Annotated Checklist of Vascular Flora Capitol Reef National Park

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Natural Resource Technical Report NPS/NCPN/NRTR–2009/154 ON THE COVER Last Chance townsendia (Townsendia aprica), Capitol Reef National Park. Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Capitol Reef National Park

Natural Resource Technical Report NPS/NCPN/NRTR–2009/154

Author Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741

Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532

January 2009

U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The Natural Resource Publication series addresses natural resource topics that are of interest and applicability to a broad readership in the National Park Service and to others in the management of natural resources, including the scientiﬁ c community, the public, and the NPS conservation and environmental constituencies. Manuscripts are peer-reviewed to ensure that the information is scientiﬁ cally credible, technically accurate, appropriately written for the intended audience, and is designed and published in a professional manner.

The Natural Resource Technical Report series is used to disseminate the peer-reviewed results of scientiﬁ c studies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service’s mission. The reports provide contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. Current examples of such reports include the results of research that addresses natural resource management issues; natural resource inventory and monitoring activities; resource assessment reports; scientiﬁ c literature reviews; and peer- reviewed proceedings of technical workshops, conferences, or symposia.

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Please cite this publication as: Fertig, W. 2009. Annotated checklist of vascular ﬂ ora: Capitol Reef National Park. Natural Resource Technical Report NPS/NCPN/NRTR–2009/154. National Park Service, Fort Collins, Colorado.

NPS D-243, January 2009 ii Annotated Checklist of Vascular Flora: Capitol Reef National Park Contents

Figures ...... v Tables ...... v Abstract ...... vii Acknowledgements ...... ix 1 Introduction ...... 1 1.1 Background and justiﬁ cation ...... 1 1.2 Objectives ...... 1 1.3 Study area ...... 1 1.3.1 History and setting ...... 1 1.3.2 Geology ...... 2 1.3.3 Vegetation ...... 4 1.3.4 Previous botanical studies ...... 4

2 Methods ...... 9

3 Results ...... 11 3.1 Summary of the ﬂ ora of Capitol Reef National Park ...... 11 3.2 Life form and biogeographic diversity ...... 12 3.3 Plants of special concern ...... 12 3.4 Non-native species and noxious weeds ...... 12 4 Discussion ...... 17

5 Literature Cited ...... 19

Appendices ...... 23 Appendix A. Confi rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family and scientifi c name with taxonomic and biogeographic notes...... 26 Appendix B. Potential (unconfi rmed) taxa of Capitol Reef National Park...... 105 Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park. .117 Appendix D. Confi rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes...... 129

Contents iii iv Annotated Checklist of Vascular Flora: Capitol Reef National Park Figures

Figure 1.3.1. Location of Capitol Reef National Park (Wayne, Garﬁ eld, Emery, and Sevier counties, Utah). .3 Figure 1.3.4. Increase in the number of plant taxa conﬁ rmed as present in Capitol Reef National Park by decade, 1930–2006...... 7

Tables

Table 1.3.4. Chronology of plant collecting efforts in Capitol Reef National Park...... 5 Table 3.1. Statistical summary of the ﬂ ora of Capitol Reef National Park...... 11 Table 3.3. Plant species of conservation concern from Capitol Reef National Park...... 13

Contents v vi Annotated Checklist of Vascular Flora: Capitol Reef National Park Abstract

In 2005–2006, the National Park Service’s Northern Colorado Plateau Network received funding to develop a revised checklist of the vascular plant fl ora of Capitol Reef National Park. This entailed a thorough review of existing literature, re-examination of specimens from the Capitol Reef National Park herbarium and other state and national museum collections, and additional fi eld work to cor- roborate unverifi ed species reports or locate new species. Based on this study, 887 vascular plant taxa have been confi rmed or reliably reported as occuring in the park (835 are known from validated herbarium specimens and 52 from unvouchered literature reports). Another 117 species formerly considered part of the park fl ora have now been shown to be falsely reported. The current fl ora of Capitol Reef National Park represents about 25% of the known fl ora of Utah, and 52.4% of the fl ora of the Colorado Plateau ecoregion within Utah. At least 7 of these species are federally listed as threatened or endangered, and 54 more are considered species of concern by the state natural heritage program. Introduced plants make up 13.8% of the park fl ora (45 species), of which only 3 species are considered noxious weeds by the State of Utah. Further surveys in Capitol Reef National Park are likely to fi nd additional new species (about 192 are known from similar habitats outside the park boundary), especially among late- season fl owering, weedy, or wetland taxa.

Contents vii viii Annotated Checklist of Vascular Flora: Capitol Reef National Park Acknowledgements

I would like to thank the following individuals for their assistance with this project. Botanist Dr. Angela Evenden, former leader of the Northern Colorado Plateau Network in Moab, Utah, was an active proponent of this project and helped secure funding for ﬁ eld work and herbarium studies in 2006. Tom Clark, natural resources chief at Capitol Reef National Park, and Debi Clark, interagency botanist, provided access to all mounted and unmounted CARE herbarium specimens, research permits, reprints, and advice on botanical issues related to the park. Amy Tendick and Liz Ballenger recorded specimen-label data from all mounted collections in the CARE herbarium. Dr. Mary Barkworth, of Utah State University, corroborated the identiﬁ cation of Stipa scribneri in the park. Libby Nance, former database specialist with the NCPN, assisted with development and quality control of the park’s NPSpecies database. Libby Nance, Janet Coles, and Helen Thomas also provided data from recent vegetation mapping and wetland inventories in the park. Drs. Stanley Welsh and Duane Atwood provided access to herbarium collections at Brigham Young University. Dr. Michael Piep provided access to collections at the Utah State University (Intermountain) herbarium. Dr. Ron Hartman and Ernie Nelson provided access and specimen loans from the Rocky Mountain Herbarium at the University of Wyoming. Thanks to the technical reviewers (Dr. Angela Evenden, Dr. Dusty Perkins, and Libby Nance) for their comments and suggestions. To any others whom I may have inadvertently forgotten, please accept my apologies (and thanks).

Research was conducted under permit # CARE-2006-SCI-0002 for study number CARE-06002. Funding was provided by the National Park Service through the University of Wyoming.

Contents ix x Annotated Checklist of Vascular Flora: Capitol Reef National Park (Evenden et al. 2002). This eff ort has entailed re- 1 Introduction viewing existing species checklists, corroborating the identifi cation of specimens in park museum In 2006, the National Park Service (NPS) estab- and herbarium collections, locating additional lished a cooperative agreement with the Univer- specimen records from other museums, mining sity of Wyoming, through the Rocky Mountains relevant literature and datasets, and conducting Cooperative Ecosystem Studies Unit, to produce fi eld surveys to locate new taxa or confi rm un- a series of technical reports summarizing the vas- substantiated reports.† All of these data have been cular plant fl ora of each of the 16 national park synthesized in NPSpecies, the park service’s stan- units of the Northern Colorado Plateau Network dardized database for documenting park status (NCPN). These reports are based on existing in- (present, probably present, unconfi rmed, falsely formation in the park service’s NPSpecies data- reported), abundance, residency, nativity, and base system and augmented by further data min- rarity. With the production of a technical series of ing, reviews of state and regional herbaria, and reports on the fl ora of each park, the NPSpecies new fi eld surveys to resolve the status of many data will now be available to a broader audience. reported and unconfi rmed species. The objective of these reports is to summarize the current state of knowledge of the fl ora of each park for the 1.2 Objectives benefi t of NPS biologists and interpretive staff , The objectives of this project were to: outside researchers, and the general public. The following report addresses the vascular fl ora of 1. Document the current status (present, Capitol Reef National Park (NP), in south-central reported, potential, or falsely reported) of all Utah. Other NCPN park units are addressed in vascular plant species that have been cited separate reports. as occurring in Capitol Reef NP, based on a review of herbarium collections and past literature; and 1.1 Background and justifi cation One of the primary missions of the NPS is “to 2. Develop an annotated checklist of the fl ora conserve unimpaired the natural and cultural of Capitol Reef NP, with supplemental in- resources and values of the national park system formation on taxonomic synonyms, distri- for the enjoyment of this and future generations” bution, within-park abundance, fl owering (NPS 2000). Due to their high degree of protec- period, growth form, nativity, habitat, and tion and distribution across a wide variety of hab- sources for each record. itat types, the nation’s national park units make a signifi cant contribution to the protection of bio- 1.3 Study area logical diversity (Noss and Cooperrider 1994). Unfortunately, the full contribution of the NPS 1.3.1 History and setting system is not known, as many park units have The oldest evidence of human occupation of the traditionally lacked comprehensive biological in- Capitol Reef and Waterpocket Fold area comes ventories or monitoring programs to identify the from dart points dating from 2,000 to 8,000 years composition and condition of their biota. Know- before present. Sedentary farmers and hunter- ing the status of park fl ora and fauna is critical for gatherers of the Ancestral Puebloan and Fremont eff ective park management and for systematically cultures inhabited areas near perennial water identifying and fi lling gaps in the nation’s protect- sources within the park during the period from ed area network (Margules and Pressey 2000). 600 to 1275 A.D. Following periods of extensive drought, native cultures adopted the more no- To rectify this shortcoming, Congress passed the madic lifestyle that existed at the time of Euro- National Parks Omnibus Management Act in 1998 pean contact (Frye 1998). to provide guidance and funding for an inventory and monitoring program across the entire na- Waterpocket Fold presented a signifi cant bar- tional park system. Since 2000, the NCPN Inven- rier to early European explorers, who largely by- tory and Monitoring Program has been develop- passed the park area until the mid-19th century. ing and verifying species lists of vertebrates and Almon Thompson, of the Powell survey, made vascular plants for each park unit in the network

†Taxa refer to all named species, subspecies, and varieties.

Chapter 1: Introduction 1 the fi rst scientifi c explorations through the future traffi c. Waterpocket Fold, an east-dipping mono- park in 1872, en route to the Henry Mountains. cline extending from Thousand Lake Mountain Geologists Grove Karl Gilbert and Clarence Dut- nearly to Lake Powell, was uplifted during a pe- ton mapped the Waterpocket Fold in detail in riod of tectonic activity in the late Cretaceous or the late 1870s. Mormon pioneers began settling early Tertiary, along with much of the Colorado the area in the vicinity of Capitol Reef in the late Plateau and Rocky Mountains. Diff erent rates of 1870s and early 1880s. A small farming commu- erosion (depending on the nature and position nity began at Fruita in 1886, but struggled due to of sedimentary beds) over time carved the deep its remoteness (Frye 1998). canyons, domes, arches, and bedrock holes (“wa- terpockets”) characteristic of the Capitol Reef Wayne County civic leaders and boosters began area (Chronic 1988; Morris et al. 2003). espousing protection of a portion of the Capitol Reef area around Fruita (the “Wayne Wonder- Rocks exposed in Capitol Reef NP range in age land”) as a state or national park in the early 1920s. from Permian sandstone to Quaternary volcanic Such a designation would result in improved road boulders. The oldest strata tend to occur in the access to the area and greater economic pros- western part of the park, becoming progressively perity from tourism. These eff orts reached frui- younger as one travels eastward, following the tion in August 1937, with the creation of Capitol tilt of the Waterpocket Fold monocline (Chronic Reef National Monument by President Franklin 1988). Permian-age outcrops of the Cutler Group Roosevelt, under the Antiquities Act. The monu- (including the Cedar Mesa and White Rim sand- ment’s original boundaries were enlarged by stones), dating to about 265 million years ago, are President Eisenhower in 1958, and again in 1969 exposed in the bottom of the Fremont River canby President Johnson, who increased the area by yon and Sulphur Creek near the park’s western nearly 600%. In December 1971, Congress fi xed boundary. These strata were originally deposited the current boundaries at 241,904 acres (97,895 as beach and wind-blown sand dunes, and are ha) and upgraded Capitol Reef to full national capped by the marine-derived Kaibab limestone. park status (Evenden et al. 2002, Frye 1998). With In the early Triassic, the Capitol Reef area was a the additions in 1969, the park currently stretches coastal tidal fl at that was periodically refl ooded nearly 70 miles, from Thousand Lake Mountain by marine waters, forming the alternating brown almost to Lake Powell. and red mudstones and siltstones of the Moen- kopi Formation. During the late Triassic, the park Capitol Reef National Park is located in west-cen- area was buried by river-borne volcanic ash and tral Wayne, north-central Garfi eld, southwestern clay of the Chinle Formation. Emery, and southeastern Sevier counties in central Utah (Figure 1.3.1), about 75 road miles east Continuing eastward, the Chinle is covered by ear- of Richfi eld. Elevation within the park ranges from ly Jurassic Wingate Sandstone (derived from wind- 3,880 feet (1,183 m) at the bottom of Halls Creek blown sand dunes) and sandstone-claystone con- to 8,960 feet (2,731 m) on the fl ank of Thousand glomerates of the fl uvial Kayenta Formation. Above Lake Mountain. Annual precipitation averages 5 the Kayenta are the thick beds of Navajo Sandstone inches (12.5 cm) at the south end of the park and that form the park’s characteristic monolithic cliff s 12 inches (30 cm) at the higher, north end. Fruita and domes. The Navajo was part of a Sahara-like has an annual precipitation of 7.2 inches (18 cm), dune system that extended from Nevada to south- with 30% coming in the summer monsoon sea- ern Wyoming. Later in the Jurassic and early Cre- son. The average minimum temperature at Fruita taceous, the Capitol Reef area was again inundated in January is 18°F (–8° C), and in July, 92°F (33°C) by marine intrusions or exposed as a tidal fl at. Al- (Heil et al. 1993). ternating beds of sandstone, siltstone, mudstone, gypsum, and limestone were laid down during this 1.3.2 Geology time as the San Rafael Group (Page Sandstone, Carmel, Entrada, Curtis, and Summerville forma- The “Capitol” in Capitol Reef comes from the tions), Morrison Formation, and Dakota Sand- dome-like outcrops of Navajo Sandstone in the stone. Deep marine waters, alternating with delta Fruita area that resemble the rotunda shape of and coastal plain depositional environments, pre- many capitol buildings. Early geographers applied dominated during the early to mid-Cretaceous, the term “reef” to the nearly 100-mile-long Water- giving rise to the various members of the Mancos pocket Fold, which created a formidable barrier Shale and Mesa Verde Formation. to travelers—much like marine reefs impede ship

2 Annotated Checklist of Vascular Flora: Capitol Reef National Park Capitol Reef National ParkCapito l Reef National Park Utah Utah

Burr Trail

Torrey Visitor Center 24

02461 Kilometers

02461 Map location Northern Section Miles Southern Section

Established August 2, 1937 Legend 1,183–2,731 m elevation (3,880–8,960 ft elevation) Intermittent streams Grand Staircase-Escalante NM (BLM) 97,895 ha (241,904 ac) Perennial streams U.S. Bureau of Land Management Rivers Private Paved roads State Unpaved roads U.S. Forest Service Unimproved/4WD roads Glen Canyon NRA (NPS) Park boundary (NPS) NCPN boundary

MAP CREATED BY ANETH WIGHT, NCPN (DERIVED FROM EVENDEN ET AL. 2002) Figure 1.3.1. Location of Capitol Reef National Park (Wayne, Garﬁ eld, Emery, and Sevier counties, Utah).

Chapter 1: Introduction 3 The Cenozoic has largely been a time of erosion Mixed conifer forests, which occur at higher el- in Capitol Reef NP, interrupted by periods of vol- evations on deep, coarse-textured soils, or in canic activity about 20–25 million years ago and shady slot canyons, consist primarily of Ponde- again as recently as 3.7 million years ago (Morris rosa pine (Pinus ponderosa), bitterbrush (Purshia et al. 2003). Remnants of the last period of vol- tridentata), alder-leaf mountain mahogany (Cer- canism persist in the form of intrusive igneous cocarpus montanus), and greenleaf manzanita dikes, sills, and diatremes found sporadically in (Arctostaphylos patula). Quaking aspen (Populus the South Desert area. Large, black boulders on tremuloides) may predominate in more mesic hills near the visitor center were derived from sites with rich soils. Western bristlecone pine (Pi- lava ﬁ elds on plateaus to the west and north and nus longaeva)-cushion plant stands occur locally transported to the park as outwash from glacia- in the park, and are considered a community of tion (Chronic 1988). conservation concern by Heil et al. (1993).

Wetlands occur across all elevations and are as- 1.3.3 Vegetation sociated with perennial rivers, streams, or seeps. The plant communities of Capitol Reef Nation- Fremont cottonwood (Populus fremontii) wood- al Park have been described in detail by Meyer lands with an understory of willows (Salix spp.) (1990), Heil et al. (1993), and the Northern Colo- or rabbitbrush occur commonly along the larger rado Plateau Network (in ed.). These can be ag- watercourses. Stands of common reed (Phrag- gregated into fi ve general vegetation types that mites australis) or Baltic rush (Juncus arcticus) correlate with elevational and moisture gradients. may occur among cottonwood forests. At higher Bentonite and gypsum badlands and low-eleva- elevations, riparian woodlands may be dominat- tion, clay-rich substrates (such as the Moenkopi, ed by thinleaf alder (Alnus incana), water birch Chinle, and Mancos) are dominated by salt des- (Betula occidentalis), blue spruce (Picea pungens), ert scrub vegetation of shadscale (Atriplex confer- or red-osier dogwood (Cornus sericea). Hanging tifolia), mat saltbush (A. corrugata), Castle Valley gardens associated with shady cliff alcoves and saltbush (A. gardneri var. cuneata), spiny hopsage perennial seeps are uncommon in the park, but (Grayia spinosa), and crisp-leaf wild buckwheat are commonly occupied by southern maidenhair (Eriogonum corymbosum var. corymbosum). fern (Adiantum capillus-veneris), giant helleborine Low-elevation desert sites with shallow soils over (Epipactis gigantea), and western poison ivy (Tox- impervious bedrock support stands of black- icodendron rydbergii) (Fowler et al. 2007; Welsh brush (Coleogyne ramosissima) and rabbitbrush 1989). Many of the riverside terraces in the Fruita (Chrysothamnus spp.) (Heil et al. 1993). area were converted to orchards by Mormon pioneers. These are still maintained by the National Sagebrush shrublands and grasslands replace Park Service, and contain a number of cultivated desert scrub at higher elevations and on allu- fruit species and weeds (Heil et al. 1993). vial terraces or sandy soils. These stands may be dominated by big sagebrush (Artemisia triden- 1.3.4 Previous botanical studies tata), rubber rabbitbrush (Chrysothamnus nauseosus), or fourwing saltbush (Atriplex canescens). Lester Ward, of the Powell Survey, explored Rab- More saline terraces may be locally dominated by bit Valley and the Aquarius Plateau west of Capi- greasewood (Sarcobatus vermiculatus), and sites tol Reef in 1875, but apparently did not collect over stabilized sand dunes by sand sagebrush (Ar- botanical specimens within the future park itself. temisia fi lifolia) or perennial grasses. Likewise, Marcus E. Jones conducted fi eld work in the Henry Mountains and vicinity of Hanks- Pinyon-juniper woodlands predominate on thin, ville and Loa in 1894, Per Axel Rydberg and E. C. rocky soils at low to mid-elevations. These pyg- Carlton surveyed the Aquarius Plateau in 1905, my forests are dominated by two-needle pinyon and W. D. Stanton botanized the Henry Moun- (Pinus edulis) and Utah juniper (Juniperus os- tains in 1930, but all apparently bypassed Capitol teosperma), with understories of green ephedra Reef, itself. The earliest collections known from (Ephedra viridis) or other low shrubs or grasses. Capitol Reef National Monument were made Wind-blasted, rocky ridges are often locally dom- by Bertrand F. Harrison, Ben Markham, and El- inated by cushion plant species, while slickrock den Beck, of Brigham Young University, in 1938 exposures or ledges may be sparsely vegetated by (Table 1.3.4, Figure 1.3.4). Noteworthy among the dwarf mountain mahogany (Cercocarpus intrica- fi nds that year was the collection of the holotype tus). of Beck’s spring-parsley (Cymopterus beckii),

4 Annotated Checklist of Vascular Flora: Capitol Reef National Park Table 1.3.4. Chronology of plant collecting efforts in Capitol Reef National Park. # of new Year Collector(s) taxa Comments 1938–1940 B. F. Harrison, B. 7 Bertrand Harrison and colleagues from Brigham Young University collected Markham, E. Beck, at least 7 plant taxa in Capitol Reef National Monument over a three-year Reed, & W. B. period, including the holotype of Cymopterus beckii in 1938. NPS regional McDougall biologist W. B. McDougall made a cursory checklist of monument fl ora in 1940 based on a one-day visit (McDougall 1940). 1951 B. F. Harrison 1 B. F. Harrison collected the fi rst record of Springfi eld’s bluestem (Bothriochloa springfi eldii) while passing through the monument. 1958 F. Swancara & E. 40 Frank Swancara and Emery Lehnert made collections of 40 new taxa. These Lehnert specimens formed the foundation of the Capitol Reef herbarium (CARE). 1961 R. Barneby 1 Rupert Barneby, of the New York Botanical Garden, collected the type of Astragalus harrisonii from Capitol Reef National Monument. 1965–1968 K. Erdman & K. 11 Kimball Erdman and Kelly Taylor added 11 new species to the monument Taylor fl ora. 1970–1971 R. Nelson 79 Ruth Nelson discovered 79 new species for Capitol Reef National Park, including fi rst records for several rare or locally endemic species, such as Bicknell milkvetch (Astragalus consobrinus), Woodruff’s milkvetch (A. woodruffi i), and low hymenoxys (Hymenoxys acaulis var. nana). 1972 N. D. Atwood & L. 1 Duane Atwood and Larry Higgins collected the type of Phacelia rafaelensis Higgins from Capitol Reef National Park. 1973–1975 H. K. Harrison 90 Keith Harrison made nearly 600 collections for the CARE herbarium and documented 90 new taxa for the park fl ora. 1977–1978 P. Camp, E. Neese, 38 Pam Camp, of Utah State University, made more than 200 collections in the and S. White Halls Creek area of Capitol Reef National Park and Glen Canyon National Recreation Area. Excluding Glen Canyon records, she added 37 new taxa to the fl ora of Capitol Reef National Park. Elizabeth Neese and S. White discovered Spiranthes diluvialis in Capitol Reef in 1977 (this species was listed as threatened under the Endangered Species Act in 1992). 1980–1981 S. Meyer 186 Susan Meyer made more than 300 collections for the CARE herbarium and added 186 new species to the park fl ora. Meyer (1982) compiled a checklist of 512 plant species for the park. 1982 N. D. Atwood, 4 Stan Welsh and colleagues discovered 4 new species for the park, including S. Goodrich, S. the type of Erigeron maguirei var. harrisonii (no longer considered distinct Welsh, and E. from E. maguirei). Neese 1984–1985 J. Lichthardt, S. 7 Juanita Lichthardt, Stan Welsh, and Elizabeth Neese added 7 new species to Welsh, and E. the park fl ora. Neese 1986–1988 K. Heil, J. M. 353 Ken Heil, Mark Porter, and Rich Fleming, of San Juan College, and Porter, R. Fleming, William Romme, of Fort Lewis College, conducted a multi-year vegetation and W. Romme classifi cation, fl oristic inventory, and rare-plant survey for Capitol Reef National Park. In the course of collecting more than 650 specimens for the CARE herbarium, this group added 353 new species to the park fl ora and 33 new rare species, including fi rst records of Rabbit Valley gilia (Gilia caespitosa), Winkler’s pincushion cactus (Pediocactus winkleri), Wright’s fi shhook cactus (Sclerocactus wrightiae), Jones’ cycladenia (Cycladenia humilis var. jonesii), and Barneby’s reed-mustard (Schoenocrambe barnebyi). Heil et al. (1993) produced a checklist of the park fl ora with 759 taxa. 1989–1991 E. Haskell, J. 3 Eric Haskell, John Spence, Karen Mason, and Ken Heil added fi rst reports of Spence, K. Mason, Thunberg’s barberry (Berberis thunbergii), Dryopteris fi lix-mas, and cardinal- and K. Heil fl ower (Lobelia cardinalis). 1992–1993 J. Fowler, C. L. 0 Jim Fowler and colleagues surveyed hanging garden vegetation in the park, May, and J. Hak but added no new species to the fl ora (Fowler et al. 2007).

Chapter 1: Introduction 5 Table 1.3.4. Chronology of plant collecting efforts in Capitol Reef National Park, cont.

# of new Year Collector(s) taxa Comments 1995–1997 T. Clark, D. Clark, 2 Tom and Debi Clark discovered the ﬁ rst record of Gilia tenuis (a rare species) K. Harper, R. Van in the park. Kim Harper, Renee Van Buren, and Z. Aanderud discovered Buren, and Z. Gnaphalium wrightii in Capitol Reef NP. Aanderud 1997–2002 D. Clark 1 Debi Clark and colleagues with the Interagency Rare Plant Team surveyed Capitol Reef and adjacent BLM and USFS lands for rare plants, partly funded by a grant from Canon USA, Inc. Thelesperma subnudum var. alpinum was discovered in the park in 2000 (Clark 2002) 2004 K. Andersen, D. 3 (+ 25 Donna Barnes and Kim Harper of Utah Valley State College collected more Barnes, K. Harper, reported) than 50 specimens for the CARE herbarium, including ﬁ rst records of 2 & Northern taxa. Kim Andersen collected one new exotic species and reported 2 others Colorado Plateau during a survey of noxious weeds (Dewey and Andersen 2005). The Northern Network veg Colorado Plateau Network (in ed.) vegetation mapping team reported 23 new mapping team species for the park (without vouchers) 2005 R. Harms, W. 7 Rebecca Harms discovered 3 new species for the park during a survey of seeps Fertig, & P. Lesica and springs (Springer et al. 2006). Walter Fertig and Peter Lesica added 4 new species.

named in honor of its discoverer. National Park on the Glen Canyon side and remain uncon- Service regional biologist W. B. McDougall com- fi rmed (but potential) for Capitol Reef. Elizabeth piled the fi rst plant checklist for the new monu- Neese and S. White discovered an unusual orchid ment (mostly prominent trees and shrubs) based along the Fremont River in 1977 that was even- on a one-day visit in 1940 (Frye 1998; McDougall tually identifi ed as Ute ladies’-tresses (Spiranthes 1940). diluvialis, federally listed as threatened under the Endangered Species Act since 1992). From 1980 Botanical research and collecting was sporadic to 1981, Susan Meyer made more than 300 col- during the Capitol Reef National Monument era. lections from the park and added 186 new species Frank Swancara and Emery Lehnert initiated the to the fl ora (increasing it by nearly 40%). Meyer Capitol Reef herbarium (CARE) with their col- (1982, 1990) published a park checklist identi- lections in 1958. Rupert Barneby visited the mon- fying 512 taxa. Stan Welsh and colleagues from ument in 1961 and collected the type specimen of Brigham Young University conducted a study of a new species, Harrison’s milkvetch (Astragalus grazing impacts on the park in 1982, and discov- harrisonii). Ruth Nelson more than doubled the ered the holotype of Maguire’s fl eabane (Erigeron known fl ora of the area by adding 79 new species maguirei var. harrisonii, now considered synony- in 1970–1971. Duane Atwood and Larry Higgins, mous with var. maguirei) (Welsh 1982). graduate students from Brigham Young Universi- ty, took the type of San Rafael phacelia (Phacelia The most comprehensive survey of the fl ora and rafaelensis) from the park in 1972 (Welsh 1982). vegetation of Capitol Reef NP took place from 1986 to 1988, under the direction of Ken Heil, The pace of new discovery accelerated in the mid- Mark Porter, and Rich Fleming, of San Juan Col- 1970s and early 1980s. Keith Harrison made near- lege, and William Romme, of Fort Lewis College ly 600 collections for the CARE herbarium from (Heil et al. 1993). This group collected more than 1973 to 1975, and documented 90 new species 650 specimens for the CARE herbarium in the for the park. Pam Camp, of Utah State University, course of sampling 34 vegetation types. Heil et al. collected more than 200 specimens in the Halls (1993) produced a checklist identifying 759 taxa Creek drainage area along both sides of the Capi- for the park, of which 353 represented new species. tol Reef NP/Glen Canyon National Recreation In addition, Heil and his colleagues discovered 33 Area boundary in 1977–1978. Camp discovered new rare species for the park, including four of the at least 37 new species for Capitol Reef NP, but park’s seven listed threatened or endangered plant several additional species credited to her in later species (see Tables 1.3.4 and 3.3). checklists (Heil et al. 1993) were actually found

6 Annotated Checklist of Vascular Flora: Capitol Reef National Park In the past 20 years, the number of new species Andersen of Utah State University vouchered or discovered in Capitol Reef NP has leveled off reported three new invasive plant taxa in 2004 (Figure 1.3.4), but not stopped. Rare and uncom- (Dewey and Andersen 2005). Vegetation mapping mon species continue to be discovered, including crews from Capitol Reef NP and the Northern male fern (Dryopteris fi lix-mas), by John Spence Colorado Plateau Network (in ed.) reported 23 and Karen Mason in 1990, Mussentuchit gilia new species (without vouchers) in 2004. Rebecca (Gilia tenuis) and alpine greenthread (Thelesper- Harms discovered three new riparian species in a ma subnudum var. alpinum), by Tom and Debi 2005 survey of seeps and springs (Springer et al. Clark in 1995 and 2000, and Wright’s cudweed 2006). Montana botanist Peter Lesica and I col- (Gnaphalium wrightii), by Kim Harper and Renee lected four new species during the 2005 fi eld sea- Van Buren in 1997. More recently, Donna Barnes son (Table 1.3.4). and Kim Harper, from Utah Valley State College, added two new species while conducting gen- Additional details on plant-collecting eff orts at eral fl oristic collecting in the park in 2004. Kim Capitol Reef NP can be found in Table 1.3.4.

900

800

700

600

500

400

300

Number of vascular taxa 200

100

0 1930 1940 1950 1960 1970 1980 1990 2000 2006

Year

Figure 1.3.4. Increase in the number of plant taxa conﬁ rmed as present in Capitol Reef National Park by decade, 1930–2006.

