siskiyou, and Yolo Co., Davis (UCD). Tehama counties. New records: None. REFERENCES.—None. REFERENCES.—Shelley 1994a, Shelley and REMARKS.—Cylindroiulus truncorum is a Bauer 1997a. new record for California. Californiulus yosemitensis Chamberlin, 1941 Genus Ophyiulus Berlese, 1884 TYPE LOCALITY.—Aspen Valley, Yosemite National Park, Tuolumne County. Ophyiulus pilosus (Newport, 1843) OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—England. records: Butte, Fresno, Kern, Lassen, Madera, OCCURRENCE IN CALIFORNIA.—Published Mariposa, Modoc, Mono, Nevada, Placer, records: None. New records: Los Angeles Co., Plumas, Shasta, Sierra, Tulare, and Tuolumne Claremont (FSCA). Sacramento Co., Carmichael counties. New record: County unknown (either (UCD). Placer or El Dorado), Lake Tahoe (BMNH). REFERENCES.—None. REFERENCES.—Shelley 1994a, Shelley and REMARKS.—Ophyiulus pilosus is a new rec- Bauer 1997a. ord for California. Genus Paeromopus Karsch, 1881 Family Nemasomatidae Paeromopus angusticeps angusticeps (Wood, Genus Orinisobates 1864) Lohmander, 1932 TYPE LOCALITY.—Oakland, Alameda County (Chamberlin, 1941) Orinisobates expressus (neotype locality). TYPE LOCALITY.—Washington. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published records: Alameda, Marin, Monterey, Santa record: Glenn County. New records: Shasta Co., Clara, and Santa Cruz counties. New records: along California highway 89 near Bartle (UCD). None. Trinity Co., 4 mi (6.4 km) S Hayfork (UCB). REFERENCES.—Shelley 1994a, Shelley and REFERENCE.—Enghoff 1985. Bauer 1997a. Paeromopus angusticeps buttensis Chamberlin, Family Paeromopodidae 1954 Genus Californiulus Verhoeff, 1938 TYPE LOCALITY.—Big Bar Mountain E Pulga, Shelley and Californiulus blechrostriatus Butte County. Bauer, 1997 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Saline Valley, Death Val- records: Amador, Butte, Calaveras, El Dorado, ley National Park, Inyo County. Humboldt, Lake, Mendocino, Nevada, Placer, 2002] CALIFORNIA MILLIPEDS 95

Plumas, Shasta, Sierra, Solano, Sonoma, Trin- Bollmaniulus pachysomus (Chamberlin, ity, Tuolumne, Yolo, and Yuba counties. New 1940) records: None. TYPE LOCALITY.—Yosemite National Park, REFERENCES.—Shelley 1994a, Shelley and Mariposa/Tuolumne counties. Bauer 1997a. OCCURRENCE IN CALIFORNIA.—Published Paeromopus angusticeps intergrades record: Mariposa/Tuolumne counties (known only from the type locality). OCCURRENCE IN CALIFORNIA.—Published records: Solano and Sonoma counties. New REFERENCE.—Chamberlin 1940. records: None. Bollmaniulus pearcei (Chamberlin, 1943) REFERENCE.—Shelley and Bauer 1997. TYPE LOCALITY.—South Fork of Bishop Paeromopus cavicolens Chamberlin, 1949 Creek, Inyo County. OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Windeler Cavern, ca 8 mi (12.8 km) N Columbia, Tuolumne County. record: Inyo County (known only from the type locality). OCCURRENCE IN CALIFORNIA.—Published record: Tuolumne County (known definitely REFERENCE.—Chamberlin 1943a. only from the type locality). Bollmaniulus rhodogeus (Chamberlin, 1940) REFERENCE.—Shelley 1994a. TYPE LOCALITY.—Redlands, San Bernardino Paeromopus eldoradus Chamberlin, 1941 County. OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Coloma, El Dorado County. record: San Bernardino County (known only OCCURRENCE IN CALIFORNIA.—Published record: El Dorado County (known only from from the type locality). the type locality). REFERENCE.—Chamberlin 1940. REFERENCE.—Shelley 1994a. Genus Codiulus Chamberlin, 1940 Paeromopus paniculus Shelley and Bauer, 1997 Codiulus etirus Chamberlin, 1941

TYPE LOCALITY.—1.5 mi (2.4 km) W El Por- TYPE LOCALITY.—Cardiff-by-the-Sea, San tal, Mariposa County. Diego County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published record: Mariposa County. New records: None. record: San Diego County (known only from REFERENCE.—Shelley and Bauer 1997a. the type locality). REFERENCE.—Chamberlin 1941a. Family Parajulidae Codiulus oulogon Chamberlin, 1940 Genus Bollmaniulus Verhoeff, 1926 TYPE LOCALITY.—Box Springs/Riverside, Bollmaniulus catalinae (Chamberlin, 1940) Riverside County. TYPE LOCALITY.—Santa Catalina Island, Los OCCURRENCE IN CALIFORNIA.—Published Angeles County. record: Riverside County (known only from OCCURRENCE IN CALIFORNIA.—Published the type locality). record: Santa Catalina Island, Los Angeles REFERENCE.—Chamberlin 1940. County (known only from the type locality). REFERENCE.—Chamberlin 1940. Genus Mulaikiulus Chamberlin, 1941 Bollmaniulus concolor (Chamberlin, 1940) Mulaikiulus stanleius Chamberlin, 1941

TYPE LOCALITY.—Redlands, San Bernardino TYPE LOCALITY.—12 mi (19.2 km) NE Ham- County. mond, Tulare County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published record: San Bernardino County (known only record: Tulare County (known only from the from the type locality). type locality). REFERENCE.—Chamberlin 1940. REFERENCE.—Chamberlin 1941b. 96 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

Genus Parajulus Humbert and Genus Spathiulus Chamberlin, 1940 DeSaussure, 1869 Spathiulus elegantulus Causey, 1950 Parajulus varius Bollman, 1887 TYPE LOCALITY.—Happy Isle, Yosemite Na- TYPE LOCALITY.—San Diego, San Diego tional Park, Mariposa County. County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published record: Mariposa County (known only from records: Mendocino and San Diego counties, the type locality). and “Rosario Mission,” county unknown. New REFERENCE.—Causey 1950. records: None. Spathiulus leptus Chamberlin, 1940 REFERENCES.—Bollman 1887a, 1887b. REMARKS.—This species, listed under “Para- TYPE LOCALITY.—Yosemite National Park, julidae of Uncertain Systematic Position” by Mariposa/Tuolumne counties. Chamberlin and Hoffman (1958) and “Parajul- OCCURRENCE IN CALIFORNIA.—Published idae of Uncertain Generic Position or Validity” record: Mariposa/Tuolumne counties (known by Hoffman (1999), was described as new twice only from the type locality). by Bollman (1887a, 1887b). I cite it here under REFERENCE.—Chamberlin 1940. Parajulus, but undoubtedly it belongs to a dif- Chamberlin, 1941 ferent genus, a decision that must await com- Spathiulus tribolus prehensive studies on western components of TYPE LOCALITY.—Sequoia National Park, 12 the family. In the original descriptions, Boll- mi (19.2 km) N Hammond, Tulare County. man reported it from San Diego; Ukiah, Men- OCCURRENCE IN CALIFORNIA.—Published docino County; and “Rosario Mission,” which record: Tulare County (known only from the was reported to be in Southern California and type locality). “Lower California,” i.e., Baja California. Cham- REFERENCE.—Chamberlin 1941b. berlin and Hoffman (1958) suggested San Diego (Chamberlin, 1950) as a restricted type locality but did not desig- Spathiulus tuolumnus nate a lectotype. TYPE LOCALITY.—Pinecrest, Tuolumne County. Genus Simiulus Chamberlin, 1940 OCCURRENCE IN CALIFORNIA.—Published record: Tuolumne County (known only from Simiulus arius (Chamberlin, 1918) the type locality). TYPE LOCALITY.—Stanford, Santa Clara REFERENCE.—Chamberlin 1950. County. REMARKS.—Chamberlin and Hoffman OCCURRENCE IN CALIFORNIA.—Published (1958) and Hoffman (1999) show this species record: Santa Clara County (known only from as having been originally included in Spathiu- the type locality). lus, but it was actually described in Sophiulus. REFERENCE.—Chamberlin 1940. Genus Tuniulus Chamberlin, 1941 Genus Sophiulus Chamberlin, 1940 Tuniulus milpitanus (Chamberlin, 1940) Sophiulus lomondus Chamberlin, 1941 TYPE LOCALITY.—Milpitas, Santa Clara TYPE LOCALITY.—Ben Lomond, Santa Cruz County. County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published record: Santa Clara County (known only from record: Santa Cruz County (known only from the type locality). the type locality). REFERENCE.—Chamberlin 1940. REFERENCE.—Chamberlin 1941a. Order Spirobolida (Chamberlin, 1912) Sophiulus tivius Family Atopetholidae TYPE LOCALITY.—Mill Valley, Marin County. Subfamily Arinolinae OCCURRENCE IN CALIFORNIA.—Published Loomis, 1953 records: Marin, San Francisco, and Santa Clara Arinolus latus counties. New records: None. TYPE LOCALITY.—Antelope Valley between REFERENCE.—Chamberlin 1940. Lancaster and Palmdale, Los Angeles County. 2002] CALIFORNIA MILLIPEDS 97

OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Hoffman and Orcutt 1960. record: Los Angeles County. New record: San Chamberlin, 1918 Bernardino Co., Pisgah Crater (LACMNH). Atopetholus fraternus REFERENCE.—Hoffman and Orcutt 1960. TYPE LOCALITY.—Friant, Fresno County. OCCURRENCE IN CALIFORNIA.—Published Genus Piedolus Chamberlin, 1930 record: Fresno County. New record: Kings Co., Avenal (AMNH). Piedolus utus Chamberlin, 1930 REFERENCE.—Hoffman and Orcutt 1960. TYPE LOCALITY.—Utah. OCCURRENCE IN CALIFORNIA.—Published Atopetholus paroicus Chamberlin, 1941 record: Inyo County. New records: None. TYPE LOCALITY.—Mountain Springs, Impe- REFERENCE.—Shelley 2000b. rial County. OCCURRENCE IN CALIFORNIA.—Published Subfamily Atopetholinae record: Imperial County (known only from the Genus Atopetholus Chamberlin, 1918 type locality). Atopetholus angelus Chamberlin, 1920 REFERENCE.—Hoffman and Orcutt 1960.

