A Pangolin Also Scaly Anteater Or Trenggiling, Is a Mammal of the Order Pholidota

Total Page:16

File Type:pdf, Size:1020Kb

A Pangolin Also Scaly Anteater Or Trenggiling, Is a Mammal of the Order Pholidota A pangolin also scaly anteater or Trenggiling, is a mammal of the order Pholidota. There is only one extant family (Manidae) and one genus (Manis) of pangolins, comprising eight species. There are also a number of extinct taxa. Pangolins have large keratin scales covering their skin and are the only mammals with this adaptation. They are found in tropical regions of Africa and Asia. The name "pangolin" derives from the Malay word pengguling ("something that rolls up"). Pangolins are nocturnal animals, and use their well-developed sense of smell to find insects. The long-tailed pangolin is also active by day. Pangolins spend most of the daytime sleeping, curled up into a ball.[3] Pangolins were classified with various other orders, for example Xenarthra, which includes the ordinary anteaters, sloths, and the similar-looking armadillos. But newer genetic evidence[4] indicates that their closest living relatives are the Carnivora, with which they form the clade Ferae.[5] Some paleontologists have classified the pangolins in the order Cimolesta, together with several extinct groups. Laurasiath Eulipotyphla eria Scrotife Chiroptera ra Fereuungul Fera Pholidota ata e Carnivora Perissodacty la Cetartiodact yla Contents [hide] • 1 Physical description and behaviour • 2 Diet • 3 Reproduction • 4 Threats • 5 Taxonomy [citation needed] • 6 References • 7 External links [edit] Physical description and behaviour The physical appearance of pangolins is marked by large, hardened, plate-like scales. The scales, which are soft on newborn pangolins but harden as the animal matures, are made of keratin, the same material of which human fingernails and tetrapod claws are made. The pangolin is often compared to a walking pine cone or globe artichoke. It can curl up into a ball when threatened, with its overlapping scales acting as armour and its face tucked under its tail. The scales are razor-sharp, providing extra defence. The front claws are so long that they are unsuited for walking, and so the animal walks with its fore paws curled over to protect them. Pangolins can also emit a noxious smelling acid from glands near the anus, similar to the spray of a skunk. Pangolins have short legs, with sharp claws which they use for burrowing into termite and ant mounds, as well as climbing. Tree Pangolin and Chinese Pangolin skeletons on exhibit at The Museum of Osteology. The size of pangolins varies by species, ranging from 30 cm to 100 cm (12 to 39 inches). Females are generally smaller than males. The tongues of pangolins are extremely elongated and extend into the abdominal cavity. By convergent evolution pangolins, the giant anteater, and the tube-lipped nectar bat, all have tongues which are unattached to their hyoid bone and extend past their pharynx deep into the thorax.[6] This extension lies between the sternum and the trachea. Large pangolins can extend their tongues as much as 40 cm (16 inches), with a diameter of only 0.5 cm (1/4 inch).[3] In pangolins, the section of the brain that relates to problem solving is highly developed. Although their problem solving ability is primarily used to find food in obscure locations, when kept in captivity pangolins are remarkable escape artists.[citation needed]. Arboreal pangolins live in hollow trees, whereas the ground dwelling species dig tunnels underground, up to a depth of 3.5 m (11 feet).[3] Pangolins are also good swimmers.[3] [edit] Diet A drawing of a pangolin Pangolins lack teeth and the ability to chew. Instead, they tear open anthills or termite mounds with their powerful front claws and probe deep into them with their very long tongues. Pangolins have glands in their chests to lubricate the tongue with sticky, ant-catching saliva. Some species, such as the Tree Pangolin, use their strong, prehensile tails to hang from tree branches and strip away bark from the trunk, exposing insect nests inside. [edit] Reproduction Gestation is 120–150 days. African pangolin females usually give birth to a single offspring at a time, but the Asiatic species can give birth from one to three.