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Home , Achimenes, Acmella, Adoxa, Aegiphila, Aiouea, Alfaroa

Vascular List Excluding: Pteriodophyte ( & Allies) & Orchidaceae (Orchids)

Higher Classification1 Kingdom: Plantae, Phylum: Tracheophyta

Class (C:), (O:) and (F:) Scientific Name1 English Name2 C: Equisetopsida (Horsetails) See & Allies Species List3 C: Filicopsida See Fern & Allies Species List3 (Leptosporangiate Ferns) C: Isoetopsida (Isotopsids) See Fern & Allies Species List3 C: Liliopsida (Monocots) O: F: Araceae (Arum) Anthurium concinnatum (Flamingo Plant) Anthurium salvinii ’s-nest Anthurium Colocasia sp. Monstera deliciosa Willowleaf Monstera dissecta Monstera epipremnoides Syngonium sp. O: F: (Palm) Chamaedorea pittieri Chamaedorea sp. sp. Geonoma undata subsp.

edulis Prestoea longepetiolata O: F: Orchidaceae (Orchid) See Orchid Species List4 O: Commelinaceae F: Commelinaceae Commelina sp. (Dayflower) (Spiderwort) Tradescantia poelliae (Spiderwort) Tradescantia zanonia (Spiderwort) O: F: Cyclanthaceae Cyclanthaceae sp. (-hat) O: F: Liliaceae (Lily) Lilium longiflorum Easter Lily O: (Grasses) F: nutans Nodding Strap Airplant (Bromeliad) Tillandsia leiboldiana (Airplant) Tillandsia punctulata (Airplant) spp. F: (Sedge) Cyperus papyrus Papyrus Kyllinga sp. Rhynchospora sp. F: Poaceae Brachiaria cf. ruziziensis Congo Grass (True Grasses) Digitaria costaricensis

Page 1 of 10 Cloudbridge Nature Reserve, Last Updated: March 9, 2017 Species List Excluding: Pteriodophyte (Ferns & Allies) & Orchidaceae (Orchids)

Class (C:), Order (O:) and Family (F:) Scientific Name1 English Name2 C: Liliopsida (Monocots) (cont’d) O: Poales (Grasses) (cont’d) F: Poaceae Guadua sp. (True Grasses) (cont’d) O: F: Cannaceae Canna tuerckheimii Maraca Roja (Indian Shot) F: Costaceae Costus wilsonii Spiral Flag F: Heliconiaceae beckneri (Heliconia) (Lobster-Claw) Heliconia lankesteri (Heliconia) Heliconia rostrata Lobster Claw Heliconia tortuosa (Heliconia) Heliconia wagneriana (Heliconia) F: Maranthaceae sp. F: Zingiberaceae (Ginger) Etlingera elatior Kantan Renealmia alpinia Jenjibre-de-jardin C: Lycopodiopsida See Fern & Allies Species List3 (Clubmosses) C: Magnoliopsida (Dicots) O: F: Apiaceae (Umbel) Sanicula liberta Pacific Blacksnakeroot F: Araliaceae (Ivy) Dendropanax arboreus Angelica Oreopanax sp. Oreopanax standleyi Oreopanax xalapensis Schefflera rodriguesiana Schefflera, Umbrella Tree O: F: costaricensis O: F: sp. (Spotflower) (Aster & Daisy) Ageratum conyzoides Tropical Whiteweed schiedeana Bidens pilosa Beggar's Tick mollis Soft Elephantsfoot fosbergii Florida Tasselflower Emilia sonchifolia Lilac Tasselflower sp. (Gallant-soldier) Heliopsis buphthalmoides (Sun Glory) hirta Lasianthaea fruticosa (Lasianthaea) Koanophyllon pittieri (Throughwort) Melanthera nivea Snow Squarestem Senecio copeyensis (Ragwort) Senecio grandifolius (Ragwort)

Page 2 of 10 Cloudbridge Nature Reserve, Costa Rica Last Updated: March 9, 2017 Vascular Plant Species List Excluding: Pteriodophyte (Ferns & Allies) & Orchidaceae (Orchids)

Class (C:), Order (O:) and Family (F:) Scientific Name1 English Name2 C: Magnoliopsida (Dicots) (cont’d) O: Asterales (cont’d) F: Asteraceae Verbesina gigantea (Crownbeard) (Aster & Daisy) (cont’d) Verbesina turbacensis (Crownbeard) F: Campanulaceae Centropogon granulosus (Bellflower) Centropogon gutierrezii O: F: () latifolia sp.