TYPE LOCALITY.—Silver Lake (Edendale sub- Atopetholus pearcei Chamberlin, 1950 urb), Los Angeles, Los Angeles County. TYPE LOCALITY.—Oildale, Kern County. OCCURRENCE IN CALIFORNIA.— Published OCCURRENCE IN CALIFORNIA.—Published record: Los Angeles County. New records: San record: Kern County (known only from the type Diego Co., U.S. International University, Elliott locality). Campus and Scripps Ranch (NCSM, UCD). REFERENCE.—Hoffman and Orcutt 1960. REFERENCE.—Hoffman and Orcutt 1960. REMARKS.—This name may prove to be a Atopetholus wheeleri (Chamberlin, 1918), junior synonym of A. californicus. new combination Atopetholus barbaranus Chamberlin, 1949 TYPE LOCALITY.—Cold Spring Canyon, Santa Ynez Mountains, Santa Barbara County. TYPE LOCALITY.—Orcutt, Santa Barbara OCCURRENCE IN CALIFORNIA.—Published County. record: Santa Barbara County (known only from OCCURRENCE IN CALIFORNIA.—Published the type locality). record: Santa Barbara County (known only from the type locality). REFERENCES.—Hoffman and Orcutt 1960, Hoffman 1998. REFERENCE.—Hoffman and Orcutt 1960 REMARKS.—Hoffman (1999) considered Atopetholus californicus Chamberlin, 1918 Hesperolus a questionable synonym of Atope- TYPE LOCALITY.—Claremont, Los Angeles/ tholus but nevertheless listed H. wheeleri, mak- San Bernardino counties. ing the combination with Atopetholus new. OCCURRENCE IN CALIFORNIA.—Published Atopetholus sp. record: Los Angeles/San Bernardino counties. New records: Kern Co., Tehachapi Mountains, OCCURRENCE IN CALIFORNIA.—Kern County, along California highway 58 at Clear Creek Erskine Canyon SE of Kernville (LACMNH). (NMNH). Orange Co., Newport Beach (UCD). Riverside County, Joshua Tree National Park, San Diego Co., Torrey Pines State Beach and Pinyon Wells (LACMNH). Reserve (NMNH). REMARKS.—For completion I list here coun- REFERENCES.—Hoffman and Orcutt 1960, ties from which unidentified species of Atope- Hoffman 1998. tholus have been taken.

Atopetholus carmelitus Chamberlin, 1940 Genus Orthichelus Chamberlin TYPE LOCALITY.—Hastings Reservation, Mon- and Hoffman, 1950 terey County. Orthichelus michelbacheri (Verhoeff, 1938) OCCURRENCE IN CALIFORNIA.—Published record: Monterey County (known only from TYPE LOCALITY.—Walker Pass, 6 mi (9.6 km) the type locality). NW Freeman, Kern County. 98 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Colorado Desert, Riverside record: Kern County. New records: Inyo Co., County. Argus Mountains (FSCA). Riverside Co. (UCD). OCCURRENCE IN CALIFORNIA.—Published San Bernardino Co., 5 mi (8 km) S Atolia records: San Bernardino, San Diego, and (FSCA). Riverside counties. New records: None. REFERENCE.—Hoffman and Orcutt 1960. REFERENCE.—Shelley 1998.

Genus Tidolus Chamberlin, 1949 Family Spirobolidae Subfamily Tylobolinae Tidolus parvus Chamberlin, 1918 Genus Hiltonius Chamberlin, 1918 TYPE LOCALITY.—Claremont, Los Angeles (Grinnell, 1908) County. Hiltonius australis OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Rancho La Brea tar pits, record: Los Angeles County. New records: Los Angeles, Los Angeles County. None. OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Hoffman and Orcutt 1960. record: Los Angeles County (known only from the type locality). Genus Watichelus Chamberlin, 1949 REFERENCE.—Grinnell 1908. Watichelus edentatus Loomis, 1949 REMARKS.—I follow Hoffman (1999) in plac- ing this species in Hiltonius. He stated that it TYPE LOCALITY.—Between San Diego and is “almost certainly a senior synonym of either El Centro, San Diego/Imperial counties. H. pulchrus or Tylobolus claremontus.” OCCURRENCE IN CALIFORNIA.—Published record: San Diego/Imperial counties (known Hiltonius hebes (Bollman, 1887) only from the type locality). TYPE LOCALITY.—San Diego, San Diego REFERENCE.—Hoffman and Orcutt 1960. County. Watichelus parallelus Loomis, 1949 OCCURRENCE IN CALIFORNIA.—Published record: San Diego County. New records: None. TYPE LOCALITY.—Chula Vista, San Diego REFERENCE.—Keeton 1960. County. OCCURRENCE IN CALIFORNIA.—Published Hiltonius mimus Chamberlin, 1941 record: San Diego County (known only from TYPE LOCALITY.—Mountain Springs, San the type locality). Diego County. REFERENCE.—Hoffman and Orcutt 1960. OCCURRENCE IN CALIFORNIA.—Published Watichelus robustus Loomis, 1949 record: San Diego County (known only from the type locality). TYPE LOCALITY.—Chula Vista, San Diego REFERENCE.—Keeton 1960. County. OCCURRENCE IN CALIFORNIA.—Published Hiltonius pulchrus Chamberlin, 1918 record: San Diego County (known only from TYPE LOCALITY.—Claremont, Los Angeles the type locality). County. REFERENCE.—Hoffman and Orcutt 1960. OCCURRENCE IN CALIFORNIA.—Published Watichelus smithi (Chamberlin, 1947) records: Kern, Los Angeles, and Riverside coun- ties. New records: None. TYPE LOCALITY.—Murray Canyon, 3 mi (4.8 REFERENCE.—Keeton 1960. km) N Palm Springs, Riverside County. OCCURRENCE IN CALIFORNIA.—Published Genus Tylobolus Cook, 1904 record: Riverside County (known only from the type locality). Tylobolus castaneus Chamberlin, 1918 REFERENCE.—Hoffman and Orcutt 1960. TYPE LOCALITY.—Brookdale, Santa Cruz County. Subfamily Onychelinae OCCURRENCE IN CALIFORNIA.—Published Genus Onychelus Cook, 1904 records: Alameda, Amador, Contra Costa, Onychelus obustus Cook, 1904 Fresno, Mariposa, Sacramento, San Joaquin, 2002] CALIFORNIA MILLIPEDS 99

Santa Clara, Santa Cruz, and Tulare counties. Tylobolus uncigerus (Wood, 1864) New record: El Dorado Co., 1 mi (1.6 km) E TYPE LOCALITY.—California without further Pacific House (NCSM). specification; restricted by Keeton (1960) to REFERENCES.—Keeton 1960, 1966. Solano County. Tylobolus claremontus Chamberlin, 1918 OCCURRENCE IN CALIFORNIA.—Published records: Contra Costa, Humboldt, Mariposa, TYPE LOCALITY.—Claremont, Los Angeles Sacramento, San Joaquin, Santa Cruz, and County. Solano counties. New records: Butte, Colusa, OCCURRENCE IN CALIFORNIA.—Published Glenn, Humboldt, Lake, Marin, Mendocino, records: Los Angeles, Riverside, and San Diego Napa, Santa Clara, Shasta, Sonoma, Tehama, counties. New records: None. Trinity, and Yolo cos. (CDFA, FSCA, MCZ, REFERENCES.—Keeton 1960, 1966. UCB, UCD) Tylobolus deses deses Cook, 1904 REFERENCES.—Keeton 1960, 1966.

TYPE LOCALITY.—California without further Tylobolus utahensis Chamberlin, 1925 specification; stated to be Stanford University, TYPE LOCALITY.—Utah Santa Clara County, by Buckett and Gardner OCCURRENCE IN CALIFORNIA.—Published (1966). record: Inyo County. New records: None. OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Shelley and Bauer 1997b. records: Marin, San Mateo, Santa Cruz, and Sonoma counties. New records: None. Order Spirostreptida REFERENCES.—Keeton 1960, 1966, Buckett Family Cambalidae and Gardner 1966. Genus Alakene Chamberlin, 1941 Tylobolus deses magnificus Buckett and Alakene simplex Chamberlin, 1941 Gardner, 1966 TYPE LOCALITY.—5 mi (8 km) NE Lemon- TYPE LOCALITY.—2 mi (3.2 km) W Cordelia, cove, Tulare County. Solano County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published record: Tulare County (known only from the records: Merced, Napa, and Solano counties. type locality). New records: Contra Costa Co., 2 mi (3.2 km) REFERENCE.—Chamberlin 1941b. E Clayton Valley (UCD). Sacramento Co., 5 mi (8 km) SW Folsom Dam (UCD). Stanislaus Genus Buwatia Chamberlin, 1912 Co., 15 mi (24 km) W Patterson (CAS). Buwatia monterea Chamberlin, 1912 REFERENCES.—Keeton 1960, 1966, Buckett and Gardner 1966. TYPE LOCALITY.—Pacific Grove, Monterey County. Tylobolus loomisi Keeton, 1966 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—South of Ft. Tejon, Kern record: Monterey County (known only from County. the type locality). OCCURRENCE IN CALIFORNIA.—Published REFERENCES.—Chamberlin 1912, Loomis records: Kern, Santa Barbara, and Ventura 1938. counties. New records: None. REFERENCE.—Keeton 1966. Genus Doilene Chamberlin, 1941 Tylobolus monachus (Chamberlin, 1949) Doilene carmela Chamberlin, 1941

TYPE LOCALITY.—Hastings Reservation, Mon- TYPE LOCALITY.—Hastings Reservation, Mon- terey County. terey County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published records: Fresno and Monterey counties. New record: Monterey County (known only from records: None. the type locality). REFERENCE.—Keeton 1960. REFERENCE.—Chamberlin 1941a. 100 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