[3] Weight at birth is 80–450 g (3–18 ounces), and the scales are initially soft. The young cling to the mother's tail as she moves about, although in burrowing species, they remain in the burrow for the first 2–4 weeks of life. Weaning takes place at around three months of age, and pangolins become sexually mature at two years.[7] [edit] Threats A coat of armor made of pangolin scales, an unusual object presented to George III in 1820. Pangolins are hunted and eaten in many parts of Africa and are one of the more popular types of bush meat. Pangolins are also in great demand in China because their meat is considered a delicacy and some Chinese believe pangolin scales reduce swelling, promote blood circulation and help breast- feeding women produce milk. This, coupled with deforestation, has led to a large decrease in the numbers of Giant Pangolins. In November 2010, pangolins were added to the Zoological Society of London's list of genetically distinct and endangered mammals.[8] Pangolin populations have suffered from illegal trafficking. In May 2007, for example, 31 pangolins were found aboard an abandoned vessel off the coast of China. The boat contained some 5,000 endangered animals.[9] The Guardian provided a description of the killing and eating of pangolins: "A Guangdong chef interviewed last year in the Beijing Science and Technology Daily described how to cook a pangolin: 'We keep them alive in cages until the customer makes an order. Then we hammer them unconscious, cut their throats and drain the blood. It is a slow death. We then boil them to remove the scales. We cut the meat into small pieces and use it to make a number of dishes, including braised meat and soup. Usually the customers take the blood home with them afterwards.'"[9] [edit] Taxonomy[citation needed] The pangolin raids a termite cathedral mound for a meal. • ORDER PHOLIDOTA o Family †Epoicotheriidae o Family †Metacheiromyidae o Family Manidae . Subfamily †Eurotamanduinae . Genus †Eurotamandua . Subfamily Maninae . Genus †Cryptomanis . Genus †Eomanis . Genus †Necromanis . Genus †Patriomanis . Genus Manis . Subgenus Manis . Indian Pangolin (M. crassicaudata) . Chinese Pangolin (M. pentadactyla) . Asian Giant Pangolin (M. paleojavanica) † . Subgenus Paramanis . Sunda Pangolin (M. javanica) . Philippine Pangolin (M. culionensis) . Subgenus Smutsia . Giant Pangolin (M. gigantea) . Ground Pangolin (M. temmincki) . Subgenus Phataginus . Tree Pangolin (M. tricuspis) . Subgenus Uromanis . Long-tailed Pangolin (M. tetradactyla) [edit] References 1. ^ Schlitter, Duane A. (16 November 2005). Wilson, Don E., and Reeder, DeeAnn M., eds. ed. Mammal Species of the World (3rd ed.). Baltimore: Johns Hopkins University Press, 2 vols. (2142 pp.). pp. 530–531. ISBN 978-0-8018- 8221-0. OCLC 62265494. 2. ^ The Encyclopedia of World Wildlife. Paragon Books. 2006. pp. 63. 3. ^ a b c d e Mondadori, Arnoldo Ed., ed (1988). Great Book of the Animal Kingdom. New York: Arch Cape Press. pp. 252. 4. ^ Murphy, Willian J. et al. (2001-12-14). "Resolution of the Early Placental Mammal Radiation Using Bayesian Phylogenetics". Science 294 (5550): 2348–2351. doi:10.1126/science.1067179. PMID 11743200. 5. ^ BioMed Central | Full text | A higher-level MRP supertree of placental mammals 6. ^ Chan, Lap-Ki (1995). "Extrinsic Lingual Musculature of Two Pangolins (Pholidota: Manidae)". Journal of Mammalogy (Journal of Mammalogy, Vol. 76, No. 2) 76 (2): 472–480. doi:10.2307/1382356. JSTOR 1382356. 7. ^ Dickman, Christopher R. (1984). Macdonald, D.. ed. The Encyclopedia of Mammals. New York: Facts on File. pp. 780–781. ISBN 0-87196-871-1. 8. ^ 'Asian unicorn' and scaly anteater make endangered list 9. ^ a b Watts, Johnathan (May 20007). "'Noah's Ark' of 5,000 rare animals found floating off the coast of China". The Guardian. Retrieved 13 August 2011. [edit] External links Wikimedia Commons has media related to: Pholidota Wikispecies has information related to: Pholidota Look up pangolin in Wiktionary, the free dictionary. • Pangolin: Wildlife summary from the African Wildlife Foundation • A photograph of a pangolin • Tree of Life of Pholidota • National Geographic video of a Pangolin • Pangolin rituals in Tanzania • Proceedings of the Workshop on Trade and Conservation of Pangolins Native to south and Southeast Asia (PDF).