Wigandia urens Caracus , Wigandia O: F: Amaranthaceae Pleuropetalum sprucei (Pigweed) F: Arenaria lanuginosa (Chickweed) Drymaria cf. cordata West Indian Drymary Drymaria multiflora Slender Drymary F: Phytolaccaceae Ledenbergia peruviana (Pokeweed) O: F: Celastraceae Salacia petensis Guava (Stackhousia) Zinowiewia integerrima Golden Root O: F: Cornaceae (Dogwood) Cornus disciflora (Dogwood) F: Loasaceae (Loasa) Nasa triphylla O: F: Begoniaceae (Begonia) Begonia convallariodora (Begonia) Begonia involucrata (Begonia) Begonia semiovata (Begonia) F: (Gourd) Cyclanthera sp. O: F: (Adoxa) Viburnum costaricanum (Viburnum) O: F: Actinidiaceae Saurauia montana (Kiwi- & Chinese Saurauia pittieri Moquillo, Snot Tree Gooseberry) Saurauia rubiformis Snot Tree F: Balsaminaceae (Balsam) walleriana Buzzy Lizzy, Busy Lizzie F: Clethraceae Clethra mexicana (Clethra) F: (Heather) Cavendishia callista meiantha F: Pentaphylacaceae Freziera candicans Coral F: (Primrose) Ardisia compressa (Marlberry) Clavija cf. biborrana Myrsine coriacea Leathery Colicwood, Little Mouse

Page 3 of 10 Cloudbridge Nature Reserve, Costa Rica Last Updated: March 9, 2017 Vascular Plant Species List Excluding: Pteriodophyte (Ferns & Allies) & Orchidaceae (Orchids)

Class (C:), Order (O:) and Family (F:) Scientific Name1 English Name2 C: Magnoliopsida (Dicots) (cont’d) O: Ericales (cont’d) F: Primulaceae (Primrose) Myrsine dependens (Colicwood) (cont’d) Parathesis cartagoana F: Sapotaceae (Milkwood) Manilkara chicle Manilkara zapota Sapote

Pouteria fossicola F: Styracaceae (Storax) Styrax warscewiczii (Snowbell) F: (Tea) fruticosa (Gordonia) O: F: (Pea) pintoi Pinto Dalbergia sp. (Indian Rosewood) Desmodium sp. (Ticktrefoil) Diphysa americana Erythrina cf. poeppigiana Mountain Immortelle Erythrina costaricensis (Sykes Coral Tree) Erythrina lanceolata Machete sepium Gliricidia Inga oerstediana Guaba Inga sierrae (Ice Cream Bean), Wild Avocado Lysiloma divaricatum Mauto Mimosa pudica Herbe Sensible Mucuna urens Horse-eye Beans, Ox Eye Phaseolus cf. lunatus Lima Bean Styphnolobium monteviridis Vigna sp. (Cowpea) F: (Milkwort) Monnina sp. O: F: Betulaceae () Alnus acuminata Alder F: Fagaceae () Quercus benthamii () Quercus bumelioides White Oak Quercus copeyensis Black Oak Quercus costaricensis White Oak Quercus gulielmitreleasei (Oak) Quercus insignis (Oak) Quercus oleoides (Oak) Quercus oocarpa (Oak) Quercus salicifolia (Oak) Quercus seemannii (Oak) F: (Walnut) Alfaroa costaricensis F: Myricaceae (Waxberry) Morella sp. (Bayberry)

Page 4 of 10 Cloudbridge Nature Reserve, Costa Rica Last Updated: March 9, 2017 Vascular Plant Species List Excluding: Pteriodophyte (Ferns & Allies) & Orchidaceae (Orchids)