Genus Endere Loomis, 1938 published drawings of a male from an unknown locality. As noted by Hoffman (1999), there is Endere disora Loomis, 1938 no reason to believe that this species is either TYPE LOCALITY.—Sunnyside Mine near conspecific or congeneric with the type speci- Seneca, Plumas County. mens. OCCURRENCE IN CALIFORNIA.—Published record: Plumas County (known only from the Nannolene catalina Chamberlin, 1941 type locality). TYPE LOCALITY.—Hamilton Canyon, Santa REFERENCE.—Loomis 1938. Catalina Island, Los Angeles County. OCCURRENCE IN CALIFORNIA.—Published Genus Leiodere Loomis, 1938 record: Santa Catalina Island, Los Angeles Leiodere angelorum Chamberlin, 1943 County. New records: None. REFERENCE.—Chamberlin 1941a. TYPE LOCALITY.—Los Angeles, Los Angeles County. Nannolene corticolens Chamberlin, 1951 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Base of Mt. Diablo, Contra record: Los Angeles County (known only from Costa County. the type locality. OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Chamberlin 1943a. record: Contra Costa County (known only from Leiodere dasyura Loomis, 1938 the type locality). REFERENCE.—Chamberlin 1951. TYPE LOCALITY.—Tajiguas, Santa Barbara County. Nannolene dorothea Chamberlin, 1941 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—One of 3 sites in Kern record: Santa Barbara County (known only County (Isabella, Kernville, or Squaw Valley) from the type locality). or 1 mi (1.6 km) E Hammond, Tulare County. REFERENCE.—Loomis 1938. OCCURRENCE IN CALIFORNIA.—Published Leiodere nana Loomis, 1938 records: Kern and Tulare counties. New records: None. TYPE LOCALITY.—Between Vallejo and Cor- REFERENCE.—Chamberlin 1941b. delia, Solano County. OCCURRENCE IN CALIFORNIA.—Published Nannolene keiferi Chamberlin, 1943 record: Solano County (known only from the type locality). TYPE LOCALITY.—Near Sacramento, Sacra- Reference.—Loomis 1938. mento County. OCCURRENCE IN CALIFORNIA.—Published Leiodere torreyana Loomis, 1938 record: Sacramento County (known only from TYPE LOCALITY.—Torrey Pines State Beach the type locality). and Reserve, San Diego County. REFERENCE.—Chamberlin 1943b. OCCURRENCE IN CALIFORNIA.—Published Nannolene minor Loomis, 1938 record: San Diego County (known only from the type locality). TYPE LOCALITY.—Near Bakersfield, Kern Reference.—Loomis 1938. County. OCCURRENCE IN CALIFORNIA.—Published Genus Nannolene Bollman, 1887 record: Kern County (known only from the type locality). Nannolene burkei (Bollman, 1887) REFERENCE.—Loomis 1938. TYPE LOCALITY.—Ukiah, Mendocino County. Nannolene personifer Chamberlin, 1941 OCCURRENCE IN CALIFORNIA.—Published record: Mendocino County (known only from TYPE LOCALITY.—9 mi (14.4 km) N Placer- the type locality). ville, El Dorado County. REFERENCES.—Chamberlin 1922, Loomis OCCURRENCE IN CALIFORNIA.—Published 1938. record: El Dorado County (known only from REMARKS.—Without providing a written the type locality). account, Chamberlin (1922, pl. 1, figs. 4–10) REFERENCE.—Chamberlin 1941b. 2002] CALIFORNIA MILLIPEDS 101

Nannolene violacea Loomis, 1938 REFERENCE.—Chamberlin 1941a.

TYPE LOCALITY.—South of Atascadero, San Genus Titsona Chamberlin, 1912 Luis Obispo County. OCCURRENCE IN CALIFORNIA.—Published Titsona sima Chamberlin, 1912 records: Kern and San Luis Obispo counties. TYPE LOCALITY.—Oroville, Butte County. New records: None. OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Loomis 1938. records: Butte and Yolo counties. New records: Genus Odachurus Loomis, 1938 None. REFERENCES.—Chamberlin 1912, Loomis Odachurus petasatus Loomis, 1938 1938, Chamberlin and Hoffman 1958. TYPE LOCALITY.—Torrey Pines State Beach Genus Tridere Loomis, 1938 and Reserve, San Diego County. OCCURRENCE IN CALIFORNIA.—Published Tridere chelopa Loomis, 1938 record: San Diego County (known only from TYPE LOCALITY.—2 mi (3.2 km) “above” the type locality). Mountain Springs along the road from San REFERENCE.—Loomis 1938. Diego to El Centro; inferred by Hoffman Genus Paiteya Chamberlin, 1910 (1999) to be in the southwestern corner of Imperial County. Paiteya errans Chamberlin, 1910 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Southern California with- record: Imperial County (known only from the out further specification. type locality). OCCURRENCE IN CALIFORNIA.—No definite REFERENCE.—Loomis 1938. localities known. Unknown Cambalidae REFERENCES.—Chamberlin 1910, Loomis 1938. OCCURRENCE IN CALIFORNIA.—Alameda, Colusa, Del Norte, Glenn, Los Angeles, Mon- Genus Pharodere Loomis, 1938 terey, Napa, Orange, Plumas, Riverside, San Bernardino, Shasta, Stanislaus, Tehama, Trinity, Loomis, 1938 Pharodere radiata Yolo, and Yuba counties (AMNH, CAS, CDFA, TYPE LOCALITY.—Torrey Pines State Beach FSCA, LACMNH, NCSM, NMNH, UCB). and Reserve, San Diego County. REMARKS.—As western Cambalidae have OCCURRENCE IN CALIFORNIA.—Published not been revised and the distinctions between record: San Diego County. New records: None. genera are unclear, I list here unidentified REFERENCE.—Loomis 1938. samples I have seen from other counties in California. Genus Platydere Loomis, 1938 Superorder Nematophora Platydere caeca Loomis, 1938 Order Chordeumatida TYPE LOCALITY.—Tajiguas, Santa Barbara Family Apterouridae County. Genus Apterourus Loomis, 1966 OCCURRENCE IN CALIFORNIA.—Published record: Santa Barbara County (known only Apterourus horizontalis Loomis, 1966 from the type locality). TYPE LOCALITY.—Cajon Pass, San Bernar- REFERENCE.—Loomis 1938. dino County. OCCURRENCE IN CALIFORNIA.—Published Genus Tigolene Chamberlin, 1941 record: San Bernardino County (known only Tigolene clementinus Chamberlin, 1941 from the type locality). REFERENCES.—Loomis 1966, Shear 1977. TYPE LOCALITY.—San Clemente Island, Los Angeles County. Family Caseyidae OCCURRENCE IN CALIFORNIA.—Published Genus Caseya Cook and Collins, 1895 record: San Clemente Island, Los Angeles County (known only from the type locality). Caseya bucketti Gardner and Shelley, 1989 102 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

TYPE LOCALITY.—22 mi (35.2 km) W Red OCCURRENCE IN CALIFORNIA.—Published Bluff, Tehama County. records: Amador, Butte, Calaveras, Colusa, El OCCURRENCE IN CALIFORNIA.—Published Dorado, Mariposa, and Tuolumne counties. records: Shasta and Tehama counties. New New records: None. records: None. REFERENCE.—Gardner and Shelley 1989. REFERENCE.—Gardner and Shelley 1989. Caseya heteropa oraria Gardner and Shelley, Caseya coxalis Loomis, 1966 1989 TYPE LOCALITY.—Davenport, Santa Cruz TYPE LOCALITY.—5 mi (8 km) NE Mendo- County. cino, Mendocino County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published records: San Mateo and Santa Cruz counties. New records: None. records: Humboldt and Mendocino counties. REFERENCE.—Gardner and Shelley 1989. New records: None. REFERENCE.—Gardner and Shelley 1989. Caseya dorada (Chamberlin, 1941) Caseya longiloba Gardner and Shelley, 1989 TYPE LOCALITY.—9 mi (14.4 km) N Placer- ville, El Dorado County. TYPE LOCALITY.—1.5 mi (2.4 km) SW Dou- OCCURRENCE IN CALIFORNIA.—Published glas City, Trinity County. records: Amador, Calaveras, El Dorado, Ne- OCCURRENCE IN CALIFORNIA.—Published vada, Placer, and Sacramento counties. New record: Trinity County. New records: None. records: None. REFERENCE.—Gardner and Shelley 1989. REFERENCE.—Gardner and Shelley 1989. Caseya occidentalis Gardner and Shelley, Caseya guttata Gardner and Shelley, 1989 1989

TYPE LOCALITY.—4 mi (6.4 km) E Sunny TYPE LOCALITY.—Del Puerto Canyon, 12 mi Brae, Humboldt County. (19.2 km) W Patterson, Stanislaus County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published record: Humboldt County. New records: None. record: Stanislaus County (known only from REFERENCE.—Gardner and Shelley 1989. the type locality). Caseya heteropa disjuncta Gardner and REFERENCE.—Gardner and Shelley 1989. Shelley, 1989 Caseya paradoxa Gardner and Shelley, 1989 TYPE LOCALITY.—3 mi (4.8 km) W Forest- ville, Sonoma County. TYPE LOCALITY.—2 mi (3.2 km) E Brice- OCCURRENCE IN CALIFORNIA.—Published burg, Mariposa County. records: Napa, Solano, Sonoma, and Yolo OCCURRENCE IN CALIFORNIA.—Published counties. New record: Lake Co., Cobb Moun- record: Mariposa County. New records: None. tain, Kelsey Creek (NMNH). REFERENCE.—Gardner and Shelley 1989. REFERENCE.—Gardner and Shelley 1989. Caseya prionota Gardner and Shelley, 1989 Caseya heteropa heteropa Cook and Collins, 1895 TYPE LOCALITY.—14 mi (22.4 km) E Brice- burg, Mariposa County. TYPE LOCALITY.—An unspecified locality in OCCURRENCE IN CALIFORNIA.—Published Marin County. records: Fresno, Madera, and Mariposa coun- OCCURRENCE IN CALIFORNIA.—Published ties. New records: None. records: Alameda, Marin, Monterey, San Benito, San Mateo, Santa Clara, Santa Cruz, Solano, REFERENCE.—Gardner and Shelley 1989. and Sonoma counties. New records: None. Caseya sequoia Chamberlin, 1941 REFERENCE.—Gardner and Shelley 1989. TYPE LOCALITY.—12 mi (19.2 km) NE Ham- Caseya heteropa montana Gardner and mond, Tulare County. Shelley, 1989 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—6 mi (9.6 km) S El Dora- record: Tulare County. New records: None. do, El Dorado County. REFERENCE.—Gardner and Shelley 1989. 2002] CALIFORNIA MILLIPEDS 103