Chapter 1: Introduction 7 8 Annotated Checklist of Vascular Flora: Capitol Reef National Park the context of Utah), population size in the park 2 Methods (derived from Heil et al. 1993, NCPN in ed., and fi eld observations), fl owering period, distribu- In January and February 2005, I reviewed all tion across general vegetation types, nativity, data mounted specimens in the Capitol Reef National source (collector and collection number from Park herbarium (CARE) to determine whether a sample specimen or literature citation), year each was correctly identifi ed to species, subspe- of earliest collection (from herbarium records), cies, or variety following the nomenclature of and relevant synonymy and taxonomic prob- Welsh et al. (2003). As necessary, I made correc- lems (based on review of Barkworth et al. 2003, tions and annotations to each specimen and to 2007; Barneby 1989; Cronquist 1994; Cronquist the voucher records in the park’s Automated Na- et al. 1972, 1977, 1984, 1997; Dorn 2001; Flora of tional Cataloging System (ANCS+) and NPSpe- North America Editorial Committee 1993, 1997, cies database. Based on this herbarium work and 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, a review of park references (Heil et al. 1993; Mey- 2006c; Holmgren et al. 2005; and Weber and er 1982; Dewey and Andersen 2005; Springer et Wittmann 2001). I then incorporated the annota- al. 2006), I developed four lists: tions to these lists and label data from the CARE herbarium review into the NPSpecies database 1. A revised checklist of species that were pres- for the park. ent in the park (confi rmed by an authenticated voucher; included in Appendices A Work in 2005–2007 focused on confi rmation of and D); reported and potential species for the park fl ora through fi eld surveys and review of collections 2. A list of species that were reported in the lit- from state and regional herbaria. To target miss- erature but not corroborated with a voucher ing species, I made collecting trips to Capitol Reef (equivalent to “probably present” in NPSpe- NP in May 2005 and September 2006, emphasiz- cies terminology; included in Appendices A ing under-sampled wetland, weedy, and late- and D); fl owering taxa. I undertook herbarium studies 3. A list of potential species (“unconfi rmed” at Brigham Young University (BRY), Utah State in NPSpecies) known from the vicinity University (USU), and the University of Wyoming of Capitol Reef National Park, using an (RM) in March, June, and October 2006 and Feb- unpublished list generated by Dr. N. Duane ruary and March 2007, and documented park Atwood of Brigham Young University and specimens from the Virtual Herbarium website Albee and others (1988) (Appendix B); and of the New York Botanical Garden (http://sciweb. nybg.org/science2/hcol/intf/index.asp). Finally, I 4. A list of species that are now known to be located additional new species reports by exam- misidentifi ed or highly questionable (Ap- ining unmounted collections by NCPN research- pendix C). ers and reviewing plot data from recent vegeta- I annotated each list with supplemental infor- tion mapping and wetland inventories (NCPN in mation on life form, geographic range (within ed.; Springer et al. 2006).

Chapter 2: Methods 9 10 Annotated Checklist of Vascular Flora: Capitol Reef National Park in 2005 netted another seven new species from 3 Results my fi eld collections and those of Rebecca Harms (Springer et al. 2006). Overall, the documented 3.1 Summary of the fl ora of Capitol fl ora of Capitol Reef NP has increased by 41 species (4.9%), bringing the total number of taxa to Reef National Park 835 (Table 3.1, Appendices A, D). Of the 2,028 vascular plant vouchers in the CARE herbarium examined in 2005, 221 were misiden- Currently, 52 additional taxa have been reported tifi ed (10.9%), 81 were revised to a more current for Capitol Reef NP but have yet to be corroborat- name (4%), 288 had variety or subspecies names ed with voucher specimens (Table 3.1, Appendix added (14.2%), and 1,438 were found to be cor- A). Of these, 43 were reported by Heil et al. (1993) rectly determined (70.9%). Based solely on these or the Northern Colorado Plateau Network (in collections, the confi rmed fl ora of Capitol Reef ed.). Taken together, the number of confi rmed National Park stood at 794 taxa (Fertig 2006). and reported vascular plant taxa in Capitol Reef From 2005 to 2007, I located an additional 34 NP stands at 887 taxa (Table 3.1, Appendix A). taxa from collections at Brigham Young Univer- sity (many of which had been reported by Heil At the start of the 2006 fi eld season, I recognized et al. 1993, but without a duplicate voucher de- 216 taxa as potentially occurring in Capitol Reef posited in the CARE herbarium). Field surveys NP, based on the proximity of the park to known

Table 3.1. Statistical summary of the fl ora of Capitol Reef National Park. Number Number Category confi rmed additionally Total in park reported for park Taxonomic diversity Total taxa 835 52 887 (including varieties and subspecies) Full species 786 45 831 (excluding varieties and subspecies) Families 90 1 91 Life form diversity Tree taxa 49 2 51 Shrub taxa 115 8 123 Perennial forb taxa 396 31 427 Annual forb taxa 135 4 139 Perennial graminoid taxa 108 6 114 Annual graminoid taxa 22 1 23 Fern taxa 10 0 10 Biogeographic diversity Introduced taxa 119 3 122 Native taxa Locally endemic taxa 26 1 27 Regionally endemic taxa 110 5 115 Disjunct taxa 1 0 1 Peripheral taxa 8 0 8 Sparse taxa 14 1 15 Widespread taxa 557 42 599 Total native taxa 716 49 765 The number of taxa and families is based on taxonomic concepts of Welsh et al. (2003). See Appendices, “Codes and Defi nitions”, for an explanation of the various categories.

Chapter 3: Results 11 populations (Albee et al. 1988, Fertig 2006). At prise an additional 15% of the park’s native fl ora least 24 of these species were found to occur in (Table 3.1). Disjunct, peripheral, and sparse spe- the park based on fi eld collections or relocated cies make up just 3.1% of the total native fl ora. herbarium specimens in 2005–2007. The current number of potential species that I am recognizing 3.3 Plants of special concern for Capitol Reef NP is 192 (Appendix B). With seven listed species, Capitol Reef National Of the original pool of species documented or re- Park has the largest number of threatened or ported for Capitol Reef NP, I found 117 taxa to endangered plant species of any park unit in the be falsely reported based on misidentifi ed vouch- Northern Colorado Plateau Network (Table 3.3). er specimens in the CARE and BRY herbaria. I At least 18 additional park taxa were formerly consider an additional six species cited (but not candidates for potential listing (Category 2 or vouchered) by the NCPN vegetation mapping 3C) under the Endangered Species Act (UDWR team (NCPN in ed.) to be questionable based on 1998). Eight of these taxa are currently listed as the absence of suitable habitat or the distance of sensitive by the Utah Bureau of Land Manage- Capitol Reef NP to the known or expected range ment and U.S. Forest Service. The Utah Division of each species (Appendix C). of Wildlife Resources Conservation Data Center (UTCDC) lists 61 taxa from Capitol Reef NP as The confi rmed and reported fl ora of Capitol Reef species of concern (UDWR 1998). NP represents 24.7% of the 3,594 native and naturalized taxa documented for the state of Utah Heil’s wild buckwheat (Eriogonum corymbosum by Welsh et al. (2003) and 60.3% of the state’s var. heilii) was described as a new taxon by James 151 plant families (Table 3.1). Within Wayne and Reveal in 2004 (Flora of North America Editorial Garfi eld counties, the park fl ora captures 57.3% Committee 2005) and is apparently restricted to of the 1,547 taxa reported by Welsh et al. (2003). Thousand Lake Mountain in Capitol Reef NP At the ecoregional scale, Capitol Reef NP con- and adjacent Fishlake National Forest. It present- tains 52.4% of the 1,694 species reported for the ly has no legal status but research on its distribu- Utah portion of the Colorado Plateau (Shultz et tion and potential management needs should be al. 2006). considered a high priority. Several recent studies have focused on mapping the distribution of the 3.2 Life form and biogeographic park’s threatened, endangered and sensitive species (Clark 1999a, 1999b, 2001a, 2001b, 2002; Heil diversity et al. 1993), monitoring populations (Fertig et al. Although ecologically dominant, tree and shrub 2005 and Capitol Reef references cited therein), species make up only 19.6% of the confi rmed and and researching pollination biology (Tepedino et reported plant taxa of Capitol Reef National Park al. 2003). (Table 3.1). Perennial forbs are the most species- rich life form in the park’s fl ora, with 427 known 3.4 Non-native species and noxious and reported taxa (48.1% of the total fl ora). About 18% of the fl ora is comprised of annual weeds forbs and grass-like plants. Like trees and shrubs, Introduced plant species (not historically native perennial graminoids are ecologically important to Utah or North America) make up 13.8% of the but relatively species-poor, representing 12.2% confi rmed and reported fl ora of Capitol Reef Na- of the total fl ora. The diversity of ferns and fern- tional Park (Table 3.1). This fi gure is comparable allies in Capitol Reef NP is especially low (1.1%) to the statewide average of 13.5% non-native spe- (Table 3.1). cies (Fertig 2007; Welsh et al. 2003). Of the 122 introduced plant species confi rmed or reported for Of the park’s 765 native taxa, 599 (78.3%) are Capitol Reef NP (Table 3.1), only three—Russian widespread across Utah and western North knapweed (Centaurea repens), fi eld bindweed America. Only 27 species (3.5% of the total native (Convolvulus arvensis), and quackgrass (Elymus fl ora of the park) have extremely limited global repens)—are listed as noxious by the State of Utah distributions (local endemics found within an (State of Utah Department of Agriculture and area of less than 1° of latitude × 2° degrees of lon- Food 2007). Heil et al. (1993) and Dewey and An- gitude; see Appendix A for complete defi nitions dersen (2005) conducted surveys of non-native of biogeographic categories). Regional endemics and noxious weeds and mapped the distribution (largely restricted to the Colorado Plateau) com- of selected species. Since the mid-1980s, several

12 Annotated Checklist of Vascular Flora: Capitol Reef National Park eradication eﬀ orts have been implemented to con- and tree-of-heaven (Ailanthus altissima) along the tain the spread of ﬁ ve-stamen tamarisk (Tamarix Fremont River, Sulphur Creek, and Halls Creek chinensis), Russian olive (Elaeagnus angustifolia), Narrows (Evenden et al. 2002).

Table 3.3. Plant species of conservation concern from Capitol Reef National Park. TNC global UTCDC Family Species Common name rank Federal legal status status Apocynaceae Cycladenia humilis var. Jones’ cycladenia G3G4T2 USFWS: Threatened Rare, S2 jonesii Betulaceae Ostrya knowltonii western hop-hornbeam G3G4 Watch, S2 Boraginaceae Cryptantha capitata head cryptanth G4 Watch, S2? Cactaceae Opuntia basilaris var. Heil’s beavertail G5T2T3 Rare, S1 heilii Cactaceae Opuntia phaeacantha large pricklypear G5T5 Periph., var. major S2S3 Cactaceae Pediocactus winkleri Winkler’s pincushion G2 USFWS: Threatened Rare, S1 cactus Cactaceae Sclerocactus wrightiae Wright’s ﬁ shhook G2 USFWS: Endangered Rare, S2 cactus Campanulaceae Lobelia cardinalis ssp. cardinal-ﬂ ower G5 Periph., S2 graminea Chenopodiaceae Chenopodium graveolens New Mexico goosefoot G5T5 Periph., S1 var. neomexicanum Compositae Aster welshii Welsh’s aster G2 Watch, S3? (Asteraceae) Compositae Chrysothamnus Bigelow’s rabbitbrush G5 Periph., S2 (Asteraceae) nauseosus var. bigelovii* T3T5Q Compositae Cirsium calcareum var. Caineville thistle G4TNR Tax. Prob., (Asteraceae) calcareum SNR Compositae Cirsium neomexicanum Utah thistle G5T3T4 Add. Data, (Asteraceae) var. utahense S3? Compositae Erigeron abajoensis Abajo daisy G1G2 USFS Region 4: Sensitive Add. Data, (Asteraceae) S1S2 Compositae Erigeron maguirei Maguire’s daisy G2 USFWS: Threatened Rare, S2 (Asteraceae) Compositae Erigeron pumilus var. vernal daisy G4G5 Infreq., S2? (Asteraceae) condensatus T3T4 Compositae Gnaphalium wrightii Wright’s cudweed G5 Periph., S2? (Asteraceae) Compositae Hymenoxys acaulis var. low hymenoxys G5T1T2 USFWS: former C2 Rare, S1S2 (Asteraceae) nana candidate Compositae Thelesperma subnudum alpine greenthread G5T2 USFS Region 4: Sensitive; Rare, S1 (Asteraceae) var. alpinum BLM (UT): Sensitive; USFWS: former C2 candidate Compositae Townsendia aprica Last Chance G2 USFWS: Threatened Rare, S1 (Asteraceae) townsendia Compositae Xylorhiza confertifolia Henrieville woodyaster G2G3 USFWS: former 3C Watch, (Asteraceae) S2S3 Cruciferae Arabis demissa nodding rock cress G5T4 USFWS: former 3C Watch, S2? (Brassicaceae)

Chapter 3: Results 13 Table 3.3. Plant species of conservation concern from Capitol Reef National Park, cont.

TNC global UTCDC Family Species Common name rank Federal legal status status Cruciferae Physaria acutifolia var. purple twinpod G5T2 Rare, S2 (Brassicaceae) purpurea Cruciferae Schoenocrambe barnebyi Barneby’s reed-mustard G1 USFWS: Endangered Rare, S1 (Brassicaceae) Euphorbiaceae Tragia ramosa* noseburn G5 Periph., S1 Gramineae Andropogon glomeratus bushy bluestem G5 Periph., S2 (Poaceae) var. scabriglumis Gramineae Festuca sororia ravine fescue G4 Add. Data, (Poaceae) S2 Gramineae Panicum oligosanthes var. Scribner panicgrass G5T5 Infreq., S2? (Poaceae) scribnerianum Gramineae Schizachne purpurascens false melic G5 Periph., S1 (Poaceae) Gramineae Stipa scribneri Scribner needlegrass G4 Add. Data, (Poaceae) SR Hydrophyllaceae Nama retrorsum* Howell’s nama G3 USFWS: former 3C Watch, S2 Leguminosae Astragalus barnebyi Barneby’s milkvetch G5T3 USFWS: former 3C Watch, S2 (Fabaceae) Leguminosae Astragalus brandegei Brandegee’s milkvetch G3G4 USFWS: former C2 Watch, S2 (Fabaceae) candidate Leguminosae Astragalus consobrinus Bicknell’s milkvetch G2G3 USFS Region 4: Sensitive Watch, (Fabaceae) S2S3 Leguminosae Astragalus desperatus rockloving milkvetch G5T3 Watch, S3 (Fabaceae) var. petrophilus Leguminosae Astragalus ﬂ avus var. clay milkvetch G5T3 Tax. Prob., (Fabaceae) argillosus SNR Leguminosae Astragalus harrisonii Harrison’s milkvetch G2G3 USFWS: former 3C Watch, (Fabaceae) S2S3 Leguminosae Astragalus laccoliticus laccolite milkvetch G5T2? Rare, S2? (Fabaceae) Leguminosae Astragalus malacoides Kaiparowits milkvetch G3 USFWS: former 3C Watch, S3 (Fabaceae) Leguminosae Astragalus pardalinus panther milkvetch G3? USFWS: former 3C Watch, S3? (Fabaceae) Leguminosae Astragalus wardii* Ward’s milkvetch G3 Watch, S3 (Fabaceae) Leguminosae Astragalus woodrufﬁ i Woodruff’s milkvetch G3 USFWS: former 3C Watch, S3 (Fabaceae) Leguminosae Psoralidium junceum rush scurfpea G3 Watch, S3 (Fabaceae) Leguminosae Psoralidium lanceolatum slenderleaf scurfpea G5TNR Tax. Prob., (Fabaceae) var. stenophyllum S3? Leguminosae Psorothamnus Thompson’s indigo- G3?T3? USFWS: former 3C Watch, S3? (Fabaceae) thompsoniae var. bush thompsoniae* Nyctaginaceae Mirabilis linearis var. New Mexico G5T5 Tax. Prob., decipiens umbrellawort S2?

14 Annotated Checklist of Vascular Flora: Capitol Reef National Park Table 3.3. Plant species of conservation concern from Capitol Reef National Park, cont.

TNC global UTCDC Family Species Common name rank Federal legal status status Orchidaceae Habenaria zothecina alcove bog-orchid G2 BLM (UT): Sensitive; Rare, S2 USFWS: former C2 candidate Orchidaceae Spiranthes roman- Ute ladies’-tresses G2 USFWS: Threatened Rare, S1 zofﬁ ana var. diluvialis (S. diluvialis)** Polemoniaceae Gilia caespitosa Rabbit Valley gilia G2 USFS Region 4: Sensitive; Rare, S1 USFWS: former C1 candidate Polemoniaceae Gilia imperialis cataract gilia G4T2 BLM (UT): Sensitive Rare, S2 Polemoniaceae Gilia tenuis Mussentuchit gilia G1 BLM (UT): Sensitive; Rare, S1 USFWS: former C2 candidate Polygonaceae Eriogonum corymbosum Heil’s wild buckwheat G5T? var. heilii Polygonaceae Eriogonum leptophyllum slenderleaf wild buck- G3G4 Watch, S1 wheat Polygonaceae Eriogonum tumulosum woodside wild G3Q USFWS: former 3C Watch, S2 buckwheat Polypodiaceae Pellaea limitanea border cloak-fern G4G5 Periph., S1 (Adiantaceae, Sinopteridaceae) Polypodiaceae Woodsia oregana Oregon woodsia G5T4? Periph., S1 (Aspleniaceae, Woodsiaceae) Primulaceae Primula specuicola cave primrose G4Q USFWS: former 3C Watch, S3 Scrophulariaceae Penstemon caespitosus painted penstemon G5T3? Watch, S2? var. desertipicti Scrophulariaceae Penstemon scariosus var. plateau penstemon G4T3T4 Watch, scariosus S3S4 Umbelliferae Cymopterus beckii Beck’s spring-parsley G1 USFS Region 4: Sensitive; Rare, S1 (Apiaceae) BLM (UT): Sensitive; USFWS: former C2 candidate Umbelliferae Lomatium junceum rush lomatium G3 Watch, (Apiaceae) S2S3 Vitaceae Parthenocissus vitacea thicket-creeper G5 Periph., S1 This list is derived from Utah Division of Wildlife Resources (1998). All species are conﬁ rmed as present and extant unless indicated by “*” (reported) or “**” (historical or extirpated).

Codes: TNC global rank assesses abundance and conservation priority on a scale of 1–5 (1 being extremely vulnerable and 5 being secure) for full species (G) and varieties or subspecies (T) across their entire range. A “?” indicates uncertainty in the rank, Q = taxonomic questions, U = unknown, and NR = not ranked. A comparable scoring system is used to measure state abundance (S) in Utah under the column UTCDC status.

Under legal status, USFWS = US Fish and Wildlife Service. C2 = category 2 candidate (a former category used for taxa that might warrant being proposed for Threatened or Endangered status following additional research). C3 = category 3 candidates (species dropped from consideration for listing). USFS = U.S. Forest Service. BLM = U.S. Bureau of Land Management.

UTCDC status includes conservation categories adopted by the state natural heritage program to prioritize endemic and rare plant taxa (Utah Division of Wildlife Resources 1998). These categories include: Historical (presumed extinct or only known historically through their range), Rare (plants with rangewide viability concerns), Watch (regional endemics without rangewide viability concerns), Peripheral (rare or uncommon in Utah, but more common rangewide), Infrequent (plants occur infrequently over western US), Taxonomic Problems (validity of species, subspecies, or variety has been questioned), Additional Data (species appear rare, but more information needed before being assigned to a different category).

Chapter 3: Results 15 16 Annotated Checklist of Vascular Flora: Capitol Reef National Park nearly 18%. Many of the “missing” potential or 4 Discussion reported taxa consist of cryptic species that fl ower late in the growing season, or occur in under- Data on the fl ora of Capitol Reef National Park sampled wetland or weedy habitats. Confi rmation developed slowly in the early decades of the of reported species should remain a high priority, park’s existence, but greatly increased starting in as reports for some of these taxa may ultimately 1970, with the eff orts of Ruth Nelson and, later, prove to be erroneous. Work is especially needed Susan Meyer. Over 40% of the park’s fl ora (and a in the upper Deep Creek and Halls Creek drain- large percentage of its rarest species) were discov- ages, at the extreme northern and southern ends ered during the fl oristic and vegetation studies of of the park, to confi rm species reported by Heil Heil et al. (1993) in the late 1980s. Presently, the et al. (1993) and Camp (1978) that may actually documented and reported fl ora of Capitol Reef occur outside the park. Additional studies would National Park stands at 887 taxa (Tables 3.1 and also be useful to better refi ne the abundance 3.3, Figure 1.3.4, Appendix A). Based on available and distribution of species of high management habitats and proximity, at least another 192 plant interest (especially rare plants and invasive non- taxa may occur within the park (Appendix B). natives). Should all of these potential species be found, the documented fl ora of the park would increase by

Chapter 4: Discussion 17 18 Annotated Checklist of Vascular Flora: Capitol Reef National Park Dixie National Forest, Bureau of Land 5 Literature Cited Management Richfi eld Field Offi ce, Capitol Reef National Park, and Fishlake National Albee, B. J., L. M. Shultz, and S. Goodrich. 1988. Forest. Atlas of the vascular plants of Utah. +Occa- sional Publ. 7. Salt Lake City: Utah Museum Cronquist, A. 1994. Volume 5, Asterales, Cron- of Natural History. 670 pp. quist, A., A. H. Holmgren, N. H. Holmgren, J. L. Reveal, and P. K. Holmgren, eds., Barkworth, M. E., K. M. Capels, S. Long, and M. Intermountain fl ora: Vascular plants of the B. Piep, eds. 2003. Magnoliophyta: Com- Intermountain West, U.S.A. Bronx, N.Y.: melinidae (in part): Poaceae, part 2. Flora New York Botanical Garden. 496 pp. of North America north of Mexico, volume 25. New York: Oxford University Press. 783 Cronquist, A., A. H. Holmgren, N. H. Holmgren, pp. and J. L. Reveal, eds. 1972. Volume 1, Geo- logical and botanical history of the region, Barkworth, M. E., K. M. Capels, S. Long, L. its plant geography and a glossary. The K. Anderton, and M. B. Piep, eds. 2007. vascular cryptogams and the gymnosperms. Magnoliophyta: Commelinidae (in part): Intermountain fl ora: Vascular plants of the Poaceae, part 1. Flora of North America Intermountain West, U.S.A. Bronx, N.Y.: north of Mexico, volume 24. New York: New York Botanical Garden. 270 pp. Oxford University Press. 911 pp. Cronquist, A., A. H. Holmgren, N. H. Hol- Barneby, R. C. 1989. Volume 3, Part B. Fabales, mgren, J. L. Reveal, and P. K. Holmgren, in Cronquist, A., A. H. Holmgren, N. H. eds. 1977. Volume 6, The monocotyledons. Holmgren, J. L. Reveal, and P. K. Holmgren, Intermountain fl ora: Vascular plants of the eds., Intermountain fl ora: Vascular plants Intermountain West, U.S.A. Bronx, N.Y.: of the Intermountain West, U.S.A. Bronx, New York Botanical Garden. 584 pp. N.Y.: New York Botanical Garden. 279 pp. —— . 1984. Volume 4, subclass Asteridae (except Camp, P. A. 1978. Ecological reconnaissance, Asteraceae). Intermountain fl ora: Vascular Halls Creek, Capitol Reef National Park. plants of the Intermountain West, U.S.A. Department of Botany, Utah State Univer- Bronx, N.Y.: New York Botanical Garden. sity, Logan, Ut. 573 pp. Chronic, H. 1988. Pages of stone, geology of Cronquist, A., N. H. Holmgren, and P. K. western national parks and parks: Grand Holmgren, eds. 1997. Volume 3, Part A, Canyon and the Plateau Country. Seattle, subclass Rosidae (except Fabales). In- Wa.: The Mountaineers. 158 pp. termountain fl ora: Vascular plants of the Clark, D. J. 1999a. 1999 fi eld survey and fi nal re- Intermountain West, U.S.A. Bronx, N.Y.: sults of a two-year Winkler and San Rafael New York Botanical Garden. 446 pp. cactus study. Capitol Reef National Park, Dewey, S., and K. Andersen. 2005. An inventory Resource Management Division. 55 pp. of invasive non-native plants in Capitol —— . 1999b. Survey results for Barneby’s reed- Reef National Park (2004): Final report. mustard (Schoenocrambe barnebyi). Capitol Utah State University, Logan, Ut. Weed Sci- Reef National Park, Resource Management ence Research Project Report # SD0514A. Division. 3 pp. Dorn, R. D. 2001. Vascular plants of Wyoming. —— . 2001a. 2001 survey results for Rabbit Valley Third edition. Cheyenne, Wy.: Mountain gilia (Gilia caespitosa = Aliciella caespitosa). West Publishing. 412 pp. Fishlake National Forest and Capitol Reef Evenden, A., M. Miller, M. Beer, E. Nance, S. National Park. 5 pp. Daw, A. Wight, M. Estenson, and L. Cudlip. —— . 2001b. 2001 survey results for Mussen- 2002. Northern Colorado Plateau Vital tuchit gilia (Gilia tenuis). Capitol Reef Signs Network and Prototype Cluster, plan National Park and Bureau of Land Manage- for natural resources monitoring: Phase ment Richfi eld Field Offi ce. 7 pp. I report, October 1, 2002 [two volumes]. National Park Service, Northern Colorado —— . 2002. Summary of the Interagency Rare Plateau Network, Moab, UT. 138 pp + app. Plant Inventory Project, 1999 through 2002.

Chapter 5: Literature Cited 19 Fertig, W. 2006. Herbarium and fi eld studies of north of Mexico. Volume 20. New York: vascular plant fl ora of CARE for National Oxford University Press. 666 pp. Park Service Inventory and Monitoring —— . 2006c. Magnoliophyta: Asteridae, part 8: program. Invesigator’s Annual Report, Asteraceae, part 3. Flora of North America National Park Service. 2 pp. north of Mexico. Volume 21. New York: Fertig, W. 2007. Introduced and naturalized Oxford University Press. 616 pp. plants of Utah. Sego Lily 30(5):7–11. Fowler, J. F., N. L. Stanton, and R. L. Hartman. Fertig, W., M. Beer, and R. DenBleyker. 2005. 2007. Distribution of hanging garden Monitoring protocol for selected threat- vegetation associations of the Colorado ened, endangered, and sensitive plant Plateau, USA. Journal Botanical Research species of Capitol Reef National Park and Institute Texas 1(1):585-607. partner federal land management agen- Frye, B. J. 1998. From barrier to crossroads: cies. Northern Colorado Plateau Network, An administrative history of Capitol Reef Moab, UT. 28 pp. + 12 appendices. National Park. Volumes I & II. U.S. Depart- Flora of North America Editorial Committee. ment of Interior, National Park Service, 1993. Pteridophytes and Gymnosperms. Intermountain Region, Cultural Resources Flora of North America north of Mexico, Selections, Denver, Co. volume 2. New York: Oxford University Heil, K. D., J. M. Porter, R. Fleming, and W. H. Press. 475 pp. Romme. 1993. Vascular fl ora and vegeta- —— . 1997. Magnoliophyta: Magnoliidae and tion of Capitol Reef National Park, Utah. Hamamelidae. Flora of North America Technical Report NPS/NAUCARE/NRTR- north of Mexico, volume 3. New York: 93/01. Oxford University Press. 590 pp. Holmgren, N. H., P. K. Holmgren, and A. —— . 2000. Magnoliophyta: Alismatidae, Are- Cronquist. 2005. Volume 2, Part B, subclass cidae, Commelinidae (in part), and Zingi- Dilleniidae. Intermountain fl ora: Vascular beridae. Flora of North America north of plants of the Intermountain West, U.S.A. Mexico, volume 22. New York: Oxford Bronx, N.Y.: New York Botanical Garden. University Press. 352 pp. 488 pp. —— . 2002a. Magnoliophyta: Commelinidae (in Margules, C. R., and R. L. Pressey. 2000. Sys- part): Cyperaceae. Flora of North America tematic conservation planning. Nature north of Mexico, volume 23. New York: 405:243–253. Oxford University Press. 608 pp. McDougall, W. B. 1940. Special report: Capitol —— . 2002b. Magnoliophyta: Liliidae: Liliales Reef National Monument. Capitol Reef Na- and Orchidales. Flora of North America tional Park archives. north of Mexico, volume 26. New York: Meyer, S. E. 1982. Plant checklist: Capitol Reef Oxford University Press. 723 pp. National Park. Capitol Reef Natural History —— . 2003. Magnoliophyta: Caryophyllidae, part Association, Torrey, Ut. 1. Flora of North America north of Mexico. —— . 1990. Places in the Sun: Story of Capitol Volume 4. New York: Oxford University Reef Plants. Capitol Reef Natural History Press. 559 pp. Association, Torrey, Ut. —— . 2005. Magnoliophyta: Caryophyllidae, part Morris, T. H., V. Wood Manning, and S. M. Rit- 2. Flora of North America north of Mexico. ter. 2003. Geology of Capitol Reef National Volume 5. New York: Oxford University Park. Pages 84–105 in Sprinkel, D. A., T. Press. 656 pp. C. Chidsey, Jr., and P. B. Anderson, eds., —— . 2006a. Magnoliophyta: Asteridae, part 6: Geology of Utah’s parks and monuments. Asteraceae, part 1. Flora of North America Utah Geological Association Publication 28 north of Mexico. Volume 19. New York: (second edition). Oxford University Press. 579 pp. National Park Service. 2000. Management poli- —— . 2006b. Magnoliophyta: Asteridae, part 7: cies. U.S. Department of Interior, National Asteraceae, part 2. Flora of North America Park Service, Washington, DC. Publication

20 Annotated Checklist of Vascular Flora: Capitol Reef National Park # NPS D1416. 137 pp. Utah Division of Wildlife Resources (UDWR). 1998. Inventory of sensitive species and Northern Colorado Plateau Network (NCPN). ecosystems in Utah. Endemic and rare in ed. Capitol Reef National Park vegetation plants of Utah: An overview of their distri- classifi cation and mapping project. Natural bution and status. Report prepared for the Resource Technical Report. National Park Utah Reclamation Mitigation and Conser- Service, Denver, Co. vation Commission and U.S. Department of Noss, R. F., and A. Y. Cooperrider. 1994. Saving the Interior. 566 pp + app. Nature’s legacy: Protecting and restoring Weber, W. A., and R. C. Wittmann. 2001. Colo- biodiversity. Washington, D.C.: Island Press. rado fl ora: Western Slope. Third edition. 416 pp. Boulder: University Press of Colorado. 488 Shultz, L. M., R. D. Ramsey, and W. Lindquist. pp. 2006. Revised atlas of Utah plants. College Welsh, S. L. 1982. Utah plant types: Historical of Natural Resources, Utah State University, perspective, 1840 to 1981. Annotated list Logan, Ut. http://earth.gis.usu.edu/plants. and bibliography. Great Basin Naturalist Springer, A. E., L. E. Stevens, and R. Harms. 42:1–44. 2006. Inventory and classifi cation of —— . 1989. On the distribution of Utah’s hanging selected National Park Service springs on gardens. Great Basin Naturalist 49:1–30. the Colorado Plateau. Northern Arizona University, Flagstaff , Az. NPS Coopera- Welsh, S. L., N. D. Atwood, S. Goodrich, and L. tive Agreement # CA 1200-99-009; Task # C. Higgins. 2003. A Utah fl ora. Third edition, NAU-118. revised. Provo, Ut.: Brigham Young Univer- sity. 912 pp. State of Utah Department of Agriculture and Food. 2007. Utah noxious weed list. hhttp:// ag.utah.gov/plantind/noxious_weeds.html. Tepedino, V. J., D. Alston, and T. Toler. 2003. Integrated pest management: T&E pollina- tors. Interim Report, December. USDA- ARS Bee Biology and Systematics Labora- tory, Department of Biology, Utah State University, Logan, Ut.