Recommended publications
  • Mitochondrial Genomes of African Pangolins and Insights Into Evolutionary Patterns and Phylogeny of the Family Manidae Zelda Du Toit1,2, Morné Du Plessis2, Desiré L
    du Toit et al. BMC Genomics (2017) 18:746 DOI 10.1186/s12864-017-4140-5 RESEARCH ARTICLE Open Access Mitochondrial genomes of African pangolins and insights into evolutionary patterns and phylogeny of the family Manidae Zelda du Toit1,2, Morné du Plessis2, Desiré L. Dalton1,2,3*, Raymond Jansen4, J. Paul Grobler1 and Antoinette Kotzé1,2,4 Abstract Background: This study used next generation sequencing to generate the mitogenomes of four African pangolin species; Temminck’s ground pangolin (Smutsia temminckii), giant ground pangolin (S. gigantea), white-bellied pangolin (Phataginus tricuspis) and black-bellied pangolin (P. tetradactyla). Results: The results indicate that the mitogenomes of the African pangolins are 16,558 bp for S. temminckii, 16,540 bp for S. gigantea, 16,649 bp for P. tetradactyla and 16,565 bp for P. tricuspis. Phylogenetic comparisons of the African pangolins indicated two lineages with high posterior probabilities providing evidence to support the classification of two genera; Smutsia and Phataginus. The total GC content between African pangolins was observed to be similar between species (36.5% – 37.3%). The most frequent codon was found to be A or C at the 3rd codon position. Significant variations in GC-content and codon usage were observed for several regions between African and Asian pangolin species which may be attributed to mutation pressure and/or natural selection. Lastly, a total of two insertions of 80 bp and 28 bp in size respectively was observed in the control region of the black-bellied pangolin which were absent in the other African pangolin species. Conclusions: The current study presents reference mitogenomes of all four African pangolin species and thus expands on the current set of reference genomes available for six of the eight extant pangolin species globally and represents the first phylogenetic analysis with six pangolin species using full mitochondrial genomes.
    [Show full text]
  • The World at the Time of Messel: Conference Volume
    T. Lehmann & S.F.K. Schaal (eds) The World at the Time of Messel - Conference Volume Time at the The World The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment and the History of Early Primates 22nd International Senckenberg Conference 2011 Frankfurt am Main, 15th - 19th November 2011 ISBN 978-3-929907-86-5 Conference Volume SENCKENBERG Gesellschaft für Naturforschung THOMAS LEHMANN & STEPHAN F.K. SCHAAL (eds) The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment, and the History of Early Primates 22nd International Senckenberg Conference Frankfurt am Main, 15th – 19th November 2011 Conference Volume Senckenberg Gesellschaft für Naturforschung IMPRINT The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment, and the History of Early Primates 22nd International Senckenberg Conference 15th – 19th November 2011, Frankfurt am Main, Germany Conference Volume Publisher PROF. DR. DR. H.C. VOLKER MOSBRUGGER Senckenberg Gesellschaft für Naturforschung Senckenberganlage 25, 60325 Frankfurt am Main, Germany Editors DR. THOMAS LEHMANN & DR. STEPHAN F.K. SCHAAL Senckenberg Research Institute and Natural History Museum Frankfurt Senckenberganlage 25, 60325 Frankfurt am Main, Germany [email protected]; [email protected] Language editors JOSEPH E.B. HOGAN & DR. KRISTER T. SMITH Layout JULIANE EBERHARDT & ANIKA VOGEL Cover Illustration EVELINE JUNQUEIRA Print Rhein-Main-Geschäftsdrucke, Hofheim-Wallau, Germany Citation LEHMANN, T. & SCHAAL, S.F.K. (eds) (2011). The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment, and the History of Early Primates. 22nd International Senckenberg Conference. 15th – 19th November 2011, Frankfurt am Main. Conference Volume. Senckenberg Gesellschaft für Naturforschung, Frankfurt am Main. pp. 203.