Caseya shastensis Gardner and Shelley, 1989 Opiona bifurcata Gardner and Shelley, 1989

TYPE LOCALITY.—18 mi (28.8 km) W Red- TYPE LOCALITY.—Russian Gulch State Park, ding, Shasta County. Mendocino County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published record: Shasta County. New records: None. record: Mendocino County. New records: None. REFERENCE.—Gardner and Shelley 1989. REFERENCE.—Gardner and Shelley 1989. Caseya similis Causey, 1952 Opiona communis angusta Gardner and TYPE LOCALITY.—Tehachapi Pass, Kern Shelley, 1989 County. TYPE LOCALITY.—6 mi (9.6 km) SSE Santa OCCURRENCE IN CALIFORNIA.—Published Rosa, Sonoma County. records: Kern and Los Angeles counties. New OCCURRENCE IN CALIFORNIA.—Published records: None. records: Marin and Sonoma counties. New REFERENCE.—Gardner and Shelley 1989. records: None. Caseya taliae Gardner and Shelley, 1989 REFERENCE.—Gardner and Shelley 1989.

TYPE LOCALITY.—10 mi (16 km) NE Auberry, Opiona communis communis Gardner and Fresno County. Shelley, 1989 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—3 mi (4.8 km) N Kenwood, records: Fresno and Madera counties. New Sonoma County. records: None. OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Gardner and Shelley 1989. records: Santa Cruz, Solano, and Sonoma Caseya sp. counties. New records: None. REFERENCE.—Gardner and Shelley 1989. OCCURRENCE IN CALIFORNIA.—New records: Riverside Co., Rubidoux (LACMNH). Santa Opiona communis prolixa Gardner and Barbara Co., San Marcos Pass (NMNH). Ven- Shelley, 1989 tura Co., Wheeler Springs (LACMNH). TYPE LOCALITY.—7 mi (11.2 km) W Oak- REMARKS.—For completion I cite here addi- ville, Napa County. tional counties from which unidentifiable speci- OCCURRENCE IN CALIFORNIA.—Published mens of Caseya have been collected. records: Napa and Sonoma counties. New records: None. Genus Ochrogramma Gardner and REFERENCE.—Gardner and Shelley 1989. Shelley, 1989 Ochrogramma haigi Gardner and Shelley, Opiona distincta Gardner and Shelley, 1989 1989 TYPE LOCALITY.—2.5 mi (4.0 km) SE Guerne- TYPE LOCALITY.—5 mi (8 km) SW Gasquet, ville, Sonoma County. Del Norte County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published record: Sonoma County (known only from the record: Del Norte County. New records: None. type locality). REFERENCE.—Gardner and Shelley 1989. REFERENCE.—Gardner and Shelley 1989. Opiona exigua Gardner and Shelley, 1989 Genus Opiona Chamberlin, 1951 TYPE LOCALITY.—Russian Gulch State Park, Opiona berryessae Gardner and Shelley, Mendocino County. 1989 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—12 mi (19.2 km) W Winters, records: Mendocino and Sonoma counties. New Napa County. records: None. OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Gardner and Shelley 1989. record: Napa County (known only from the Opiona fisheri Gardner and Shelley, 1989 type locality). REFERENCE.—Gardner and Shelley 1989. TYPE LOCALITY.—Oregon. 104 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

OCCURRENCE IN CALIFORNIA.—Published Taiyutyla napa Shear, 1971 records: None. New record: Siskiyou Co., 8 mi TYPE LOCALITY.—Mount St. Helena, Napa (12.8 km) S Dunsmuir (UCB). County. REFERENCE.—Gardner and Shelley 1989. OCCURRENCE IN CALIFORNIA.—Published REMARKS.—Opiona fisheri is a new record record: Napa County (known only from the for California. type locality). Opiona siliquae Causey, 1963 REFERENCES.—Shear 1971, 1976.

TYPE LOCALITY.—Fault Rock Cave near Spy Family Rhiscosomididae Rock, Mendocino County. Genus Rhiscosomides Silvestri, 1909 OCCURRENCE IN CALIFORNIA.—Published record: Mendocino County (known only from Rhiscosomides acovescor Shear, 1972 the type locality). TYPE LOCALITY.—Samuel P. Taylor State Park, REFERENCE.—Gardner and Shelley 1989. Marin County. OCCURRENCE IN CALIFORNIA.—Published Genus Speoseya Causey, 1963 record: Marin County (known only from the Speoseya grahami Causey, 1963 type locality). REFERENCE.—Shear 1973. TYPE LOCALITY.—Shaw’s Cave, Calaveras County. Rhiscosomides montereum Chamberlin, 1910 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Pacific Grove, Monterey record: Calaveras County (known only from County. the type locality). OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Causey 1963. record: Monterey County (known only from the type locality). Family Conotylidae REFERENCE.—Shear 1973. Subfamily Conotylinae Genus Plumatyla Shear, 1971 Rhiscosomides trinitarium Shear, 1973 Plumatyla humerosa (Loomis, 1943) TYPE LOCALITY.—Butter Creek, 12 mi (19.2 km) SE Hyampom, Trinity County. TYPE LOCALITY.—Sunnyside Mine, 3 mi (4.8 OCCURRENCE IN CALIFORNIA.—Published km) SW Seneca, Plumas County. record: Trinity County (known only from the OCCURRENCE IN CALIFORNIA.—Published type locality). records: Plumas and Siskiyou counties. New REFERENCE.—Shear 1973. records: None. REFERENCE.—Shear 1971. Rhiscosomides sp. OCCURRENCE IN CALIFORNIA.—Del Norte Genus Taiyutyla Chamberlin, 1952 County, Jeddediah Smith State Park (UID). Taiyutyla extorris (Shear, 1971) San Mateo County, summit of La Honda grade (FSCA) and 6 mi (9.6 km) SE Half Moon Bay TYPE LOCALITY.—1 mi (1.6 km) S Grass Val- (NMNH). Santa Clara County, 5.2 mi (8.3 km) ley, Nevada County. SW Stevens Creek Dam (UCD). Santa Cruz OCCURRENCE IN CALIFORNIA.—Published County, Empire Cave (FSCA) and mountains record: Nevada County (known only from the between Santa Cruz and Holy City (NMNH). type locality). REFERENCE.—None. REFERENCES.—Shear 1971, 1976. REMARKS.—I have not attempted to iden- Taiyutyla francisca Shear, 1971 tify these tiny millipeds, and so I combine under this category all samples of Rhiscoso- TYPE LOCALITY.—1 mi (1.6 km) SE Inverness, mides that I have seen from California. Marin County. OCCURRENCE IN CALIFORNIA.—Published Family Striariidae record: Marin County (known only from the Genus Striaria Bollman, 1888 type locality). REFERENCES.—Shear 1971, 1976. Striaria californica Cook, 1899 2002] CALIFORNIA MILLIPEDS 105