Chapter 5: Literature Cited 21 22 Annotated Checklist of Vascular Flora: Capitol Reef National Park Synonyms/Taxonomic notes (Appendices A, B, Appendices and C) Pertinent synonyms are included for species with The following appendices contain annotated alternative scientifi c names in other recent state checklists of the vascular fl ora of Capitol Reef and regional fl oras (Barkworth et al. 2003, 2007; National Park. Barneby 1989; Cronquist 1994; Cronquist et Appendix A lists all species that have been con- al. 1972, 1977, 1984, 1997; Dorn 2001; Flora of fi rmed for the fl ora with an authenticated vouch- North America Editorial Committee 1993, 1997, er specimen (including extant and historical 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, records) or that have been reliably reported for 2006c; Holmgren et al. 2005; Weber and Witt- the park in the literature (Dewey and Ander- mann 2001). No eff ort has been made to identify sen 2005; Heil et al. 1993; Meyer 1982; NCPN in additional synonyms from monographs and fl or- ed.). Vouchers were examined from the CARE as preceding the early 1970s. herbarium and collections from Brigham Young Common name (Appendices A, B, C, and D) University (BRY), Utah State University (USU), the University of Wyoming (RM), and the digi- Common names are derived primarily from tal museum database of the New York Botanical Welsh et al. (2003) and the Intermountain Flora Garden (NY). Species entries in Appendix A are (Barneby 1989; Cronquist 1994; Cronquist et al. organized alphabetically by family and genus. 1972, 1977, 1984, 1997; Holmgren et al. 2005).

Appendix B includes species that may potentially Life form (Appendices A, B, C, and D) occur within Capitol Reef National Park based on Life form is based on the stature and growth habit the proximity of the park to other populations or of typical members of a species at reproductive the presence of suitable habitat. maturity. Trees are woody perennials with a single stem or trunk and are typically over 3.5 m tall. Appendix C lists species that have been previ- Shrubs are woody perennials with one to many ously reported for Capitol Reef National Park but trunks arising from near the base of the plant and are now considered erroneous because they are are usually less than 3.5 m tall. Perennial Forbs based on misidentifi ed specimens or question- (PerF) are broad-leaved dicots or monocots able literature records. with non-woody stems (at least above ground level) that live for multiple years. Annual Forbs Appendix D contains the same confi rmed, his- (AnnF) are broad-leaved dicots or monocots torical, and reported taxa as in Appendix A, but with non-woody stems that complete their life organized by life form rather than by family in or- cycle (mature, fl ower, and die) in one year. Peren- der to better meet the needs of NPS interpretive nial Graminoids (PerG) are linear-leaved, grass- specialists. like monocots that live for multiple years. Annual Graminoids (AnnG) are linear-leaved, grass-like The codes and references for each fi eld in the ap- monocots that complete their life cycle (mature, propriate appendices are summarized below. fl ower, and die) in one year. Ferns and fern-allies Family (Appendices A, B, C, and D) (Fern) are non-fl owering vascular plants that re- produce by spores. Family concepts and nomenclature follow A Utah Flora (Welsh et al. 2003). Alternative family names Range (Appendices A, B, and C) and treatments from other regional fl oras (Dorn Range represents the distribution of a species 2001; Flora of North America Editorial Commit- within Utah and the state’s contribution to the tee 1993, 2002b; Holmgren et al. 2005; Weber and plant’s overall global range. Disjunct (Disj) taxa Wittman 2001) are listed in parentheses. have their Utah distribution separated from the Species name (Appendices A, B, C, and D) main, contiguous portion of their range by a gap of more than 800 km (ca 500 miles). Introduced Nomenclature for scientifi c names and taxonom- (Intro) taxa are those that are not native to Utah ic concepts is derived from Welsh et al. (2003). or North America but that have become natu- Scientifi c names are listed in alphabetical rather ralized (breeding on their own without human than phylogenetic order. assistance). Local Endemics (LocEn) are taxa

Appendices 23 whose entire global range is restricted to an area Rare taxa usually have low population numbers of less than 16,500 km2 (ca 6,370 mi2, or 1° of lati- and are restricted to a single location within the tude × 2° of longitude). Peripheral (Periph) taxa park. A “?” indicates that population size is un- have a widespread global distribution but occur known. at the margin of their contiguous range in Utah and occupy less than 5% of the state’s area (usu- Source (Appendix A) ally only within a few miles of the state border). Sources are the evidence for the assigned park Regional Endemics (RegEn) are taxa with a glob- status. In the case of Present and Historical taxa, al range of 16,500–250,000 km2 (an area about the sources are herbarium records (with collector, size of the state of Wyoming). Sparse taxa occur collection number, and repository) that were cor- widely across Utah or North America, but their roborated during this study. For reported taxa, the range within Utah is small and patchy, with popu- source is a published or unpublished reference lations restricted to specialized or uncommon citing the species within the park. Specimen re- habitats. Widespread (Wide) taxa have global positories include the Capitol Reef National Park ranges exceeding 250,000 km2 and occur over at herbarium (CARE), Brigham Young University least 10% of the state. (BRY), Utah State University (USU), New York Botanical Garden (NY), and the University of Park status (Appendices A, C, and D) Wyoming’s Rocky Mountain Herbarium (RM). Park status identifi es whether a species is Pres- ent (Pres), Historical (Hist), or Reported (Rep) Year doc. (Appendix A) for Capitol Reef National Park. Species are con- Year documented indicates the fi rst year in which sidered Present if a confi rmed voucher or docu- a species was confi rmed or reported to occur mented observation has been made within park in Capitol Reef National Park. This year does boundaries since 1970. Historical species have not necessarily correspond with the date of the been confi rmed with a voucher from the park but voucher specimen cited under Source. “NA” have not been relocated since before 1970. Re- indicates that the fi rst year is not known (used ported species are listed for the park in the litera- mostly for “reported” taxa). ture (Dewey and Andersen 2005; Heil et al. 1993; Meyer 1982; NCPN in ed.), but have not been Comments (Appendices A, B, C, and D) corroborated with a voucher specimen. Species This column includes additional comments on that may potentially occur in CARE are listed in the status of a particular species in the park, the Appendix B. Rejected taxa (listed in Appendix C) place of origin for introduced species, legal sta- have either been falsely reported (FalsRep) in the tus, and other data. literature based on misidentifi ed herbarium specimens or are considered Questionable (Ques?) Nativity (Appendix D) because park reports are from well outside their Nativity indicates whether a species is native known or expected range. to Utah or Introduced (Intro), based on Fertig Pop. size (Appendix A) (2007). Population size refers to the abundance and fre- Flower time (Appendix D) quency with which a species occurs within Capitol Flower time refers to the months in which a spe- Reef National Park. Abundance data are derived cies is most likely to be in fl ower. NA indicates from Heil et al. (1993), Northern Colorado Pla- a species that is a non-fl owering fern or gymno- teau Network (in ed.), and personal observations. sperm. Abundant (Abun) taxa have large populations, have a broad ecological amplitude (often being Habitat (Appendix D) dominant in one to several vegetation types), The distribution of species across generalized or occur extensively across the park. Common habitats in Capitol Reef National Park is indi- (Com) taxa have large local populations but may cated by an “x” in the columns SaltDes through be restricted to a single vegetation type or do not Wet and is derived from the Capitol Reef Na- occur across the entire park. Uncommon (Unc) tional Park vegetation classifi cation (Northern taxa have low-to-medium-sized populations, Colorado Plateau Network, in ed.), Heil et al. typically have a narrow ecological amplitude (or (1993), Meyer (1990), and Welsh et al. (2003). at least contribute relatively little to total vegeta- SaltDes=Salt desert scrub dominated by shad- tive cover), or have a small range across the park. scale (Atriplex confertifolia), Castle Valley salt-

24 Annotated Checklist of Vascular Flora: Capitol Reef National Park bush (A. gardneri var. cuneata), and blackbrush rock. MixCon=Mixed conifer forests of ponde- (Coleogyne ramosissima). Sage=sagebrush grass- rosa pine (Pinus ponderosa) and quaking aspen lands dominated by big sagebrush (Artemisia tri- (Populus tremuloides). Wet=wetlands associated dentata), fourwing saltbush (Atriplex canescens), with perennial streams, seeps, and hanging gar- and rubber rabbitbrush (Chrysothamnus nauseo- dens. Additional comments are included for spe- sus). PJ=two-needle pinyon (Pinus edulis)/Utah cies that are restricted to a particular soil/geologic juniper (Juniperus osteosperma) woodlands/rim- type or are found mostly in disturbed sites.

Appendices 25 A. is c name with America America Helper, Carbon Co, Helper, s.n. MO) UT (Trelease eastwoodiana Grand Moab vicinity, Co, UT (Rydberg & Garrett 8468 NY) Comments doc. Year Year 1958 1966 1986 Native to Central 1986 1980 Native to tropical 1986 1986 locality is Type 1981 locality of Type 1978 1987 1958 1987 1986 (CARE) (CARE) (CARE) (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Rare Heil 2747 PerF Wide Pres Unc Meyer 4727 Shrub RegEn Pres Unc Camp 548 Shrub Wide Pres Unc Heil 2617 Life form squawbush Shrub Wide Pres Abun Swancara 19 squawbush Shrub Wide Pres Unc Heil 2553 redroot pigweed AnnF Intro Pres Unc Porter 4372 prostrate pigweed AnnF Wide Pres Unc Porter 4118 tumble pigweed AnnF Intro Pres Unc Meyer 4629 northern arrowhead Harriman’s yuccaHarriman’s Shrub Wide Pres Com Porter 3828 tomentose amsonia narrowleaved yucca Jones’ amsonia PerF RegEn Pres Unc Porter 4666 box elder Tree Wide Pres Unc Harrison 1290 western poison ivy Shrub Wide Pres Unc Fleming 368 Rocky Mountain maple Common name

var.

var. var. & ssp. Rhus radicans, T. Rhus radicans, T. radicans R. trilobata trilobata R. trilobata simplicifolia A. graecizans A. tomentosa, eastwoodiana Y. angustissima Y. angustissima interius interius, Negundo aceroides tripartitum Includes vars. neomexicanum, douglasii, Synonyms/ notes Taxonomic

var. var. exus trilobata simplicifolia harimaniae Toxicodendron Toxicodendron rydbergii Rhus aromatica var. Rhus aromatica var. Amaranthus retrofl Amaranthus blitoides Amaranthus albus Sagittaria cuneata Yucca harrimaniae Yucca var. Amsonia tomentosa stenophylla Yucca Yucca angustissima Amsonia jonesii Acer negundo A negundo glabrum Acer glabrum vascular plant taxa of Capitol Reef National Park, organized by family and scientifi rmed, historical, and reported Anacardiaceae Anacardiaceae Anacardiaceae Amaranthaceae Amaranthaceae Amaranthaceae Alismataceae Agavaceae Apocynaceae Agavaceae Apocynaceae Aceraceae FamilyAceraceae Species name taxonomic and biogeographic notes. Appendix A. Confi

26 Annotated Checklist of Vascular Flora: Capitol Reef National Park Herbarium based on collections from outside the park. Spring Canyon SE of Castledale, Emery Co, UT (Maguire 18310 UTC) Capitol Wash, Capitol Wash, Co, UT (Jones Wayne 5650 POM) San Rafael Swell, Emery Co, UT (Jones s.n. CAS). USFWS: Threatened Comments NA Reports from CARE doc. Year Year 1989 Native to Japan 1958 1981 locality is Calf Type 1971 1987 1958 locality is Type 1973 1974 1987 locality is Type 1973 c name with taxonomic and (1993) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range Shrub Intro Pres Unc Haskell s.n. PerF Wide Pres Rare Porter 5201 PerF RegEn Pres Unc Nelson 9175 Life form whorled milkweed PerF Wide Rep Thunberg’s barberry NA Heil et al. showy milkweed PerF Wide Pres Unc Porter 3973 Ruth’s milkweedRuth’s PerF RegEn Pres Unc Porter 3770 bigseed milkweed PerF RegEn Pres Unc Meyer 4522 broadleaf milkweed Circle Cliffs Circle Cliffs milkweed pallid milkweed PerF Wide Pres Unc Harrison 935 spider milkweed PerF Wide Pres Rare Harrison 1402 Jones’ cycladenia PerF RegEn Pres Rare Heil 3111 common dogbane PerF Wide Pres Unc Harrison 1185 Common name

& by some A. ssp. A. galioides B. thungbergii ruthiae Included in involucrata authors A. cryptoceras C. jonesii angustifolium Includes vars. cannabinum, glaberrimum, Synonyms/ notes Taxonomic

var. asperula jonesii Asclepias subverticillata Berberis thunbergii Asclepias speciosa Asclepias ruthiae A. uncialis Asclepias macrosperma Asclepias latifolia Asclepias labriformis Asclepias cryptoceras cryptoceras Asclepias asperula var. Cycladenia humilis var. Apocynum cannabinum rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Asclepiadaceae Berberidaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Apocynaceae biogeographic notes, cont. Appendix A. Conﬁ FamilyApocynaceae Species name

Appendices 27 Mts, San Juan Co, UT (Rydberg & Garrett 9320 NY) America Comments doc. Year Year 1980 locality is Elk Type 1971 1981 1988 1971 1974 Native to E North 1977 1985 Native to Eurasia 1973 1980 1980 1980 1958 1981 1987 Native to Eurasia c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range Tree RegEn Pres Rare Camp s.n. Tree Intro Pres Unc Heil s.n. PerF Wide Pres Com Fleming 355 Life form San Juan cryptanth PerF RegEn Pres Unc Meyer 4547 James’ cryptanth PerF Wide Pres Unc Camp 444 ashy cryptanth PerF Wide Pres Com Heil 3265 head cryptanth PerF RegEn Pres Unc Heil 4423 low cryptanth PerF RegEn Pres Com Meyer 4760 showy catalpa Tree Intro Pres Unc Harrison 1356 western hop- hornbeam European white birch water birch Tree Wide Pres Unc Harrison 1190 thinleaf alder Shrub Wide Pres Unc Meyer 4677 Oregon-grape Shrub Wide Pres Unc Meyer 4533 basin yellow cryptanth Fremont’s mahoniaFremont’s Shrub Wide Pres Com Harrison 1377 cushion cryptanth AnnF Wide Pres Unc Porter 2396 common barberry Shrub Intro Pres Unc Fleming 259 Common name

ssp. C. atwoodii C. pustulosa C. jamesii, Oreocarya suffruticosa Includes A. incana tenuifolia, A. tenuifolia Berberis fremontii Synonyms/ notes Taxonomic

var.

ora var. var. var. Cryptantha circumscissa Cryptantha cinerea pustulosa Cryptantha cinerea jamesii Cryptantha cinerea cinerea Cryptantha capitata Cryptantha abata Catalpa speciosa Ostrya knowltonii Betula pendula Betula occidentalis B. fontinalis Alnus incana occidentalis Mahonia repens Berberis repens Cryptantha confertifl Mahonia fremontii Berberis vulgaris rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Bignoniaceae Betulaceae Betulaceae Betulaceae Betulaceae Berberidaceae Boraginaceae Berberidaceae biogeographic notes, cont. Appendix A. Confi FamilyBerberidaceae Species name

28 Annotated Checklist of Vascular Flora: Capitol Reef National Park eld is Paria Monument Park (Colorado NM), near Grand Junction, Mesa Co., CO (Osterhout 6138 RM) “Southern Utah” (Parry 164 ) Emery, Emery Co, UT Emery, (Jones 5445p POM) echinoides Canyon, Garﬁ Co, UT (Jones 5297p POM) Comments doc. Year Year 1982 locality is Type 1987 locality is Type 1973 locality is Type 1987 1980 1973 locality of var. Type 1971 1971 1987 1980 1981 1986 c name with taxonomic and Goodrich 8643 (BRY) (CARE) (CARE) BC318 (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF RegEn Pres Unc Atwood & PerF Wide Pres Com Meyer 4483 PerF Wide Pres Abun Harrison 909 AnnF Wide Pres Unc Porter 5157 AnnF Wide Pres Unc Meyer 4523 Life form Osterhout’s Osterhout’s cryptanth redroot cryptanth AnnF Wide Pres Rare Porter 4676 carbon cryptanth PerF RegEn Pres Unc Heil 3275 dwarf cryptanth PerF Wide Pres Unc Fleming slender cryptanth AnnF Wide Pres Com Meyer 4527 plateau cryptanth PerF RegEn Pres Abun Harrison 828 yellow-eye cryptanth plateau yellow cryptanth sand dune cryptanth recurved cryptanth AnnF Wide Pres Com Meyer 4475 thick-sepaled cryptanth wing-nut cryptanth AnnF Wide Pres Unc Porter 2513 Common name

& ava nana avoculata Oreocarya osterhoutii Oreocarya mensana Oreocarya humilis ssp. echinoides Oreocarya fulvocanescens. Includes vars. fulvocanescens Oreocarya ﬂ Synonyms/ notes Taxonomic

avafl Oreocarya var. var. avoculata Cryptantha pterocarya Cryptantha osterhoutii Cryptantha micrantha Cryptantha mensana Cryptantha humilis Cryptantha gracilis Cryptantha fulvocanescens Cryptantha fl Cryptantha fl Cryptantha fendleri Cryptantha recurvata Cryptantha crassisepala elachantha pterocarya rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae biogeographic notes, cont. Appendix A. Confi FamilyBoraginaceae Species name

Appendices 29 Lake, Sevier Co, UT s.n. GH) (Ward miles S of Ouray, miles S of Ouray, Uintah Co, UT (Rollins 1715 GH). USFWS: former C2 candidate Hanksville, Wayne Hanksville, Wayne Co, UT (Neese 5938 BRY) Comments doc. Year Year 1987 1981 1981 1971 1980 1987 1986 locality is Fish Type 1986 1988 locality is 22 Type 1988 locality is sw of Type c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Romme 674 (BRY) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Meyer 4770 AnnF Wide Pres Unc Fleming 512 Life form Arizona pincushion PerF Wide Pres Unc Heil 1 (CARE) 1986 claretcup PerF Wide Pres Abun Fleming 646 matted tiquilia PerF RegEn Pres Unc Heil 2859 southwestern stoneseed showy stoneseed PerF Wide Pres Com Harrison 840 cupseed stickseed AnnF Wide Pres Com Meyer 4466 bindweed heliotrope bristly cryptanth PerF RegEn Pres Unc Porter 3859 brittle pricklypear PerF Wide Pres Com Heil 3008 Rollins’ cryptanth PerF RegEn Pres Unc Fleming & Heil’s beavertailHeil’s PerF RegEn Pres Rare Fleming 630 Common name

ssp. Vars. Vars.

inermis. var. var. var. (2003) (2003). arizonica. Flora of North Flora of North O. basilaris Escobaria vivipara var. not recognized in America Includes var. Includes var. Vars. not recognized Vars. in America hispidissima L. texana, redowskii cupulata, L. marginata californica Euploca convolvulacea brachyartha Oreocarya rollinsii longiareolata Synonyms/ notes Taxonomic

var. var.

var. orum heilii Opuntia basilaris var. Coryphantha vivipara arizonica Echinocereus triglochidiatus melanacanthus Tiquilia latiorTiquilia Coldenia Lithospermum multifl Lithospermum incisum Lappula occidentalis cupulata Heliotropium convolvulaceum Cryptantha setosissima Opuntia fragilis O. fragilis Cryptantha rollinsii rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Cactaceae Cactaceae Cactaceae Boraginaceae (Ehretiaceae) Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Cactaceae biogeographic notes, cont. Appendix A. Confi FamilyBoraginaceae Species name

30 Annotated Checklist of Vascular Flora: Capitol Reef National Park S. is Glen orus parvifl Capitol Reef, Wayne Capitol Reef, Wayne Co, UT (Winkler s.n. UNM). USFWS: Threatened Rafael Ridge, Emery Co, UT (Benson & Benson 16595 POM). USFWS: Endangered Canyon, San Juan Co, UT (Clover & Jotter 2398 US) Comments doc. Year Year 1973 locality of Type 1986 locality is E of Type 1986 1975 1988 1987 1986 1991 1987 locality is San Type 1988 c name with taxonomic and (CARE) (CARE) (CARE) (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Heil 3105 PerF LocEn Pres Rare Heil 2578 PerF Wide Pres Unc Heil 2593 PerF Wide Pres Unc Harrison 1950 PerF LocEn Pres Unc Fleming 310 Life form shhook ower PerF Periph Pres Rare2932 Heil ower shhook cactus shhook small-fl fi Winkler’s Winkler’s pincushion cactus Simpson’s Simpson’s hedgehog cactus rufous-spine pricklypear cardinal-fl common hops PerF Wide Pres Unc Fleming 696 plains pricklypear PerF Wide Pres Abun Fleming 376 Parry’s harebellParry’s PerF Wide Pres Rare Fleming 168 Wright’s fi Wright’s cactus large pricklypear PerF Wide Pres Com Fleming 631 Common name

O. Flora

minor parryi var. var. in

var. ssp. var. & (2003) (2003) S. contortus orus, spina Flora of North S. terrae- simpsonii intermedius. Includes & canyonae S. parviﬂ S. whipplei Includes vars. & polyacantha polyacantha (2003) Included within of North America H. lupulus americanus Flora of North America includes vars. ruﬁ trichophora Vars. not recognized Vars. in America Synonyms/ notes Taxonomic

var. var. var.

var. graminea neomexicanus spina Sclerocactus whipplei roseus Pediocactus winkleri Pediocactus simpsonii rufi Opuntia polyacantha Lobelia cardinalis ssp. Humulus lupulus var. polyacantha Campanula parryi C. parryi Opuntia polyacantha Sclerocactus wrightiae major Opuntia phaeacantha rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Cactaceae Cactaceae Cactaceae Cactaceae Campanulaceae Cannabaceae Campanulaceae Cactaceae Cactaceae biogeographic notes, cont. Appendix A. Confi FamilyCactaceae Species name

Appendices 31 and Africa America River, Emery Co, UT River, (Jones s.n. POM) been relocated. locality is Type Cottonwood Canyon, Salt Lake 477 Co, UT (Watson US) Herbarium based on collections from outside the park. Comments NA Specimen has not NA Reports from CARE doc. Year Year 1981 1987 Native to Eurasia 1980 1987 Native to E North 1971 1986 1971 locality is Green Type 1986 1958 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (1993) (CARE) (BRY) (CARE) (CARE) (1993) Source Pop. size Park status Range PerF Wide Pres Com Meyer 4749 Shrub Wide Pres Unc Meyer 4520 Shrub Wide Pres Unc Nelson 9592 AnnF Wide Pres Rare Porter 4416 Life form ower Eastwood’s Eastwood’s sandwort snowball-bush Shrub Intro Pres Unc Fleming 260 mountain snowberry coralberry Shrub Intro Pres Unc Fleming 413 long-ﬂ snowberry Utah honeysuckle Shrub Wide Rep NA Heil et al. blue elderberry Shrub Wide Pres Unc Porter 3958 Palmer’s stinkweedPalmer’s AnnF RegEn Pres Rare Porter 2434 Rocky Mountain beeplant yellow beeplant AnnF Wide Pres Unc Swancara 12 basin sandwort PerF Wide Rep NA Heil et al. Common name

var.

var. oreophilus ssp. ssp. var. ssp. & plateauensis, glabrescens, Eremogone eastwoodiae eastwoodiae, A. eastwoodiae americanum S. rotundifolius, Includes vars. parishii S. nigra caerulea, S. cerulea A. kingii uintahensis, A. kingii compacta, A. kingii ssp. Eremogone kingii var. E. kingii uintahensis Synonyms/ notes Taxonomic

var. var. orus eastwoodiae glabrescens Arenaria fendleri var. Viburnum opulus opulus V. utahensis Symphoricarpos oreophilus Symphoricarpos orbiculatus Symphoricarpos longifl Lonicera utahensis Sambucus caerulea Cleomella palmeriana palmeriana Cleome serrulata Cleome lutea Arenaria fendleri var. rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caprifoliaceae (Adoxaceae) Caprifoliaceae Caprifoliaceae Caprifoliaceae Caprifoliaceae Caprifoliaceae (Adoxaceae) Capparaceae (Cleomaceae) Capparaceae (Cleomaceae) biogeographic notes, cont. Appendix A. Confi FamilyCapparaceae (Cleomaceae) Species name

32 Annotated Checklist of Vascular Flora: Capitol Reef National Park River Valley, Box River Valley, Elder Co, UT (Watson 995 US) Herbarium based on specimen that may have been collected outside the park. Comments doc. Year Year 2004 Report from CARE 1985 locality is Raft Type 1987 1986 1980 Native to Europe 1981 1986 1987 1987 Native to Eurasia c name with taxonomic and (1993), Northern Colorado Plateau Network (in ed.) 505 (CARE) (CARE) (BRY) (CARE) (CARE) (BRY) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Rare Heil 3559 PerF Intro Pres Rare Heil 3567 Life form y AnnF Wide Pres Unc2516 Porter y silver orach AnnF Wide Rep Rare Heil et al. iodine bush Shrub Wide Pres Rare Lichthardt mountain-lover Shrub Wide Pres Unc Heil 3287 long-stalk starwort PerF Wide Pres Rare Heil 2644 sleepy catchﬂ bouncing-bet PerF Intro Pres Unc Meyer 4658 creeping nailwort PerF Wide Pres Unc Porter 4340 Drummond’s Drummond’s catchﬂ mouse-ear chickweed Common name

ssp.

Flora

ssp. in ssp. includes striata, longipes. drummondii, drummondii, A. argentea Paxistima myrsinites, Pachistima myrsinites Includes var. monantha stricta, S. longipes ssp. of North America (2005) S. longipes striata, vars. kruckebergii, & Gastrolychnis drummondii Silene drummondii ssp. S. drummondii vulgare C. vulgatum, fontanum Synonyms/ notes Taxonomic ora argentea longipes cinalis Atriplex argentea var. Allenrolfea occidentalis Pachystima myrsinites Silene antirrhina Stellaria longipes var. Saponaria offi Paronychia sessilifl Lychnis Lychnis drummondii Cerastium fontanum rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Chenopodiaceae Chenopodiaceae Celastraceae Caryophyllaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae biogeographic notes, cont. Appendix A. Confi FamilyCaryophyllaceae (Alsinaceae) Species name

Appendices 33 from berlandieri Valley, west of Valley, Hanksville, Wayne Co, UT (Jones 5697 POM) Emery, Emery Co, UT Emery, (Jones 5443 RM) Salt Lake (near mouth of Weber Co, UT River), Weber (Fremont 761 NY) Includes report of var. CARE herbarium. Comments doc. Year Year 1980 Native to Eurasia 1986 Native to Eurasia 1973 1986 1987 1987 1987 1986 locality is Blue Type 1965 locality is near Type 1971 1958 locality is Great Type 1986 1958 1986 Native to Eurasia. c name with taxonomic and (BRY, CARE) (BRY, (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (BRY) (CARE) Source Pop. size Park status Range AnnF Intro Pres Rare Fleming 201 Shrub RegEn Pres Abun Erdman 2868 AnnF Wide Pres Unc Porter 4185 Life form ve-horn lambsquarter AnnF Intro Pres Unc Porter 4363 summer-cypress AnnF Intro Pres Com Meyer 4645 ﬁ smotherweed greenmolly PerF Wide Pres Unc Harrison 1122 wedge orach AnnF Wide Pres Rare Porter 4114 stalked orach AnnF RegEn Pres Unc Heil 3525 thinleaf orach AnnF Wide Pres Rare Heil 3528 Powell’s orachPowell’s AnnF Wide Pres Unc Heil 3426 Blue Valley orachBlue Valley AnnF RegEn Pres Rare Fleming 679 Castle Valley Castle Valley saltbush mat saltbush Shrub RegEn Pres Unc Nelson 9563 shadscale Shrub Wide Pres Abun Harrison 1142 mountain goosefoot fourwing saltbush Shrub Wide Pres Abun Harrison 1196 Common name

var. var. var. var. C. album sieversiana A. saccaria caput-medusae triangularis A. powellii A. cuneata C. fremontii atrovirens A. canescens occidentalis Synonyms/ notes Taxonomic album var. ora saccaria powellii cuneata canescens Chenopodium album Bassia scoparia Kochia scoparia, B. Bassia hyssopifolia Bassia americana Kochia americana Atriplex truncata Atriplex saccaria var. Atriplex prostrata A. patula Atriplex powellii var. Atriplex gracilifl Atriplex gardneri var. Atriplex corrugata Atriplex confertifolia Chenopodium atrovirens Atriplex canescens var. rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae biogeographic notes, cont. Appendix A. Confi FamilyChenopodiaceae Species name

34 Annotated Checklist of Vascular Flora: Capitol Reef National Park C. by Northern Herbarium based on specimen that may have been collected outside the park. villosum Colorado Plateau Network (in ed.) Comments doc. Year Year 1980 1973 Native to Eurasia 2004 Report from CARE 1988 Includes report of 1977 1986 1986 1980 1973 Native to Eurasia 1986 1986 c name with taxonomic and (CARE) (CARE) (1993), Northern Colorado Plateau Network (in ed.) (CARE) (CARE) (CARE) (CARE) (CARE) (BRY) (CARE) (CARE) Source Pop. size Park status Range AnnF Wide Pres Unc Camp 449 AnnF Periph Pres Rare Fleming 171 AnnF Intro Pres Unc Harrison 1334 AnnF Wide Pres Unc Porter 3461 Life form winterfat Shrub Wide Pres Abun Meyer 4652 halogeton AnnF Intro Pres Unc Harrison 1170 spiny hopsage Shrub Wide Rep Unc Heil et al. American bugseed AnnF Wide Pres Unc Fleming 663 narrow-leaf goosefoot New Mexico goosefoot oakleaf goosefoot AnnF Wide Pres Unc Porter 4116 silvery goosefoot AnnF Wide Pres Abun Meyer 4607 barbwire Russian thistle desert goosefoot AnnF Sparse Pres Unc Porter 4185 Nuttall’s poverty- Nuttall’s weed Common name

Vars. Vars.