    [Show full text]
  • Learning About Mammals
    Learning About Mammals The mammals (Class Mammalia) includes everything from mice to elephants, bats to whales and, of course, man. The amazing diversity of mammals is what has allowed them to live in any habitat from desert to arctic to the deep ocean. They live in trees, they live on the ground, they live underground, and in caves. Some are active during the day (diurnal), while some are active at night (nocturnal) and some are just active at dawn and dusk (crepuscular). They live alone (solitary) or in great herds (gregarious). They mate for life (monogamous) or form harems (polygamous). They eat meat (carnivores), they eat plants (herbivores) and they eat both (omnivores). They fill every niche imaginable. Mammals come in all shapes and sizes from the tiny pygmy shrew, weighing 1/10 of an ounce (2.8 grams), to the blue whale, weighing more than 300,000 pounds! They have a huge variation in life span from a small rodent living one year to an elephant living 70 years. Generally, the bigger the mammal, the longer the life span, except for bats, which are as small as rodents, but can live for up to 20 years. Though huge variation exists in mammals, there are a few physical traits that unite them. 1) Mammals are covered with body hair (fur). Though marine mammals, like dolphins and whales, have traded the benefits of body hair for better aerodynamics for traveling in water, they do still have some bristly hair on their faces (and embryonically - before birth). Hair is important for keeping mammals warm in cold climates, protecting them from sunburn and scratches, and used to warn off others, like when a dog raises the hair on its neck.
    [Show full text]
  • Identificado El Primer Fósil De Pangolín En La Península Ibérica
    TIERRA Identificado el primer fósil de pangolín en la península ibérica El análisis del fémur de un pangolín de hace 16 millones de años, hallado en los años 70 en el yacimiento de Can Cerdà, en la cuenca del Vallès-Penedès (Cataluña), ha permitido identificar el primer registro de pangolines fósiles en la península ibérica. Los restos conservan características primitivas y se han asignado a un género ya extinto. Estos animales actualmente viven en Asia y África, pero hace más de cinco millones de años estuvieron también presentes en Europa. SINC 21/3/2018 10:30 CEST Ejemplar joven de pangolín asiático (Smutsia temminckii). / Maria Diekmann of Rare and Endangered Species Trust / CC BY 2.0 En la actualidad, los pangolines (u osos hormigueros escamosos) están representados por tres géneros, que difieren en su distribución geográfica y adaptaciones locomotoras. Los pangolines asiáticos se incluyen en el género Manis, mientras que los africanos están incluidos en Smutsia (pangolines de tierra) y Phataginus (pangolines arbóreos). Pero durante el Eoceno (hace más de 33 millones de años, los pangolines se extendían TIERRA desde Asia hasta América del Norte. El fémur fue excavado hace varias décadas en el yacimiento de Can Cerdà, pero el material nunca fue dibujado o descrito, y aparentemente fue olvidado En el Mioceno se encontraron exclusivamente en Europa, donde estuvieron representados por el género extinto Necromanis, de relaciones inciertas con las especies actuales. En Europa se han identificado distintas especies de este género durante el Oligoceno (hace más de 23 millones de años) y el Mioceno (hace entre 23 y 5 millones de años), generalmente representadas por restos fósiles fragmentarios.
