Gardening for Bees with the Cgs

Total Page:16

File Type:pdf, Size:1020Kb

Gardening for Bees with the Cgs GARDENING FOR BEES WITH THE CGS Bees are currently under heavy pressure from disease and habitat change, and numbers have declined drastically. This is bad news for plants and humans alike, since it affects plants' ability to set fruit and seed, and thus has a negative impact on our food sources and incomes. The poor apple harvest of 2012, resulting from a cold snap at flowering time which deterred bees, showed how important they are to us! Cottage gardeners are well placed to help bees, as by definition cottage gardens contain a wide range of plants, and the gardeners often prefer the type of plant bees like. The main thing we can do to encourage bees is to select garden plants which produce plenty of nectar. As a general rule, these are the simpler, more old- fashioned types of plant. A guide is that any plant which produces SEEDS (not just fruits, as these can be sterile) is probably a good bee plant. Highly-bred cultivars, especially doubles, are often sterile and do not produce much nectar, if any. Some good plants for bees are listed overleaf. Another point to bear in mind is that although bees hibernate during the winter, they (bumble bees especially) often appear quite early during warm spells, and at this time badly need a source of food. They will also work long into a mild autumn if food is available. Good early-flowering plants include Pulmonaria and other members in the same family such as Trachystemon orientalis and some types of Symphytum ('Hidcote Blue' and S. ibericum particularly), and the shrub Lonicera pileata, which has scarcely noticeable greenish flowers in February - but the bees find them! In cold areas Lonicera fragrantissima and L. x purpusii may still be in flower at this time. Primroses flower intermittently from autumn, and winter heathers are good. The vital autumn/winter nectar plant is ivy, often not allowed to reach flowering size in gardens as it can be a weed and a menace to trees. However, if you have an old wall (not the house!) or a strong fence or dead tree, do encourage it - trim it but let it form a bushy core that will flower. The bees will show their appreciation in your fruit and vegetable beds in summer. Parthenocissus, Boston ivy, is house-safe and nectar-rich, but summer-flowering. Bee plants Bees love the labiates (lavender, mint, thyme, oregano, salvias, agastache, stachys etc.) and plants in the Boraginaceae family (borage, pulmonaria, symphytum, anchusa, forget-me-not, echium etc). Almost all plants of the two families will attract them. Other good plants include the following: Annuals/biennials Scabious & Knautia Foxgloves Sedums Californian poppy Clover Cosmos Verbena bonariensis Sunflowers Veronica spp. (Speedwell) Sweet rocket Candytuft Shrubs & Trees Honesty Acers Mignonette Arbutus unedo (Strawberry tree) Berberis Perennials Buddleja globosa Alliums Cotoneasters Michaelmas daisies Cytisus spp. (Broom) Campanulas (singles) Hebes Thistles Bay Teasels Mahonia Globe thistles Malus spp. (Apple & crabapple) Eryngium (Sea holly) Pyracantha Snowdrops Ribes spp. (Currants) Hellebores (singles) Sarcococca confusa (Christmas box) Catmint Sorbus aucuparia (Rowan) Polemonium spp. (Jacob's Ladder) Willow Rudbeckia & Echinacea There are many others – see, for instance, the RHS website and check your seed catalogues, which now often give bee information. .
Recommended publications
  • Urban Flora and Ecological Characteristics of the Kartal District (Istanbul): a Contribution to Urban Ecology in Turkey
    Scientific Research and Essay Vol. 5(2), pp. 183-200, 18 January, 2010 Available online at http://www.academicjournals.org/SRE ISSN 1992-2248 © 2010 Academic Journals Full Length Research Paper Urban flora and ecological characteristics of the Kartal District (Istanbul): A contribution to urban ecology in Turkey Volkan Altay1*, brahim lker Özyiit2 and Celal Yarci2 1Mustafa Kemal University, Science and Arts Faculty, Department of Biology, 31000, Antakya/Hatay/Turkey. 2Marmara University, Science and Arts Faculty, Department of Biology, 34722, Göztepe/Istanbul/Turkey. Accepted 22 October, 2009 For years, ecologists who have been trying to understand the relationship between the organisms with each other and/or their environments, have carried out their researches sometimes far from civilization, sometimes on a desolate island or in a tropical rainforest. Today, about half of the world’s population lives in urban areas. Therefore, most of the ecological problems have been brought to these areas. Nevertheless, in cities, preserving and maintaining natural habitats, providing a place not only to live but also to enjoy and to relax, are possible only by applying the principles and concepts of urban ecology in planning. This study presents the outcomes of unplanned urbanization and possible preventive measures, which could be taken in the Kartal District, Istanbul-Turkey. Moreover, in this study, different kinds of urban habitats within the frontiers of Kartal were described and an inventorial study containing native, exotic and cultivated plant taxa were realized. For this plant inventory of the Kartal District, all the greenery in the area were explored in different seasons. Plant samples were collected, dried, labelled and then determined according to standard herbarium procedures.