Flora var. in var. (2003). Flora of North Ceratoides lanata, Eurotia lanata. not recognized in America grayi Includes vars. americanum rydbergii of North America (2003) Teloxys graveolens, Teloxys Dysphania graveolens C. salinum occidentale C. incanum incanum, C. incanum C. pratericola Synonyms/ notes Taxonomic

var. lanata var. var. var. Krascheninnikovia lanata Halogeton glomeratus Grayia spinosa Atriplex spinosa, A. Corispermum americanum Chenopodium leptophyllum Chenopodium graveolens neomexicanum salinum Chenopodium glaucum Chenopodium fremontii incanum Salsola paulsenii Chenopodium desiccatum Monolepis nuttalliana rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae biogeographic notes, cont. Appendix A. Conﬁ FamilyChenopodiaceae Species name Chenopodiaceae

Appendices 35

ramosissima miles E of Roosevelt, Uintah Co, UT (Stutz 9325 BRY) Juan River, San Juan Juan River, Co, UT (Brandegee s.n. GH) var. var. from Northern Colorado Plateau Network (in ed.) Comments doc. Year Year 1987 1973 locality is 15 Type 1973 locality is San Type 1980 1986 1977 Includes report of 1970 1987 1988 1973 1973 Native to Asia 1986 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) CARE V-7 CARE V-7 (CARE) (CARE) Source Pop. size Park status Range Shrub RegEn Pres Unc Harrison 1119 Life form common yarrow PerF Wide Pres Unc Fleming 176 western spiderwort PerF Wide Pres Rare Heil 3580 Plummer’s siltbushPlummer’s Shrub RegEn Pres Rare Porter 5066 Brandegee’s Brandegee’s siltbush Arizona siltbush Shrub RegEn Pres Rare Meyer 4653 small-leaf pussyoes PerF Wide Pres Rare Porter 3865 Torrey’s seepweedTorrey’s AnnF Wide Pres Com Camp 357 bur ragweed AnnF Wide Pres Com Fleming 193 broom seepweed AnnF Wide Pres Unc Heil 3497 cutleaf agoseris PerF Wide Pres Rare Heil 4427 greasewood Shrub Wide Pres Com Harrison 960 Russian thistle AnnF Intro Pres Abun Andersen Common name

var. var. var. var.

var. . Vars. . Vars. var. ora (2006) A. rosea Flora of North A. lanulosa, millefolium not recognized in occidentalis America T. occidentalis T. occidentalis Z. brandegeei plummeri, Grayia brandegei plummeri Z. brandegeei brandegeei, Atriplex brandegei Z. brandegeei arizonica Includes S. moquinii, nigra, S. fruticosa Franseria acanthicarpa S. depressa A. parviﬂ pestifer, S. australis pestifer, Synonyms/ notes Taxonomic

var. plummeri brandegei arizonica torreyana laciniata Achillea millefolium lanulosa Tradescantia Tradescantia occidentalis Zuckia brandegei var. Zuckia brandegei var. Zuckia brandegei var. Antennaria microphylla Suaeda torreyana var. Ambrosia acanthicarpa Suaeda calceoliformis Sarcobatus vermiculatus Agoseris glauca var. Salsola tragus S. kali, iberica, rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Compositae (Asteraceae) Commelinaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Chenopodiaceae Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae (Sarcobataceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyChenopodiaceae Species name

36 Annotated Checklist of Vascular Flora: Capitol Reef National Park Herbarium based on specimen that may have been collected outside the park. Comments NA Report from CARE doc. Year Year 2005 1980 1974 1986 1980 1966 1980 Native to Eurasia 1981 1978 c name with taxonomic and Lesica 21822 (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (1993) (CARE) (CARE) (BRY) Source Pop. size Park status Range PerF Wide Pres Unc Fertig & Shrub Wide Pres Abun Harrison 1108 PerF Wide Pres Unc Neese 6961 Life form Louisiana wormwood fringed sagebrush Shrub Wide Pres Unc Meyer 4637 sand sagebrush Shrub Wide Pres Unc Camp 369 tarragon PerF Wide Pres Abun Porter 4133 sand wormwood PerF Wide Pres Unc Meyer 4633 Bigelow’s Bigelow’s sagebrush breaks pussytoes PerF Wide Repburdock NA Heil et al. PerF Intro Pres Unc Meyer 4672 common pussytoes PerF Wide Pres Unc Meyer 4766 Louisiana wormwood Common name

lifolius Var. ssp. ssp. (2006) cus, O. cus, A. cus, ca Flora of North Oligosporus dracunculus. not recognized in Cronquist (1994) and America Oligosporus campestris pacifi pacifi campestris pacifi Synonyms/ notes Taxonomic

var. var. var. var. lifoliafi Oligosporus Artemisia ludoviciana Artemisia ludoviciana albula Artemisia frigida Artemisia fi Artemisia dracunculus glauca Artemisia campestris scouleriana Artemisia bigelovii Antennaria rosulata Arctium minus Antennaria parvifolia mexicana rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name

Appendices 37 Comments doc. Year Year 1980 2004 2004 1973 1980 1986 1986 1986 1986 c name with taxonomic and (CARE) Colorado Plateau Network (in ed.) (1993), Northern Colorado Plateau Network (in ed.) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range Shrub Wide Rep Unc Northern Shrub Wide Rep Unc Heil et al. Shrub Wide Pres Abun Harrison 1121 Life form c aster PerF Wide Pres Abun Meyer 4667 Paciﬁ Wyoming big sagebrush mountain big sagebrush basin big sagebrush Parry’s asterParry’s PerF Wide Pres Unc Meyer 4613 budsage Shrub Wide Pres Unc Porter 3744 pygmy sagebrush Shrub Wide Pres Unc Porter 3747 rayless alkali aster AnnF Wide Pres Rare Fleming 192 black sagebrush Shrub Wide Pres Unc Heil 2606 Common name

Flora parryi var. ssp. ssp. ssp. var. var. angusta A. Vars. not Vars. ssp. ora, adscendens, A. adscendens, Symphyotrichum ascendens, Virgulaster ascendens wyomingense Seriphidium tridentatum Includes tridentata pauciﬂ Seriphidium vaseyanum tridentatum Seriphidium tridentatum Symphyotrichum foliaceum Picrothamnus desertorum Seriphidium pygmaeum A. arbuscula (2006) of North America nova, Seriphidium novum. recognized in ciliatum, Brachyactis ciliata Synonyms/ notes Taxonomic

var.

var. var. var. var. nova Aster ascendens A. chilensis Artemisia tridentata wyomingensis Artemisia tridentata vaseyana Artemisia tridentata tridentata parryi Artemisia spinescens Aster foliaceus Artemisia pygmaea Artemisia nova var. Aster brachyactis Symphyotrichum rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyCompositae (Asteraceae) Species name

38 Annotated Checklist of Vascular Flora: Capitol Reef National Park “Rio Colorado, UT” (Palmer s.n. GH), but is probably from NV et al. or AZ (Welsh 2003) Herbarium based on specimen that may have been collected outside the park. Canyon, Zion NP, Canyon, Zion NP, Co., UT Washington & Hays 24379 (Welsh BRY). Comments NA Report from CARE doc. Year Year 1978 1970 1986 locality is Type 1986 1987 1986 locality is Zion Type 1980 1977 1980 1977 c name with taxonomic and (CARE) (CARE) (CARE) (1993) (CARE) (CARE) (BRY) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Camp 438 Shrub Wide Pres Com Camp 399 PerF Wide Pres Unc Porter 4160 Shrub Wide Pres Unc Heil 2942 Life form ower tassel-ﬂ brickellbush California brickellbush spiny brickellbush Shrub Wide Pres Rare Porter 3767 cutleafnodding bur- PerFmarigold Wide Rep NA Heil et al. Rio Grande seepwillow Welsh’s asterWelsh’s PerF LocEn Pres Rare Porter 4096 western aster PerF Wide Pres Unc Meyer 4655 alkali aster PerF Wide Pres Com Camp 464 blueleaf aster PerF Wide Pres Unc Meyer 4681 leafy aster AnnF Wide Pres Rare Camp 462 Common name

var. var. var. B. emoryi orus glauca, Eurybia B. atractyloides atractyloides Includes welshii Symphyotrichum spathulatum spathulatum paucifl var. glauca, Eucephalus glaucus frondosum, Brachyactis frondosa Synonyms/ notes Taxonomic orus Almutaster ora Brickellia grandifl Brickellia californica Brickellia atractyloides Bahia dissectadissecta Amauriopsis Bidens cernua Baccharis salicina Aster welshii Symphyotrichum Aster spathulatus A. occidentalis, Aster paucifl Aster glaucodes Herrickia glauca Aster frondosus Symphyotrichum rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name

Appendices 39 is American CARE Herbarium is too immature to determine to variety. weed. Native to Eurasia linifolia Fork, Jordan Valley, Utah Co, UT (Watson 493 YU) Comments NA Specimen from doc. Year Year 1981 1973 1981 1981 1986 2004 UT state noxious 1973 1973 locality of var. Type 1986 1970 1970 1970 c name with taxonomic and (CARE) (CARE) (CARE) (1993) (CARE) (CARE) CARE V-1 CARE V-1 (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range AnnF Wide Pres Unc Meyer 4720 Shrub RegEn Rep NA Heil et al. PerF Wide Pres Unc Meyer 4767 Shrub Wide Pres Unc Fleming 137 Shrub Wide Pres Unc Harrison 1306 Shrub Wide Pres Com Harrison 918 Shrub Wide Pres Unc Camp 361 Life form Eastwood-plant PerF Wide Pres Unc Porter 3829 rose-heath PerF Wide Pres Com Meyer 4477 stevia dusty- maiden Bigelow’s Bigelow’s rabbitbrush hoary dusty- maiden spear-leaf spear-leaf rabbitbrush Russian knapweed PerF Intro Pres Unc Andersen Greene’s Greene’s rabbitbrush Mohave brickellbush dwarf rabbitbrush Shrub Wide Pres Unc Porter 4140 rough brickellbush Shrub Wide Pres Com Camp 265 hispid golden-aster PerF Wide Pres Abun Harrison 1488 longleaf brickellbush Common name

parva var.

var. minor & var. hispida, hispida, H. bigelovii C. villosa var. Heterotheca villosa var. villosa C. scaposa Leucelene ericoides, Aster arenosus Ericameria nauseosa var. Includes vars. douglasii montana Lorandersonia linifolia B. oblongifolia B. longifolia longifolia Synonyms/ notes Taxonomic

var. var. var. minor Chrysopsis villosa var. Chamaechaenactis scaposa Chaetopappa ericoides Chaenactis stevioides Chrysothamnus nauseosus bigelovii Chaenactis douglasii Chrysothamnus linifolius Centaurea repens Acroptilon repens Chrysothamnus greenei Brickellia oblongifolia linifolia Chrysothamnus depressus Brickellia microphylla scabra Brickellia longifolia rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyCompositae (Asteraceae) Species name

40 Annotated Checklist of Vascular Flora: Capitol Reef National Park ed var. var. George, Washington George, Washington Co, UT (Palmer s.n. NY) Canaan Ranch, Kane Co, UT (Jones 6066c POM) Herbarium based on collections from outside the park or misidentiﬁ specimens of C. parryi nevadensis. Comments NA Reports from CARE doc. Year Year 1986 locality is St. Type 1986 1986 1986 1970 1977 1986 locality is Type c name with taxonomic and (CARE) (CARE) (BRY) (CARE) (CARE) (CARE) (1993) (CARE) Source Pop. size Park status Range Shrub RegEn Pres Unc Fleming 629 Shrub Wide Pres Com Porter 4249 Shrub Wide Pres Unc Fleming 101 Shrub Wide Pres Com Porter 4442 Shrub Wide Pres Com Heil 2861 Shrub Wide Rep NA Heil et al. Life form whitestem rabbitbrush smoothseed rabbitbrush rush rabbitbrush Shrub RegEn Pres Unc Heil 3599 glabrate rabbitbrush Nevada rabbitbrush graystem rabbitbrush greenish rabbitbrush dune rabbitbrush Shrub RegEn Pres Unc Fleming 148 Common name

var. var. var. albicaulis, speciosa leiosperma juncea graveolens, nevadensis hololeucus, hololeuca consimilis, turbinata C. nauseosus var. Ericameria nauseosa var. Ericameria nauseosa var. Ericameria nauseosa var. Ericameria nauseosa var. E. nauseosa glabrata Ericameria parryi var. C. nauseosus var. Ericameria nauseosa var. Ericameria nauseosa var. E. nauseosa oreophila, C. nauseosus oreophilus Ericameria nauseosa var. Synonyms/ notes Taxonomic

var. var. var. var. var. var. var.

var. Chrysothamnus nauseosus speciosus Chrysothamnus nauseosus leiospermus Chrysothamnus nauseosus junceus Chrysothamnus nauseosus graveolens nevadensis Chrysothamnus nauseosus gnaphalodes Chrysothamnus parryi consimilis Chrysothamnus nauseosus Chrysothamnus nauseosus turbinatus rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae)

Appendices 41 var. var. by D.J. eld Co, eld C. calcareum Bromide Pass, Henry Mtns, Garﬁ UT (Jones 5695bh US) considered “glabrous phase” of bipinnatum Keil. Comments doc. Year Year 1970 locality is Type 1986 1986 CARE material 1986 1986 1987 1980 1986 c name with taxonomic and (CARE) (BRY) (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range Shrub Wide Pres Abun Fleming 128 Shrub Wide Pres Unc Fleming 147 Shrub Wide Pres Unc Meyer 4666 Life form Caineville thistle PerF LocEn Pres Abun Meyer 4598 wingless thistle PerF RegEn Pres Unc Porter 4440 green rabbitbrush Shrub Wide Pres Com Fleming 137 slenderleaf rabbitbrush puberulent rabbitbrush New Mexico thistle PerF Wide Pres Unc Heil 3364 lanceleaf rabbitbrush pretty thistle PerF Wide Pres Unc Porter 4232 Common name

Flora var. Flora Var. ( in C. in C. var. var. var. var. (2006). (2006) orus Flora of North Flora of North Included in arizonicum 2006) bipinnatum of North America C. arizonicum bipinnatum. not recognized in Cronquist (1994) Cronquist (1994). not recognized Var. in America Included in var. Included in var. viscidiﬂ Vars. not recognized Vars. in America Var. not recognized Var. in Cronquist (1994), included in arizonicum 2006) bipinnatum of North America Synonyms/ notes Taxonomic

var. var. var. var. var. orus orus orus orus orus bipinnatum

calcareum pulchellum Cirsium calcareum var. Cirsium calcareum var. Chrysothamnus viscidifl viscidifl Chrysothamnus viscidifl stenophyllus Chrysothamnus viscidifl puberulus Cirsium neomexicanum var. neomexicanum Chrysothamnus viscidifl lanceolatus Cirsium calcareum var. rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name

42 Annotated Checklist of Vascular Flora: Capitol Reef National Park America Reef, Washington Reef, Washington Co, UT (Jones 5163q GH) “Southern Utah” (Bishop s.n. GH) Comments doc. Year Year 1986 Native to W North 1987 1986 1980 1980 Native to Europe 1973 2004 1987 locality is Silver Type 1973 locality is Type c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Colorado Plateau Network (in ed.) (CARE) (CARE) Source Pop. size Park status Range AnnF Intro Pres Unc Fleming 116 AnnF RegEn Pres Rare Fleming 508 PerF Wide Pres Unc Porter 3969 AnnF Wide Pres Unc Meyer 4676 Life form pappose glandweed Brandegee’s Brandegee’s sandplant glaucous hawksbeard Canadian horseweed bull thistle PerF Intro Pres Unc Meyer 4621 wavy-leaved thistle PerF Wide Pres Unc Harrison 1174 elk thistle PerF Wide Rep Rare Northern Utah thistle PerF RegEn Pres Unc Porter 5167 nakedstem PerF Wide Pres Unc Harrison 816 Common name

and var. Flora C. ssp. not Vars. not Vars. (2006) (2006), Flora of North (2006) brandegei. recognized in of North America Vars. not recognized Vars. in America C. undulatum C. utahense recognized in Flora of North America with elements split between neomexicanum C. inamoenum inamoenum. E. nudicaulis Synonyms/ notes Taxonomic

var. var. glauca glabrata scariosum utahense Enceliopsis Dyssodia papposa Dicoria brandegei D. canescens Crepis runcinata var. Conyza canadensis var. Cirsium vulgare undulatum Cirsium undulatum Cirsium scariosum var. Cirsium neomexicanum var. nudicaulis nudicaulis rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyCompositae (Asteraceae) Species name

Appendices 43 is 1 mile Mtns, San Juan Co, UT (Rydberg & Garrett 9755 NY). USFS: Region 4 Sensitive Rafael Swell, Emery Co, UT (Maguire 18459 NY). Type locality of var. harrisonii NE of Fruita, Capitol Co, Wayne Reef NP, & Welsh UT (Welsh USFWS: 21178 BRY). Threatened Comments doc. Year Year 1986 1980 1986 1980 1980 1987 1986 1980 1980 1987 locality is Abajo Type 1982 locality is San Type c name with taxonomic and (CARE) (CARE) (CARE) (CARE) Harper 4007 (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Welsh 21178 ( Welsh BRY) Source Pop. size Park status Range PerF Disj Pres Rare Heil 4378 Life form eabane PerF Wide Pres Uncs.n. Clark longleaf daisy PerF Wide Pres Unc Porter 4159 trailing daisy PerF Wide Pres Unc Meyer 4463 Eaton’s daisyEaton’s PerF Wide Pres Rare Porter 3886 spreading daisy PerF Wide Pres Abun Meyer 4481 tufted ﬂ San Rafael daisy PerF RegEn Pres Unc Barnes & pretty daisy AnnF Wide Pres Rare Fleming 502 silver daisy PerF Wide Pres Rare Meyer 4545 congested hairy daisy Abajo daisy PerF LocEn Pres Rare Heil 3515 Maguire’s daisyMaguire’s PerF LocEn Pres Unc & Welsh Common name

var. according var. var. E. Includes var. Includes var. Trimorpha Trimorpha lonchophylla eatonii E. tracyi, colomexicanus E. consimilis E. bellidiastrum bellidiastrum Includes awapensis to some authors harrisonii Synonyms/ notes Taxonomic

var. var. cinereus agellaris Erigeron maguirei Erigeron lonchophyllus Erigeron fl Erigeron eatonii E. eatonii Erigeron divergens var. Erigeron caespitosus Erigeron compactus consimilis Erigeron bellidiastrum Erigeron argentatus Erigeron aphanactis congestus Erigeron abajoensis rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name

44 Annotated Checklist of Vascular Flora: Capitol Reef National Park Rabbit Valley, Wayne Wayne Rabbit Valley, 401 Co, UT (Ward GH) Kanab, Kane Co, UT (Thompson s.n. GH) Willow Creek, San Juan Co, UT (Eastwood 48 CAS) Comments doc. Year Year 1970 locality is Type 1980 1987 1971 locality is Type 1986 locality is Type 1986 1986 1973 1981 1997 1981 1978 1981 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Buren, & Aanderud 97- 25 (BRY) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF RegEn Pres Abun Camp 454 PerF Wide Pres Unc Meyer 4505 PerF Periph Pres Unc Harrison 1194 Life form ower ower eabane PerF Wide Pres Abun3557 Porter basin blanketfl Hopi blanketfl western goldenrod PerF Wide Pres Unc Fleming 509 Utah daisy PerF RegEn Pres Abun Harrison 1347 slenderleaf daisy PerF RegEn Pres Rare Heil 2937 thread snakeweed Shrub Wide Pres Unc Fleming 131 aspen daisy PerF Wide Pres Rare Heil 2609 curly-cup gumweed vernal daisy PerF Wide Pres Unc Meyer 4709 Wright’s cudweedWright’s PerF Sparse Pres Rare Van Harper, Navajo fl crustweed AnnF Wide Pres Rare Camp 542 basin daisy PerF Wide Pres Unc Meyer 4741 Common name

var. var. Flora G. includes in orus subglaber Solidago occidentalis E. utahensis E. sparsiﬂ E. speciosus Included in hirsutula of North America (2006) E. concinnus condensatus G. canescens, Pseudognaphalium canescens var. E. concinnus concinnus, Synonyms/ notes Taxonomic

orus var. da utahensis sparsifl macranthus condensatus concinnus Glyptopleura marginata Gaillardia spathulata Gaillardia pinnatifi Euthamia occidentalis Erigeron utahensis var. Erigeron utahensis var. Gutierrezia microcephala Erigeron speciosus var. Grindelia squarrosa quasiperennis Erigeron pumilus var. Gnaphalium wrightii Erigeron pumilus var. Erigeron pulcherrimus rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name

Appendices 45 Mtns, San Juan Co, UT (Newberry s.n. GH) Herbarium based on specimen that may have been collected outside the park. locality is desert Type S of Hanksville, Co, UT Wayne (Stanton 328 US). Comments NA Report from CARE doc. Year Year 2005 locality is Abajo Type 1958 1980 1987 1973 1987 1973 1973 1973 c name with taxonomic and (1993) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF RegEn Pres Unc Harms s.n. Shrub Wide Pres Rare Fleming 506 PerF Wide Pres Rare Porter 5170 PerF Wide Pres Abun Camp 418 Life form ower AnnF Wide Pres Unc47 Swancara ower PerF Wide Pres Rare4654 Meyer ower AnnF Wide Pres Uncs.n. Harrison ower AnnF RegEn Rep NA Heil et al. ower sand sunfl common sunfl Nuttall’s sunfl Nuttall’s purpledisk sunfl Rusby’s Rusby’s goldenbush thrift goldenweed PerF Wide Pres Com Harrison 989 stemless goldenweed common hyalineherb prairie sunfl broom snakeweed Shrub Wide Pres Abun Harrison 1219 Common name

var.

var. var. var. lifolius ssp. orus, (2006) H. paucifl lifolius lifolius lifolius Flora of North Includes deserticola Ssp. not recognized in America H. nuttallii nuttallii H. drummondii, Isocoma rusbyi, I. drummondii Stenotus armerioides armerioides Stenotus acaulis acaulis H. fi lugens, H. fi var. H. fi megacephalus, H. fi tomentosus Includes var. petiolaris Synonyms/ notes Taxonomic

var.

acaulis ssp. var. var. lenticularis lifolius Helianthus anomalus Helianthus annuus var. Helianthus petiolaris Helianthus nuttallii Helianthella microcephala Haplopappus rusbyi Haplopappus armerioides armerioides Haplopappus acaulis Hymenopappus fi cinereus fallax Gutierrezia sarothrae rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name

46 Annotated Checklist of Vascular Flora: Capitol Reef National Park Salt Lake islands, UT 707 GH) (Watson the Muddy River, San the Muddy River, Rafael Reef, Emery Co, UT (Atwood & Goodrich 8652 BRY). USFWS: former C2 candidate been relocated Comments NA Specimen has not doc. Year Year 1980 Native to Europe 1988 1974 2004 locality is Great Type 1971 1971 locality is E of Type 1973 1986 1971 c name with taxonomic and (CARE) (CARE) (CARE) Colorado Plateau Network (in ed.) (CARE) (CARE) (1993) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Abun Harrison 988 PerF Wide Pres Unc Meyer 4509 PerF Wide Pres Abun Meyer 4558 Life form prickly lettuce AnnF Intro Pres Com Meyer 4673 marsh-elder AnnF Wide Pres Rare Fleming 698 poverty-weed PerF Wide Pres Unc Meyer 4683 western rush-pink PerF RegEn Rep Unc Northern Colorado rubber- plant low hymenoxys PerF LocEn Pres Unc Heil 2685 lanate hymenoxys PerF Wide Rep NA Heil et al. Canyonlands hymenoxys Arizona rush-pink PerF RegEn Pres Unc Porter 3773 Arizona hymenoxys Common name

Flora

var. var.

var. var. var. Vars. not Vars. var. nana arizonica oribunda xanthifolia (2006). of North America axillaris. recognized in L. dianthopsis Picardenia richardsonii ﬂ H. depressa, acaulis Tetraneuris var. H. acaulis lanata, Tetraneuris lanata, Tetraneuris acaulis caespitosa, T. brevifolia H. argentea ivesiana, Tetraneuris ivesiana L. arizonica Tetraneuris acaulis Tetraneuris var. Synonyms/ notes Taxonomic

var. var. var. ora ora nana caespitosa ivesiana arizonica oribunda Lactuca serriola Iva xanthifolia Cyclachaena Iva axillaris I. axillaris dianthopsis Hymenoxys richardsonii fl Hymenoxys acaulis var. Lygodesmia grandifl Hymenoxys acaulis var. Hymenoxys acaulis var. arizonica Hymenoxys acaulis var. Lygodesmia grandifl rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae)

Appendices 47 Willow Creek, San Juan Co, UT (Eastwood 45 CAS) miles S of Theodore, Duchesne Co, UT (Jones s.n. POM) Comments doc. Year Year 1958 1988 2004 1971 1970 1986 locality is Type 1970 1986 1973 1980 locality is 10 Type 1974 c name with taxonomic and (CARE) Lesica 21820 (CARE) Harper 4000 (CARE) (CARE) (CARE) (CARE) (CARE) 4655 (BRY) (CARE) (CARE) (CARE) Source Pop. size Park status Range AnnF Wide Pres Unc Harrison 1063 AnnF Wide Pres Unc Fertig & Life form liform desert- liform sowthistle desert- dandelion ﬁ dandelion tansyleaf aster AnnF Wide Pres Com Barnes & gumweed aster PerF Wide Pres Unc Meyer 4579 hoary aster PerF Wide Pres Com Camp 419 aristate aster PerF RegEn Pres Unc Porter 4428 thorny wirelettuce AnnF Wide Pres Unc Meyer 4650 rock-goldenrod PerF Wide Pres Unc Heil & Mietty showy rush-pink PerF Wide Pres Unc Heil 3359 low feverfew PerF RegEn Pres Unc Meyer 4560 copperweed Shrub Wide Pres Unc Harrison 1461 Common name

var. var. var. var. var. ora canescens vacans, Dieteria M. californica glabrata Haplopappus nuttallii, Xanthisma grindelioides grindelioides Aster canescens, Dieteria canescens var. M. linearis, rigida, M. canescens var. canescens aristata Stephanomeria spinosa, Pleiacanthus spinosus Solidago petradoria Bolophyta ligulata, alpinum P. ligulatum L. grandiﬂ Synonyms/ notes Taxonomic

var. var. var. var. ora ora pumila Oxytenia acerosa Iva acerosa Malacothrix sonchoides Malacothrix glabrata Machaeranthera tanacetifolia Machaeranthera grindelioides grindelioides Machaeranthera canescens canescens Machaeranthera canescens aristata Lygodesmia Lygodesmia spinosa Petradoria pumila var. Parthenium ligulatum grandifl Lygodesmia Lygodesmia grandifl rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name

48 Annotated Checklist of Vascular Flora: Capitol Reef National Park River, Uintah Co, UT River, (Fremont 549 GH) Utah” (Thompson s.n. GH) Grand Co, UT (Jones s.n. US) Herbarium based on collections from outside the park. Comments NA Reports from CARE doc. Year Year 1988 1977 1970 locality is Uinta Type 1973 1986 1966 locality is “S Type 1986 1980 1973 locality is Cisco, Type c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (1993) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Camp 456 PerF Wide Pres Unc Heil 2938 PerF RegEn Pres Abun Meyer 4747 PerF Wide Rep NA Heil et al. PerF Wide Pres Com Meyer 4620 Life form ower broom groundsel PerF Wide Pres Unc Heil 4615 many-headed groundsel basin groundsel PerF Wide Pres Com Porter 4664 Douglas’ groundsel PerF Wide Pres Unc Harrison 1126 three-nerve goldenrod greenstem paperﬂ coast goldenrod PerF Wide Pres Unc Porter 3951 Rocky Mountain groundsel Canadian goldenrod desert bahia PerF RegEn Pres Unc Porter 5080 Common name

var. var.

Vars. Vars. nana accidus

var. var. var. ora, (2006) (2006) (2006) ora douglasii accidus Flora of North Flora of North Flora of North simplex accidus, S. fl Packera streptanthifolia streptanthifola. not recognized Vars. in America S. spartioides. not recognized in America S. multicapitatus. not recognized Vars. in America Packera multilobata S. longilobus, S. fl fl var. includes vars. & S. velutina sparsifl S. lepida salebrosa Bahia nudicaulis desertorum Synonyms/ notes Taxonomic

var. var. var. var. ora

longilobus Senecio streptanthifolius var. streptanthifolius Senecio spartioides spartioides Senecio spartioides multicapitatus Senecio multilobatus Senecio douglasii var. Solidago velutina S. sparsifl Solidago simplex S. spathulata, Psilostrophe sparsifl Solidago canadensis salebrosa Platyschkuhria integrifolia desertorum rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name

Appendices 49 “Southern Utah” (Parry 109 GH) is 3 miles E of Bicknell, Wayne Co, UT (Atwood & Thompson 7646a USFS: Region BRY). 4 Sensitive, UT BLM: Sensitive, USFWS: former C2 candidate Comments doc. Year Year 1986 1987 1985 1976 Native to Eurasia 1980 1970 1971 locality is Type 1987 1980 2000 locality Type 1986 Native to Europe c name with taxonomic and (CARE) (BRY) 504 (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range Shrub Wide Pres Unc Porter 3817 Shrub Wide Pres Unc Porter 5039 Shrub Wide Pres Unc Lichthardt PerF Wide Pres Abun Harrison 911 AnnF Wide Pres Com Heil 3332 AnnF Intro Pres Unc Porter 4100 Life form shortspine horsebrush littleleaf horsebrush spineless horsebrush common dandelion PerF Intro Pres Unc Lindahl s.n. scale glandweed PerF Wide Pres Unc Meyer 4723 slender wirelettuce PerF Wide Pres Com Harrison 1493 scapose greenthread desert wirelettuce PerF Wide Pres Rare Heil s.n. white-plume wirelettuce alpine greenthread PerF LocEn Pres Rare Clark s.n. spiny-leaf sowthistle Common name

Flora

var. exigua minor. minor. T. var. by some ssp. var. (2006) (2006). Flora of North Flora of North (2006). of North America T. windhamii, T. Included in not authors. Vars. recognized in pubscens T spinosa spinosa Dyssodia pentachaeta belenidium Vars. not recognized Vars. in America S. minor Vars. not recognized Vars. in America S. exigua Synonyms/ notes Taxonomic

var. var. var. var. cinale alpinum Thelesperma subnudum Tetradymia Tetradymia spinosa Tetradymia Tetradymia glabrata Tetradymia Tetradymia canescens Taraxacum Taraxacum offi Thymophylla pentachaeta belenidium Stephanomeria tenuifolia tenuifolia subnudum Stephanomeria runcinata Thelesperma subnudum Stephanomeria exigua Sonchus asper rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name

50 Annotated Checklist of Vascular Flora: Capitol Reef National Park America near Epsom Creek, San Juan Co, UT (Eastwood 36 CAS) miles S of Fremont Jct., Sevier Co, UT (Reveal & Welsh USFWS: 721 BRY). Threatened Cottonwood Wash, Cottonwood Wash, 1 1/2 mi N of Bluff, San Juan Co, UT (Maguire 13509 GH) Comments doc. Year Year 1970 1986 1988 Native to C North 1986 locality is Type 1987 Native to Europe 1970 1988 1981 locality is 6 Type 1981 locality is Type 1986 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Heil 4369 PerF LocEn Pres Unc Meyer 4702 Life form rough mule’s-ears PerF Wide Pres Com Harrison 937 showy goldeneye PerF Wide Pres Rare Fleming 113 plains ironweed PerF Intro Pres Rare Fleming 693 resinbush Shrub RegEn Pres Unc Heil 2929 yellow salsify PerF Intro Pres Unc Fleming 383 hoary townsendia PerF Wide Pres Com Harrison 1033 stemless townsendia Last Chance townsendia annual townsendia AnnF Wide Pres Unc Meyer 4694 rough cocklebur AnnF Wide Pres Unc Fleming 87 Common name

major var. according var. (2006). T. jonesii T. ora ora scabra lutea Flora of North X. pensylvanicum, X. saccharatum. ssp. Heliomeris multiﬂ multiﬂ stylosa T. dubius T. to some authors Includes var. Vars. not recognized Vars. in America Synonyms/ notes Taxonomic

var. var. ora ora Xanthium strumarium canadense Wyethia scabra Scabrethia scabra multifl Viguiera multifl Vernonia Vernonia marginata Vanclevea stylosaVanclevea Chrysothamnus Tragopogon Tragopogon dubius Townsendia Townsendia incana Townsendia Townsendia exscapa Townsendia aprica Townsendia Townsendia Townsendia annua rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi FamilyCompositae (Asteraceae) Species name

Appendices 51

eld meionandrum var. var. weed. Native to Eurasia Grand Co, UT (Jones s.n. POM) bank of Green River W of Moab (Cronquist 8994 NY) 11 miles NE of Henrieville, Garfi Co, UT (Cronquist 9164 NY). USFWS: 3C from Northern Colorado Plateau Network (in ed.). locality is Left Type Fork Sawtooth Canyon, House Range, Millard Co, UT (Holmgren & Holmgren 11292 NY). Comments doc. Year Year 1987 Includes reports of 1987 1980 1980 1958 1974 Utah state noxious 1986 locality is Cisco, Type 1978 locality is E Type 1973 locality is Type c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range Shrub RegEn Pres Unc Porter 5180 PerF Wide Pres Unc Heil 2530 PerF RegEn Pres Unc Porter 2376 PerF LocEn Pres Rare Harrison 877 Life form eld dodder AnnF Wide Pres Unc Lehnert s.n. eld bindweed PerF Intro Pres Unc Harrison 1373 small-leaved greasebush lance-leaved stonecrop red-osier dogwood Shrub Wide Pres Unc Meyer 4678 evolvulus PerF Wide Pres Unc Meyer 4476 fi fi Cisco woodyaster PerF RegEn Pres Rare Porter 3480 hurtleaf woodyaster Henrieville woodyaster Common name

var. subalpinum Amerosedum lanceolatum, Includes ssp. lanceolatum C. stolonifera, Swida sericea Includes reports of C. glabrior venusta Machaeranthera tortifolia imberbis Machaeranthera confertifolia Synonyms/ notes Taxonomic

var.

var. var. Glossopetalon spinescens microphyllum Sedum lanceolatum Cornus sericea sericea Evolvulus nuttallianus Cuscuta pentagona Convolvulus arvensis Xylorhiza venusta Machaeranthera Xylorhiza tortifolia imberbis Xylorhiza confertifolia rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Crossosomataceae Crassulaceae Cornaceae Convolvulaceae Convolvulaceae Convolvulaceae Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyCompositae (Asteraceae) Species name

52 Annotated Checklist of Vascular Flora: Capitol Reef National Park is 18 miles N A. drummondii. of a mixed collection with russeola Uintah Co, of Vernal, UT (Rollins 1757 GH). USFWS: 3C Westwater, Grand Westwater, Co, UT (Jones s.n. US) Comments doc. Year Year 1973 1987 1982 1986 CARE specimen part 1981 1981 1988 locality of var. Type 1980 Native to Eurasia 1987 1971 locality is Type c name with taxonomic and (CARE) BC320 (CARE) & Atwood 16571 (BRY) (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Rare Meyer 4778 Life form perennial rockcress PerF Wide Pres Unc Meyer 4530 Holboell’s rockcressHolboell’s PerF Wide Pres Rare Fleming Holboell’s rockcressHolboell’s PerF Wide Pres Rare Goodrich Fendler’s rockcressFendler’s PerF Wide Pres Unc Porter 3869 hairy rockcress PerF Wide Pres Unc Meyer 4759 Drummond’s Drummond’s rockcress nodding rock cress PerF Wide Pres Unc Heil 4365 European madwort AnnF Intro Pres Rare Meyer 4537 Selby’s rockcressSelby’s PerF Wide Pres Com Porter 4658 desert rockcress PerF Wide Pres Unc Harrison 837 Common name

var. var.

var.