TYPE LOCALITY.—California without further Genus Speostriaria Causey, 1960 specification; said to be near Sausolito, Marin County, by Hoffman (1999). Speostriaria shastae (Causey, 1958) OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Samwel Cave, Shasta records: Marin, Santa Cruz, and Solano coun- County. ties. New records: None. OCCURRENCE IN CALIFORNIA.—Published REFERENCES.—Cook 1899, Loomis 1936. record: Shasta County (known only from the Striaria carmela Chamberlin, 1947 type locality). REFERENCES.—Causey 1958, 1960. TYPE LOCALITY.—Hastings Reservation, Mon- terey County. Genus Vaferaria Causey, 1958 OCCURRENCE IN CALIFORNIA.—Published record: Monterey County (known only from Vaferaria imberbis (Loomis, 1936) the type locality). TYPE LOCALITY.—South of Atascadero, San REFERENCE.—Chamberlin 1947. Luis Obispo County. Striaria eldora Chamberlin, 1953 OCCURRENCE IN CALIFORNIA.—Published record: San Luis Obispo County (known only TYPE LOCALITY.—Crystal Cosumnes Cave, El Dorado County. from the type locality). REFERENCES.—Loomis 1936, Causey 1958. OCCURRENCE IN CALIFORNIA.—Published record: El Dorado County (known only from the type locality). Order Callipodida REFERENCE.—Chamberlin 1953. Family Schizopetalidae Subfamily Tynommatinae Striaria eutypa (Chamberlin, 1941) Tribe Colactidini TYPE LOCALITY.—9 mi (14.4 km) N Placer- Genus Colactis Loomis, 1937 ville, El Dorado County. Colactis protenta Loomis, 1937 OCCURRENCE IN CALIFORNIA.—Published record: El Dorado County (known only from TYPE LOCALITY.—Baja California Norte, Mex- the type locality). ico. REFERENCE.—Chamberlin 1941b. OCCURRENCE IN CALIFORNIA.—Published Striaria nana Loomis, 1936 record: San Diego County. New records: None. REFERENCE.—Shelley 1996b. TYPE LOCALITY.—Altamont Pass above Niles, Alameda County. Colactis tiburona (Chamberlin, 1923) OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Sonora, Mexico. record: Alameda and Santa Cruz counties. OCCURRENCE IN CALIFORNIA.—Published New records: None. record: Imperial County. New records: None. REFERENCE.—Loomis 1936. REFERENCE.—Shelley 1996b. Striaria sp. Genus Heptium Loomis, 1937 OCCURRENCE IN CALIFORNIA.—Published record: Contra Costa County. New records: Heptium carinellum Loomis, 1937 Alameda, El Dorado, Humboldt, Los Angeles, Marin, Mendocino, Sacramento, San Francisco, TYPE LOCALITY.—2 mi (3.2 km) E “Indian San Mateo, Santa Cruz, Shasta, Solano, Sonoma, Head,” presumably along California highway Stanislaus, and Trinity cos. (AMNH, CAS, 86, Imperial County. CDFA, FSCA, LACMNH, NCSM, NMNH, OCCURRENCE IN CALIFORNIA.—Published UCB, UCD). records: Imperial and Riverside counties. New REFERENCE.—Causey 1958. records: None. REMARKS.—As Striaria has not been revised, REFERENCE.—Shelley 1996b. it is impossible to identify forms at the species Heptium scamillatum Loomis, 1937 level. I therefore combine under this category all samples of Striaria that I have seen from TYPE LOCALITY.—Between Perris and Elsi- California. nore, Riverside County. 106 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Shelley 1996b. records: Riverside and San Bernardino coun- ties. New records: None. Diactis soleata Loomis, 1937 REFERENCE.—Shelley 1996b. TYPE LOCALITY.—Along Temescal Canyon Road ca 10.2 mi (16.3 km) S Corona, Riverside Tribe Diactidini County. Genus Caliactis Shelley, 1996 OCCURRENCE IN CALIFORNIA.—Published Caliactis bistolata Shelley, 1996 records: Los Angeles and Riverside counties. New records: None. TYPE LOCALITY.—Mission Viejo, Orange REFERENCE.—Shelley 1996b. County. OCCURRENCE IN CALIFORNIA.—Published Diactis strumella Shelley, 1996 record: Orange County (known only from the TYPE LOCALITY.—Laguna Beach, Orange type locality). County. REFERENCE.—Shelley 1996b. OCCURRENCE IN CALIFORNIA.—Published Genus Diactis Loomis, 1937 record: Orange County (known only from the type locality). Diactis amniscela Shelley, 1996 REFERENCE.—Shelley 1996b. TYPE LOCALITY.—Banning, Riverside County. Diactis triangula Loomis, 1937 OCCURRENCE IN CALIFORNIA.—Published record: Riverside County (known only from TYPE LOCALITY.—Ca 46 mi (73.6 km) E San the type locality). Diego, San Diego County. REFERENCE.—Shelley 1996b. OCCURRENCE IN CALIFORNIA.—Published records: Los Angeles, San Bernardino, and San Diactis cupola Shelley, 1996 Diego counties. New records: None. TYPE LOCALITY.—3 mi (4.8 km) from Santa REFERENCE.—Shelley 1996b. Monica, Los Angeles County. OCCURRENCE IN CALIFORNIA.—Published Genus Florea Shelley, 1996 record: Los Angeles County. New records: None. Florea sinuata Shelley, 1996 REFERENCE.—Shelley 1996b. TYPE LOCALITY.—Trabuco Ranger District, Diactis frondifera Loomis, 1937 Cleveland National Forest, Orange County. OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Torrey Pines State Beach record: Orange County (known definitely only and Reserve, San Diego County. from the paratype locality). OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Shelley 1996b. records: Los Angeles and San Diego counties. New records: None. Tribe Tynommatini REFERENCE.—Shelley 1996b. Genus Tynomma Loomis, 1937 Diactis jacinto Shelley, 1996 Tynomma magnum Buckett and Gardner, TYPE LOCALITY.—San Jacinto Mountains 1969 south of Banning, Riverside County. OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—7 mi (11.8 km) E Monterey, record: Riverside County (known only from Monterey County. the type locality). OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Shelley 1996b. record: Monterey County. New records: None. REFERENCE.—Shelley 1996b. Diactis procera Shelley, 1996 Tynomna mutans (Chamberlin, 1910) TYPE LOCALITY.—Silverado Canyon ca 11 mi (17.6 km) N Mission Viejo, Orange County. TYPE LOCALITY.—Stanford, Santa Clara OCCURRENCE IN CALIFORNIA.—Published County. record: Orange County (known only from the OCCURRENCE IN CALIFORNIA.—Published type locality). records: Alameda, Contra Costa, Marin, San 2002] CALIFORNIA MILLIPEDS 107

Mateo, Santa Clara, Santa Cruz, Solano, and Phreatodesmus torreyanus Loomis, 1960 Sonoma counties. New records: Monterey/San TYPE LOCALITY.—Torrey Pines State Beach Benito cos., between Salinas and San Juan and Reserve, San Diego County. Bautista (UCD). Napa Co., 7 mi (11.2 km) W OCCURRENCE IN CALIFORNIA.—Published Oakville (UCD). records: Riverside and San Diego counties. REFERENCE.—Shelley 1996b. New records: None. REFERENCE.—Loomis 1960. Tynomma schlingeri Gardner, 1973 TYPE LOCALITY.—Frank Raines Park, Del Genus Tidesmus Chamberlin, 1943 Puerto Canyon, Stanislaus County. Tidesmus episcopus Chamberlin, 1943 OCCURRENCE IN CALIFORNIA.—Published record: Santa Clara and Stanislaus counties. TYPE LOCALITY.—Bishop’s Road, Reservoir New records: None. Hill, Los Angeles County. REFERENCES.—Gardner 1973, Shelley 1996b. OCCURRENCE IN CALIFORNIA.—Published REMARKS.—As explained by Hoffman (1999), record: Los Angeles County (known only from Shelley (1996b) redescribed this species based the type locality). on the type specimens of T. schlingeri and REFERENCE.—Chamberlin 1943c. renamed it T. gardneri, a rare example of an Family Nearctodesmidae objective synonym. Genus Harpogonopus Loomis, 1960 Superorder Merocheta Harpogonopus confluentus Loomis, 1960 Order Polydesmida TYPE LOCALITY.—45 mi (72 km) E San Diego, Family Fuhrmannodesmidae San Diego County. Genus Phreatodesmus Loomis, 1960 OCCURRENCE IN CALIFORNIA.—Published Phreatodesmus cooki Loomis, 1960 records: Los Angeles, Orange, San Diego, Santa Barbara, and Ventura counties. New records: TYPE LOCALITY.—Tehachapi Pass, Kern None. County. REFERENCES.—Loomis 1960, Shelley 1993a, OCCURRENCE IN CALIFORNIA.—Published 1994b. record: Kern County (known only from the REMARKS.—Buckett (1964) spelled the genus type locality). correctly, but Hoffman (1980, 1999) and Shel- REFERENCE.—Loomis 1960. ley (1993a) misspelled it as “Harpagonopus.” Shelley (1993a) followed Hoffman (1980) in Phreatodesmus dentatus Loomis, 1960 considering the genus of uncertain family TYPE LOCALITY.—Cajon Pass, San Bernar- position in the Trichopolydesmoidea, but the dino County. following year he (Shelley 1994b) assigned it OCCURRENCE IN CALIFORNIA.—Published to the Nearctodesmidae. record: San Bernardino County (known only from the type locality). Genus Nearctodesmus Silvestri, 1910 REFERENCE.—Loomis 1960. Nearctodesmus cerasinus (Wood, 1864) Phreatodesmus hastingsus (Chamberlin 1941) TYPE LOCALITY.—Oregon. OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Hastings Reservation, Mon- records: Del Norte, Humboldt, Mendocino, terey County. and Siskiyou counties. New records: None. OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Shelley 1994b. record: Monterey County (known only from the type locality). Nearctodesmus salix Chamberlin, 1949 REFERENCES.—Loomis 1960, Shelley 1997b. TYPE LOCALITY.—12 mi (19.2 km) W Willow REMARKS.—Shelley (1997b) outlined the Creek (town), Humboldt County. confusion that surrounds this species, and, as OCCURRENCE IN CALIFORNIA.—Published noted by Hoffman (1999), placement in Phreat- records: Humboldt, Marin, Mendocino, Sono- odesmus is entirely speculative. ma, and Trinity counties. New records: None. 108 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

REFERENCE.—Shelley 1994b. OCCURRENCE IN CALIFORNIA.—Published record: Sacramento County (known only from Family Paradoxosomatidae the type locality). Tribe Australiosomatini REFERENCE.—Shear 1972. Genus Akamptogonus Attems, 1914 Bidentogon helferorum Buckett and Gardner, Akamptogonus novarae (Humbert and 1968 DeSaussure, 1869) TYPE LOCALITY.—1 mi (1.6 km) N Mendo- TYPE LOCALITY.—New Zealand. cino, Mendocino County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published records: Mendocino, Napa, San Diego, San records: Alameda and Mendocino counties. Francisco, Santa Barbara, and Sonoma coun- New records: Marin Co., Stinson Beach ties. New records: Mendocino Co., Mendocino (NCSM), 1 mi (1.6 km) W Olema (UCB), and Woodlands (CAS). Napa/Sonoma cos., locality Point Reyes Station and 2 mi (3.2 km) E Point not specified (CAS). San Diego Co., Torrey Reyes Station (FSCA, NCSM). Santa Clara Pines State Beach and Reserve (NMNH). Co., Milpitas (NMNH), Stanford University Santa Barbara Co., Accli Garden (NMNH). (NMNH), and Mt. Madonna (FSCA). Santa REFERENCES.—Hoffman 1979, Shelley et Cruz Co., Ben Lomond (UCB). San Mateo Co., al. 1998. near junction of Portola and San Gregorio roads REMARKS.—Shelley et al. (1998) recorded (FSCA). California counties for this species, to which I REFERENCES.—Buckett and Gardner 1968a, now add locality details. Shear 1972.