Class (C:), Order (O:) and Family (F:) Scientific Name1 English Name2 C: Magnoliopsida (Dicots) (cont’d) O: F: Apocynaceae Apocynaceae sp. (Dogbane) Asclepias curassavica Tropical Milkweed F: (Coffee) Chione sylvicola pubescens Cosmibuena macrocarpa Elaeagia auriculata Gonzalagunia rosea Gonzalagunia ovatifolia Guettarda crispiflora poasana Hoffmannia cf. congesta Jicarita Palicourea purpurea (Cappel) Posoqueria latifolia Psychotria aubletiana Psychotria horizontalis Psychotria sylvivaga Randia sp. (Indigoberry) Rogiera amoena Spermacoce alata Broadleaf Buttonweed Spermacoce remota Woodland False Buttonweed O: F: Geraniaceae Geranium guatemalense (Crane’s-Bill) (Geranium) O: F: Gunneraceae Gunnera insignis Poor Man’s Umbrella O: F: Aphelandra cf. scabra (Black Mangrove) Hansteinia blepharorhachis Polkadot Plant, Measles Plant, Freckle Face aurea (Water-) Megaskepasma Brazilian Red-cloak erythrochlamys Pachystachys lutea Lollipop-plant Poikilacanthus macranthus Pseuderanthemum sp. Razisea spicata Stenostephanus sp. F: Amphilophium crucigerum Monkey Comb (Jacaranda) Amphitecna sessilifolia Black Calabash Spathodea campanulata African Tulip Tree, Flame of the Forest Trumpetflower, Yellow Cedar, Ginger Tecoma stans Thomas F: Calceolariaceae Calceolaria tripartita Splitleaf Slipperwort

Page 5 of 10 Cloudbridge Nature Reserve, Costa Rica Last Updated: March 9, 2017 Vascular Plant Species List Excluding: Pteriodophyte (Ferns & Allies) & Orchidaceae (Orchids)

Class (C:), Order (O:) and Family (F:) Scientific Name1 English Name2 C: Magnoliopsida (Dicots) (cont’d) O: Lamiales (cont’d) F: Achimenes sp. (African Violet) Besleria cf formosa Columnea polyantha Diastema racemiferum Drymonia sp. spicata (Kohleria) Moussonia deppeana F: Aegiphila sp. (Spiritweed) (Mint & Basil) Salvia cacaliifolia (Sage) F: Phrymaceae (Phrymas) Hemichaena fruticosa F: Plantaginaceae Plantago australis Mexican Plantain (Plantain) F: (Verbena) Citharexylum caudatum Citharexylum donnell-smithii Lantana camara Largeleaf Lantana Lippia umbellata Stachytarpheta frantzii Purple Porterweed Verbena litoralis Seashore Vervain O: F: (Laurel) Aiouea costaricensis Beilschmiedia sp. brenesii (Cinnamon) Cinnamomum triplinerve Mexican Cinnamon Nectandra sp. (Sweetwood) Ocotea insularis Laurel Sassafras, Little Avocado Ocotea laetevirens (Sweetwood), Little Avocado Ocotea stenoneura (Sweetwood), Little Avocado Persea americana Avocado, Alligator Pear Persea caerulea Wild Avocado Persea schiedeana (Bay) F: Monimiaceae Mollinedia sp. (Lemonwood) F: Siparunaceae Siparuna sp. Limoncita O: F: Annonaceae Annona muricata Graviola, Soursop (Custard-apple) Guatteria recurvisepala Guatteria talamancana F: Magnoliaceae Magnolia poasana (Magnolia) Myristica fragrans Nutmeg F: Myristicaceae (Nutmeg) Otoba sp.