    [Show full text]
  • Autecology of the Sunda Pangolin (Manis Javanica) in Singapore
    AUTECOLOGY OF THE SUNDA PANGOLIN (MANIS JAVANICA) IN SINGAPORE LIM T-LON, NORMAN (B.Sc. (Hons.), NUS) A THESIS SUBMITTED FOR THE DEGREE OF MASTER OF SCIENCE DEPARTMENT OF BIOLOGICAL SCIENCES NATIONAL UNIVERSITY OF SINGAPORE 2007 An adult male Manis javanica (MJ17) raiding an arboreal Oceophylla smaradgina nest. By shutting its nostrils and eyes, the Sunda Pangolin is able to protect its vulnerable parts from the powerful bites of this ant speces. The scales and thick skin further reduce the impacts of the ants’ attack. ii ACKNOWLEDGEMENTS My supervisor Professor Peter Ng Kee Lin is a wonderful mentor who provides the perfect combination of support and freedom that every graduate student should have. Despite his busy schedule, he always makes time for his students and provides the appropriate advice needed. His insightful comments and innovative ideas never fail to impress and inspire me throughout my entire time in the University. Lastly, I am most grateful to Prof. Ng for seeing promise in me and accepting me into the family of the Systematics and Ecology Laboratory. I would also like to thank Benjamin Lee for introducing me to the subject of pangolins, and subsequently introducing me to Melvin Gumal. They have guided me along tremendously during the preliminary phase of the project and provided wonderful comments throughout the entire course. The Wildlife Conservation Society (WCS) provided funding to undertake this research. In addition, field biologists from the various WCS offices in Southeast Asia have helped tremendously throughout the project, especially Anthony Lynam who has taken time off to conduct a camera-trapping workshop.
    [Show full text]
  • Transfer of Manis Crassicaudata, M. Pentadactyla, M. Javanica from Appendix II to Appendix I
    Prop. 11.13 CONSIDERATION OF PROPOSALS FOR AMENDMENT OF APPENDICES I AND II Other proposals A. Proposal Transfer of Manis crassicaudata, M. pentadactyla, M. javanica from Appendix II to Appendix I. B. Proponents India, Nepal, Sri Lanka and the United States of America C. Supporting Statement 1. Taxonomy 1.1 Class: Mammalia 1.2 Order: Pholidota 1.3 Family: Manidae 1.4 Genus: Manis crassicaudata Gray, 1827 Manis javanica Desmarest, 1822 Manis pentadactyla Linneaus, 1758 1.5 Scientific synonyms: 1.6 Common names: English: (Manis crassicaudata) - Indian pangolin (Manis javanica) - Malayan pangolin (Manis pentadactyla) - Chinese pangolin French: (Manis crassicaudata) - Grand pangolin de l’Inde (Manis javanica) - Pangolin malais (Manis pentadactyla) - Pangolin de Chino Spanish: (Manis crassicaudata) - Pangolín indio (Manis javanica) - Pangolín malayo (Manis pentadactyla) - Pangolín Chino 1.7 Code numbers: Manis crassicaudata: A-108.001.001.001 Manis javanica: A-108.001.001.003 Manis pentadactyla: A-108.001.001.005 2. Biological Parameters 2.1 Distribution Manis crassicaudata occurs in the Indian sub-continent from eastern Pakistan, through much of India (south of the Himalayas), Bangladesh, and Sri Lanka, and, possibly, Myanmar and extreme western China (IUCN 1996, WCMC et al. 1999). Additional details on the distribution of this species are provided in Appendix A. Manis javanica occurs in tropical Southeast Asia. Although the northern and western limits of its range are very poorly defined, it has been recorded in much of Indonesia, Malaysia, the Philippines (Palawan Province), the southern half of Indo-China, much of Thailand and southern Myanmar (Nowak 1991, WCMC et al. 1999). It may also occur in Bangladesh and southwest Prop.