    [Show full text]
  • Ornithogalum Sigmoideum Ve Trachystemon Orientalis' in Acanthamoeba Castellanii KİSTLERİ Ve TROFOZOİTLERİ ÜZERİNE İN VİTRO AMOEBİSİDAL AKTİVİTELERİNİN ARAŞTIRILMASI
    T.C. ORDU ÜNİVERSİTESİ FEN BİLİMLERİ ENSTİTÜSÜ Ornithogalum sigmoideum ve Trachystemon orientalis' in Acanthamoeba castellanii KİSTLERİ ve TROFOZOİTLERİ ÜZERİNE İN VİTRO AMOEBİSİDAL AKTİVİTELERİNİN ARAŞTIRILMASI BÜLENT KAYNAK YÜKSEK LİSANS TEZİ ORDU 2017 ÖZET Ornithogalum sigmoideum ve Trachystemon orientalis' in Acanthamoeba castellanii KİSTLERİ ve TROFOZOİTLERİ ÜZERİNE İN VİTRO AMOEBİSİDAL AKTİVİTELERİNİN ARAŞTIRILMASI Bülent KAYNAK Ordu Üniversitesi Fen Bilimleri Enstitüsü Moleküler Biyoloji ve Genetik Anabilim Dalı, 2017 Yüksek Lisans Tezi, 69s. Danışman: Doç. Dr. Zeynep KOLÖREN Serbest yaşayan amiplerden Acanthamoeba türleri, Acanthamoeba keratiti, Granülomatöz amibik ensefalit, Kutanöz acanthamoebiasis gibi önemli hastalıkların etkenidir. Serbest yaşayan amip kaynaklı enfeksiyon olgularının tamamen tedavi edilmesi oldukça zordur. Trofozoitler, olumsuz şartlarda kistleştiklerinden yeterli ve etkili olmayan tedavilerde enfeksiyon genellikle tekrarlayabilmektedir. Günümüze kadar aşılagelen tedavi yöntemlerine karşı kistlerin trofozoitlerden daha fazla direnç göstermesi, Acanthamoeba enfeksiyonlarında etkili ilaç kombinasyolarının uygulandığı tedavilerde ciddi yan etkilerin görülmesi, mevcut ilaçların istenilen aktivite ve selektiviteye sahip olmaması gibi nedenler tıp dünyasını yeni ve daha etkili ilaç arayışına yönlendirmiştir. Bu çalışmada, O. sigmoideum ve T. orientalis bitkilerinden elde edilen ringer ve metanol özütlerinin Acanthamoeba castellanii kist ve trofozoitleri üzerindeki yüzde (%) canlılık etkisi ve IC50 değeri araştırılmıştır. A. castellanii trofozoit formu üzerindeki IC50 değeri 72., 48., 24., 8., 6., 3. ve 1. saatlerde sırasıyla, O. sigmoideum’un metanol özütünde, 2.9, 6.6, 8.3, 10.1, 12.1, 15.6 ve 23.2 mg/ml, O. sigmoideum’un ringer özütünde, 7.3, 8.4, 8.9, 11.9, 13.3, 16.7 ve 24.2 mg/ml, T.orientalis’in metanol özütünde, 4, 7.2, 8.7, 11.1, 14.1, 21.4 ve 23.8 mg/ml, T.