Page 6 of 10 Cloudbridge Nature Reserve, Costa Rica Last Updated: March 9, 2017 Vascular Plant Species List Excluding: Pteriodophyte (Ferns & Allies) & Orchidaceae (Orchids)

Class (C:), Order (O:) and Family (F:) Scientific Name1 English Name2 C: Magnoliopsida (Dicots) (cont’d) O: F: Chrysobalanaceae sp. (Coco-plum) F: Chrysochlamys allenii (Mangosteen) sp. (Attorney), Autograph Tree intermedia Symphonia globulifera Ojoru, Boar , Hog Doctor F: Codiaeum variegatum Garden () Croton draco Mexican Dragon’s Blood Euphorbia pulcherrima Poinsettia Pausandra trianae

Sapium glandulosum Gumtree Sapium laurifolium Hichahuevos Sapium pachystachys (Milktree) F: Hypericaceae Vismia baccifera F: Lacistemataceae Lozania mutisiana F: Passifloraceae Grenadia, Sweet Passion Fruit, Sweet Passiflora ligularis (Passion Flower) Granadilla F: Hieronyma alchorneoides Hieronyma oblonga F: (Willow) sylvestris Crackopen Hasseltia floribunda Macrohasseltia

macroterantha Xylosma intermedium F: (Violet) Viola capillaris O: F: (Lindens) Apeiba sp. americanus White Moho Malvaviscus arboreus Turkscap Malvaviscus concinnus Turkscap Mortoniodendron

anisophyllum Pavonia penduliflora (Swampmallow) Sida sp. (Fanpetal) Triumfetta semitriloba Sacramento Burrbark Waltheria glomerata False Matico Wercklea cocleana O: F: Combretaceae Terminalia oblonga Peruvian Almond, Sura (White Mangrove) F: Lythraceae Cuphea cf. appendiculata (Waxweed) (Mangrove Apple) Cuphea sp. (Waxweed)

Page 7 of 10 Cloudbridge Nature Reserve, Costa Rica Last Updated: March 9, 2017 Vascular Plant Species List Excluding: Pteriodophyte (Ferns & Allies) & Orchidaceae (Orchids)

Class (C:), Order (O:) and Family (F:) Scientific Name1 English Name2 C: Magnoliopsida (Dicots) (cont’d) O: Myrtales (cont’d) F: Arthrostemma ciliatum Pinkfringe (Melastoma) Blakea gracilis Centradenia grandifolia Conostegia oerstediana (Snailwood) Conostegia rhodopetala (Snailwood) Conostegia xalapensis (Snailwood) Heterotis sp. (African Arowanas) Leandra subseriata costaricensis (Johnnyberry) Miconia donaeana (Johnnyberry) Miconia doriana (Johnnyberry) Miconia neomicrantha (Johnnyberry) Monochaetum floribundum Tibouchina heteromalla Silverleafed Princess Flower Tibouchina longifolia Longleaf Glorytree F: (Myrtle) stipitata (Stopper) Myrcianthes fragrans Twinberry Psidium guajava Guava Syzygium jambos Rose Apple F: paniculata Shrubby Fuchsia (Evening Primrose) Ludwigia peruviana Primrose Willow O: F: Brunelliaceae Brunellia costaricensis F: Elaeocarpaceae ampla Monkey Comb O: F: Piperaceae (Pepper) Peperomia sp. (Radiator Plant) Piper pseudolanceifolium (Pepper) O: F: Proteaceae (Protea) Panopsis suaveolens F: Sabiaceae Meliosma idiopoda Diamond Meliosma vernicosa O: F: (Poppy) frutescens Tree Poppy O: F: Cannabaceae (Dagga) Celtis trinervia Almex Trema micrantha Jamaican Nettle Tree F: (Fig) Brosimum costaricanum hartwegii (Fig) Ficus pertusa (Fig) Ficus tuerckheimii Florida Strangler Fig sp.