    [Show full text]
  • Report on Biodiversity and Tropical Forests in Indonesia
    Report on Biodiversity and Tropical Forests in Indonesia Submitted in accordance with Foreign Assistance Act Sections 118/119 February 20, 2004 Prepared for USAID/Indonesia Jl. Medan Merdeka Selatan No. 3-5 Jakarta 10110 Indonesia Prepared by Steve Rhee, M.E.Sc. Darrell Kitchener, Ph.D. Tim Brown, Ph.D. Reed Merrill, M.Sc. Russ Dilts, Ph.D. Stacey Tighe, Ph.D. Table of Contents Table of Contents............................................................................................................................. i List of Tables .................................................................................................................................. v List of Figures............................................................................................................................... vii Acronyms....................................................................................................................................... ix Executive Summary.................................................................................................................... xvii 1. Introduction............................................................................................................................1- 1 2. Legislative and Institutional Structure Affecting Biological Resources...............................2 - 1 2.1 Government of Indonesia................................................................................................2 - 2 2.1.1 Legislative Basis for Protection and Management of Biodiversity and
    [Show full text]
  • PROCEEDINGS of the WORKSHOP on TRADE and CONSERVATION of PANGOLINS NATIVE to SOUTH and SOUTHEAST ASIA 30 June – 2 July 2008, Singapore Zoo Edited by S
    PROCEEDINGS OF THE WORKSHOP ON TRADE AND CONSERVATION OF PANGOLINS NATIVE TO SOUTH AND SOUTHEAST ASIA 30 June – 2 July 2008, Singapore Zoo Edited by S. Pantel and S.Y. Chin Wildlife Reserves Singapore Group PROCEEDINGS OF THE WORKSHOP ON TRADE AND CONSERVATION OF PANGOLINS NATIVE TO SOUTH AND SOUTHEAST ASIA 30 JUNE –2JULY 2008, SINGAPORE ZOO EDITED BY S. PANTEL AND S. Y. CHIN 1 Published by TRAFFIC Southeast Asia, Petaling Jaya, Selangor, Malaysia © 2009 TRAFFIC Southeast Asia All rights reserved. All material appearing in these proceedings is copyrighted and may be reproduced with permission. Any reproduction, in full or in part, of this publication must credit TRAFFIC Southeast Asia as the copyright owner. The views of the authors expressed in these proceedings do not necessarily reflect those of the TRAFFIC Network, WWF or IUCN. The designations of geographical entities in this publication, and the presentation of the material, do not imply the expression of any opinion whatsoever on the part of TRAFFIC or its supporting organizations concerning the legal status of any country, territory, or area, or its authorities, or concerning the delimitation of its frontiers or boundaries. The TRAFFIC symbol copyright and Registered Trademark ownership is held by WWF. TRAFFIC is a joint programme of WWF and IUCN. Layout by Sandrine Pantel, TRAFFIC Southeast Asia Suggested citation: Sandrine Pantel and Chin Sing Yun (ed.). 2009. Proceedings of the Workshop on Trade and Conservation of Pangolins Native to South and Southeast Asia, 30 June-2 July
    [Show full text]
  • Supplemental File 1: Addressing Claims of “Zombie” Lineages on Phillips’ (2016) Timetree
    Supplemental File 1: Addressing claims of “zombie” lineages on Phillips’ (2016) timetree The soft explosive model of placental mammal evolution Matthew J. Phillips*,1 and Carmelo Fruciano1 1School of Earth, Environmental and Biological Sciences, Queensland University of Technology, Brisbane, Australia *Corresponding author: E-mail: [email protected] Contents Addressing claims of “zombie” lineages on Phillips’ (2016) timetree ................................................... 1 Incorrect or poorly supported fossil placements ................................................................................. 1 Figure S1 ............................................................................................................................................ 4 Table S1 .............................................................................................................................................. 6 References .............................................................................................................................................. 7 Addressing claims of “zombie lineages” on Phillips’ (2016) timetree Phillips [1] found extreme divergence underestimation among large, long-lived taxa that were not calibrated, and argued that calibrating these taxa instead shifted the impact of the underlying rate model misspecification to inflating dates deeper in the tree. To avoid this “error-shift inflation”, Phillips [1] first inferred divergences with dos Reis et al.’s [2] calibrations, most of which are set among taxa