    [Show full text]
  • The Aromatic-Medicinal Plant Taxa of Pure Scots Pine Stands in Sürmene - Camburnu (Trabzon)
    Int. J. Sec. Metabolite, Vol. 4: 3 (2017) pp. 517-529 Special Issue 2: Research Article ISSN: 2148-6905 online Journal homepage: http://www.ijate.net/index.php/ijsm The Aromatic-Medicinal Plant Taxa of pure Scots pine stands in Sürmene - Camburnu (Trabzon) Arzu ERGÜL BOZKURT *1, Salih TERZİOĞLU2 1Artvin Çoruh University, Faculty of Forestry, Artvin, Turkey 2Karadeniz Technical University, Faculty of Forestry, Trabzon, Turkey Received: 04 May 2017 – Revised: 21 September 2017 - Accepted: 03 November 2017 Abstract: Forests not only produce wood raw material ecosystems, but also that have rich medicinal and aromatic plants. In addition to this situation, forests have been produced many goods and services. Regulation of protection- utilization balance with wood and non-wood herbal products is very important to these ecosystems. In order to ensure sustainable utilization of forest ecosystems, first of all, it is necessary to identify the natural resource components in these ecosystems. In this study, medicinal-aromatic plants of natural Scots pine (Pinus sylvestris L.) stands, which are a sensitive ecosystem, were investigated in 2013-2014. Pinus sylvestris has special ecological conditions in Sürmene-Çamburnu (Trabzon) region because in this region Pinus sylvestris is descending down to the beach. In the study, 81 (77%) out of 105 vascular plant taxa were found to have medicinal-aromatic potential. The parts of the identified plant taxa used for different medical and aromatic purposes are explained in detail. In addition, recommendations were made about regulation of utilization in sensitive ecosystems. Keywords: Pinewood, medicinal-aromatic plant, flora 1. INTRODUCTION Forests are the ecosystems that produce not only wood raw material but as well non-wood forest products.
    [Show full text]
  • The Bees (Apidae, Hymenoptera) of the Botanic Garden in Graz, an Annotated List 19-68 Mitteilungen Des Naturwissenschaftlichen Vereines Für Steiermark Bd
    ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Mitteilungen des naturwissenschaftlichen Vereins für Steiermark Jahr/Year: 2016 Band/Volume: 146 Autor(en)/Author(s): Teppner Herwig, Ebmer Andreas Werner, Gusenleitner Fritz Josef [Friedrich], Schwarz Maximilian Artikel/Article: The bees (Apidae, Hymenoptera) of the Botanic Garden in Graz, an annotated list 19-68 Mitteilungen des Naturwissenschaftlichen Vereines für Steiermark Bd. 146 S. 19–68 Graz 2016 The bees (Apidae, Hymenoptera) of the Botanic Garden in Graz, an annotated list Herwig Teppner1, Andreas W. Ebmer2, Fritz Gusenleitner3 and Maximilian Schwarz4 With 65 Figures Accepted: 28. October 2016 Summary: During studies in floral ecology 151 bee (Apidae) species from 25 genera were recorded in the Botanic Garden of the Karl-Franzens-Universität Graz since 1981. The garden covers an area of c. 3.6 ha (buildings included). The voucher specimens are listed by date, gender and plant species visited. For a part of the bee species additional notes are presented. The most elaborated notes concern Hylaeus styriacus, three species of Andrena subg. Taeniandrena (opening of floral buds for pollen harvest,slicing calyx or corolla for reaching nectar), Andrena rufula, Andrena susterai, Megachile nigriventris on Glau­ cium, behaviour of Megachile willughbiella, Eucera nigrescens (collecting on Symphytum officinale), Xylocopa violacea (vibratory pollen collection, Xylocopa-blossoms, nectar robbing), Bombus haematurus, Nomada trapeziformis, behaviour of Lasioglossum females, honeydew and bumblebees as well as the flowers ofViscum , Forsythia and Lysimachia. Andrena gelriae and Lasioglossum setulosum are first records for Styria. This inventory is put in a broader context by the addition of publications with enumerations of bees for 23 other botanic gardens of Central Europe, of which few are briefly discussed.
    [Show full text]
  • Reproductive Characteristics As Drivers of Alien Plant Naturalization and Invasion
    Reproductive characteristics as drivers of alien plant naturalization and invasion Dissertation submitted for the degree of Doctor of Natural Sciences presented by Mialy Harindra Razanajatovo at the Faculty of Sciences Department of Biology Date of the oral examination: 12 February 2016 First referee: Prof. Dr. Mark van Kleunen Second referee: Prof. Dr. Markus Fischer Konstanzer Online-Publikations-System (KOPS) URL: http://nbn-resolving.de/urn:nbn:de:bsz:352-0-324483 Summary Due to human activity and global movements, many plant species have been introduced to non-native regions where they experience novel abiotic and biotic conditions. Some of these alien species manage to establish reproducing naturalized populations, and some naturalized alien species subsequently become invasive. Invasion by alien plant species can negatively affect native communities and ecosystems, but what gives the alien species an advantage under novel conditions is still not clear. Therefore, identifying the drivers of invasions has become a major goal in invasion ecology. Reproduction is crucial in plant invasions, because propagule supply is required for founding new populations, population maintenance and spread in non-native regions. Baker’s Law, referring to the superior advantage of species capable of uniparental reproduction in establishing after long distance dispersal, has received major interest in explaining plant invasions. However, previous findings regarding Baker’s Law are contradicting. Moreover, there has been an increasing interest in understanding the integration of alien plant species into native plant-pollinator networks but few studies have looked at the pollination ecology of successful (naturalized and invasive) and unsuccessful (non-naturalized and non-invasive) alien plant species.