Page 8 of 10 Cloudbridge Nature Reserve, Costa Rica Last Updated: March 9, 2017 Vascular Plant Species List Excluding: Pteriodophyte (Ferns & Allies) & Orchidaceae (Orchids)

Class (C:), Order (O:) and Family (F:) Scientific Name1 English Name2 C: Magnoliopsida (Dicots) (cont’d) O: Rosales (cont’d) F: Moraceae (Fig) (cont’d) affinis F: Roasceae (Rose) Eriobotrya japonica () Prunus annularis (Cherries & Allies) Rubus cf. urticifolius Mora Silvestre Rubus eriocarpus Horse Blackberry Rubus niveus Kala Hinsalu Rubus rosifolius West Indian Raspberry F: Ulmaceae () Ulmus Mexicana Mexican Elm F: (Nettle) angustifolia (Pumpwood) Cecropia obtusifolia Trumpet Tree Cecropia peltata (Pumpwood) Cecropia polyphlebia (Pumpwood) Pilea cf. dauciodora (Clearweed) Urera sp. O: F: Loranthaceae Struthanthus sp. (Showy ) O: F: Anacardiaceae (Rhus) Mauria heterophylla Spondias purpurea Purple Mombin, Red Mombin, Spanish Plum F: Hippocastanaceae Hippocastanaceae sp. (Horse-) F: () tonduzii Spanish Cedar, Sweet Cedar glabra Alligatorwood Guarea kunthiana Mahogany calantas (Redcedar) Trichilia cf. havanensis F: Rutaceae (Citrus) Citrus aurantium Seville Orange Zanthoxylum sp. (Pricklyashes) F: Allophylus psilospermus (Soapberry) Billia rosea Nephelium lappaceum O: F: Convolvulaceae Ipomoea cf. alba Moon Flower (Morning-glory) F: (Potato) americana Bush Violet grandiflora Largeflower Brunfelsia Brugmansia spp. (Angel Trumpets) Cestrum racemosum (Flowering Jessamine)

Cestrum schlechtendalii Poisonous-berry Jaltomata sp. (False )

Page 9 of 10 Cloudbridge Nature Reserve, Costa Rica Last Updated: March 9, 2017 Vascular Plant Species List Excluding: Pteriodophyte (Ferns & Allies) & Orchidaceae (Orchids)

Class (C:), Order (O:) and Family (F:) Scientific Name1 English Name2 C: Magnoliopsida (Dicots) (cont’d) O: Solanales (cont’d) F: Solanaceae (Potato) Solanum aphyodendron (Nightshade) (cont’d) Solanum quitoense Lulo Solanum rovirosanum (Nightshade) Solanum storkii (Nightshade) Solanum torvum Turkey Berry, Devilsfig Witheringia cf. solanacea C: Pinopsida (Conifers) O: Pinales (Conifers) F: Cupressaceae (Cypress) Cupressus lusitanica Mexican Cypress, Cedar of Goa C: Psilotopsida (Whisk Ferns) See Fern & Allies Species List3

NOTES: Short-forms: cf., indicates the individual was difficult to identify and the full species identification is uncertain; sp., indicates one or an unknown number of species of the was identified; spp. indicates multiple species of the taxon were identified. 1, Classifications and scientific names based on The Catalogue of Life (www.catalogueoflife.org). 2, English names in plain text are from the (www.eol.org); names in italics are local usage names or gleaned from other sources; names in brackets are the common name of the species’ as taken from the Encyclopedia of Life. 3, See the Ferns & Allies Species List at www.cloudbridge.org/publications/species-lists/. 4, See the Orchid Species List at www.cloudbridge.org/publications/species-lists/.

Contributors: Laura Antonaru, Marian Barz, Matthijs Bol, Kendall DeLyser, Dario Elliott, Clarice Esch, Tom Gode, Neža Gorše, Robert Fraser, Marloes Fröling, Tijmen Hoogendijk, Olli Isotupa, Stephen Lehmann, Ana Mazzolari, Jacqueline Medrano, Wouter Meijboom, Kate Moss, Duncan Nicol, Ivo Polach, Mario García Quesada, Anna Rabone, Michiel Spek, Christian Staunton, Joseph Taylor, Debbie Valliere, Britt van Engelen, Lara Van Meter, Dennis Vroomen, Sama Winder, and Cara Wolinsky.

Page 10 of 10 Cloudbridge Nature Reserve, Costa Rica Last Updated: March 9, 2017