    [Show full text]
  • Hyaenodontidae (Creodonta, Mammalia) and the Position of Systematics in Evolutionary Biology
    Hyaenodontidae (Creodonta, Mammalia) and the Position of Systematics in Evolutionary Biology by Paul David Polly B.A. (University of Texas at Austin) 1987 A dissertation submitted in partial satisfaction of the requirements for the degree of Doctor of Philosophy in Paleontology in the GRADUATE DIVISION of the UNIVERSITY of CALIFORNIA at BERKELEY Committee in charge: Professor William A. Clemens, Chair Professor Kevin Padian Professor James L. Patton Professor F. Clark Howell 1993 Hyaenodontidae (Creodonta, Mammalia) and the Position of Systematics in Evolutionary Biology © 1993 by Paul David Polly To P. Reid Hamilton, in memory. iii TABLE OF CONTENTS Introduction ix Acknowledgments xi Chapter One--Revolution and Evolution in Taxonomy: Mammalian Classification Before and After Darwin 1 Introduction 2 The Beginning of Modern Taxonomy: Linnaeus and his Predecessors 5 Cuvier's Classification 10 Owen's Classification 18 Post-Darwinian Taxonomy: Revolution and Evolution in Classification 24 Kovalevskii's Classification 25 Huxley's Classification 28 Cope's Classification 33 Early 20th Century Taxonomy 42 Simpson and the Evolutionary Synthesis 46 A Box Model of Classification 48 The Content of Simpson's 1945 Classification 50 Conclusion 52 Acknowledgments 56 Bibliography 56 Figures 69 Chapter Two: Hyaenodontidae (Creodonta, Mammalia) from the Early Eocene Four Mile Fauna and Their Biostratigraphic Implications 78 Abstract 79 Introduction 79 Materials and Methods 80 iv Systematic Paleontology 80 The Four Mile Fauna and Wasatchian Biostratigraphic Zonation 84 Conclusion 86 Acknowledgments 86 Bibliography 86 Figures 87 Chapter Three: A New Genus Eurotherium (Creodonta, Mammalia) in Reference to Taxonomic Problems with Some Eocene Hyaenodontids from Eurasia (With B. Lange-Badré) 89 Résumé 90 Abstract 90 Version française abrégéé 90 Introduction 93 Acknowledgments 96 Bibliography 96 Table 3.1: Original and Current Usages of Genera and Species 99 Table 3.2: Species Currently Included in Genera Discussed in Text 101 Chapter Four: The skeleton of Gazinocyon vulpeculus n.
    [Show full text]
  • 71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
    ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas.
    [Show full text]
  • Mammal and Plant Localities of the Fort Union, Willwood, and Iktman Formations, Southern Bighorn Basin* Wyoming
    Distribution and Stratigraphip Correlation of Upper:UB_ • Ju Paleocene and Lower Eocene Fossil Mammal and Plant Localities of the Fort Union, Willwood, and Iktman Formations, Southern Bighorn Basin* Wyoming U,S. GEOLOGICAL SURVEY PROFESS IONAL PAPER 1540 Cover. A member of the American Museum of Natural History 1896 expedition enter­ ing the badlands of the Willwood Formation on Dorsey Creek, Wyoming, near what is now U.S. Geological Survey fossil vertebrate locality D1691 (Wardel Reservoir quadran­ gle). View to the southwest. Photograph by Walter Granger, courtesy of the Department of Library Services, American Museum of Natural History, New York, negative no. 35957. DISTRIBUTION AND STRATIGRAPHIC CORRELATION OF UPPER PALEOCENE AND LOWER EOCENE FOSSIL MAMMAL AND PLANT LOCALITIES OF THE FORT UNION, WILLWOOD, AND TATMAN FORMATIONS, SOUTHERN BIGHORN BASIN, WYOMING Upper part of the Will wood Formation on East Ridge, Middle Fork of Fifteenmile Creek, southern Bighorn Basin, Wyoming. The Kirwin intrusive complex of the Absaroka Range is in the background. View to the west. Distribution and Stratigraphic Correlation of Upper Paleocene and Lower Eocene Fossil Mammal and Plant Localities of the Fort Union, Willwood, and Tatman Formations, Southern Bighorn Basin, Wyoming By Thomas M. Down, Kenneth D. Rose, Elwyn L. Simons, and Scott L. Wing U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1540 UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1994 U.S. DEPARTMENT OF THE INTERIOR BRUCE BABBITT, Secretary U.S. GEOLOGICAL SURVEY Robert M. Hirsch, Acting Director For sale by U.S. Geological Survey, Map Distribution Box 25286, MS 306, Federal Center Denver, CO 80225 Any use of trade, product, or firm names in this publication is for descriptive purposes only and does not imply endorsement by the U.S.
    [Show full text]