    [Show full text]
  • T.C. Kastamonu Üniversitesi Fen Bilimleri Enstitüsü Orman Mühendisliği Ana Bilim Dali
    T.C. KASTAMONU ÜNİVERSİTESİ FEN BİLİMLERİ ENSTİTÜSÜ ORMAN MÜHENDİSLİĞİ ANA BİLİM DALI YENİCE YABAN HAYATI GELİŞTİRME SAHASI’NIN BİTKİ SOSYOLOJİSİ YÖNÜNDEN ARAŞTIRILMASI AYŞE ÖZTÜRK DOKTORA TEZİ DANIŞMAN Dr. Öğr. Üyesi KERİM GÜNEY KASTAMONU 2018 T.C. KASTAMONU ÜNİVERSİTESİ FEN BİLİMLERİ ENSTİTÜSÜ YENİCE YABAN HAYATI GELİŞTİRME SAHASI’NIN BİTKİ SOSYOLOJİSİ YÖNÜNDEN ARAŞTIRILMASI Ayşe ÖZTÜRK Danışman Dr. Öğr. Üyesi Kerim GÜNEY Jüri Üyesi Prof. Dr. Fatmagül GEVEN Jüri Üyesi Prof. Dr. Sibel ATASAĞUN Jüri Üyesi Doç. Dr. Barış BANİ Jüri Üyesi Dr. Öğr. Üyesi Nurcan YİĞİT DOKTORA TEZİ ORMAN MÜHENDİSLİĞİ ANA BİLİM DALI KASTAMONU – 2018 ÖZET Doktora Tezi YENİCE YABAN HAYATI GELİŞTİRME SAHASI’NIN BİTKİ SOSYOLOJİSİ YÖNÜNDEN ARAŞTIRILMASI Ayşe ÖZTÜRK Kastamonu Üniversitesi Fen Bilimleri Enstitüsü Orman Mühendisliği Ana Bilim Dalı Danışman: Dr. Öğr. Üyesi Kerim GÜNEY Araştırma alanı, coğrafi açıdan Batı Karadeniz Bölgesi’nde, Karabük ili sınırları içerisinde Karabük’ün en batıdaki ilçesi konumunda olan Yenice’de bulunan Yenice Yaban Hayatı Geliştirme Sahası’dır. Yenice YHGS, biyocoğrafik açıdan Avrupa Sibirya Fitocoğrafik Bölgesi’nin Öksin (Euxine) provensinin batı sektörü içerisinde yer almaktadır. Çalışma alanının bulunduğu iklim Sub-Akdeniz’li olarak belirlenmiş olup Karadeniz öncesi bölge ile Asıl Karadeniz bölgesi arasında bir geçiş bölgesi olarak tanımlanabilir. Yenice YHGS’nda 445 takson tespit edilmiştir. Araştırma alanı biyocoğrafik konumu itibariyle orman, pseudomaki ve akarsu ekosistemi gibi çok sayıda ekosistemi içinde barındırmakla birlikte hakim vejetasyon tipi orman formasyonudur. Braun-Blanquet metoduna göre orman vejetasyonuna ait toplamda 4 bitki birliği tespit edilmiştir. Tespit edilen birliklerden Corno mas-Quercetum ibericae ass. nova bilim dünyası için yenidir. Sintaksonlar ve bağlı oldukları sosyolojik birimler şu şekildedir: Sınıf : Querco-Fagetea (Br.-Bl and Vlieger 1937) Fuk.