& orum retrofracta, secunda, B. pinetorum. languida, pallens Boechera perennans A. holboellii var. Boechera holboellii var. retrofracta Boechera holboellii var. Name may be misapplied, according to Holmgren et al. (2005) Boechera fendleri, A. fendleri pycnocarpa Includes vars. glabrata Boechera angustifolia, B. drummondii, B. stricta var. B. demissa demissa, B. oxylobula A. parviﬂ micranthum, A. simplex selbyi Boechera pulchra var. Synonyms/ notes Taxonomic

var. var. perennans secunda pinetorum fendleri Arabis pulchra Arabis perennans var. Arabis holboellii var. Arabis holboellii var. Arabis holboellii var. Arabis hirsuta Arabis drummondii Arabis demissa Boechera demissa micranthum Arabis selbyi Boechera selbyi Alyssum minus pallens rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyCruciferae (Brassicaceae) Species name

Appendices 53 been relocated Valley (near Alton), Valley Kane Co, UT (Jones s.n. POM) been relocated Comments doc. Year Year 1938 1938 Specimen has not 1986 1981 1987 1980 Native to Asia 1978 1987 locality is Sink Type 1980 Native to Europe 1980 Native to Europe 1980 Native to Europe 1973 c name with taxonomic and (BRY) (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range AnnF Wide Pres Unc Harrison 1801 AnnF Wide Pres Unc Harrison 1801 PerF Wide Pres Rare Meyer 4774 PerF RegEn Pres Rare Heil 3430 PerF Wide Rep NA Meyer (1982) NA Specimen has not AnnF Wide Pres Unc Harrison 868 Life form ixweed AnnF Intro Pres Abun4461 Meyer western tansy- mustard mountain tansy- mustard wedgeleaf draba AnnF Wide Pres Unc Porter 3469 California tansy- mustard golden draba PerF Wide Pres Rare Heil 3571 blue mustard AnnF Intro Pres Unc Meyer 4451 spectacle-pod AnnF Wide Pres Unc Camp 550 smooth spindlestem heartleaf bittercress ﬂ whitetop PerF Intro Pres Rare Meyer 4526 shepherd’s-purse AnnF Intro Pres Unc Meyer 4462 western tansy- mustard Common name

var. var. ssp. var.

ssp. by some draba var var. Included in D. pinata osmiarum authors incisa, D. incana incana D. richardsonii sonnei, D. incisa incisa, D. incana D. cuneifolia wislizenii C. cordifolia draba D. pinnata halictorum Synonyms/ notes Taxonomic

var. var. var. var. sonnei cuneifolia Descurainia pinnata intermedia Descurainia incana var. Draba cuneifolia var. Descurainia californica Draba aurea Chorispora tenella Dithyrea wislizenii Dimorphocarpa Caulanthus crassicaulis glaber Cardamine cordifolia cordifolia Cardaria draba Lepidium draba, C. Descurainia sophia Capsella bursa- pastoris Descurainia pinnata osmiarum rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Cruciferae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyCruciferae (Brassicaceae) Species name (Brassicaceae)

54 Annotated Checklist of Vascular Flora: Capitol Reef National Park George, Washington George, Washington Co, UT (Jones 1636 NY) the Great Salt Lake, Salt Lake Co, UT (Carrington s.n.) Comments doc. Year Year 1958 locality is St. Type 1988 1980 1988 1981 1988 locality is near Type 1986 Native to Europe 1988 1980 Native to Africa 2004 1988 Native to Europe c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Colorado Plateau Network (2007, in ed.) (CARE) Source Pop. size Park status Range PerF Wide Pres Rare Heil 4442 PerF Wide Pres Rare Heil 4373 AnnF Intro Pres Rare Fleming 559 Life form ower PerF Wide Pres Unc4779 Meyer ower clasping pepperwort Jones’ pepperwort PerF Wide Pres Com Swancara 35 prairie pepperwort AnnF Wide Pres Unc Fleming 622 prairie pepperwort AnnF Wide Pres Unc Meyer 4540 ovalpurse AnnF Wide Pres Rare Fleming 588 shy wallﬂ sand dune wallﬂ watercress PerF Intro Pres Unc Porter 4409 slender whitlow- grass African mustard AnnF Intro Pres Unc Meyer 4460 dwarf draba AnnF Wide Rep Unc Northern Common name

var. var. Included capitatum Includes var. Includes var. macrocarpum Hornungia procumbens, Hymenolobus procumbens E. asperum asperum, E. capitatum purshii. in var. in Holmgren et al. (2005) Rorippa nasturtium- aquaticum D. albertina Synonyms/ notes Taxonomic

var. var. var. var. orum orum orum nana cinale Lepidium perfoliatum Lepidium montanum jonesii ramosum Lepidium densifl densifl Lepidium densifl Hutchinsia procumbens Erysimum inconspicuum purshii Erysimum asperum Nasturtium offi Draba stenoloba var. Malcolmia africana Draba reptans rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) biogeographic notes, cont. Appendix A. Confi FamilyCruciferae (Brassicaceae) Species name

Appendices 55 eld Co, UT Aquarius Plateau, Garﬁ 589 GH) (Ward Herbarium based on collections from outside the park. Sego Canyon, about 5 miles N of Thompson, Grand 6902 Co, UT (Welsh USFWS: 3C BRY). Butte, San Rafael Swell, Emery Co, UT 20345 BRY). (Welsh USFWS: Endangered Comments NA Reports from CARE doc. Year Year 1986 1988 locality is Type 1980 1988 1978 1971 1986 Native to Europe 1988 locality is Type 1971 1971 1987 locality is Sye’s Type c name with taxonomic and (1993) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF RegEn Pres Abun Meyer 4548 PerF RegEn Pres Unc Fleming s.n. PerF Wide Pres Abun Meyer 4754 PerF Wide Pres Rare Meyer 4561 PerF LocEn Pres Unc Fleming 324 Life form ax-leaved smooth yellowcress PerF Wide Rep NA Heil et al. low yellowcress AnnF Wide Pres Unc Porter 4119 Ward’s bladderpodWard’s PerF RegEn Pres Rare Fleming 610 Colorado Plateau bladderpod Newberry’s Newberry’s twinpod silver bladderpod PerF Wide Pres Unc Camp 534 Watson’s Watson’s bladderpod tumble mustard AnnF Intro Pres Com Porter 4134 purple twinpod PerF LocEn Pres Rare Heil 4371 ﬂ plainsmustard sharp-leaf twinpod PerF Wide Pres Com Harrison 959 reed- Barneby’s mustard Common name

by var. glabra P. kingii P. P. var. by var. parvifolia intermedia R. palustris fernaldiana, R. palustris Included in var. Holmgren et al. (2005) P. newberryi P. racemosa Lesquerella ludoviciana Lesquerella intermedia, L. alpina var. Holmgren et al. (2005) Included in grahamii Sisymbrium linifolium P. australis P. Thelypodiopsis barnebyi Synonyms/ notes Taxonomic glabra purpurea acutifolia Rorippa islandica var. Rorippa tenerrima Physaria wardii Lesquerella wardii. Physaria rectipes Lesquerella rectipes Physaria newberryi Physaria ludoviciana Physaria intermedia Sisymbrium altissimum Physaria acutifolia var. Schoenocrambe linifolia Physaria acutifolia var. Schoenocrambe barnebyi rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyCruciferae (Brassicaceae) Species name Cruciferae (Brassicaceae)

56 Annotated Checklist of Vascular Flora: Capitol Reef National Park miles E of Hite and ca 75 miles W of Blanding, San Juan Co, UT (Cronquist 9033 GH) America Comments doc. Year Year 1973 1986 1986 1986 locality is 10 Type 1980 1986 1980 1971 1987 1980 1987 Native to E North 1958 1981 1986 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Porter 4290 PerF RegEn Pres Unc Porter 2421 PerF Wide Pres Unc Porter 3473 AnnF Wide Pres Abun Meyer 4695 PerG Wide Pres Unc Fleming s.n. PerF Wide Pres Com Meyer 4610 Tree Wide Pres Unc Meyer 4698 PerF Wide Pres Rare Fertig 22196 Life form ddle- ower Utah juniper Shrub Wide Pres Abun Harrison 803 common juniper Shrub Wide Pres Unc Porter 3847 entire-leaf thelypody candelabra thelypody Douglas’ sedge PerG Wide Pres Rare Meyer 4452 heart-leaf jewelﬂ golden sedge PerG Wide Pres Abun Meyer 4574 long-beak ﬁ mustard slender-beaked slender-beaked sedge green prince’s- plume white cedar Tree Intro Pres Unc Fleming 516 prince’s-plume PerF Wide Pres Abun Nelson 9172 Rocky Mountain juniper smoothleaf prince’s-plume Common name

ssp. var. var. var. Sabina scopulorum Sabina osteosperma J. communis alpina S. cordatus cordatus pinnata S. pinnata integrifolia Synonyms/ notes Taxonomic

ora

var. var. Juniperus scopulorum Juniperus osteosperma Juniperus communis depressa Thelypodium integrifolium integrifolium Thelypodiopsis divaricata Carex douglasii Streptanthus cordatus Carex aurea C. garberi Streptanthella longirostris Carex athrostachya Stanleya viridifl Thuja occidentalis Stanleya pinnata S. pinnata Stanleya integrifolia rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Cupressaceae Cupressaceae Cupressaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cyperaceae Cruciferae (Brassicaceae) Cyperaceae Cruciferae (Brassicaceae) Cyperaceae Cruciferae (Brassicaceae) Cupressaceae Cruciferae (Brassicaceae) biogeographic notes, cont. Appendix A. Confi FamilyCruciferae (Brassicaceae) Species name

Appendices 57 Herbarium based on collections from outside the park. Comments doc. Year Year 1987 1987 1980 1987 1980 1986 1981 1987 1987 2004 Reports from CARE c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) BC314 (BRY) (CARE) (1993), Northern Colorado Plateau Network (in ed.) Source Pop. size Park status Range PerG Sparse Pres Unc Fleming Life form Ross’ sedge PerG Wide Pres Unc Heil 3543 blackcreeper sedge PerG Wide Pres Unc Heil 3303 woolly sedge PerG Wide Pres Com Meyer 4715 western sedge PerG Wide Pres Rare Heil 3320 Nebraska sedge PerG Wide Pres Unc Meyer 4465 small-wing sedge PerG Wide Pres Unc Heil 3327 bottlebrush sedge PerG Wide Pres Rare Meyer 4728 White Mountain sedge threadleaf sedge PerG Wide Pres Unc Heil 3512 narrowleaf sedge PerG Wide Rep Unc Heil et al. Common name

includes ssp. var. var. var. (2002) for exa microptera, lifolia lifolia boottii, C. brevipes, C. pityophila var. C. festivella, macloviana microptera, C. limnophila Not included in Flora of North America UT fi eleocharis stenophylla, C. stenophylla Synonyms/ notes Taxonomic lifoliafi C. Carex rossii C. defl Carex praegracilis Carex pellita C. lanuginosa Carex occidentalis Carex nebrascensis Carex microptera C. microptera Carex hystericina C. hystricina Carex geophila Carex fi Carex duriuscula C. eleocharis, rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae biogeographic notes, cont. Appendix A. Confi FamilyCyperaceae Species name

58 Annotated Checklist of Vascular Flora: Capitol Reef National Park been relocated Herbarium based on collections from outside the park. Comments doc. Year Year 1980 1980 1986 1986 1980 1987 1977 2004 Reports from CARE c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (1993), Northern Colorado Plateau Network (in ed.) Source Pop. size Park status Range PerG Wide Pres Rare Porter 3979 Life form at-sedge AnnG Wide Pres Rare363 Camp ower panicled bulrush PerG Wide Rep NA Meyer (1982) NA Specimen has not alkali bulrush PerG Wide Pres Rare Meyer 4569 American bulrush PerG Wide Pres Unc Meyer 4606 hardstem bulrush PerG Wide Pres Unc Fleming 200 few-ﬂ spikerush common spikerush PerG Wide Pres Abun Meyer 4596 slender spikerush PerG Wide Pres Com Heil 3401 bearded ﬂ valley sedge PerG Wide Rep Unc Heil et al. Common name E.

& acutus var.

acutus, E. ssp. ora, ana E. var. var. paludosus, var. Schoenoplectus maritimus, Bolboschoenus maritimus paludosus Schoenoplectus americanus acutus occidentalis, Schoenoplectus acutus Schoenoplectus lacustris includes E. paucifl suksdorfi uniglumis Includes macrostachya, E. erythropoda, C. aristatus Synonyms/ notes Taxonomic ora Scirpus microcarpus Scirpus maritimus Scirpus maritimus Scirpus americanus Scirpus olneyi, Scirpus acutus Schoenoplectus Eleocharis quinquefl Eleocharis palustris Eleocharis acicularis Cyperus squarrosus Carex vallicola rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae biogeographic notes, cont. Appendix A. Confi FamilyCyperaceae Species name

Appendices 59 valley of the Virgin, Kane Co, UT (Siler s.n. ISC) America Comments doc. Year Year 1973 1973 1958 1973 1958 locality is upper Type 1987 1986 1973 Native to Europe 1986 Native to E North 1986 1971 1971 c name with taxonomic and (BRY) (CARE) (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) 337 (CARE) Source Pop. size Park status Range Shrub RegEn Pres Abun Harrison 820 Shrub Wide Pres Unc Heil 3285 Tree Intro Pres Unc Porter 4355 PerG Wide Pres Com Porter 3540 Fern Wide Pres Unc Fleming BC- Life form eld horsetail Fern Wide Pres Unc Harrison 854 tall scouring-rush Fern Wide Pres Unc Heil 2752 ﬁ Cutler’s ephedraCutler’s Shrub RegEn Pres Unc Harrison 890 green ephedra Shrub Wide Pres Abun Swancara 32 Torrey’s ephedraTorrey’s Shrub Wide Pres Abun Harrison 812 roundleaf buffaloberry Canada buffaloberry silver buffaloberry Shrub Wide Pres Unc Fleming 173 Russian olive Tree Intro Pres Unc Harrison 1123 common persimmon three-square bulrush smooth scouring- rush Common name nis, afﬁ

ssp. var. var. var. polyphyllus, Hippochaete hyemalis E. praealtum E. cutleri E. viridis pungens polyphyllus, Schoenoplectus pungens longispicatus, Schoenoplectus pungens Hippochaeta laevigata, E. kansanum Scirpus pungens var. Schoenoplectus pungens Synonyms/ notes Taxonomic

var. viscida viridis longispicatus ne Equisetum hyemale affi Equisetum arvense Ephedra viridis var. Ephedra viridis var. Ephedra torreyana Shepherdia rotundifolia Shepherdia canadensis argentea Elaeagnus angustifolia Diospyros virginiana Equisetum laevigatum Scirpus pungens var. rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Equisetaceae Equisetaceae Ephedraceae Ephedraceae Ephedraceae Elaegnaceae Elaegnaceae Elaegnaceae Shepherdia Elaegnaceae Ebenaceae Equisetaceae biogeographic notes, cont. Appendix A. Confi FamilyCyperaceae Species name

60 Annotated Checklist of Vascular Flora: Capitol Reef National Park

Q. tuckeri var. var. is Low Pass E of Kens Lake, San Juan Co, & Higgins UT (Welsh 23630 BRY) havardii been relocated George, Washington George, Washington Co, UT (Parry 247 MO) northern slope Uinta Mtns, UT (Engelmann s.n. MO) Comments doc. Year Year 1980 1987 1980 locality of Type 1986 1980 1977 locality is St. Type 1980 1987 locality is Type 1970 1986 1958 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Rare Meyer 4641 AnnF Wide Pres Abun Meyer 4636 PerF Wide Pres Rare Heil 3562 PerF Wide Pres Rare Fleming 103 Shrub Wide Pres Unc Lehnert s.n. Life form Rocky Mountain Great Basin centaury golden corydalis PerF Wide Pres Unc Heil 3324 shinnery oak Shrub RegEn Pres Unc Meyer 4510 noseburnGambel oak PerF Tree Sparse Rep Wide Pres NA Com Meyer (1982) Fleming 177 NA Specimen has not shorthorn spurge PerF Wide Pres Abun Meyer 4580 Parry’s spurgeParry’s AnnF Wide Pres Rare Camp 377 ridge-seeded spurge Engelmann’s Engelmann’s gentian Fendler’s spurgeFendler’s PerF Wide Pres Abun Harrison 1180 gentian greenleaf manzanita Common name

ssp.

var. var. var. ssp. nis Pneumonanthe affi C. curvisiliqua occidentalis tuckeri, Q. undulata Q. gambelii E. robusta, brachycera robusta, Tithymalus brachyceras Euphorbia glyptosperma G. amarella heterosepala Euphorbia fendleri A. patula platyphylla Synonyms/ notes Taxonomic nis nis affi occidentalis gambelii Gentiana affi var. Centaurium exaltatum Corydalis aurea var. Quercus welshii Q. havardii Tragia ramosaTragia Quercus gambelii stylaris T. var. Euphorbia brachycera Chamaesyce parryi Euphorbia parryi Chamaesyce glyptosperma Gentianella heterosepala Chamaesyce fendleri Arctostaphylos patula rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Gentianaceae Gentianaceae Fumariaceae Fagaceae Euphorbiaceae Fagaceae Euphorbiaceae Euphorbiaceae Euphorbiaceae Gentianaceae Euphorbiaceae biogeographic notes, cont. Appendix A. Confi FamilyEricaceae Species name

Appendices 61 is eld marginale and Africa var. var. Aquarius Plateau, head of Poison Creek, Garﬁ Co, UT (Rydberg & Carlton 7401 NY) near St. George, Co, UT Washington (Palmer s.n. US) Comments doc. Year Year 1988 1974 Native to Eurasia 1980 1973 Native to Eurasia. 2004 1980 locality of Type 1988 Native to Europe 1987 1981 locality is Type 1986 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) Colorado Plateau Network (in ed.) (CARE) (CARE) (CARE) (CARE) (BRY) Source Pop. size Park status Range PerF Wide Rep Unc Northern PerF Wide Pres Unc Meyer 4581 PerF Wide Pres Unc Meyer 4740 Life form shortawn foxtail PerG Wide Pres Rare Heil 4433 redtop PerG Intro Pres Com Porter 5188 spike bentgrass PerG Wide Pres Rare Meyer 4587 crested wheatgrass PerG Intro Pres Unc Harrison 1127 Richardson’s Richardson’s crane’s-bill small-leaf geranium stork’s-bill AnnF Intro Pres Abun Fleming 554 elkweed PerF Wide Pres Rare Heil 3563 white-margined swertia Hall’s bluestemHall’s PerG Wide Pres Unc Porter 4408 Common name

ssp. var. var. ssp. cowenii, G. A. aequalis aequalis A. alba, gigantea A cristatum cristatum, A cristatum pectinatum, A. pectinatum, A. sibiricum, A. desertorum, A. fragile, mongolicum atropurpureum, G. parryi G. atropurpureum var. atropurpureum, G. caespitosum Frasera albomarginata A. hallii Synonyms/ notes Taxonomic

var.

var. Andropogon gerardii chrysocomus Alopecurus aequalis Agrostis stolonifera Agrostis exarata Agropyron cristatum Geranium richardsonii marginale Geranium caespitosum Erodium cicutarium Swertia radiata Frasera speciosa Swertia albomarginata rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Geraniaceae Geraniaceae Geraniaceae Gentianaceae biogeographic notes, cont. Appendix A. Conﬁ FamilyGentianaceae Species name

62 Annotated Checklist of Vascular Flora: Capitol Reef National Park been relocated. Native to Eurasia Comments NA Specimen has not doc. Year Year 1940 1986 1951 1958 1984 2004 1987 1958 1938 1977 c name with taxonomic and (CARE) (BRY) (CARE) 11881 (BRY) (1993) (CARE) 401 (CARE) Colorado Plateau Network (in ed.) (CARE) (CARE) (BRY) Source Pop. size Park status Range PerG Sparse Pres Rare Harrison AnnG Periph Pres Rare Lichthardt Life form eld’s eld’s black grama PerG Wide Pres Unc Camp 352 sideoats grama PerG Wide Pres Unc Heil 2699 sixweeks grama AnnG Wide Pres Unc Fleming 143 Springﬁ bluestem oats AnnG Intro Rep NA Heil et al. purple three-awn PerG Wide Pres Abun Nelson 9170 sixweeks three- awn nodding brome PerG Wide Rep Unc Northern bushy bluestem PerG Periph Pres Rare Fleming 504 blue grama PerG Wide Pres Abun Swancara 6 big bluestem PerG Wide Pres Unc Markham s.n. Common name

or in

Includes var. var. oras is oras B. porteri barbata B. anomalus Chondrosum eriopodum caespitosa Includes vars. curtipendula rothrockii Chondrosum barbatum, vars. A. sativa purpurea fendleriana, A. purpurea longiseta, A. fendleriana Bromopsis porteri. Material ascribed to mostly B. lanatipes. regional ﬂ A. gerardii Synonyms/ notes Taxonomic

var.

var. var. eldii Bouteloua eriopoda Bouteloua curtipendula Bouteloua barbata Avena fatua Avena sativa Bothriochloa springfi Aristida purpurea A. glauca, Aristida adscensionis Bromus anomalus Bromus frondosus, Andropogon glomeratus scabriglumis Bouteloua gracilis Chondrosum gracile gerardii Andropogon gerardii rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Confi FamilyGramineae (Poaceae) Species name

Appendices 63 and Africa is Springdale, Co, UT Washington (Jones 6075 POM) America Comments doc. Year Year 1965 1974 Native to Eurasia 1980 1986 1987 locality Type 1986 Native to Eurasia 2005 Native to Eurasia 1986 Native to Eurasia 1980 Native to Eurasia 2004 1980 1980 Native to South 1980 Native to Eurasia c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Lesica 21821 (CARE) (CARE) (CARE) Colorado Plateau Network (in ed.) & Hak 458 (CARE) (CARE) (CARE) Source Pop. size Park status Range Life form eld sandbur AnnG Wide Pres Unc Meyer 4685 desert saltgrass PerG Wide Pres Unc Erdman 2870 orchard grass PerG Intro Pres Unc Harrison 1352 ﬁ slimstem reedgrass PerG Wide Pres Rare Fleming 666 Jones’ reedgrass PerG Wide Pres Unc Porter 5197 cheatgrass AnnG Intro Pres Abun Porter 2506 red brome AnnG Intro Pres Unc Fertig & Japanese chess AnnG Intro Pres Unc Porter 3983 smooth brome PerG Intro Pres Abun Meyer 4453 fringed brome PerG Wide Rep Unc Northern Canada wildrye PerG Wide Pres Unc May, Fowler, rescue grass AnnG Intro Pres Unc Meyer 4457 barnyard-grass AnnG Intro Pres Rare Meyer 4625 Common name

)

var. & Includes Includes var. var. E. muricata ( E. micro- ciliatus microstachya stricta, D. stricta C. neglecta, inexpansa B. commutatus Bromopsis inermis canadensis. vars. richardsonii canadensis B. willdenowii, catharticus catharticus Welsh et al. (2003) Welsh include stachya var. var. Synonyms/ notes Taxonomic inermis Echinochloa crus- galli Distichlis spicata D. spicata Dactylis glomerata Cenchrus longispinus Calamagrostis stricta Calamagrostis scopulorum Bromus tectorum Anisantha tectorum Bromus rubens Bromus japonicus B. arvensis. Bromus inermis var. Bromus ciliatus Bromopsis Elymus canadensis E. canadensis Bromus catharticus rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyGramineae (Poaceae) Species name

64 Annotated Checklist of Vascular Flora: Capitol Reef National Park of Salina Pass, Sevier Co, UT (Jones 5447 POM) weed. Native to Eurasia Comments doc. Year Year 1986 1980 1981 locality is top Type 1986 Utah state noxious 1986 1987 Native to Russia 1973 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) BC326 (CARE) (CARE) Source Pop. size Park status Range PerG Wide Pres Unc Porter 3965 PerG Wide Pres Unc Meyer 4665 PerG Wide Pres Unc Heil 2590 Life form bluebunch wheatgrass western wheatgrass Salina wildrye PerG Wide Pres Abun Meyer 4751 quackgrass PerG Intro Pres Unc Porter 4003 thickspike wheatgrass Russian wildrye PerG Intro Pres Unc Fleming squirreltail PerG Wide Pres Abun Harrison 874 Common name

var. smithii var. var. spicata Includes ssp. & thsii lanceolatus, albicans, palmeri spicatum, Pseuodroegneria spicata. inermis Pascopyrum smithii. Includes vars. & ambiguus, Leymus salinus, L. salina, ambiguus Elytrigia repens griffi var. Agropyron dasystachyum dasystachyum. Includes var. Includes var. riparium, A. albicans var. A. albicans juncea elymoides, Sitanion hystrix Synonyms/ notes Taxonomic Elymus spicatus Agropyron Elymus smithii Agropyron smithii, Elymus salinus E. salina, Elymus repens Agropyron repens Elymus lanceolatus E. lanceolatus Elymus junceus Psathyrostachys Elymus elymoides E. elymoides rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Confi FamilyGramineae (Poaceae) Species name

Appendices 65 Comments doc. Year Year 1986 1980 Native to Europe 1988 1980 1987 1988 1977 1987 1980 1980 1980 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerG Sparse Pres Rare Meyer 4472 PerG Wide Pres Unc Meyer 4575 Life form uffgrass PerG Wide Pres Unc5356 Porter Rocky Mountain fescue sixweeks fescue AnnG Wide Pres Unc Porter 2519 tall fescue PerG Intro Pres Unc Meyer 4576 ﬂ hairy tridens PerG Wide Pres Unc Meyer 4497 tufted lovegrass AnnG Wide Pres Rare Heil 3582 fowl mannagrass PerG Wide Pres Rare Heil 4432 spike pappusgrass PerG Sparse Pres Unc Camp 402 ravine fescue PerG Sparse Pres Rare Porter 5104 Saunders’ wheatgrass slender wheatgrass PerG Wide Pres Com Meyer 4464 Common name

E. & x var. var.

Hybrid of G. elata var. ora, Vulpia Vulpia ora, ora latiglumis, trachycaulus, Agrositanion F. ovina F. rydbergii, F. saximontana saximontana octoﬂ octoﬂ Includes vars. glauca, hirtella, Lolium arundinaceum, Schedonorus phoenix, S. arundinaceus Dasyochloa pulchella Includes x saundersii. E. trachycaulus elymoides E. trachycaulus var. E. alaskanus ssp. Agropyron caninum, A. subsecundum, A. trachycaulum, latiglume Synonyms/ notes Taxonomic

var. ora

x Festuca ovina saximontana Festuca octofl Festuca arundinacea Erioneuron pulchellum Erioneuron pilosum Eragrostis pectinacea Glyceria striata Elymus saundersii Enneapogon desvauxii Festuca sororia Elymus trachycaulus rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Confi FamilyGramineae (Poaceae) Species name

66 Annotated Checklist of Vascular Flora: Capitol Reef National Park M. is between and Africa curtifolia Kanab & Carmel, Kane Co, UT (Jones 6077j US) Comments doc. Year Year 1986 1977 1980 1986 1977 1987 1986 Native to Eurasia 1981 1986 1987 Native to Europe 1987 1977 1970 1980 locality of Type c name with taxonomic and (CARE) (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) BC313 (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerG Wide Pres Unc Meyer 4671 Life form Thurber’s muhlyThurber’s PerG Wide Pres Unc Meyer 4442 green muhly PerG Wide Pres Rare Porter 4265 sandhill muhly PerG Wide Pres Abun Camp 423 New Mexican muhly scratchgrass PerG Wide Pres Com Porter 4073 foxtail muhly PerG Wide Pres Abun Camp 433 wolftail PerG Wide Pres Unc Fleming 526 perennial ryegrass PerG Intro Pres Unc Heil 3400 junegrass PerG Wide Pres Unc Meyer 4773 little barley AnnG Wide Pres Unc Porter 2508 rabbit barley AnnG Intro Pres Unc Fleming foxtail barley PerG Wide Pres Abun Heil 3354 false buffalograss AnnG Wide Pres Unc Camp 396 galleta PerG Wide Pres Abun Nelson 9555 Common name

ssp. M. Includes curtifolia M. arsenei Includes reports of K. cristata, pyramidata, K. nitida H. murinum leporinum, H. leporinum, H. glaucum, Critesion glaucum Synonyms/ notes Taxonomic ora Muhlenbergia thurberi Muhlenbergia racemosa Muhlenbergia pungens Muhlenbergia paucifl Muhlenbergia asperifolia Muhlenbergia andina Lycurus phleoidesLycurus L. setosus Lolium perenne Koeleria macrantha Hordeum pusillum Critesion pusillum Hordeum murinum Hordeum jubatum Critesion jubatum Munroa squarrosa Monroa squarrosa Hilaria jamesii Pleuraphis jamesii rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Confi FamilyGramineae (Poaceae) Species name