Tribe Sulciferini Genus Calianotus Shelley, 1997 Genus Oxidus Cook, 1911 Calianotus bituberculatus (Loomis, 1960) Oxidus gracilis (C. L. Koch, 1847) TYPE LOCALITY.—Niles Canyon, Alameda TYPE LOCALITY.—Austria (from a green- County. house). OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published records: Alameda and Contra Costa counties. records: None. New records: Alameda, Contra New records: None. Costa, Fresno, Los Angeles, Nevada, Orange, REFERENCE.—Shelley 1997b. Riverside, Sacramento, San Diego, San Fran- (Chamberlin, 1910) cisco, San Joaquin, San Luis Obispo, San Mateo, Calianotus sastianus Santa Barbara, Santa Clara, Shasta, and Yolo TYPE LOCALITY.—8 mi (12.8 km) S Duns- cos. (AMNH, BMNH, CAS, CDFA, LACMNH, muir, Shasta County (neotype locality). MCZ, NMNH, TMM, UCB, UCD, UCR). OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Buckett 1964. records: Shasta and Siskiyou counties. New REMARKS.—Buckett (1964) stated that this records: None. introduced, synanthropic diplopod occurs REFERENCE.—Shelley 1997b. throughout the western United States. It is one of the few millipeds with a common name, Calianotus yosemitensis (Causey, 1954) the hothouse or greenhouse milliped, and TYPE LOCALITY.—Vernal Falls, Yosemite unquestionably occurs in urban environments National Park, Mariposa County. in all counties of California. OCCURRENCE IN CALIFORNIA.—Published records: Calaveras, Madera, Mariposa, Tulare, Family Polydesmidae and Tuolumne counties. New records: None. Genus Bidentogon Buckett and REFERENCE.—Shelley 1997b. Gardner, 1968 Genus Scytonotus C. L. Koch, 1847 Bidentogon californicus (Chamberlin, 1918) Scytonotus simplex Chamberlin, 1941 TYPE LOCALITY.—Sacramento, Sacramento County. TYPE LOCALITY.—Oregon. 2002] CALIFORNIA MILLIPEDS 109

OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published records: Del Norte, Humboldt, Marin, Mendo- records: Calaveras, Mariposa, and Tuolumne cino, San Joaquin, Sonoma, and Trinity coun- counties. New records: None. ties. New records: None. REFERENCE.—Shelley 1995b. REFERENCE.—Shelley 1993b. Sigmocheir furcata Shelley, 1995

Family Xystodesmidae TYPE LOCALITY.—Williams Cave, ca 1 mi Subfamily Xystodesminae (1.6 km) N Murphys, Calaveras County. Tribe Chonaphini OCCURRENCE IN CALIFORNIA.—Published Genus Selenocheir Shelley, 1994 records: Amador, Calaveras, El Dorado, Placer, and Sacramento counties. New records: None. Selenocheir arcuata Shelley, 1994 REFERENCE.—Shelley 1995b. TYPE LOCALITY.—18.8 mi (30.1 km) NE Red Bluff, Tehama County. Sigmocheir maculifer (Chamberlin, 1941) OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—9 mi (14.4 km) N Wood- records: Humboldt, Lake, Mendocino, and lake, Tulare County. Tehama counties. New records: None. OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Shelley 1994c. records: Fresno and Tulare counties. New records: None. Selenocheir directa Shelley, 1994 REFERENCE.—Shelley 1995b. TYPE LOCALITY.—7 mi (11.2 km) NE Gas- quet, Del Norte County. Tribe Xystocheirini OCCURRENCE IN CALIFORNIA.—Published Genus Anombrocheir Buckett and record: Del Norte County. New records: None. Gardner, 1969 REFERENCE.—Shelley 1994c. Anombrocheir bifurcata Gardner and Selenocheir sinuata Shelley, 1994 Buckett, 1969 TYPE LOCALITY.—2 mi (3.2 km) SW Dales, TYPE LOCALITY.—8.5 mi (13.6 km) NW Rum- Tehama County. sey, Colusa County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published records: Butte, El Dorado, Humboldt, Placer, records: Colusa and Yolo counties. New records: Shasta, and Tehama counties. New records: None. None. REFERENCE.—Gardner and Buckett 1969. REFERENCE.—Shelley 1994c. Anombrocheir spinosa Buckett and Gardner, Tribe Sigmocheirini 1969 Genus Ochthocelata Shelley, 1995 TYPE LOCALITY.—3.5 mi (5.6 km) N Lees- ville, Colusa County. Ochthocelata adynata Shelley, 1995 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—4 mi (6.4 km) E Glenville, records: Colusa and Glenn counties. New Kern County. records: None. OCCURRENCE IN CALIFORNIA.—Published REFERENCES.—Buckett and Gardner 1969, record: Kern County. New records: Kern Co., Gardner and Buckett 1969. Live Oak Picnic Grounds (FSCA), and Kern R. Canyon, County Picnic Grounds, along Genus Motyxia Chamberlin, 1941 Stork Creek (FSCA, NCSM). Motyxia kerna Chamberlin, 1941 REFERENCE.—Shelley 1995b. TYPE LOCALITY.—7 mi (11.2 km) N Glenville, Genus Sigmocheir Chamberlin, 1951 Kern County. OCCURRENCE IN CALIFORNIA.—Published Chamberlin, 1951 Sigmocheir calaveras records: Kern and Tulare counties. New records: TYPE LOCALITY.—Crystal-Stanislaus Cave, ca None. 3 mi (4.8 km) W Columbia, Calaveras County. REFERENCE.—Shelley 1997c. 110 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

Motyxia monica Chamberlin, 1944 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Madelia Canyon, Sherman records: Kern County. New records: None. Oaks, Los Angeles County. REFERENCE.—Shelley 1997c. OCCURRENCE IN CALIFORNIA.—Published Motyxia tularea ollae Causey and Tiemann, records: Kern and Los Angeles counties. New 1969 records: None. REFERENCE.—Shelley 1997c. TYPE LOCALITY.—5.4 mi (8.6 km) W of Tule River Indian Reservation, Tulare County. Motyxia pior Chamberlin, 1941 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—12 mi (19.2 km) NE Ham- records: Tulare County. New records: None. mond, Tulare County. REFERENCE.—Shelley 1997c. OCCURRENCE IN CALIFORNIA.—Published records: Tulare County. New records: None. Motyxia tularea tularea (Chamberlin, 1949) REFERENCE.—Shelley 1997c. TYPE LOCALITY.—Sugarloaf Mountain, Tulare County. Motyxia porrecta Causey and Tiemann, 1969 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Stork Creek picnic site, records: Tulare County. New records: None. Kern River Canyon, Kern County. REFERENCE.—Shelley 1997c. OCCURRENCE IN CALIFORNIA.—Published records: Kern County. New records: None. Genus Parcipromus Shelley, 1995 REFERENCE.—Shelley 1997c. Parcipromus cooki (Causey, 1955) Motyxia sequoia alia Causey and Tiemann, 1969 TYPE LOCALITY.—Kings River Canyon, Fresno County (neotype locality). YPE LOCALITY T .—Along California highway OCCURRENCE IN CALIFORNIA.—Published 198 at Horseshoe Creek, Tulare County. records: Fresno and Tulare counties. New rec- OCCURRENCE IN CALIFORNIA.—Published ords: None. records: Tulare County. New records: None. REFERENCE.—Shelley 1995c. REFERENCE.—Shelley 1997c. Parcipromus gigantoarboricolus Shelley, 1995 Motyxia sequoia sequoia (Chamberlin, 1941) TYPE LOCALITY.—Giant Forest, Sequoia Na- TYPE LOCALITY.—10 mi (16 km) E Ham- tional Park, Tulare County. mond, Tulare County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published records: Tulare County. New records: None. records: Tulare County. New records: None. REFERENCE.—Shelley 1995c. REFERENCE.—Shelley 1997c. Motyxia sequoia intergrades Parcipromus tiemanni Shelley, 1995 TYPE LOCALITY.—11.2 mi (17.9 km) S Camp OCCURRENCE IN CALIFORNIA.—Published records: Tulare County. New records: None. Nelson, Tulare County. CCURRENCE IN ALIFORNIA REFERENCE.—Shelley 1997c. O C .—Published records: Tulare County. New records: None. Motyxia sequoiae (Loomis and Davenport, REFERENCE.—Shelley 1995c. 1951)

TYPE LOCALITY.—Above Camp Nelson, Genus Wamokia Chamberlin, 1941 Tulare County. Wamokia dentata Buckett and Gardner, 1968 OCCURRENCE IN CALIFORNIA.—Published records: Tulare County. New records: None. TYPE LOCALITY.—3 mi (4.8 km) NW Somer- REFERENCE.—Shelley 1997c. set, El Dorado County. OCCURRENCE IN CALIFORNIA.—Published Motyxia tiemanni Causey, 1960 record: El Dorado County (known only from TYPE LOCALITY.—Shirley Meadows, Kern the type locality). County. REFERENCE.—Buckett and Gardner 1968b. 2002] CALIFORNIA MILLIPEDS 111

Wamokia discordis Buckett and Gardner, OCCURRENCE IN CALIFORNIA.—Published 1968 record: San Luis Obispo County (known only from the type locality). TYPE LOCALITY.—1 mi (1.6 km) E Auburn, REFERENCE.—Shelley 1996c. Placer County. OCCURRENCE IN CALIFORNIA.—Published Xystocheir brachymacris Shelley, 1996 records: El Dorado and Placer counties. New TYPE LOCALITY.—1 mi (1.6 km) NE Pacific records: None. House, El Dorado County. REFERENCE.—Buckett and Gardner 1968b. OCCURRENCE IN CALIFORNIA.—Published Wamokia falcata Buckett and Gardner, 1968 records: El Dorado County. New records: None. TYPE LOCALITY.—Camino, El Dorado County. REFERENCE.—Shelley 1996c. OCCURRENCE IN CALIFORNIA.—Published record: El Dorado County (known only from Xystocheir dissecta dissecta (Wood, 1867) the type locality). TYPE LOCALITY.—Probably near San Fran- REFERENCE.—Buckett and Gardner 1968b. cisco or San Jose, San Francisco or Santa Clara Wamokia hoffmani Buckett and Gardner, counties. 1968 OCCURRENCE IN CALIFORNIA.—Published records: Alameda, Contra Costa, Marin, San TYPE LOCALITY.—1 mi (1.6 km) N Shingle Francisco, San Mateo, Santa Clara, and Sonoma Springs, El Dorado County. counties. New records: None. CCURRENCE IN ALIFORNIA O C .—Published REFERENCE.—Shelley 1996c. records: Amador and El Dorado counties. New records: None. Xystocheir dissecta microrama Shelley, 1996 REFERENCE.—Buckett and Gardner 1968b. TYPE LOCALITY.—2 mi (3.2 km) W Cordelia, Wamokia placera Chamberlin, 1941 Solano County. OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—9 mi (14.4 km) N Placer- records: Alameda, Contra Costa, Sacramento, ville, El Dorado County. and Solano counties. New records: None. OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Shelley 1996c. records: El Dorado County. New records: None. Xystocheir dissecta taibona Chamberlin, REFERENCE.—Buckett and Gardner 1968b. 1912 Wamokia remota Buckett and Gardner, 1968 TYPE LOCALITY.—Pacific Grove, Monterey County. TYPE LOCALITY.—8 mi (12.8 km) NE Auburn, OCCURRENCE IN CALIFORNIA.—Published Placer County. records: Monterey, San Benito, Santa Clara, OCCURRENCE IN CALIFORNIA.—Published and Santa Cruz counties. New records: None. records: Placer County. New records: None. REFERENCE.—Shelley 1996c. REFERENCE.—Buckett and Gardner 1968b. Xystocheir dissecta intergrades Wamokia sierrae Buckett and Gardner, 1968 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—1 mi (1.6 km) SW Grass records: Alameda, Contra Costa, Lake, Mendo- Valley, Nevada County. cino, Napa, Solano, Sonoma, Stanislaus, and OCCURRENCE IN CALIFORNIA.—Published Yolo counties. New records: None. record: Nevada County (known only from the REFERENCE.—Shelley 1996c. type locality). Xystocheir modestior haerens Shelley, 1996 REFERENCE.—Buckett and Gardner 1968b. TYPE LOCALITY.—Along California highway Genus Xystocheir Cook, 1904 180, ca 28 mi (44.8 km) E Fresno, Fresno County. Xystocheir bistipita Shelley, 1996 OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—1 mi (1.6 km) N San Luis record: Fresno County. New records: None. Obispo, San Luis Obispo County. REFERENCE.—Shelley 1996c. 112 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