    [Show full text]
  • Aizoales 3-663.20.00
    Aizoales 3-663.20.00 Taxonomy Introduction In the Apg2 classifcation Te suborder is recognised with Lo- Tey have a tendency to give a lot in order to be accepted. Tey phiocarpaceae, Barbeuiaceae, Aizoaceae, Gisekiaceae, Nyctag- want to belong to the group, the family and in order to do so inaceae, Phytolaccaceae and Sarcobataceae. they have a tendency to adapt, to give in. Because they are high- ly sensitive they accurately feel what the others want and need Plant theory and can easily adapt to that. It is only when they feel placed In the Plant theory the above clade is given the name Aizoales. outside of the group that they can become angry. It feels like a Aizoales is placed in Phase 2 of the Caryophyllidae. basic need of life to be accepted but it is difcult for them to feel In the frst version the above Families were placed in the sever- completely accepted as their inner life is ofen felt as peculiar al Subphases. and strange. Tey feel weird in a strange world. Tey feel very In Plant theory 2 only Aizoaceae is lef inPhase 2. religious, a connection with the spiritual world and God and Te other Families are transferred toPhase 3. that connection is ofen not very well accepted in society. Due to their inner convictions they can get in confict with society. Subphases Mostly their solution is to keep their opinions and feelings to 1. Sesuvioideae Aizoaceae themselves; they prefer to avoid the conficts. Tey hope to be 2. Drosanthemoideae Aizoaceae be able to stay with their own inner convictions and that they 3.
    [Show full text]
  • Annual Report 2017–2018
    Annual Report 2017 –2018 Cambridge University Botanic Garden (CUBG) supports University teaching and research while also being a place of enjoyment and inspiration to visitors of all ages. The Botanic Garden is one of the largest University-owned botanic gardens in the world. Opened to the public in 1846, the 40-acre Garden has an unparalleled living collection of over 8,000 species, including nine National Collections, with glasshouses, experimental plots, lake, herbarium and botanical library. The Garden offers year-round inspiration for gardeners as well as an exciting introduction to the natural world through a programme of family, school and adults’ activities and events. CUBG collaborates with national and international researchers from a wide range of partner organisations, including universities, conservation bodies and botanic gardens worldwide, to promote the conservation and scientific understanding of global plant biodiversity. N i g e l L u c k h u r s Director’s Report t The core goals of Cambridge University Botanic Garden stay constant from year to year – to support a globally excellent network of research and teaching around plants while providing stimulating opportunities for educational groups and visitors to engage with the collections. But in some years our focus is on particular aspects of those goals, and in some years life has other plans for us! This year our focus has been on the fruition of our“Understanding Plant Diversity” project, to reinvigorate the scientific value, horticultural quality, interpretation and engagement value of our wonderful Systematic Beds. But at the same time, the hottest and driest summer for several decades pulled our attention firmly back to our horticultural standards and our responsibility to lead by example when it comes to sustainable horticulture.
    [Show full text]
  • BSBI News April 2017 No
    BSBI News April 2017 No. 135 Edited by Trevor James & Gwynn Ellis ISSN 2397-8813 Small form of Conyza floribunda with corymbose inflorescence and near simple main stem leaves. Sprowston, Norfolk (v.c.27). Photo Bob Leaney © 2012 (p. 7) Tiny form of Conyza canadensis (c.20 cm high) with atypical inflorescence structure and simple lower stem leaves. Wroxham, Norfolk (v.c.27). Photo Bob Leaney © 2006 (p. 7) Carex oederi in Devil’s Hole, Sefton Coast Platanthera ×hybrida, Ard Dorch, Skye (v.c.104). (v.c.59). Photo P.H. Smith © 2016 (p. 22) Photo T. Swainbank © 2016 (p. 51) CONTENTS BSBI Atlas 2020 Extracting Records from the Scottish Coordinator’s Corner.......................P. Stroh 2 Saltmarsh Survey.................S. Bungard 63 Important Notices Digital plant photography.........J. Presland 64 From the President.....................J. Faulkner 4 Adventives & Aliens News, 11....M. Berry 67 New Journal of Botany...............I. Denholm 5 Phyla nodiflora var. minor discovered in v.c.5 Notes from the Editors......T. James & G. Ellis 6 .........................................R. FitzGerald 69 Notes.......................................................... 7-78 Dorycnium hirsutum (Canary Clover) in Common problems with identification in Britain and Ireland...................M. Berry 71 Conyza.....................................B. Leaney 7 Cardamine occulta, another small white- New Altitudinal Limit for Taxus baccata flowered weedy brassica ............................................R.A. Dalton 18 .......................E. Cooke & J. Heathcote 73 Hieracium sowadeense re-discovered in Pachyphragma macrophyllum naturalised by Orkney..................................J. Crossley 18 streamsides in v.c.64..............K. Walker 75 Unusual feature of Carex lepidocarpa Lotus ornithopodioides in Surrey (v.c.17) ........................M. Duffell & D. Wallace 20 .........................................G. Hounsome 77 Carex oederi on the Sefton Coast (v.c.59) Baccharis halmifolia – a response.