Appendices 67 Comments doc. Year Year 1986 Native to Eurasia 1988 1973 1980 Native to Eurasia 1977 1987 1984 Native to Asia 1986 1980 Native to Europe 1981 1986 1987 1973 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) 402 (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range AnnG Intro Pres Unc Lichthardt Life form Canada bluegrass PerG Intro Pres Unc Porter 4271 Bigelow’s bluegrassBigelow’s AnnG Wide Pres Rare Fleming 584a common reed PerG Wide Pres Com Harrison 1192 timothy PerG Intro Pres Com Meyer 4688 switchgrass PerG Wide Pres Unc Camp 259 Scribner panicgrass PerG Sparse Pres Unc Fleming 528 broomcorn panicgrass witchgrass AnnG Wide Pres Unc Fleming 117 Kentucky bluegrass PerG Intro Pres Com Meyer 4663 bundle panicgrass PerG Wide Pres Rare Meyer 4716 fowl bluegrass PerG Wide Pres Rare Porter 4170 littleseed ricegrass PerG Wide Pres Unc Porter 5116 muttongrass PerG Wide Pres Abun Harrison 836 Common name

var. var. ssp. ssp. ssp includes var. P. cusickii P. P. epilis P. longiligula; fendleriana P. communis P. Dichanthelium oligosanthes scribnerianum P. miliaceum P. ruderale occidentale P. occidentale, P. tennessense, P. huachucae, P. Dichanthelium acuminatum fasciculatum, D. lanuginosum fasciculatum Piptatherum micranthum reports of & Synonyms/ notes Taxonomic

var. Poa fendleriana longiligula, P. P. Poa compressa Poa bigelovii Phragmites australis Phleum pratense Panicum virgatum Panicum oligosanthes scribnerianum Panicum miliaceum Panicum capillare capillare P. Poa pratensis agassizensis P. Panicum acuminatum Poa palustris Oryzopsis micrantha rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyGramineae (Poaceae) Species name

68 Annotated Checklist of Vascular Flora: Capitol Reef National Park and Africa and Africa Comments doc. Year Year 1980 1980 Native to Eurasia 1980 Native to Eurasia 1986 1986 1986 1980 1987 Native to Eurasia 1981 Native to Eurasia 1980 Native to Eurasia 1973 1980 1977 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerG Intro Pres Unc Porter 5162 PerG Wide Pres Unc Meyer 4507 Life form alkali cordgrass PerG Wide Pres Unc Meyer 4682 green bristlegrass AnnG Intro Pres Com Meyer 4644 yellow bristlegrass AnnG Intro Pres Com Meyer 4670 little bluestem PerG Wide Pres Unc Fleming 167 false melic PerG Sparse Pres Rare Fleming 156 tumblegrass PerG Sparse Pres Unc Fleming 119 Nuttall’s alkaligrassNuttall’s PerG Wide Pres Unc Meyer 4570 weeping alkaligrass water polypogon PerG Intro Pres Unc Meyer 4718 rabbitfoot-grass AnnG Intro Pres Unc Meyer 4600 alkali sacaton PerG Wide Pres Abun Harrison 1010 Sandberg’s Sandberg’s bluegrass Common name prairie wedgegrass PerG Wide Pres Rare Camp 1677

var. var. ssp. P. and all vars. P. secunda P. S. obtusata pumila, Pennisetum glaucum Andropogon scoparius, S. scoparium neomexicanum P. airoides P. Agrostis semiverticillata, P. viridis S. airoides Includes gracillima, P. gracillima, P. juncifolia, P. sandbergii, P. scabrella, canbyi, P. nevadensis, P. P. ampla, of Synonyms/ notes Taxonomic obtusata

var. Sphenopholis obtusata Spartina gracilis Setaria viridis Setaria glauca S. pumila Schizachyrium scoparium Schizachne purpurascens Schedonnardus paniculatus Puccinellia nuttalliana Puccinellia distans Polypogon semiverticillatus Polypogon monspeliensis Sporobolus airoides airoides Poa secunda rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyGramineae (Poaceae) Species name

Appendices 69 eld Co, UT Panguitch Lake, Milford, Beaver Co, UT (Jones 2106) Garﬁ (Jones 6023 POM) Comments doc. Year Year 1985 locality is Type 1973 2004 1958 1986 1938 1981 locality is near Type 1977 1986 1986 1980 c name with taxonomic and 503 (CARE) (CARE) Colorado Plateau Network (in ed.) (CARE) (CARE) Lesica 21813 (CARE) (CARE) (CARE) (BRY) (CARE) (CARE) Source Pop. size Park status Range PerG Wide Pres Rare Harrison 914 PerG Wide Rep Unc Northern PerG Wide Pres Rare Porter 5168 PerG Wide Pres Unc Meyer 4746 Life form pine needlegrass PerG Wide Pres Unc Lichthardt New Mexico feathergrass Letterman’s Letterman’s needlegrass Indian ricegrass PerG Wide Pres Abun Swancara 7 crested needlegrass needle-and-thread PerG Wide Pres Abun Fertig & Mormon needlegrass giant dropseed PerG Wide Pres Unc Camp 349 mesa dropseed PerG Wide Pres Com Heil 2948 sand dropseed PerG Wide Pres Abun Fleming 165 spike dropseed PerG Wide Pres Com Meyer 4589 Common name comata parishii, var. ssp. pinetorum neomexicana lettermanii hymenoides, Achnatherum hymenoides Achnatherum coronatum, A. parishii S. parishii Hesperostipa comata aridum Synonyms/ notes Taxonomic

var. var. exuosus Stipa pinetorum Achnatherum Stipa neomexicana Hesperostipa Stipa lettermanii Achnatherum Stipa hymenoides Oryzopsis Stipa coronata parishii Stipa comata comata Stipa arida Achnatherum Sporobolus giganteus Sporobolus fl Sporobolus cryptandrus Sporobolus contractus rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Confi FamilyGramineae (Poaceae) Species name

70 Annotated Checklist of Vascular Flora: Capitol Reef National Park rmed by Mary Herbarium based on collections from outside the park. US FWS: Category 3C America from CARE conﬁ Barkworth, 2005. First reported for UT in Cronquist et al. (1977), but not recognized in Welsh et al. (2003). Comments NA Report from CARE doc. Year Year 1980 1971 1973 1986 1986 Native to Central 1986 1978 1986 Photos of specimen c name with taxonomic and (CARE) (CARE) (1993) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Rare Porter 4107 PerG Periph Pres Unc Porter 4083 Life form Ives’ phacelia AnnF Wide Pres Com Meyer 4444 brittle phacelia AnnF RegEn Pres Com Nelson 9567 Howell’s namaHowell’s AnnF RegEn Repcorrugate phacelia AnnF NA Wide Heil et al. Pres Com Harrison 1143 common mare’s- tail maize AnnG Intro Pres Unc Fleming 194 Bloomer’s ricegrassBloomer’s PerG Wide Pres Rare Porter 3531 desert needlegrass PerG Wide Pres Com Camp 545 Scribner needlegrass Common name

x and species Hybrids S. S. comata Stipa bloomeri, P. demissa P. Var. not recognized Var. in Cronquist et al. (1984) except between hymenoides other bloomeri. x Achnatherum speciosum, Jarava speciosa scribneri Synonyms/ notes Taxonomic bloomeri Stiporyzopsis x demissa corrugata Phacelia ivesiana Phacelia demissa var. Nama retrorsum Phacelia crenulata var. Hippuris vulgaris Zea mays Stipa Stipa speciosa Achnatherum Stipa scribneri Achnatherum rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Hydrophyllaceae Hydrophyllaceae Hydrophyllaceae Hydrophyllaceae Hippuridaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyGramineae (Poaceae) Species name

Appendices 71 America Fruita, Capitol Reef Co, UT Wayne NP, (Atwood & Higgins 1834 BRY) Comments doc. Year Year 1986 1980 1980 1986 1986 1986 1958 Native to Eurasia 1974 Native to E North 1974 1986 1972 locality is Type c name with taxonomic and (CARE) (CARE) (CARE) BC363 (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Rare Heil 2753 Life form longstyle rush PerG Wide Pres Unc Porter 3980 mountain rush PerG Wide Pres Unc Meyer 4605 Tracy’s rushTracy’s PerG Wide Pres Unc Meyer 4635 toad rush AnnG Wide Pres Unc Fleming Baltic rush PerG Wide Pres Abun Porter 2517 northern rush PerG Wide Pres Unc Porter 4274 English walnut Tree Intro Pres Unc Swancara 34 black walnut Tree Intro Pres Unc Harrison 1382 blue-eyed grass PerF Wide Pres Unc Harrison 1452 Rocky Mountain iris San Rafael phacelia PerF RegEn Pres Unc Meyer 4708 Common name

forma J.

J. P. & by some by var. var. J. tracyi, ater, J. balticus ater, montanus; J. saximontanus, Includes xiphoides Included in ensifolius montanus saximontanus some authors; brunnescens ambiguus Includes vars. bufonius, occidentalis, halophilus, var. includes var. mexicanus balticus, J. arcticus ssp. Included in constancei authors Synonyms/ notes Taxonomic montanus brunnescens Juncus longistylus Juncus ensifolius var. Juncus ensifolius var. Juncus bufonius Juncus arcticus J. arcticus Juncus alpinus J. alpinoarticulatus Juglans regia Juglans nigra Sisyrinchium demissum Iris missouriensis Phacelia rafaelensis rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Juglandaceae Juglandaceae Iridaceae Iridaceae biogeographic notes, cont. Appendix A. Conﬁ FamilyHydrophyllaceae Species name

72 Annotated Checklist of Vascular Flora: Capitol Reef National Park is Johnson debilis Canyon, Kane Co, UT (Jones 5289 POM) Herbarium based on collections from outside the park. Comments NA Reports from CARE doc. Year Year 1958 Native to Europe 1980 Native to Europe. 1980 1988 Native to Eurasia 1987 1987 1980 2005 locality of var. Type 1980 1986 1978 1986 1980 Native to Europe c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (1993) (CARE) Source Pop. size Park status Range PerF Intro Pres Unc Heil 4430 Life form eld mint PerF Wide Pres Unc Meyer 4615 spearmint PerF Intro Pres Unc Lehnert s.n. peppermint PerF Intro Pres Unc Meyer 4643 ﬁ common horehound rough bugleweed PerF Wide Pres Rare Heil 3491 mock pennyroyal PerF Wide Pres Unc Porter 5094 seaside arrowgrass PerG Wide Pres Unc Meyer 4597 low arrowgrass PerG Wide Pres Unc Harms 192 annual sage PerF Sparse Pres Unc Meyer 4642 Torrey’s rushTorrey’s PerG Wide Pres Unc Porter 4268 purple sage Shrub Wide Pres Unc Camp 556 poverty rush PerG Wide Pres Unc Porter 4284 stinking horsemint PerF Wide Rep NA Heil et al. catnip PerF Intro Pres Unc Meyer 4680 Common name

M. var.

T. x . Hybrid var. T. var. var. by some Includes piperita x M. aquatica tenuis, J. dudleyi spicata. of M. arvensis canadensis T. maritima T. elata. concinnum Included in maritima recent authors Includes vars. dudleyi, congestus, & M. odoratissima glauca, M. glauca Synonyms/ notes Taxonomic

var. exa glabrata Monardella odoratissima Mentha spicata Mentha piperita M. Mentha arvensis var. Marrubium vulgare Lycopus asper Lycopus Hedeoma drummondii Triglochin Triglochin maritima Triglochin Triglochin concinnum debilis Salvia refl Juncus torreyi Poliomintha incana Juncus tenuis Nepeta cataria rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Juncaginaceae Juncaceae Labiatae (Lamiaceae) Juncaceae Labiatae (Lamiaceae) biogeographic notes, cont. Appendix A. Confi FamilyJuncaceae Species name Labiatae (Lamiaceae)

Appendices 73 candidate Grand Co, UT (Jones s.n. POM) Bicknell, Wayne Co, Bicknell, Wayne UT (Ripley & Barneby 8605 CAS). USFS: Region 4 Sensitive Gate, Carbon Co, UT (Jones s.n. POM) Comments doc. Year Year 1988 1986 USFWS: former C2 1988 USFWS: 3C 1973 locality is Cisco, Type 1982 1973 1971 1987 1971 locality is near Type 1980 1980 locality is Castle Type 1987 1971 c name with taxonomic and Harper 3880 (CARE) (CARE) (CARE) (CARE) (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF RegEn Pres Unc Fleming 640 PerF RegEn Pres Rare Fleming 620 PerF RegEn Pres Unc Harrison 1027 PerF RegEn Pres Unc Neese 11317 PerF Wide Pres Unc Porter 3967 Life form eld milkvetch PerF Wide Pres Rare Heil 3569 Torrey’s milkvetchTorrey’s PerF Wide Pres Rare Barnes & Brandegee’s Brandegee’s milkvetch Barneby’s Barneby’s milkvetch milkweed milkvetch Sheldon’s Sheldon’s milkvetch crescent milkvetch PerF Wide Pres Unc Porter 2364 lesser rushy milkvetch ﬁ Bicknell milkvetch PerF LocEn Pres Unc Nelson 9581 hairy hedge-nettle PerF Wide Pres Rare Meyer 4614 Colton’s milkvetchColton’s PerF RegEn Pres Unc Meyer 4501 little skullcap PerF Wide Pres Rare Porter 5099 painted milkvetch PerF Wide Pres Com Meyer 4445 Common name

var. var. var. pilosa var. consobrinus A. calycosus A. desperatus conspectus campestris A. diversifolius A. castaneiformis var. S. pilosa S. galericulata epilobifolia Synonyms/ notes Taxonomic

var. var. var. var. var. pilosa coltonii Astragalus ceramicus Astragalus calycosus calycosus Astragalus brandegei Astragalus barnebyi Astragalus asclepiadoides Astragalus amphioxys vespertinus Astragalus amphioxys amphioxys Astragalus convallarius convallarius Astragalus agrestis A. dasyglottis Astragalus consobrinus Stachys palustris var. Astragalus coltonii var. Scutellaria galericulata ceramicus rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Labiatae (Lamiaceae) Leguminosae (Fabaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyLabiatae (Lamiaceae) Species name

74 Annotated Checklist of Vascular Flora: Capitol Reef National Park is near eld, Sevier Co, eld Co, UT Cottrel’s Ranch, Cottrel’s Henry Mtns., Garﬁ (Jones 5658q POM) Fruita, Capitol Reef Co, UT Wayne NP, (Barneby 13131 CAS). USFWS: 3C candicans Richﬁ 246 GH) UT (Ward River, Emery Co, UT River, (Jones s.n. POM) “southern Utah”, Kane Co?, UT (Bishop s.n. GH) “southern Utah”, (Parry 32 ISC) Rafael Swell, Emery Co, UT (Jones s.n. POM) Montezuma Canyon, San Juan Co, UT (Eastwood s.n. POM) Comments doc. Year Year 1988 locality is Type 1988 1961 locality is near Type 1988 locality of var. Type 1971 locality is Green Type 1971 locality is Type 1980 locality is Type 1971 locality is San Type 1973 locality is Type c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF RegEn Pres Unc Harrison 830 PerF LocEn Pres Unc Meyer 4529 PerF Wide Pres Com Meyer 4467 PerF LocEn Pres Unc Meyer 4440 Life form straggling milkvetch laccolite milkvetch PerF LocEn Pres Rare Fleming 617 tall kentrophyta PerF Wide Pres Com Heil 4649 Harrison’s Harrison’s milkvetch yellow milkvetch PerF Wide Pres Unc Heil 4356 clay milkvetch PerF LocEn Pres Rare Fleming 309 bishop’s milkvetchbishop’s PerF RegEn Pres Unc Nelson 9570 great rushy milkvetch rockloving milkvetch Common name

var. var.

var. avus A. chamaeleuce laccoliticus A. ﬂ candicans Not recognized by many authorities A. episcopus episcopus Synonyms/ notes Taxonomic

var. var. avus avus var. avus fl argillosus Astragalus lentiginosus palans Astragalus laccoliticus Astragalus kentrophyta elatus Astragalus harrisonii Astragalus fl var. Astragalus fl var. Astragalus episcopus Astragalus lonchocarpus Astragalus desperatus petrophilus rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) biogeographic notes, cont. Appendix A. Confi FamilyLeguminosae (Fabaceae) Species name

Appendices 75 “southern Utah” (near Kanab), Kane Co, UT (Bishop s.n. GH) Mountain, Emery Co, UT (Jones s.n. NY). USFWS: 3C miles S of Ferron, Emery Co, UT (Jones 5454a POM) Kanab, Kane Co, UT (Thompson s.n. GH) is Kaiparowits Plateau, S of Willow Kane Co, UT Tank, (Harrison 9069 US). USFWS: 3C Comments doc. Year Year 1986 locality is Type 1988 1971 1985 1986 locality is Cedar Type 1977 1973 locality is 2 Type 1971 locality is Type 1986 1980 locality Type 1981 c name with taxonomic and (CARE) (CARE) (CARE) Neese 23445 (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF RegEn Pres Unc Porter 3796 PerF RegEn Pres Unc & Welsh AnnF RegEn Pres Rare Camp 533 PerF RegEn Pres Unc Porter 3527 PerF LocEn Pres Unc Meyer 4518 Life form owered pulse milkvetch PerF Wide Pres Unc Meyer 4776 sandstone milkvetch gravel milkvetch AnnF Wide Pres Unc Fleming 645 stinking milkvetch PerF Wide Pres Unc Nelson 9495 Patterson’s Patterson’s milkvetch panther milkvetch PerF LocEn Pres Unc Porter 3812 small-ﬂ milkvetch Ferron milkvetch PerF RegEn Pres Unc Harrison 822 woolly milkvetch PerF Wide Pres Com Nelson 9481 Moenkopi milkvetch Kaiparowits milkvetch Common name Synonyms/ notes Taxonomic

var. var. var. orus Astragalus tenellus Astragalus sesquifl Astragalus sabulonum Astragalus praelongus praelongus Astragalus pattersonii Astragalus pardalinus Astragalus nuttallianus micranthiformis Astragalus musiniensis Astragalus mollissimus thompsoniae Astragalus moencoppensis Astragalus malacoides rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) biogeographic notes, cont. Appendix A. Confi FamilyLeguminosae (Fabaceae) Species name

76 Annotated Checklist of Vascular Flora: Capitol Reef National Park eld Co, eld America “Kanab” (but probably from farther east (Thompson s.n. GH) is Springdale, Co, UT Washington (Jones 5259 POM) Rafael Swell, Emery Co, UT (Jones s.n. POM). USFWS: 3C CARE Herbarium based on specimen that may have been collected outside of the park. Type locality is Aquarius Plateau, Garﬁ 424 GH) UT (Ward Comments NA Report from the doc. Year Year 1973 1980 1958 Native to E North 1986 1981 locality is Type 1986 locality Type 1971 locality is San Type c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (BRY) (1993) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Harrison 898 PerF Wide Pres Unc Heil 2857 PerF RegEn Pres Unc Meyer 4735 PerF LocEn Pres Rare Heil 2737 Life form northern sweetvetch Nuttall’s licoriceNuttall’s PerF Wide Pres Unc Meyer 4657 honey locust Tree Intro Pres Unc Swancara 22 white prairie- clover Kanab prairie- clover Zion milkvetch PerF RegEn Pres Unc Porter 2394 Ward’s milkvetchWard’s PerF RegEn Rep NA Heil et al. Woodruff’s Woodruff’s milkvetch Common name

var. var. avescens Includes var. Includes var. pabulare G. lepidota lepidota oligophylla candidum, P. candidum, P. occidentale, D. candida D. ﬂ Synonyms/ notes Taxonomic

i var. avescens avescens fl zionis Hedysarum boreale boreale Glycyrrhiza lepidota Gleditsia triacanthos Dalea oligophylla Petalostemon Dalea fl var. Astragalus zionis var. Astragalus wardii Astragalus woodruffi rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) biogeographic notes, cont. Appendix A. Confi FamilyLeguminosae (Fabaceae) Species name Leguminosae (Fabaceae)

Appendices 77 Hogan Pass, Sevier Isely, Co, UT (Welsh, & Moore 6452 BRY) miles E of Zion Kane Co, UT NP, (Hitchcock 19013 isotype NY) Comments doc. Year Year 1958 Native to Europe 1974 Native to Europe 1986 Native to Europe 1987 1971 1988 1973 1987 1973 1986 locality is E of Type 1980 locality is 10 Type 1958 Native to Europe c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Intro Pres Com Harrison 1457 Life form yellow sweet- clover white sweet-clover AnnF Intro Pres Com Swancara 24 alfalfa PerF Intro Pres Com Harrison 4664 black medick AnnF Intro Pres Unc Porter 4092 silky lupine PerF Wide Pres Unc Heil 3544 dwarf lupine AnnF Wide Pres Com Harrison 1015 Great Basin lupine AnnF Wide Pres Com Fleming 566 Utah lupine PerF Wide Pres Unc Harrison 899 nodding locoweed PerF Wide Pres Rare Heil 3570 silvery lupinesubalpine lupine PerF PerF Wide Wide Pres Pres Unc Unc Harrison 1026 1973 Meyer 4772 southern locoweed PerF RegEn Pres Unc Porter 3863 Zion sweetpea PerF RegEn Pres Unc Fleming 585 Common name by

sativa ssp. var. Included var. cinalis M. ofﬁ in some authors Vars. not recognized Vars. in Barneby (1989) L. caudatus caudatus, L. argenteus utahensis O. viscida Synonyms/ notes Taxonomic

exa var.

var. sericeus pusillus utahensis sericea argenteus rubricaulis australis cinalis Melilotus offi Melilotus alba M. albus. Medicago sativa M. sativa Medicago lupulina Lupinus sericeus var. Lupinus pusillus var. Lupinus pusillus intermontanus Lupinus caudatus var. Oxytropis defl var. Lupinus argenteus var. Lupinus argenteus var. Oxytropis borealis var. zionis Lathyrus brachycalyx rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) biogeographic notes, cont. Appendix A. Confi FamilyLeguminosae (Fabaceae) Species name

78 Annotated Checklist of Vascular Flora: Capitol Reef National Park AZ, near Kanab, or along the Colorado River in SE UT or AZ (Thompson s.n. US). USFWS: 3C Wilson Mesa, Grand Co, UT (Rydberg & Garrett 8367 NY) Epsom Creek, San Juan Co, UT (Eastwood 21 CAS) Kanab, Kane Co, UT (Thompson s.n. GH) E of Bicknell, Wayne E of Bicknell, Wayne & Co, UT (Welsh Moore 13828 BRY) America Comments doc. Year Year 1986 Native to E North 2004 locality in N Type 1981 1973 locality is near Type 2004 1981 locality is Type 1971 1986 locality is Type 1971 locality is 1 mile Type 1971 c name with taxonomic and (CARE) Colorado Plateau Network (in ed.) (CARE) (CARE) Harper 3965 (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range Shrub LocEn Rep Rare Northern Shrub Wide Pres Unc Meyer 4725 PerF RegEn Pres Rare Heil 3414 PerF Wide Pres Unc Meyer 4446 PerF RegEn Pres Rare Porter 3772 PerF Wide Pres Unc Nelson 9494 Life form black locust Tree Intro Pres Unc Porter 4346 Thompson’s Thompson’s indigo-bush Fremont’s indigo- Fremont’s bush slenderleaf scurfpea lemon scurf-pea PerF Wide Pres Com Barnes & rush scurfpea PerF RegEn Pres Rare Meyer 4719 montane golden pea Thompson’s Thompson’s peteria Bicknell locoweed PerF RegEn Pres Unc Meyer 4750 Lambert’s Lambert’s locoweed Common name

var. var.

includes Vars. not recognized Vars. in Barneby (1989) Psoralea lanceolata. not recognized Vars. in Barneby (1989) Psoralea juncea ovata montana; rhombifolia T. T. pinetorum, T. pinetorum, T. T. rhombifolia Includes var. lambertii Synonyms/ notes Taxonomic

var.

var. var. var. var. Robinia pseudoacacia thompsoniae Psorothamnus thompsoniae Psorothamnus fremontii Psoralidium lanceolatum stenophyllum Psoralidium lanceolatum lanceolatum Psoralidium junceum Peteria thompsoniae Oxytropis oreophila juniperina Thermopsis montana Oxytropis lambertii bigelovii rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyLeguminosae (Fabaceae) Species name

Appendices 79 ower of Utah Kanab, Kane Co, UT (Thompson s.n. US) Kanab, Kane Co, UT (Thompson s.n.) Green River, Emery Green River, Co, UT (Jones s.n. POM) Comments doc. Year Year 1973 locality is near Type 1958 Native to Eurasia 1980 locality is near Type 1973 1971 1986 1988 1986 Native to China 1980 1980 Native to Europe 1988 1980 Native to Europe 1987 locality is near Type 1988 Native to Europe 1968 State fl c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (BRY) (CARE) Source Pop. size Park status Range PerF Wide Pres Com Porter 5361 Life form sego-lily PerF Wide Pres Unc s.n. Taylor golden mariposa PerF RegEn Pres Unc Harrison 873 asparagus PerF Intro Pres Unc Swancara 39 pink funnel-lily PerF Wide Pres Unc Meyer 4549 Nevada onion PerF Wide Pres Rare Harrison 876 San Juan onion PerF Wide Pres Unc Porter 2366 nodding onion PerF Wide Pres Unc Fleming 152 common bladderwort Chinese wisteria PerF Intro Pres Unc Porter 4217 American vetch PerF Wide Pres Unc Meyer 4771 white clover PerF Intro Pres Unc Meyer 4679 leopard-lily PerF Wide Pres Rare Heil 4426 red clover PerF Intro Pres Unc Meyer 4687 sand lily PerF RegEn Pres Rare Porter 4673 strawberry clover PerF Intro Pres Rare Fleming 671 Common name obtusum var. var. & in Welsh in Welsh A. Brodiaea paysonii Includes atrorubens cristatum et al. (2003) Includes vars. cernuum U. macrorhiza Synonyms/ notes Taxonomic orum americana cinalis Calochortus nuttallii Calochortus aureus Asparagus offi Androstephium brevifl Allium nevadense Allium macropetalum Allium cernuum Utricularia vulgaris Wisteria sinensis Vicia americana var. Trifolium repens Trifolium Fritillaria atropurpurea Trifolium pratense Trifolium Eremocrinum albomarginatum Trifolium Trifolium fragiferum rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Liliaceae (Calochortaceae) Liliaceae (Calochortaceae) Liliaceae (Asparagaceae) Liliaceae (Themidaceae) Liliaceae (Alliaceae) Liliaceae (Alliaceae) Liliaceae (Alliaceae) Lentibulariaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae Leguminosae (Fabaceae) Liliaceae (Anthericaceae) biogeographic notes, cont. Appendix A. Confi FamilyLeguminosae (Fabaceae) Species name

80 Annotated Checklist of Vascular Flora: Capitol Reef National Park Willow Creek, San Juan Co, UT (Eastwood 31 CAS) miles E of Hite, San Juan Co, UT (Cronquist 9030 NY) Comments doc. Year Year 1986 Native to Asia 1980 locality is Type 1986 1971 1986 locality is 10 Type 1986 1971 1987 1970 1986 1987 1971 1980 1980 Native to Eurasia c name with taxonomic and (CARE) (CARE) (CARE) (BRY) (CARE) (CARE) (CARE) (CARE) (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range AnnF Wide Pres Abun Nelson 9487 PerF Wide Pres Abun Harrison 1124 PerF Wide Pres Unc Heil 4385 PerF Wide Pres Unc Heil 3550 PerF Wide Pres Abun Harrison 835 PerF Wide Pres Unc Meyer 4454 Life form ax AnnF Wide Pres Unc3783 Porter ax AnnF Wide Pres Unc4612 Meyer ax PerF Wide Pres Unc3102 Heil common mallow AnnF Intro Pres Unc Meyer 4686 rose-of-sharon Shrub Intro Pres Unc Porter 4213 wing-seed stickleaf PerF Wide Pres Unc Meyer 4504 desert stickleaf PerF Wide Pres Unc Heil 2860 Colorado stickleaf PerF RegEn Pres Unc Porter 4443 white-stem blazingstar downy fl small-leaf globemallow blue fl broomfl gooseberry-leaf globemallow mountain death camas scarlet globemallow starry false Solomon’s-seal Common name

& in var. var. lewisii, Includes ora var. ora, Nuttallia ora, ora cronquistii grossulariifolia Nuttallia pterosperma var. var. M. marginata marginata, M. cronquistii. M. multifl Welsh et al. 2003 Welsh multifl multifl Acrolasia albicaulis, A. gracilis, M. obscura puberulum L. lewisii Adenolinum lewisii S. grossulariifolia var. Includes ssp. coccinea, elata, dissecta stellatum Synonyms/ notes Taxonomic ora lewisii Malva neglecta Hibiscus syriacus Mentzelia pterosperma Mentzelia marginata Mentzelia multifl Mentzelia albicaulis Linum puberulum Mesynium Linum perenne ssp. Sphaeralcea parvifolia Linum aristatum Mesynium aristatum Sphaeralcea grossulariifolia Zigadenus elegans Anticlea elegans Sphaeralcea coccinea Smilacina stellata Maianthemum rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Malvaceae Malvaceae Loasaceae Loasaceae Loasaceae Loasaceae Linaceae Linaceae Malvaceae Linaceae Malvaceae Liliaceae (Melanthiaceae) Malvaceae biogeographic notes, cont. Appendix A. Confi FamilyLiliaceae (Convallariaceae or Ruscaceae) Species name

Appendices 81 Beaver City, Beaver Beaver City, Co, UT (Palmer 404) America Labyrinth Canyon, San Juan Co, UT (Newberry s.n. NY) America America Comments doc. Year Year 1970 1958 1987 1987 1977 1986 locality is near Type 1971 1986 Native to E North 1974 Native to China 1958 locality is Type 1987 Native to E North 1986 Native to E North 1986 1973 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Camp 439 PerF Wide Pres Abun Heil 3365 PerF Sparse Pres Rare Heil 3529 PerF Wide Pres Unc Camp 358 PerF Wide Pres Abun Harrison 1286 PerF Wide Pres Unc Heil 2599 AnnF Wide Pres Unc Harrison 1169 Life form ower small-ﬂ sandpuffs spreading four- o’clock showy four-o’clock PerF Wide Pres Abun Harrison 913 narrowleaf four- o’clock New Mexico umbrellawort trailing four- o’clock low sand-verbena PerF RegEn Pres Unc Porter 3512 fragrant sand- verbena green ash Tree Intro Pres Unc Porter 4351 white mulberry Tree Intro Pres Unc Harrison 1404 singleleaf ash Shrub Wide Pres Abun Harrison 1184 Osage orange Tree Intro Pres Unc Fleming 514 white ash Tree Intro Pres Unc Porter 4229 woodland pinedrops Common name

var.

nana & var. var. ora, harrisii Abronia micrantha, pedunculatus T. glandulosa Includes vars. multifl pubescens, Oxybaphus linearis, O. decumbens M. decipiens incarnata fragrans elliptica, A. nana var. Fraxinus pensylvanica lanceolata Synonyms/ notes Taxonomic ora linearis decipiens Tripterocalyx Tripterocalyx micranthus Mirabilis oxybaphoides Mirabilis multifl Mirabilis linearis var. Mirabilis linearis var. Allionia incarnata A. incarnata Abronia nana A. nana Abronia fragrans A. elliptica, Fraxinus pennsylvanica Morus alba Fraxinus anomala Maclura pomifera Fraxinus americana Pterospora andromedea rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Nyctaginaceae Nyctaginaceae Nyctaginaceae Nyctaginaceae Nyctaginaceae Nyctaginaceae Nyctaginaceae Nyctaginaceae Oleaceae Moraceae Oleaceae Moraceae Oleaceae biogeographic notes, cont. Appendix A. Confi FamilyMonotropaceae (Ericaceae) Species name