Xystocheir modestior modestior (Chamberlin, Xystocheir stolonifera intergrades 1941) OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—4–7 mi (6.4–11.2 km) N record: Madera County. New records: None. Badger, Fresno/Tulare counties. REFERENCE.—Shelley 1996c. OCCURRENCE IN CALIFORNIA.—Published records: Fresno, Madera, and Tulare counties. Tribe Xystodesmini New records: None. Genus Harpaphe Cook, 1904 REFERENCE.—Shelley 1996c. Harpaphe haydeniana cummingsiensis Xystocheir prolixorama Shelley, 1996 (Verhoeff, 1944)

TYPE LOCALITY.—5 mi (8 km) N Cloverdale, TYPE LOCALITY.—Cummings, Mendocino Mendocino County. County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published record: Mendocino County. New records: None. records: Humboldt, Mendocino, and Sonoma REFERENCE.—Shelley 1996c. counties. New records: None. REFERENCE.—Buckett and Gardner 1968c. Xystocheir reducta (Causey, 1955) Harpaphe haydeniana haydeniana (Wood, TYPE LOCALITY.—2.1 mi (3.4 km) E Brice- 1864) burg, Mariposa County. OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—Oregon (neotype locality). records: Amador, Calaveras, Mariposa, and OCCURRENCE IN CALIFORNIA.—Published Tuolumne counties. New records: None. records: Marin County. New records: None. REFERENCE.—Shelley 1996c. REFERENCE.—Buckett and Gardner 1968c. Xystocheir solenofurcata Shelley, 1996 Harpaphe haydeniana inlignea Chamberlin, 1949 TYPE LOCALITY.—2 mi (3.2 km) N Plymouth, Amador County. TYPE LOCALITY.—Inwood, Shasta County. OCCURRENCE IN CALIFORNIA.—Published OCCURRENCE IN CALIFORNIA.—Published record: Amador County. New records: None. records: Shasta County. New records: None. REFERENCE.—Shelley 1996c. REFERENCE.—Buckett and Gardner 1968c. Xystocheir stenomacris Shelley, 1996 Harpaphe haydeniana lanceolata Buckett and Gardner, 1968 TYPE LOCALITY.—15 mi (24 km) N Ahwah- nee, Mariposa County. TYPE LOCALITY.—Mount St. Helena, Napa OCCURRENCE IN CALIFORNIA.—Published County. record: Mariposa County. New records: None. OCCURRENCE IN CALIFORNIA.—Published REFERENCE.—Shelley 1996c. records: Lake and Napa counties. New records: None. Xystocheir stolonifera stolonifera Shelley, REFERENCE.—Buckett and Gardner 1968c. 1996 Harpaphe haydeniana maurogona Buckett TYPE LOCALITY. —2 mi (3.2 km) S Tollhouse, and Gardner, 1968 Fresno County. OCCURRENCE IN CALIFORNIA.—Published TYPE LOCALITY.—2 mi (3.2 km) E Baxter, record: Fresno County. New records: None. Nevada County. REFERENCE.—Shelley 1996c. OCCURRENCE IN CALIFORNIA.—Published record: Nevada County (known only from the Xystocheir stolonifera uncinata Shelley, 1996 type locality). TYPE LOCALITY.—3 mi (4.8 km) S Coarse- REFERENCE.—Buckett and Gardner 1968c. gold, Madera County. Harpaphe haydeniana scotia (Chamberlin, OCCURRENCE IN CALIFORNIA.—Published 1941) records: Fresno and Madera counties. New records: None. TYPE LOCALITY.—12 mi (19.2 km) S Los REFERENCE.—Shelley 1996c. Gatos, Santa Clara County. 2002] CALIFORNIA MILLIPEDS 113

OCCURRENCE IN CALIFORNIA.—Published ______. 1968b. Revision of the milliped genus Wamokia records: Monterey, Santa Clara, and Santa Chamberlin from the Sierra Nevada of central Cali- fornia (Diplopoda: Polydesmida: Xystodesmidae). Cruz counties. New records: Alameda Co., Proceedings of the Biological Society of Washington Berkeley (UCB). San Mateo Co., County 81:511–538. Memorial Park (CAS). ______. 1968c. Revision of the milliped genus Harpaphe REFERENCE.—Buckett and Gardner 1968c. Cook from western North America (Polydesmida: Xystodesmidae). California Department of Agricul- Harpaphe pottera Chamberlin, 1949 ture, Occasional Papers of the Bureau of Entomol- ogy 11:1–51. TYPE LOCALITY.—Potter Creek, Mendocino ______. 1968d. Rediscovery of the type of the milliped County. Harpaphe telodonta (Chamberlin) (Diplopoda: Eury- desmidae). Journal of the New York Entomological OCCURRENCE IN CALIFORNIA.—Published Society 76:60–63. records: Humboldt and Mendocino counties. ______. 1969. A new genus of xystodesmid milliped from New records: None. northern California. Entomological News 80:67–73. REFERENCE.—Buckett and Gardner 1968c. CAUSEY, N.B. 1950. New genera and species of millipeds Paraiulidae (Juloidea). Proceedings of the Arkansas Harpaphe telodonta Chamberlin, 1943 Academy of Science 3:45–58. ______. 1958. New records and descriptions of a new TYPE LOCALITY.—Arcata, Humboldt County. genus and a new species of millipeds of the family OCCURRENCE IN CALIFORNIA.—Published Striariidae (Chordeumida). Proceedings of the Bio- records: Del Norte and Humboldt counties. logical Society of Washington 71:179–184. New records: None. ______. 1960. Speostriaria, new genus (Diplopoda: Chor- deumida: Chordeumidea: Striariidae). Proceedings REFERENCES.—Buckett and Gardner 1968c, of the Biological Society of Washington 73:25–28. 1968d. ______. 1963. Two new caseyid millipeds from California caves (Chordeumida: Chordeumidea). Wasmann Jour- ACKNOWLEDGMENTS nal of Biology 21:193–198. CHAMBERLIN, R.V. 1910. Diplopoda from the western states. Annals of the Entomological Society of Amer- I thank my colleague W.A. Shear for loan of ica 3:233–262. material in his private collection, and the fol- ______. 1912. New North American chilopods and diplo- lowing curators and collection managers for pods. Annals of the Entomological Society of Amer- loans from or access to the indicated collec- ica 5:141–172. ______. 1914. A new Julus from California. Canadian Ento- tions: N.I. Platnick (AMNH), P. Hillyard mologist 46:314–315. (BMNH), C.E. Griswold (CAS), A.R. Hardy ______. 1922. Notes on West Indian millipeds. Proceed- (CDFA), P. Sierwald (FMNH), G.B. Edwards ings of the United States National Museum 61:1–19. (FSCA), the late C.L. Hogue (LACMNH), L. ______. 1940. New genera and species of North American Leibensperger (MCZ), J.A. Coddington Paraiulidae. Bulletin of the University of Utah 30(11) [Biological Series 5(7)]:1–39. (NMNH), J.R. Reddell (TMM), C.A. Olsen ______. 1941a. New American millipeds. Bulletin of the (UAZ), C.B. Barr (UCB), L.S. Kimsey (UCD), University of Utah 31(11) [Biological Series 6(4)]:1–39. S.I. Frommer (UCR), F.W. Merickel (UID), ______. 1941b. New western millipeds. Bulletin of the R.S. Zack (WSU), and W.D. Sissom (WTAMU). University of Utah 31(12) [Biological Series 6(5)]:1–23. ______. 1943a. On some genera and species of American millipeds. Bulletin of the University of Utah 34(6) LITERATURE CITED [Biological Series 8(2)]:1–20. ______. 1943b. A new cambalid diplopod. Entomological BOLLMAN, C.H. 1887a. Notes on North American Julidae. News 54:88–89. Annals of the New York Academy of Science ______. 1943c. On nine North American polydesmoid 4:25–44. millipeds. Proceedings of the Biological Society of ______. 1887b. Description of new genera and species of Washington 56:35–40. North American Myriopoda (Julidae). Entomologica ______. 1947. Seven new American millipeds. Proceed- Americana 1:225–228. ings of the Biological Society of Washington 60:9–16. BUCKETT, J.S. 1964. Annotated list of the Diplopoda of ______. 1950. Three new genera and eight new species of California. Simmons Publishing Co., Davis, CA. 34 western millipeds. Chicago Academy of Science pp. Natural History Miscellanea 68:1–6. BUCKETT, J.S., AND M.R. GARDNER. 1966. Rediscovery ______. 1951. Records of American millipeds and cen- and redescription of Tylobolus deses Cook, with the tipeds collected by Dr. D. Elden Beck in 1950. description of a new subspecies (Diplopoda: Great Basin Naturalist 11:27–35. Spirobolidae). Proceedings of the Biological Society ______. 1953. Two new millipeds taken in California caves. of Washington 79:41–48. Entomological News 64:93–95. ______. 1968a. A new genus and species of milliped from CHAMBERLIN, R.V., AND R.L. HOFFMAN. 1958. Checklist of northern California (Polydesmida: Vanhoeffeniidae). the millipeds of North America. United States Nation- Pan-Pacific Entomologist 44:198–202. al Museum Bulletin 212:1–236. 114 MONOGRAPHS OF THE WESTERN NORTH AMERICAN NATURALIST [No. 1