    [Show full text]
  • An Ecological Database of the British Flora
    An Ecological Database of the British Flora submitted by Helen Jacqueline Peat for examination for the degree Doctor of Philosophy Department of Biology University of York October 1992 Abstract The design and compilation of a database containing ecological information on the British Flora is described. All native and naturalised species of the Gymnospermae and Angiospermae are included. Data on c.130 characteristics concerning habitat, distribution, morphology, physiology, life history and associated organisms, were collected by both literature searching and correspondence with plant ecologists. The evolutionary history of 25 of the characteristics was investigated by looking at the amount of variance at each taxonomic level. The variation in pollination mechanisms was found at high taxonomic levels suggesting these evolved, and became fixed, early on in the evolution of flowering plants. Chromosome number, annualness, dichogamy and self-fertilization showed most variance at low taxonomic levels, suggesting these characteristics have evolved more recently and may still be subject to change. Most of the characteristics, however, eg. presence of compound leaves, height and propagule length showed variance spread over several taxonomic levels suggesting evolution has occurred at different times in different lineages. The necessity of accounting for phylogeny when conducting comparative analyses is discussed, and two methods allowing this are outlined. Using these, the questions: 'Why does stomatal distribution differ between species?' and 'Why do different species have different degrees of mycorrhizal infection?' were investigated. Amphistomaty was found to be associated with species of unshaded habitats, those with small leaves and those with hairy leaves, and hypostomaty with woody species, larger leaves and glabrous leaves.
    [Show full text]
  • Flora News Newsletter of the Hampshire & Isle of Wight Wildlife Trust’S Flora Group No
    Flora News Newsletter of the Hampshire & Isle of Wight Wildlife Trust’s Flora Group No. 58 Spring 2020 Published January 2020 In This Issue Forthcoming Events ................................................................................................................................................... 2 Reports of Recent Events .......................................................................................................................................... 5 Features Monitoring the success of the New Forest Non-Native Plants Project ...................... Catherine Chatters ........ 11 Recording What has Atlas 2020 told us about Hampshire’s flora? ............................................. Martin Rand ...................12 WANTED: A Joint BSBI Recorder for South Hampshire ............................................ Martin Rand ...................18 BSBI Atlas 2020 in Hampshire ................................................................................... Tony Mundell .................20 Bryophyte Recording in Hampshire and the Isle of Wight – January 2020 update ... John Norton ...................21 VC11 Notes and Records ........................................................................................... Martin Rand ...................24 VC12 Records ............................................................................................................ Tony Mundell .................34 Membership and Recording Information .................................................................................................................
    [Show full text]
  • Evolutionary Relationships, Taxonomy, and Patterns of Character Evolution
    Cladistics Cladistics (2013) 1–31 10.1111/cla.12036 A phylogenetic analysis of morphological and molecular characters of Boraginaceae: evolutionary relationships, taxonomy, and patterns of character evolution James I. Cohen* Texas A&M International University, 379D LBVSC, 5201 University Blvd, Laredo, TX, 78041, USA Accepted 22 April 2013 Abstract The angiosperm family Boraginaceae includes ca. 1600 species distributed among ca. 110 genera. Some floral features are con- stant within the family, but many vegetative, floral, pollen, and nutlet traits vary. Utilizing 224 species of Boraginaceae and related taxa, five matrices were constructed with various combinations of morphological characters, three chloroplast DNA regions, and one nuclear ribosomal DNA region. Phylogenetic analyses were conducted for these matrices, and patterns of char- acter evolution were examined. Boraginaceae is resolved as monophyletic, with Wellstedia as its sister. Codon is sister to Bora- ginaceae + Wellstedia. Although most of the investigated morphological characters have a low consistency index, particular character states are synapomorphies for large clades in each of the tribes of the family. In Boraginaceae, the breeding system heterostyly evolved at least 12 times, which is the largest number of origins resolved in any family; therefore Boraginaceae can serve as a model for the evolution and development of heterostyly. Nutlet ornamentation is most diverse in Cynoglosseae and Trichodesmeae, while pollen and floral features are most variable in Boragineae and Lithospermeae. Phylogenetic relationships and patterns of character evolution identified in the present study set the stage for future work creating an updated taxonomic system of Boraginaceae. © The Willi Hennig Society 2013. Introduction and Heliotropioideae) or families that are character- ized by a scorpioid cyme and two-parted gynoecium The angiosperm family Boraginaceae includes ca.
    [Show full text]