82 Annotated Checklist of Vascular Flora: Capitol Reef National Park Paradox Valley, Paradox Valley, Montrose Co, CO Herbarium based on collections from outside the park. River, Kane Co, UT River, (Cronquist 10202 MO) Comments NA Reports from CARE doc. Year Year 1986 1987 1980 1986 locality is Type 1973 1971 locality is Paria Type 1981 1978 1971 1973 1981 1986 Native to Europe c name with taxonomic and (CARE) (CARE) (1993) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range AnnF Wide Pres Unc Fleming 86 PerF Wide Pres Unc Heil 3494 PerF Wide Pres Unc Meyer 4618 AnnF Wide Pres Unc Porter 3493 PerF RegEn Pres Unc Harrison 1061 AnnF RegEn Pres Unc Meyer 4710 PerF Wide Pres Abun Camp 539 PerF Wide Pres Unc Heil 3122 AnnF Wide Pres Unc Harrison 966 AnnF Wide Pres Unc Meyer 4768 Life form owered reweed PerF Wide Rep NA Heil et al. branching small-ﬂ gaura Hornemann’s Hornemann’s willow-herb ﬁ northern willow- herb Walker’s Walker’s camissonia Paiute suncup AnnF Wide Pres Abun Harrison s.n. Paria evening- primrose Eastwood’s Eastwood’s camissonia long-tube evening- primrose lavender-leaf lavender-leaf sundrops white-stem evening-primrose groundsmoke common lilac Shrub Intro Pres Unc Porter 4353 Common name

ssp.

ssp. var. var. . Includes brachycarpa canescens, C. E. hornemannii Chamerion angustifolium var. angustifolium circumvagum, C. danielsii ciliatum, E. watsonii, E. adenocaulon Oenothera scapoidea ssp. O. cespitosa marginata lavandulifolius C. hartwegii Synonyms/ notes Taxonomic

var.

var. oramollis G. var. var. var. Gayophytum ramosissimum Gaura parvifl hornemannii Epilobium hornemannii canescens Epilobium angustifolium Epilobium ciliatum E. ciliatum Camissonia walkeri Camissonia scapoidea scapoidea Oenothera caespitosa navajoensis Camissonia eastwoodiae Oenothera caespitosa marginata Oenothera albicaulis Calylophus lavandulifolius Syringa vulgaris rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae biogeographic notes, cont. Appendix A. Confi FamilyOleaceae Species name

Appendices 83 O. ed Herbarium based on collections from outside the park. Armstrong & White canyons (Natural Bridges NM), San Juan Co, UT (Rydberg & Garrett 9410 NY) Dugway, Tooele Co, Tooele Dugway, UT (Jones s.n. RM) Herbarium based on collections from outside the park or misidentiﬁ specimens of longissima. Comments doc. Year Year 1986 1980 2004 Reports from CARE 1958 1958 locality is Type 1980 locality is Type 2004 Reports from CARE 1987 c name with taxonomic and (CARE) (CARE) (1993), Northern Colorado Plateau Network (in ed.) (CARE) (CARE) (CARE) (1993), Northern Colorado Plateau Network (in ed.) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Porter 4056 PerF Wide Pres Abun Harrison 930 PerF RegEn Pres Rare Swancara 38 PerF Wide Pres Rare Heil 3144 PerF Wide Rep Unc Heil et al. PerF Wide Pres Rare Heil 3534 Life form northern bog orchid giant helleborine PerF Wide Pres Unc Meyer 4609 spotted coral-root PerF Wide Rep Rare Heil et al. pale evening- primrose Bridges evening- primrose bronze evening- primrose western evening- primrose hairy-throat evening-primrose Common name

var.

ssp. Platanthera Limnorchis hyperborea, Includes aquilonis, P. huronensis C. maculata maculata Includes var. Includes var. runcinata O. brachycarpa hirsutissima, O. hookeri Synonyms/ notes Taxonomic pallida Habenaria hyperborea Epipactis gigantea Corallorhiza maculata Oenothera pallida var. Oenothera longissima Oenothera howardii Oenothera elata O. elata Oenothera coronopifolia rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Orchidaceae Orchidaceae Orchidaceae Onagraceae Onagraceae Onagraceae Onagraceae biogeographic notes, cont. Appendix A. Conﬁ FamilyOnagraceae Species name

84 Annotated Checklist of Vascular Flora: Capitol Reef National Park been relocated and Colorado been relocated Northern Colorado Plateau Network (in ed.). USFWS: Threatened 1 mile N of Moab, Grand Co, UT (Welsh & Higgins 23269 UT BLM: BRY). Sensitive, USFWS: former C2 candidate Comments NA Specimen has not doc. Year Year 1973 1986 State tree of Utah 1986 1980 1986 1977 Reported by 1987 locality is ca Type 1987 1987 1986 c name with taxonomic and Herbarium (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) BC322 (CARE) (CARE) Source Pop. size Park status Range PerF Wide Rep NA Meyer (1982) NA Specimen has not PerF Wide Pres Unc Meyer 4632 PerF Wide Pres Unc Porter 3777 Tree Wide Pres Unc Porter 4143 Life form limber pine Tree Wide Rep NA NAU two-needle pinyon Tree Wide Pres Abun Harrison 801 blue spruce Tree Wide Pres Unc Porter 3942 armed prickly- poppy Engelmann spruce Tree Wide Pres Unc Porter 3891 Cooper’s Cooper’s broomrape clustered broomrape Ute ladies’-tresses PerF Sparse Ext NA Porter 4112 alcove bog-orchid PerF RegEn Pres Rare Heil 3586 slender bog orchid PerF Wide Pres Rare Heil 3549 ponderosa pine Tree Wide Pres Com Fleming western bristlecone pine Common name

O. Flora from . Vars. not . Vars. O. ora ora Includes multiﬂ UT Flora (1993), cooperi Aphyllon fasciculatum S. diluvialis Platanthera zothecina, Limnorchis zothecina (2002) of North America recognized in Platanthera sparsiﬂ Synonyms/ notes Taxonomic

var. var. ana exilis ora ora rotundata diluvialis scopulorum Pinus fl Pinus edulis Picea pungens Argemone munita var. Picea engelmannii Orobanche ludoviciana cooperi Orobanche fasciculata Spiranthes romanzoffi var. Habenaria zothecina Pinus ponderosa var. Habenaria sparsifl sparsifl Pinus longaeva aristata P. rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Pinaceae Pinaceae Pinaceae Pinaceae Papaveraceae Pinaceae Orobanchaceae Orobanchaceae Orchidaceae Orchidaceae Pinaceae biogeographic notes, cont. Appendix A. Confi FamilyOrchidaceae Species name

Appendices 85 is pachyphylla Cataract Canyon N of Imperial Canyon, San Juan Co, UT 22507 BRY). (Welsh UT BLM: Sensitive Rabbit Valley, Wayne Wayne Rabbit Valley, 575 Co, UT (Ward US). USFS Region 4: Sensitive, USFWS: former Category 1 Candidate Europe. Type locality Europe. Type of the native phase var. Salt Salt Lake City, Lake Co, UT (Jones 1030 POM) Comments doc. Year Year 1980 1986 locality is Type 1986 locality is Type 1986 1980 Mostly native to 1973 1986 Native to Eurasia 1987 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) BC323 (CARE) Source Pop. size Park status Range AnnF Wide Pres Rare Porter 2410 Life form r Tree Wide Pres Unc Fleming ower shy gilia AnnF Wide Pres Abun Meyer 4493 cataract gilia AnnF LocEn Pres Rare Heil 2749 Rabbit Valley giliaRabbit Valley PerF LocEn Pres Rare Heil 2552 few-ﬂ eriastrum common plantain PerF Intro Pres Uncwoolly plantain Meyer 4668 AnnF Wide Pres Com Harrison 1065 English plantain PerF Intro Pres Unc Porter 4222 Douglas-ﬁ Common name

var.

var. & and orum ssp. pachyphylla Includes vars. inconspica sinuata, G. ophthalmoides, G. sinuata, G. clokeyi, G. transmontana imperialis, Aliciella latifolia imperialis wilcoxii, E. wilcoxii E. sparsiﬂ Includes vars. patagonica, gnaphaloides, spinulosa, breviscapa Includes “native” var. Synonyms/ notes Taxonomic

var. orum Gilia inconspicua Gilia imperialis G. latifolia Gilia caespitosa Aliciella caespitosa Eriastrum sparsifl Plantago patagonica Plantago major Plantago lanceolata Pseudotsuga menziesii glauca rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Plantaginaceae Plantaginaceae Plantaginaceae biogeographic notes, cont. Appendix A. Confi FamilyPinaceae Species name

86 Annotated Checklist of Vascular Flora: Capitol Reef National Park “sand banks of the Green River”, UT (Cruetzfeldt s.n. NY) is Springdale, Co, UT Washington (Jones 5247 NY) of Mussentuchit Creek, Sevier Co, UT (Neese, Smith, & Shaw 18025 BRY). UT BLM: Sensitive, USFWS: former C2 candidate Comments doc. Year Year 1986 1980 1980 1980 1971 1981 locality is Type 1980 locality Type 1968 1995 locality is head Type 1970 2005 1973 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) 501 (CARE) (CARE) (CARE) (CARE) s.n. (CARE) (CARE) Lesica 21814 (CARE) (CARE) Source Pop. size Park status Range PerF Sparse Pres Rare Porter 3947 Life form owered gilia owered PerF Wide Pres Unc Lichthardt matted prickly- phlox San Rafael gilia PerF RegEn Pres Unc Meyer 4491 dwarf gilia AnnF Wide Pres Unc Meyer 4565 spreading gilia AnnF Wide Pres Unc Meyer 4478 long-ﬂ Gunnison’s giliaGunnison’s AnnF RegEn Pres Unc Meyer 4729 shrubby gilia PerF RegEn Pres Unc Meyer 4594 scarlet gilia PerF Wide Pres Abun s.n. Taylor Mussentuchit gilia PerF LocEn Pres Rare Clark & carmine gilia PerF RegEn Pres Abun Nelson 9182 common gilia AnnF Wide Pres Rare Fertig & common gilia AnnF Wide Pres Abun Harrison 987 Common name

var. var. ora Linanthus caespitosus Gilia polycladon Gilia longiﬂ Gilia gunnisonii Gilia congesta frutescens Gilia aggregata aggregata G. micromeria, Aliciella micromeria Aliciella leptomeria Synonyms/ notes Taxonomic

var. var. ora micromeria leptomeria Leptodactylon caespitosum Ipomopsis roseata Gilia roseata Ipomopsis pumila Gilia pumila Ipomopsis polycladon Ipomopsis longifl Ipomopsis gunnisonii Ipomopsis congesta frutescens Ipomopsis aggregata aggregata Gilia tenuis Aliciella tenuis Gilia subnuda Aliciella subnuda Gilia leptomeria var. Gilia leptomeria var. rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae biogeographic notes, cont. Appendix A. Confi FamilyPolemoniaceae Species name

Appendices 87 Thompson’s Springs, Thompson’s Grand Co, UT (Jones s.n. POM) Beaver Dam Mtns, Co, UT Washington (Fremont 447 US) Cottonwood Canyon, Salt Lake s.n. Co, UT (Watson GH) Comments doc. Year Year 1973 locality is Type 1986 1968 2004 1971 1987 1970 locality is Type 1987 locality is Type 1980 1973 c name with taxonomic and (CARE) (CARE) (CARE) Colorado Plateau Network (in ed.) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF RegEn Pres Unc Camp 554 PerF Wide Pres Abun Porter 3948 PerF Wide Pres Rare Heil 3425 AnnF Wide Pres Abun Harrison 1183 Life form pretty wild buckwheat winged wild buckwheat cushion milkwort PerF Wide Pres Unc Harrison 875 cushion phlox PerF Wide Rep Rare Northern long-leaf phlox PerF Wide Pres Unc Meyer 4503 carpet phlox PerF Wide Pres Unc Porter 4662 desert phlox PerF Wide Pres Unc Meyer 4433 Watson’s prickly- Watson’s phlox prickly-phlox Shrub Wide Pres Unc Meyer 4546 nodding wild buckwheat Common name

var.

& ssp. var. (2005) alatum, Flora of North Vars. not recognized Vars. in America var. Pterogonum alatum P. subspinosa P. subspinosa pulvinata, P. pulvinata, P. caespitosa stansburyi Includes vars. longifolia viscidula Includes ssp. glabrata, hoodii, Var. not recognized Var. in Cronquist et al. (1984) Linanthus watsonii pungens, Linanthus pungens Leptodactylon pungens Synonyms/ notes Taxonomic

var.

Eriogonum cernuum cernuum Eriogonum bicolor Eriogonum alatum E. alatum Polygala subspinosa Phlox pulvinata sibirica P. Phlox longifolia Phlox hoodii canescens Phlox austromontana var. austromontana Leptodactylon watsonii Leptodactylon pungens rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Polygonaceae Polygonaceae Polygonaceae Polygalaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae biogeographic notes, cont. Appendix A. Conﬁ FamilyPolemoniaceae Species name

88 Annotated Checklist of Vascular Flora: Capitol Reef National Park E. Type Type var. var. (2005). Flora of North River, Emery Co, UT River, (Jones s.n. POM) previously documented as corymbosum revealianum. locality is Thousand Lake Mtn., head of Red Canyon, Wayne Co, UT (Reveal 8543 NY) George, Washington George, Washington Co, UT (Jones 6091 POM). Considered a local endemic restricted to the Shivwits area of Co Washington in America American Fork Canyon, Utah Co., 1033 Utah (Watson US) Comments doc. Year Year 1986 1986 1986 1986 locality is Green Type 1988 CARE material 1970 1970 locality is St. Type 1958 locality is Type c name with taxonomic and (CARE) (CARE) (CARE) (BRY) Groebner 3 (NY) (CARE) (CARE) (CARE) Source Pop. size Park status Range AnnF Wide Pres Unc Porter 3926 AnnF RegEn Pres Com Porter 3737 AnnF Wide Pres Unc Fleming 163 Shrub RegEn Pres Unc Fleming 184 Shrub LocEn Pres Rare Clark & Shrub Wide Pres Abun Harrison 1323 Shrub RegEn Pres Rare Harrison 1300 AnnF Wide Pres Com Harrison 1104 Life form Gordon’s wild Gordon’s buckwheat bent wild buckwheat skeletonweed wild buckwheat rimrock wild buckwheat Heil’s wild Heil’s buckwheat crisp-leaf wild buckwheat golden wild buckwheat Hooker’s wild Hooker’s buckwheat Common name

E. exum

in Welsh in Welsh var. in Welsh in Welsh and exum exum E deﬂ deﬂ lancifolium et al. 2003 Includes var. velutinum Includes var. Includes var. glutinosum et al. 2003 Synonyms/ notes Taxonomic

var. var. var. var. exumfl Stenogonum exum Eriogonum gordonii Eriogonum fl Eriogonum defl Eriogonum corymbosum orbiculatum Eriogonum corymbosum heilii Eriogonum corymbosum corymbosum Eriogonum corymbosum aureum Eriogonum hookeri rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae biogeographic notes, cont. Appendix A. Confi FamilyPolygonaceae Species name

Appendices 89 var. var. is near may belong atum E. inﬂ atum near Butler Wash, near Butler Wash, San Juan Co, UT (Eastwood 134 CAS) longilobum Price, Carbon Co, UT (Jones 5590j POM) River, Emery Co, UT River, (Cruezfeldt s.n. NY) of inﬂ here. Comments doc. Year Year 1973 1970 1986 1977 locality is Type 1973 locality of var. Type 1986 locality is Green Type 1986 1973 1978 1968 Previous reports c name with taxonomic and (CARE) (CARE) (San Juan College) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (BRY) Source Pop. size Park status Range PerF Wide Pres Unc Harrison 1227 PerF Wide Pres Abun Harrison 1304 Shrub RegEn Pres Unc Fleming 199 Shrub RegEn Pres Abun Heil 2854 PerF Wide Pres Unc Harrison 883 Shrub RegEn Pres Unc Heil 2855 PerF Wide Pres Unc Fleming 107 AnnF Wide Pres Rare Camp 5317 AnnF RegEn Pres Unc Harrison 919 Life form Palmer’s wild Palmer’s cushion wild buckwheat slender wild buckwheat slenderleaf wild buckwheat San Juan wild buckwheat Shockley’s wild Shockley’s buckwheat sand wild buckwheat redroot wild buckwheat desert trumpet PerF Wide Pres Unc Harrison 919 buckwheat Grand Valley Grand Valley desert trumpet Common name

E. var. var.

& Welsh et Welsh atum E. ovalifolium ovalifolium E. microthecum foliosum. al. (2003) include phoeniceum Includes vars. longilobum shockleyi E. racemosum E. inﬂ E. fusiforme Synonyms/ notes Taxonomic

var.

var. var. var. var. var. atum atum atum Eriogonum palmerianum Eriogonum ovalifolium Eriogonum microthecum simpsonii Eriogonum leptophyllum Eriogonum leptocladon ramosissimum Eriogonum shockleyi Eriogonum leptocladon leptocladon Eriogonum racemosum racemosum infl Eriogonum infl Eriogonum infl fusiforme rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae biogeographic notes, cont. Appendix A. Confi FamilyPolygonaceae Species name

90 Annotated Checklist of Vascular Flora: Capitol Reef National Park the San Juan River, the San Juan River, San Juan Co, UT (Eastwood 124 CAS) Red Plateau SW of Emery Co, Woodside, UT (Ripley & Barneby 8678 CAS). USFWS: 3C Comments doc. Year Year 1980 1986 1986 1980 Native to Eurasia 1980 Native to Eurasia 1987 1980 Native to Eurasia 1986 locality is along Type 1986 1988 locality is Type 1987 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (BRY) Source Pop. size Park status Range AnnF RegEn Pres Unc Fleming 189 PerF Wide Pres Unc Fleming 146 PerF RegEn Pres Rare Fleming 6398 Fern Wide Pres Unc Fleming 228 Life form southern maidenhair fern willow dock PerF Wide Pres Unc Meyer 4623 golden dock AnnF Wide Pres Unc Porter 4113 canaigre dock PerF Wide Pres Unc Porter 3532 curly dock PerF Intro Pres Com Meyer 4571 willow-weed AnnF Intro Pres Unc Meyer 4624 Sawatch knotweed AnnF Wide Pres Unc Porter 5166 yard knotweed AnnF Intro Pres Unc Meyer 4646 Wetherill’s wild Wetherill’s buckwheat arid wild buckwheat Woodside wild Woodside buckwheat Common name

var.

var. var. var. ssp. mexicanus, R. mexicanus, R. salicifolius var. R. utahensis, triangulivalvis mexicanus, Rumex triangulivalvis montigenitus R. fueginus Persicaria lapathifolia sawatchense P. sawatchense P. P. arenastrum, P. aviculare P. aviculare Synonyms/ notes Taxonomic

var. var. triangulivalvis fueginus Adiantum capillus- veneris Rumex salicifolius ssp. Rumex maritimus var. Rumex hymenosepalus Rumex crispus Polygonum lapathifolium Polygonum douglasii johnstonii Polygonum aviculare Eriogonum wetherillii subaridum Eriogonum umbellatum Eriogonum tumulosum rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Polypodiaceae (Adiantaceae) Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae biogeographic notes, cont. Appendix A. Conﬁ FamilyPolygonaceae Species name

Appendices 91 C. is Negro America utahensis Bill Canyon, Grand Co, UT (Windham & Windham 90-282) San Juan River near San Juan Co, Bluff, UT (Rydberg 9882 NY). USFWS: 3C Comments doc. Year Year 1939 1981 1980 1974 1980 Native to E North 1986 1986 1990 1987 locality of Type 1970 1977 locality is along Type c name with taxonomic and Neese 23451a (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Mason 4137 (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range AnnF Wide Pres Unc Meyer 4775 PerF Wide Pres Unc Heil 3490 PerF Wide Pres Rare Harrison 1335 Life form ower sea-milkwort PerF Wide Pres Unc & Welsh pygmy-ﬂ rock-jasmine slenderleaf pondweed sausageleaf talinum common purslane AnnF Intro Pres Unc Meyer 4647 Oregon woodsia Fern Periph Pres Unc Porter 4180 border cloak-fern Fern Periph Pres Rare Fleming 181 male fern Fern Wide Pres Rare Spence & brittle bladder fern Fern Wide Pres Rare Heil 3568 slender lip-fern Fern Wide Pres Unc Nelson 9190 cave primrose PerF RegEn Pres Rare Camp s.n. Common name

& var. & var.

C. ssp. & liformis ssp. C. occidentalis subilifera Includes ssp. puberulenta, subumbellata, Stuckenia ﬁ var. Phemeranthus brevifolius oregana limitanea limitanea, Notholanea limitanea Includes tenuis utahensis, C. reevesiana, Synonyms/ notes Taxonomic

lix- var. liformis Glaux maritima Androsace septentrionalis Potamogeton fi occidentalis Talinum Talinum brevifolium Portulaca oleracea Woodsia oregana Woodsia oregana W. Pellaea limitanea Argyrochosma Dryopteris fi mas Cystopteris fragilis Cheilanthes feei Primula specuicola rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Primulaceae Primulaceae (Myrsinaceae) Primulaceae Potamogetonaceae Portulacaceae Portulacaceae Polypodiaceae (Aspleniaceae, Woodsiaceae) Polypodiaceae (Adiantaceae, Sinopteridaceae) Polypodiaceae (Aspleniaceae) Polypodiaceae (Aspleniaceae) biogeographic notes, cont. Appendix A. Confi FamilyPolypodiaceae (Adiantaceae) Species name

92 Annotated Checklist of Vascular Flora: Capitol Reef National Park San Juan Co, UT s.n. CAS) (Wetherill Herbarium based on collections from outside the park. Comments NA Reports from CARE doc. Year Year 1980 1973 1958 1981 1987 locality is Bluff, Type 1986 2004 1987 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (BRY) (CARE) Colorado Plateau Network (in ed.) (1993) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Swancara 16 PerF Wide Pres Unc Meyer 4757 PerF Wide Pres Unc Heil 3561 PerF Wide Rep NA Heil et al. Life form marsh buttercup PerF Wide Pres Unc Meyer 4561 pale larkspur PerF Wide Pres Unc Harrison 901 white virgin’s- bower Columbian virgin’s- bower alcove columbine PerF RegEn Pres Rare Porter 4679 Colorado columbine red baneberry PerF Wide Rep Unc Northern Columbian monkshood blister buttercup AnnF Wide Pres Unc Heil 3522 Common name

includes arguta Vars. Vars. ssp. var. ssp. (1997) ochroleuca columbiana, Flora of North Halerpestes cymbalaria saximontana, R. cymbalaria cymbalaria. not recognized in America D. scaposum C. columbiana var. C. pseudoalpina, Atragene columbiana pinetorum vars. Includes var. Includes var. ochroleucum Hecatonia scelerata Synonyms/ notes Taxonomic

var.

var. var. dus Ranunculus sceleratus multifi Ranunculus cymbalaria Delphinium andersonii scaposum Clematis ligusticifolia Clematis columbiana Aquilegia micrantha Aquilegia corulea A. caerulea, Aconitum columbianum columbianum Actaea rubra A. rubra rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Ranunculaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) biogeographic notes, cont. Appendix A. Confi FamilyRanunculaceae (Helleboraceae) Species name Ranunculaceae (Helleboraceae)

Appendices 93 10 km E of Oak Millard Co, City, UT (Holmgren & Holmgren 11298 NY) American Fork Canyon, Utah Co, UT 314 US) (Watson is Toquerville, is Toquerville, Co, UT Washington (Jones 1716 GH) Comments doc. Year Year 1986 locality is ca Type 1958 locality is Type 1966 locality Type 1970 1981 1986 2004 1987 Native to SW Asia 1986 1978 1987 Native to Eurasia 1986 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Colorado Plateau Network (in ed.) 288 (CARE) (CARE) (CARE) BC282 (CARE) Source Pop. size Park status Range Shrub Wide Pres Unc Porter 3856 Shrub Wide Pres Unc Porter 3959 Shrub Wide Pres Abun Harrison 834 Shrub Wide Pres Unc Nelson 9482 Shrub Wide Pres Unc Meyer 4713 Shrub Wide Pres Unc Porter 3887 PerF Wide Rep Rare Northern PerF Wide Pres Unc Porter 3860 Life form ower alder-leaf alder-leaf mountain mahogany curl-leaf mountain mahogany dwarf mountain mahogany Utah serviceberry Shrub Wide Pres Abun Harrison 1378 Saskatoon serviceberry birchleaf buckthorn Fendler’s Fendler’s mountain-lilac few-ﬂ meadow-rue quince Tree Intro Pres Unc Fleming BC- Fendler’s meadow- Fendler’s rue blackbrush Shrub Wide Pres Unc Camp 538 bur buttercup AnnF Intro Pres Rare Fleming Common name

var. var. var. orum C. ledifolius intercedens C. ledifolius intricatus Frangula betulifolia saximontanum T. sparsiﬂ T. Ceratocephala orthoceras, C. testiculata Synonyms/ notes Taxonomic

var. orum Cercocarpus montanus Cercocarpus ledifolius intermontanus Cercocarpus intricatus Amelanchier utahensis Amelanchier alnifolia Rhamnus betulifolia Ceanothus fendleri Thalictrum sparsifl Cydonia oblonga Thalictrum fendleri Coleogyne ramosissima Ranunculus testiculatus rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rhamnaceae Rhamnaceae Ranunculaceae (Thalictraceae) Rosaceae Ranunculaceae (Thalictraceae) Rosaceae biogeographic notes, cont. Appendix A. Confi FamilyRanunculaceae Species name

94 Annotated Checklist of Vascular Flora: Capitol Reef National Park Comments doc. Year Year 1981 1987 1987 2004 1980 1977 1987 Native to Eurasia 1973 Native to Eurasia 1986 Native to Japan 1971 2004 1958 c name with taxonomic and (CARE) (CARE) BC317 (CARE) Colorado Plateau Network (in ed.) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (1993), Northern Colorado Plateau Network (in ed.) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Meyer 4769 PerF Wide Pres Unc Fleming Life form beautiful cinquefoil hair-tuft cinquefoilhair-tuft PerF Wide Pres Unc Heil 3535 bicrenate cinquefoil silverweed PerF Wide Rep Rare Northern dwarf ninebark Shrub Wide Pres Unc Meyer 4512 rock spiraea PerF Wide Pres Unc Camp 269 crabapple Tree Intro Pres Unc Fleming 515 common apple Tree Intro Pres Unc Harrison 1369 showy crabapple Tree Intro Pres Unc Porter 4216 bush oceanspray Shrub Wide Pres Unc Nelson 9839 Virginia strawberry PerF Wide Rep Unc Heil et al. Apache plume Shrub Wide Pres Abun Nelson 9589 Common name glauca platypetala P. pulcherrima P. P. bicrenata P. Spiraea caespitosa H. discolor & Includes ssp. Synonyms/ notes Taxonomic

var. oribunda pulcherrima Potentilla gracilis var. Potentilla crinita bicrenata Potentilla concinna Potentilla anserina Argentina anserina Physocarpus alternans Petrophytum caespitosum Malus sylvestris Malus pumila Pyrus malus Malus fl Holodiscus dumosus Fragaria virginiana Fallugia paradoxa rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae biogeographic notes, cont. Appendix A. Confi FamilyRosaceae Species name

Appendices 95 ed

or on Herbarium based on misidentiﬁ specimens of crinita P. collections from outside the park. Comments doc. Year Year 1971 1973 Native to Eurasia 1981 1970 1986 Native to China 1986 Native to Old World 1986 Native to Eurasia 1986 Native to Eurasia 1986 Native to Asia 1973 Native to Eurasia 1986 Native to China 1987 2004 Reports from CARE 1981 c name with taxonomic and BC375 (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (1993), Northern Colorado Plateau Network (in ed.) (CARE) Source Pop. size Park status Range Shrub Wide Pres Com Meyer 4753 PerF Wide Pres Unc Heil 3566 Life form owering plum Tree Intro Pres Unc Porter 4348 cleavers AnnF Wide Pres Unc Meyer 4711 Woods’ roseWoods’ Shrub Wide Pres Com Fleming common pear Tree Intro Pres Unc Harrison 852 antelope bitterbrush cliff-rose Shrub Wide Pres Abun Nelson 9588 peach Tree Intro Pres Unc Porter 4359 almond Tree Intro Pres Unc Porter 4358 common plum Tree Intro Pres Unc Porter 4225 sour cherry Tree Intro Pres Unc Porter 4357 ﬂ sweet cherry Tree Intro Pres Unc Harrison 853 apricot Tree Intro Pres Unc Porter 4214 Pennsylvania cinquefoil woolly cinquefoil PerF Wide Rep Unc Heil et al. Common name

, var. var. R. manca G. spurium ultramontana Includes P. stansburiana, P. Cowania mexicana P. hippiana P. hippiana Synonyms/ notes Taxonomic

var. stansburyana Galium aparine echinospermon Rosa woodsii R. woodsii Pyrus communis Purshia tridentata Purshia mexicana var. Prunus persica Prunus dulcis Prunus domestica Prunus cerasus Prunus cerasifera Prunus avium Prunus armeniaca Armeniaca vulgaris Potentilla pensylvanica Potentilla hippiana rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Rubiaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae biogeographic notes, cont. Appendix A. Conﬁ FamilyRosaceae Species name

96 Annotated Checklist of Vascular Flora: Capitol Reef National Park Ewing Canyon, Daggett Co, UT (Goodrich 24441 BRY) Comments doc. Year Year 1974 1981 2004 1986 1986 1987 locality is Jesse Type 1986 1986 Native to Eurasia 1973 1958 1980 1986 1980 1973 c name with taxonomic and (CARE) (CARE) Colorado Plateau Network (in ed.) (CARE) (BRY) BC296 (BRY BC296 (BRY Paratype) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range Tree Wide Pres Unc Fleming Tree Wide Pres Abun Harrison 1389 Tree Wide Pres Unc Nelson 9535 Tree Wide Pres Unc Meyer 4516 Life form yellow willow Shrub Wide Pres Unc Harrison 1460 strapleaf willow Shrub Wide Pres Unc Meyer 4706 Booth’s willowBooth’s Shrub Wide Rep Rare Northern weeping willow Tree Intro Pres Unc Porter 4362 peach-leaf willow Tree Wide Pres Unc Porter 4070 lance-leaf cottonwood quaking aspen Tree Wide Pres Unc Porter 3944 black poplar Tree Intro Pres Unc Porter 4361 Fremont cottonwood narrow-leaved cottonwood arroyo willow Shrub Wide Pres Unc Meyer 4443 shrubby bedstraw PerF Wide Pres Unc Porter 2424 Goodding’s black Goodding’s willow coyote willow Shrub Wide Pres Com Meyer 4455 Common name

P. x . x exigua Hybrid vallicola var var. orum var. watsonii P. fremontii P. S. lutea, rigida var. S. ligulifolia angustifolia P. acuminata, P. acuminata, P. P. acuminata. of P. deltoides P. fremontii G. multiﬂ S. exigua Synonyms/ notes Taxonomic

var. var.