COOK, O.F.1899. The diplopod family Striariidae. Proceed- PIERCE, W.D. 1940. A rare myriopod from Anacapa Island ings of the United States National Museum 21: compared with two Texas species. Bulletin of the 667–676. Southern California Academy of Science 39:158–171. ENGHOFF, H. 1985. The millipede family Nemasomatidae, REEVES, W.K. 2000. Additional California records of with the description of a new genus, and a revision Brachyiulus lusitanus (Julida: Julidae), an exotic Euro- of Orinisobates (Diplopoda: Julida). Entomologica pean millipede. Pan-Pacific Entomologist 76:251. Scandinavica 16:27–67. SHEAR, W.A. 1971. The milliped family Conotylidae in GARDNER, M.R. 1973. A new milliped of the genus Ty- North America, with a description of the new family nomma from California (Lysiopetalidea: Lysiopetali- Adritylidae (Diplopoda: Chordeumida). Bulletin of dae). Pan-Pacific Entomologist 49:270–272. the Museum of Comparative Zoology 141:55–97. ______. 1975. Revision of the millipede family Androg- ______. 1972. The milliped genus Bidentogon (Diplopoda, nathidae in the Nearctic region. Memoirs of the Polydesmida, Trichopolydesmidae). Proceedings of Pacific Coast Entomological Society 5:1–61. the Biological Society of Washington 85:489–492. ______. 1973. The milliped family Rhiscosomididae (Dip- GARDNER, M.R., AND J.S. BUCKETT. 1969. A new species of Anombrocheir Buckett and Gardner from the inner lopoda: Chordeumatida: Striarioidea). Psyche 80: Coast Ranges of northern California. Entomological 189–203. ______. 1976. The milliped family Conotylidae (Diplo- News 80:293–299. poda, Chordeumida) revision of the genus Taiyutyla, GARDNER, M.R., AND R.M. SHELLEY. 1989. New records, with notes on recently proposed taxa. American species, and genera of caseyid millipeds from the Museum Novitates 2600:1–22. Pacific Coast of North America (Diplopoda: Chor- ______. 1977. The systematic position of the milliped deumatida: Caseyidae). Pan-Pacific Entomologist family Apterouridae (Diplopoda, Chordeumida, Stri- 65:177–268. arioidea) and a redescription of Apterourus horizon- GRINNELL, F. 1908. Quaternary myriopods and insects of talis Loomis. Proceedings of the Biological Society California. Bulletin of the University of California of Washington 89:689–694. Department of Geology 5:207–215. ______. 1986. Millipeds from caves in Mexico and Central HOFFMAN, R.L. 1979. An Australian polydesmoid milliped America. V. New species and records of Glomeridae, in San Francisco (Paradoxosomatidae). Wasmann Trichopetalidae, Cleidogonidae, Fuhrmannodesmi- Journal of Biology 37:55–58. dae, Cryptodesmidae, Cambalidae, Typhlobolellidae, ______. 1980 (1979). Classification of the Diplopoda. Rhachodesmidae, and Sphaeriodesmidae. Texas Me- Museum d’Histoire Naturelle, Geneva, Switzerland. morial Museum Speleological Monograph 1:63–86. 237 pp. SHELLEY, R.M. 1988. The millipeds of eastern Canada ______. 1998. Some necessary fine-tuning in the order (Arthropoda: Diplopoda). Canadian Journal of Zool- Spirobolida (Spirobolidae, Messicobolidae, Atopethol- ogy 66:1638–1663. idae). Myriapodologica 5:63–76. ______. 1990. A new milliped of the genus Metaxycheir ______. 1999. Checklist of the millipeds of North and from the Pacific Coast of Canada (Polydesmida: Xy- Middle America. Virginia Museum of Natural His- stodesmidae), with remarks on the tribe Chonaphini tory Special Publication 8:1–584. and the western Canadian and Alaskan diplopod HOFFMAN, R.L., AND B.S. ORCUTT. 1960. A synopsis of the fauna. Canadian Journal of Zoology 68:2310–2322. Atopetholidae, a family of spiroboloid millipeds. Pro- ______. 1993a. Harpagonopus confluentus Loomis, a Pacific ceedings of the United States National Museum 111: Coast milliped of the United States and Mexico 95–166. (Polydesmida: Trichopolydesmoidea). Myriapodolog- JOHNSON, B.M. 1954. The millipeds of Michigan. Papers ica 2:73–81 of the Michigan Academy of Science, Arts, and Let- ______. 1993b. Revision of the milliped genus Scytonotus ters 39:241–252. Koch (Polydesmida: Polydesmidae). Brimleyana 19: 1–60. KEETON, W.T. 1960. A taxonomic study of the milliped family Spirobolidae (Diplopoda: Spirobolida). Mem- ______. 1994a. Revision of the milliped family Paeromo- oirs of the American Entomological Society 17:1–146. podidae, and elevation of the Aprosphylosomatinae to family status (Julida: Paeromopodoidea). Entomo- ______. 1966. The species of the milliped genus Tylobolus logica Scandinavica 25:169–214. (Diplopoda: Spirobolida), a re-examination. Transac- ______. 1994b. The milliped family Nearctodesmidae in tions of the American Entomological Society 92: northwestern North America, with accounts of Sako- 17–28. phallus and S. simplex Chamberlin (Polydesmida). LOOMIS, H.F. 1936. New millipeds of the American family Canadian Journal of Zoology 72:470–495. Striariidae. Journal of the Washington Academy of ______. 1994c. The Chonaphini, a biogeographically sig- Science 26:404–409. nificant milliped tribe in eastern and western North ______. 1938. The cambaloid millipeds of the United America (Polydesmida: Xystodesmidae). Brimleyana States, including a family new to the fauna and new 20:111–200. genera and species. Proceedings of the United States ______. 1995a. The milliped family Hirudisomatidae in National Museum 86:27–66. the New World (Polyzoniida). Brimleyana 23:103–143. ______. 1960. Millipeds of the order Polydesmida from ______. 1995b. The Sigmocheirini, a xystodesmid mil- the western states and Baja California. Journal of the liped tribe in the Sierra Nevada Mountains, Califor- Kansas Entomological Society 33:57–68. nia, U.S.A. (Polydesmida: Xystodesmidae). Entomo- ______. 1966. Two new families and other North Ameri- logica Scandinavica 26:339–360. can Diplopoda of the suborder Chordeumidea. Pro- ______. 1995c. Parcipromus, n. gen., a xystodesmid milli- ceedings of the Biological Society of Washington ped genus from the Sierra Nevada Mountains, Cali- 79:221–230. fornia (Polydesmida). Myriapodologica 3:53–70. 2002] CALIFORNIA MILLIPEDS 115

______. 1996a. The milliped order Siphonophorida in the desmida: Xystodesmidae). Journal of the Elisha United States and northern Mexico. Myriapodolog- Mitchell Scientific Society 116:177–205. ica 4:21–33. ______. 2000b. The milliped genus Piedolus Chamberlin ______. 1996b. The milliped order Callipodida in western (Spirobolida: Atopetholidae: Arinolinae). Myriapodo- North America (Schizopetalidae: Tynommatinae), logica 6:77–81. and a summary of the New World fauna. Entomolog- ______. 2001. Annotated checklist of the millipeds of ica Scandinavica 27:25–64. Florida (Arthropoda: Diplopoda). Insecta Mundi 14: ______. 1996c. Revision of the milliped genus Xystocheir 241–251. Cook (Polydesmida: Xystodesmidae). Canadian Jour- SHELLEY, R.M., AND S.B. BAUER. 1997a. New records and nal of Zoology 74:1336–1363. species, and taxonomic alterations, in the milliped ______. 1997a. The milliped family Polyzoniidae in North family Paeromopodidae (Julida). Entomological News America, with a classification of the global fauna 108:1–14. (Diplopoda Polyzoniida). Arthropoda Selecta 6(3/4): ______. 1997b. Distribution of the milliped Tylobolus uta- 3–34. hensis Chamberlin, with remarks on T. fredericksoni ______. 1997b. The identity of Polydesmus sastianus (Causey) (Spirobolida: Spirobolidae). Great Basin Chamberlin, proposal of a new milliped genus, and Naturalist 57:231–237. remarks on the identity of Phreatodesmus hastingsus SHELLEY, R.M., S.B. BAUER, AND S.F. SWIFT. 1998. The (Chamberlin) (Polydesmida: Polydesmidae). Myria- milliped family Paradoxosomatidae in the Hawaiian podologica 4:59–67. Islands (Diplopoda: Polydesmida). Bishop Museum ______. 1997c. A re-evaluation of the milliped genus Occasional Papers 56:43–53. Motyxia Chamberlin, with a re-diagnosis of the tribe SNIDER, R.M. 1991. Updated species lists and distribution Xystocheirini and remarks on the bioluminescence records for the Diplopoda and Chilopoda of Michi- (Polydesmida: Xystodesmidae). Insecta Mundi 11: gan. Michigan Academician 24:177–194. 331–351. WILLIAMS, S.R., AND R.A. HEFNER. 1928. The millipedes ______. 1998. The milliped genus Onychelus Cook (Spiro- and centipedes of Ohio. Bulletin of the Ohio Biolog- bolida: Atopetholidae: Onychelinae). Myriapodolog- ical Survey 4:93–146. ica 5:111–117. ______. 2000a. Annotated checklist of the millipeds of Received 18 September 2000 North Carolina (Arthropoda: Diplopoda), with re- Accepted 21 February 2001 marks on the genus Sigmoria Chamberlin (Poly-