x orum orum watsonii ligulifolia fremontii Salix eriocephala var. Salix eriocephala var. Salix boothii Salix babylonica Salix amygdaloides Populus intercurrens Populus tremuloides Populus nigra Populus fremontii var. Populus angustifolia Salix lasiolepis multifl Salix gooddingii S. nigra Galium multifl Salix exigua stenophylla rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae biogeographic notes, cont. Appendix A. Confi FamilyRubiaceae Species name

Appendices 97 been relocated. Kanab, Kane Co, UT (Thompson s.n. GH). Probably collected farther to the E. Comments doc. Year Year 1987 1970 1971 1980 1974 1986 1986 locality is Type 1981 1986 Native to Asia 1973 1987 1986 1980 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Source Pop. size Park status Range Shrub Wide Pres Unc Meyer 4577 PerF Wide Pres Abun Nelson 9843 PerF Wide Pres Unc Meyer 4593 Shrub Wide Pres Unc Heil 3297 Life form ax PerF Wide Pres Com908 Harrison golden currant Shrub Wide Pres Unc Porter 5093 littleleaf mock- orange Wyoming paintbrush marsh grass-of- Parnassus red alumroot PerF Wide Rep NA Meyer (1982) NA Specimen has not annual paintbrush AnnF Wide Pres Unc Camp 407 littleleaf alumroot PerF Wide Pres Rare Porter 3957 bastard toadﬂ Utah fendlerella Shrub Wide Pres Unc Heil 2684 desert paintbrush PerF Wide Pres Abun Meyer 4763 Matsuda’s willowMatsuda’s Tree Intro Pres Unc Porter 4350 whiplash willow Shrub Wide Pres Rare Porter 5182 whitestem gooseberry wax currant Shrub Wide Pres Unc Meyer 4578 Common name

var. var.

tenuis minor var. var. var. ssp. (1997) martinii occidentalis, Intermountain aureum P. microphyllus P. var. occidentalis P. occidentalis C. linariaefolia Vars. not recognized Vars. in Includes var. Flora utahensis Includes vars. parvifolia, major, C. pallida dubia, C. applegatei ssp. C. angustifolia S. lasiandra caudata R. cereum pedicellare Synonyms/ notes Taxonomic

var. ssp. var. montanensis Ribes cereum inebrians Ribes aureum R. aureum Philadelphus microphyllus Castilleja linariifolia Heuchera rubescens Parnassia palustris var. Heuchera parvifolia Castilleja exilis C. minor Comandra umbellata pallida Fendlerella utahensis Salix matsudana Castilleja chromosa caudata Ribes inerme Salix lucida rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Saxifragaceae (Hydrangeaceae) Scrophulariaceae Saxifragaceae Saxifragaceae (Parnassiaceae) Saxifragaceae Scrophulariaceae Santalaceae Saxifragaceae (Hydrangeaceae) Salicaceae Scrophulariaceae Saxifragaceae (Grossulariaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilySalicaceae Species name

98 Annotated Checklist of Vascular Flora: Capitol Reef National Park Lake, Millard Co, UT 11180) (Tidestrom Rafael Swell, Emery Co, UT (Cottam 5229 PH) Juan River near San Juan Co, Bluff, UT (Rydberg 9883 NY) Comments doc. Year Year 1978 1986 1987 1986 1987 locality is Preuss Type 1987 locality is San Type 1977 locality is San Type 1988 1986 2005 1986 1971 c name with taxonomic and (CARE) (BRY) BC362 (CARE) (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (BRY) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Porter 4281 PerF Wide Pres Rare Heil 3526 PerF RegEn Pres Rare Heil 2695 PerF RegEn Pres Rare Fleming 638 AnnF Wide Pres Unc Porter 3814 AnnF Wide Pres Unc Harms 186 PerF Wide Pres Unc Heil 2605 PerF RegEn Pres Unc Meyer 4432 Life form ower ower ower ower bird’s- ower eshy penstemon PerF RegEn Pres Unc Heil 3103 beardlip veined penstemon PerF RegEn Pres Unc Camp 546 bush penstemon PerF Wide Pres Rare Heil 2922 dwarf lousewort PerF Wide Pres Unc Fleming yellow monkeyfl glabrous monkeyfl fl Eastwood’s Eastwood’s monkey-fl painted penstemon Wright’s bird’s- Wright’s beak smallfl beak penstemon Eastwood’s Eastwood’s paintbrush Common name

var. torreyi caudatus Includes var. Includes var. Sometimes confused with ssp. of WY & Great Plains Leiostemon ambiguus M. glabratus glabratus C. scabrida Synonyms/ notes Taxonomic

var.

var. var. var. orus utahensis scabrida Penstemon barbatus barbatus Penstemon angustifolius venosus laevissimus Penstemon ambiguus Pedicularis centranthera Mimulus guttatus Mimulus glabratus var. Penstemon carnosus Mimulus eastwoodiae desertipicti Cordylanthus wrightii Penstemon caespitosus Cordylanthus parvifl Castilleja scabrida var. rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae biogeographic notes, cont. Appendix A. Confi FamilyScrophulariaceae Species name

Appendices 99 eld Co, UT Musinea Peak, Sanpete Co, UT 568 US) (Tidestrom miles N of Kanab, Kane Co, UT (Maguire, Maguire, & Piranian 12279 UTC) near Canaan Ranch, Rockville, Co, UT Washington (Jones 5262 POM) Thurber, near Loa, Thurber, Co, UT (Jones Wayne 5708 US) George, Washington George, Washington Co, UT (Jones 5110h US) Provo Canyon, Utah 776 Co, UT (Watson GH) Aquarius Plateau, Garﬁ 462 GH) (Ward Comments doc. Year Year 1986 1986 locality is Type 1986 1958 locality is 2.5 Type 1980 locality is Type 1968 locality is Type 1988 1987 locality is St. Type 2005 locality is lower Type 1980 locality is Type c name with taxonomic and (BRY) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) BC278 (CARE) Lesica 21823 (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Heil 2612 PerF RegEn Pres Unc Swancara 1 PerF Wide Pres Unc Meyer 4761 PerF RegEn Pres Unc Fleming 595 PerF Wide Pres Abun Fleming Life form recracker PerF Wide Pres Unc& Fertig Rocky Mountain penstemon plateau penstemon PerF RegEn Pres Unc Heil 3305 beaked penstemon PerF Wide Pres Unc Porter 4136 Palmer’s Palmer’s penstemon Rockville penstemon Loa penstemon PerF RegEn Pres Unc Meyer 4738 Moffatt Moffatt penstemon undulate penstemon Eaton’s ﬁ Eaton’s dusty penstemon PerF RegEn Pres Unc Meyer 4582 Common name P. bridgesii P. P. eatonii P. Synonyms/ notes Taxonomic

var.

var. var. orus undosus eatonii Penstemon strictus Penstemon scariosus scariosus Penstemon rostrifl Penstemon palmeri eglandulosus Penstemon pachyphyllus congestus Penstemon ophianthus Penstemon moffatii Penstemon eatonii var. Penstemon eatonii var. Penstemon comarrhenus rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae biogeographic notes, cont. Appendix A. Confi FamilyScrophulariaceae Species name

100 Annotated Checklist of Vascular Flora: Capitol Reef National Park America Native to Eurasia America, may be adventive in UT. between Hatch’s between Hatch’s & Monticello, Wash San Juan Co, UT (Eastwood s.n. CAS) America Comments doc. Year Year 1986 Native to C North 1958 1981 1980 1981 1958 1958 Native to China 1980 1980 Native to Europe 1986 1986 Native to C North 1971 locality is Type 1980 Native to South c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) Andersen (2005) 2004 (BRY) (CARE) (CARE) Source Pop. size Park status Range PerF Intro Pres Unc Porter 4456 PerF Wide Pres Com Porter 4373 Shrub Wide Pres Unc Meyer 4726 Fern Wide Pres Unc Meyer 4435 PerF Wide Pres Unc Porter 4285 AnnF Intro Pres Unc Meyer 4674 Life form silverleaf nightshade long-leaf ground- cherry desert tobacco AnnF Wide Pres Unc Meyer 4724 coyote tobacco AnnF Wide Pres Unc Meyer 4628 Anderson’s Anderson’s wolfberry angel’s trumpetangel’s AnnF Wide Pres Unc Swancara 18 tree-of-heaven Tree Intro Pres Unc Harrison 1367 blunt-leaf spike- moss water speedwell PerF Intro Pres Unc Meyer 4538 American brooklime woolly mullein PerF Intro Rep Unc Dewey and cut-leaf nightshade AnnF Wide Pres Rare Heil 3521 Utah penstemon PerF Wide Pres Com Meyer 4441 ground-cherry nightshade Common name

vars. var. mutica var. var.

var. & S. villosum, physalifolium sonorae palmeri, N. obtusifolia, trigonophylla trigonophylla nitidibaccatum Synonyms/ notes Taxonomic orum Solanum sarrachoides Solanum elaeagnifolium Physalis longifolia virginiana P. Nicotiana trigonophylla Nicotiana attenuata Lycium andersonii Lycium Datura wrightii D. meteloides Ailanthus altissima Selaginella mutica S. mutica Veronica Veronica anagallis-aquatica Veronica Veronica americana Verbascum Verbascum thapsus Solanum trifl Penstemon utahensis rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientifi Solanaceae Solanaceae Solanaceae Solanaceae Solanaceae Solanaceae Solanaceae Simaroubaceae Selaginellaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Solanaceae biogeographic notes, cont. Appendix A. Confi FamilyScrophulariaceae Species name

Appendices 101 Fruita, Capitol Reef Co, UT Wayne NP, USFS: (Beck s.n. BRY). Region 4 Sensitive, UT BLM: Sensitive, USFWS: former C2 candidate America Comments doc. Year Year 1971 1938 locality is Type 1980 1958 1986 Native to Asia 2004 Native to Asia 1986 Native to Asia 1970 1988 Native to E North 1980 1986 1973 1958 Native to Eurasia 1986 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) Andersen (2005) (CARE) & Hak 446 (CARE) (CARE) (CARE) (CARE) 971A (CARE) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Unc Fleming 619 PerF LocEn Pres Unc Camp 440 PerF Wide Pres Unc Meyer 4563 PerG Wide Pres Com Meyer 4660 PerF Wide Pres Unc Harrison Shrub Intro Pres Abun Swancara 20 PerF Wide Pres Rare Heil 3287 Life form ve-stamen onion spring- parsley Beck’s spring- Beck’s parsley Fendler’s Fendler’s cymopterus water-hemlock PerF Wide Pres Rare Harrison 1284 dill AnnF Intro Pres Unc Porter 4374 Siberian elm Tree Intro Rep Unc Dewey and Chinese elm Tree Intro Pres Unc Porter 4220 netleaf hackberry Tree Wide Pres Unc May, Fowler, common hackberry Tree Intro Pres Unc Heil 4431 broad-leaved cattail southern cattail PerG Wide Pres Unc Porter 4147 basin white-cup spring-parsley ﬁ tamarisk Lemmon’s spring- Lemmon’s parsley Common name

var. C. fendleri, glomeratus C. douglasii, occidentalis reticulata ramosissima Pseudocymopterus montanus Synonyms/ notes Taxonomic

var. angustifolia Cymopterus lemmonii Cymopterus bulbosus Cymopterus beckii Cymopterus acaulis fendleri Cicuta maculata var. Anethum graveolens Ulmus pumila Ulmus parvifolia Celtis reticulata C. laevigata Celtis occidentalis Typha latifolia Typha Typha Typha domingensis Cymopterus purpurascens Tamarix chinensisTamarix pentandra, T. T. rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Ulmaceae Ulmaceae Ulmaceae (Celtidaceae) Ulmaceae (Celtidaceae) Typhaceae Typhaceae Umbelliferae (Apiaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyTamaricaceae Species name

102 Annotated Checklist of Vascular Flora: Capitol Reef National Park Salt Lake (Stansbury s.n. NY) Herbarium based on collections from outside the park. Herbarium based on collections from outside the park. “Southern Utah” (Parry 85) Rafael Swell, Emery Co, UT (Holmgren, Holmgren, & Barneby 8778 NY) Herbarium based on collections from outside the park. Comments NA Reports from CARE NA NA Reports from CARE NA Reports from CARE doc. Year Year 1980 1980 1977 locality is Great Type 1980 locality is Type 1980 locality is San Type 1973 c name with taxonomic and (CARE) (CARE) (1993) Conservation Data Center database (1993) (CARE) (CARE) (1993) (CARE) (CARE) Source Pop. size Park status Range PerF Wide Rep NA Heil et al. PerF RegEn Pres Com Meyer 4551 Life form at-seed lomatium PerF Wide Pres Unc Camp 440 prostrate vervain PerF Wide Pres Unc Meyer 4684 stinging nettle PerF Wide Pres Unc Meyer 4619 blunt sweet-cicely PerF Wide Rep NA Heil et al. water parsnip PerF Wide Rep NA Heil et al. common yampah PerF Wide Rep Unc UT ﬂ Parry’s lomatiumParry’s PerF Wide Pres Rare Porter 4686 southern ligusticum rush lomatium PerF LocEn Pres Unc Heil 3502 Colorado Plateau spring-parsley Common name

ssp. ssp. ssp. holosericea P. gairdneri P. borealis U. dioica holosericea, U. gracilis L. simplex C. purpureus Synonyms/ notes Taxonomic

var. ssp. var. var. Verbena bracteata Verbena Osmorhiza depauperata Sium suave borealis Perideridia montana Urtica dioica occidentalis platycarpum Lomatium triternatum Lomatium parryi Ligusticum porteri Lomatium junceum purpureus Cymopterus purpureus rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Verbenaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Urticaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) biogeographic notes, cont. Appendix A. Conﬁ FamilyUmbelliferae (Apiaceae) Species name

Appendices 103

P. Picea and (rarely or Juniperus Pseudotsuga Abies America on osteosperma Pinus ponderosa on menziesii on species) Pinus edulis monophylla Comments doc. Year Year 1986 Native to Eurasia 1980 1986 Native to E North 1987 1986 parasitic Typically 1988 parasitic on Typically 1987 parasitic Typically 1986 parasitic on Typically 1987 1987 c name with taxonomic and (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) (CARE) BC339 (CARE) (CARE) Source Pop. size Park status Range PerF Wide Pres Rare Heil 4368 PerF Wide Pres Rare Fleming 462 PerF Wide Pres Unc Fleming 130 PerF Wide Pres Rare Fleming Life form r dwarf- r puncture vine AnnF Intro Pres Unc Fleming 82 horned pondweed PerF Wide Pres Unc Meyer 4616 northern fox grape Shrub Intro Pres Unc Porter 4224 thicket-creeper Shrub Wide Pres Rare Heil 3593 juniper mistletoe PerF Wide Pres Unc Porter 4065 southwestern dwarf-mistletoe Douglas-ﬁ mistletoe pinyon dwarf- mistletoe northern bog violet hook-spur violet PerF Wide Pres Unc Heil 3294 Common name nis afﬁ var. V. sororia V. Synonyms/ notes Taxonomic

var. Tribulus terrestris Tribulus Zannichellia palustris Vitis labrusca Parthenocissus vitacea Phoradendron juniperinum Arceuthobium vaginatum cryptopodum Arceuthobium douglasii Arceuthobium divaricatum Viola nephrophylla Viola adunca rmed, historical, and reported vascular plant taxa of Capitol Reef National Park, organized by family scientiﬁ Zygophyllaceae Zannichelliaceae Vitaceae Vitaceae Viscaceae Viscaceae Viscaceae Viscaceae Violaceae biogeographic notes, cont. Appendix A. Conﬁ FamilyViolaceae Species name

104 Annotated Checklist of Vascular Flora: Capitol Reef National Park ed specimens of . USFWS: 3C Grand Co, UT (Eastwood s.n. CAS) on collections from outside the park. based on misidentifi C. abata on collections from outside the park. road, San Rafael Swell, Emery Co, USFWS: UT (Despain 266a BRY). Endangered Range Comments Life form PerF RegEn Reports from CARE Herbarium based PerF LocEn Overlook locality is Wedge Type ower AnnF Wide ower AnnF RegEn Utah pricklypear PerF Wide Common name yuccaBailey’s Shrub Periph sea purslane PerF Wide Hall’s milkweedHall’s orange milkweed PerF PerF Sparse Wide Wetherill’s cryptanthWetherill’s showy stickseed PerF RegEn locality is Courthouse Wash, Type PerF Wide Jones’ popcorn-fl twist-spine pricklypear PerF Wide climbing milkweed cryptanth PerFBaker’s Periph Reports from CARE Herbarium based PerF RegEn Paradox cryptanth PerF RegEn Reports from CARE Herbarium are meadow popcorn-fl Glen Canyon hedgehog cactus Nichol’s pricklypearNichol’s PerF LocEn Whipple’s chollaWhipple’s Shrub RegEn Despain’s pincushion Despain’s cactus common pricklypear PerF Wide

ssp.

nicholii erinacea

var. var. var. terminalis var. Flora of North Flora of North hispidulus Vars. not Vars. ssp. Var. not Var. var. (2003) (2003) Synonyms/Taxonomic Synonyms/Taxonomic notes O. macrorhiza A. tuberosa Funastrum cynanchoides heterophyllum P. scouleri P. decumbens. recognized in America E. engelmannii O. polyacantha O. polyacantha hystricina. recognized in America

var.

var. var. var. var.

var. rmed) taxa of Capitol Reef National Park. oribunda variegatus Opuntia macrorhiza macrorhiza Yucca baileyi Yucca Sesuvium verrucosum Asclepias hallii Asclepias tuberosa interior Sarcostemma cynanchoides hartwegii Cryptantha wetherillii Hackelia ﬂ Plagiobothrys jonesii Plagiobothrys scouleri penicillatus Opuntia nicholii O. polyacantha Cryptantha bakeri Oreocarya bakeri Cryptantha paradoxa Oreocarya paradoxa Echinocereus engelmannii var. Opuntia whipplei Cylindropuntia whipplei Pediocactus despainii Opuntia erinacea erinacea Opuntia erinacea utahensis Cactaceae FamilyAgavaceae Species name Aizoaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Cactaceae Boraginaceae Boraginaceae Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Appendix B. Potential (unconﬁ

Appendices 105 . eld Co, UT (Stutz ed specimens of longipes var. var. River, Grand Co, UT (Stutz 9547 BRY) River, Spring, Hwy 95, Garfi 9575 BRY) UT (Jones 5481 POM) Grand Co, UT (Rydberg & Garrett 8465 NY) based on misidentifi S. longipes Range Comments Life form eld chickweed eld PerF Wide Common name black twinberry Shrub Wide Green River orach AnnF LocEn locality is 3 miles E of Green Type horned orach AnnF RegEn locality is ca 0.5 mi SE of Hog Type red elderberry Shrub Wide blade orach AnnF RegEn Emery Co, locality is Green River, Type Hooker’s sandwortHooker’s PerF RegEn blunt-leaf sandwort PerF Wide Rydberg’s sandwortRydberg’s PerF Wide reddish sandwort PerF Wide fi umbrella starwort PerF Wide Garrett’s saltbushGarrett’s Shrub LocEn locality is vicinity of Moab, Type Menzies’ campionnorthern starwort PerF Wide PerF Wide James’ chickweed PerF Wide Reports from CARE Herbarium are S.

by Flora ssp.

in var. ora (2005) menziesii, Included in sitchana racemosa var. hookeri Eremogone ssp. var. var. Synonyms/Taxonomic Synonyms/Taxonomic notes A. minuticarpa S. microbotrys. A. asterocarpa racemosa some authors A. cornuta Included in hookeri of North America obtusiloba Minuartia rubella, Tryphane Minuartia rubella, Tryphane rubella Strictum Anotites menziesii menziesii

var.

var. var. var. var. var. var. oralaterifl Moehringia rmed) taxa of Capitol Reef National Park, cont. minuticarpa Lonicera involucrata Distegia involucrata Sambucus racemosa microbotrys Atriplex saccaria asterocarpa Atriplex saccaria cornuta Arenaria hookeri desertorum Arenaria laterifl Arenaria obtusiloba Minuartia obtusiloba, Lidia Arenaria rubella rubella Cerastium arvense C. strictum, arvense Silene menziesii S. menziesii Stellaria umbellata Atriplex garrettii garrettii Atriplex powellii Stellaria calycanthaStellaria jamesiana S. borealis Pseudostellaria jamesiana FamilyCaprifoliaceae Species name Caprifoliaceae (Adoxaceae) Chenopodiaceae Chenopodiaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae Caryophyllaceae (Alsinaceae) Chenopodiaceae Chenopodiaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Appendix B. Potential (unconfi

106 Annotated Checklist of Vascular Flora: Capitol Reef National Park in Welsh et al. in Welsh cordata Mountains, Beaver Co, UT (Martin & 17148 BRY) Welsh on collections from outside the park. locality is W of Leeds along Type road to Silver Reef Campgrounds, Co, UT (Semple & Heard Washington 7859 NY) on collections from outside the park. 2003 on collections from outside the park. Range Comments Life form Common name hairy golden-aster PerF Sparse smallhead goosefoot AnnF Wide locality is Indian Creek, Tushar Type Zion golden-aster PerF RegEn Reports from CARE Herbarium based Fremont’s goosefootFremont’s winged pigweed AnnF Wide Reports from CARE Herbarium based AnnF Wide Wright’s pereziaWright’s PerF Periph orange agoseris PerF Wide short agoseris PerF Wide Chamisso’s arnicaChamisso’s PerF Wide heartleaf arnica PerF Wide Misspelled var. biennial wormwood PerF Wide Carruth’s wormwoodCarruth’s PerF Wide Eaton’s asterEaton’s PerF Wide Canby’s asterCanby’s PerF Wide Reports from CARE Herbarium based Siskiyou aster PerF Wide Mexican devilweed PerF Wide

Flora

var. var. andina

(2006) hesperium var. var. biennis var. var. foliosa var. canbyi Synonyms/Taxonomic Synonyms/Taxonomic notes Heterotheca villosa villosa C. overi, foliosum villosa C. fremontii A. chamissonis Vars. not recognized in Vars. of North America bracteolatus var. spinosa hesperius, Symphyotrichum lanceolatum

var.

var. canbyi Symphyotrichum foliaceum var. var. var. var. rmed) taxa of Capitol Reef National Park, cont. parvicapitatum fremontii Chrysopsis villosa villosa Chenopodium capitatum var. Chrysopsis zionensis Heterotheca zionensis, H. Chenopodium fremontii var. Cycloloma atriplicifolia C. atriplicifolium Acourtia wrightii Perezia wrightii Agoseris aurantiaca aurantiaca Agoseris glauca dasycephala Arnica chamissonis foliosa Arnica cordifolia cordifolia Artemisia biennisArtemisia carruthii A. biennis Aster eatonii Symphyotrichum eatonii, A. Aster foliaceus Aster hesperius A. lanceolatus Aster spinosus Chloracantha spinosa Appendix B. Potential (unconﬁ FamilyChenopodiaceae Species name Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Appendices 107 on collections from outside the park. on collections from outside the park. Eurasia UT (Jones 5715 NY) San Juan Co, UT (Rydberg near Bluff, & Garrett 10001 NY). USFWS: 3C Range Comments Life form Common name curly-cup gumweed PerF Wide pretty rabbitbrush Shrub Periph Vasey’s rabbitbrushVasey’s Shrub Wide Reports from CARE Herbarium based Clement’s goldenweedClement’s PerF Wide Reports from CARE Herbarium based Canada thistleFish Lake thistle thistleRydberg’s PerF PerF Intromountain hawksbeard RegEn Utah state noxious weed. Native to PerF PerF locality is Fish Lake, Sevier Co, Type RegEn Wide locality is along San Juan River Type gray hawksbeard PerF Wide noddinghead PerF RegEn hairy daisy PerF Wide cut-leaved daisy PerF Wide spreading daisybeautiful daisy PerF Wide PerF Wide lowland cudweed AnnF Wide erect gumweed PerF RegEn

& Flora

var. discoideus (2006) aphanactis var. var. horridum var. var. var. G. hirsutula Flora of North America Vars. not recognized in Vars. of North America Synonyms/Taxonomic Synonyms/Taxonomic notes Ericameria pulchella, Lorandersonia baileyi, C. baileyi clementis Includes vars. integrifolium, Breea arvensis E. concinnus E. compositus E. divergens formosissimus Included in in (2006)

var. var. var. var. var. rmed) taxa of Capitol Reef National Park, cont. baileyi serrulata Chrysothamnus pulchellus var. Haplopappus clementis Pyrrocoma clementis Chrysothamnus vaseyi Cirsium arvense Cirsium clavatum clavatum Cirsium rydbergii Crepis acuminata Psilochenia acuminata Crepis intermedia Psilochenia intermedia Enceliopsis nutans Encelia nutans Erigeron aphanactis aphanactis Erigeron compositus glabratus Erigeron divergens divergens Erigeron formosissimus E. formosissimus Gnaphalium palustre Grindelia fastigiata Grindelia squarrosa FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconﬁ

108 Annotated Checklist of Vascular Flora: Capitol Reef National Park ed specimens ed or from specimens H. helenioides based on misidentiﬁ of collected outside the park. on collections from outside the park. Lake, UT (Stansbury s.n. GH) on collections from outside the park. UT (Thompson s.n. GH) on collections from outside the park. locality is Cottonwood Canyon, Type 676 NY) Salt Lake Co, UT (Watson on collections from outside the park. Range Comments Life form PerF Wide ower PerF Sparse Common name orange sneezeweed PerF Wide Reports from CARE Herbarium are Cooper’s hymenoxysCooper’s PerF Wide sneezeweed hymenoxys PerF RegEn Reports from CARE Herbarium based Colorado rubber-plant PerF Wide tidytips AnnF Wide Torrey’s desert-dandelionTorrey’s AnnF Wideprenanthella locality is shores of Great Salt Type piliferous turtlebackprairie coneﬂ AnnF AnnF RegEnwoolly groundsel Wide locality is near Kanab, Kane Co, Type Reports from CARE Herbarium based PerF Wide cut-leaved groundsel PerF Wide Reports from CARE Herbarium based saw-toothed groundsel PerF Wide Reports from CARE Herbarium based Rocky Mountain goldenrod baby goldenrod PerF Wide brown-plume wirelettuce PerF Wide

Flora May var. (2006). Helenium [Hymenoxys] Synonyms/Taxonomic Synonyms/Taxonomic notes Dugaldia hoopesii be of hybrid origin between Hymenoxys richardsonii and hoopesii. Picardenia richardsonii richardsonii Vars. not recognized in Vars. of North America

var. ora var. serra var. rmed) taxa of Capitol Reef National Park, cont. richardsonii Helenium hoopesii Hymenoxys hoopesii, Hymenoxys cooperi Actinella cooperi Hymenoxys helenioides Picradenia helenioides. Hymenoxys richardsonii var. Layia glandulosa Malacothrix torreyi Prenanthella exiguaPsathyrotes pilifera exigua Lygodesmia Ratibida columnifera Senecio canus Packera cana Senecio eremophilus kingii Senecio serra Solidago multiradiata scopulorum Solidago nana Stephanomeria pauciﬂ Appendix B. Potential (unconﬁ FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Appendices 109 . ed specimens of glabra var. var. near St. George, Washington Co, UT near St. George, Washington (Parry 19 NY) based on misidentiﬁ R. islandica Range Comments Life form ax AnnF Intro Native to Asia Common name desert nest-straw AnnF Periph dogweed PerF Wide slate thelypody PerF Wide USFWS: 3C beautiful sedge PerG Wide soft-leaved sedge PerG Wide showy townsendia PerF Wide golden crownbeard AnnF Wide tower-mustard PerF Wide little-pod false ﬂ mountain drabaSt. George pepperwort AnnF AnnF Wide RegEn locality is valley of the Virgin Type hairy-pod pepperwort AnnF Wide poor-man’s pepperwortpoor-man’s AnnF Wide Chambers’ twinpod PerF Wide arrowhead thelypody PerF Wide blunt-leaf yellowcress PerF WideMoab thelypody Reports from CARE Herbarium are PerF RegEn

var. Vars. not Vars. tenellum var. orifera Synonyms/Taxonomic Synonyms/Taxonomic notes Ximenesia encelioides recognized by Holmgren et al. (2005) L. lasiocarpum Thelypodium sagittatum sagittatum R. curvipes

var. var. var. var.

var. orumwrightii T. glabra glabra. Turritis

var. var. var. oriferfl T. rmed) taxa of Capitol Reef National Park, cont. sagittata Stylocline micropoides Thelypodium laxifl Thymophylla acerosa Dyssodia acerosa Townsendia fl Townsendia Carex bella Carex disperma Verbesina encelioides Verbesina exauriculata Arabis glabra Camelina microcarpa Draba rectifructa georginum lasiocarpum Lepidium lasiocarpum Lepidium lasiocarpum Lepidium virginicum pubescens Physaria chambersii chambersii Rorippa curvipes Thelypodiopsis sagittata var. curvipes Thelypodium integrifolium gracilipes Appendix B. Potential (unconfi FamilyCompositae (Asteraceae) Species name Cruciferae (Brassicaceae) Compositae (Asteraceae) Compositae (Asteraceae) Cyperaceae Cyperaceae Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

110 Annotated Checklist of Vascular Flora: Capitol Reef National Park on collections from outside the park. UT CDC are based on collections from outside the park in Glen Canyon NRA. locality is Cottonwood Canyon, Type Kane Co, UT (Barneby 14434 NY). BLM: Sensitive, USFWS: 3C Kaibab Plateau “on the southern edge of Utah”, Kane Co, UT (Jones s.n. POM) on collections from outside the park. on collections from outside the park. Range Comments Life form at-sedge AnnG Wide Common name blunt sedge PerG Wide liddon sedge PerG Wide Raynolds’ sedge PerG Wide beaked sedgeredroot fl tiny spurge PerG Wide Reports from CARE Herbarium based AnnF Wide thyme-leaved spurge AnnF Wide Utah spurge AnnF centauryBuckley’s RegEn Reports from CARE Herbarium and AnnF Periph Utah swertia PerF RegEn locality is Buckskin Mountains, Type ticklegrassmeadow foxtail PerG PerG Wide Intro Reports from CARE Herbarium based Native to Eurasia needle grama AnnG Periph tall wheatgrassmountain muhly PerG Intro PerG Native to Eurasia Wide mat muhly PerG Wide vine mesquite PerG Wide Reports from CARE Herbarium based interior bluegrass PerG Wide ponticus, interior, P. P. interior, var. liculmis ssp. M. fi Synonyms/Taxonomic Synonyms/Taxonomic notes Agropyron elongatum, Thinopyrum ponticum Includes interior glauca nemoralis P. var. rmed) taxa of Capitol Reef National Park, cont. Carex obtusata Carex petasata Carex raynoldsii Carex utriculataCyperus erythrorhizos Chamaesyce micromeria C. rostrata (sensu lato) Euphorbia micromeria Chamaesyce serpyllifolia Euphorbia serpyllifolia Euphorbia nephradenia Centaurium calycosum C. arizonicum Swertia utahensis Frasera paniculata Agrostis scabraAlopecurus pratensis A. hiemalis, geminata Bouteloua aristidoides Elymus elongatus E. elongatus Muhlenbergia montana Muhlenbergia richardsonis Panicum obtusum Poa glauca Appendix B. Potential (unconfi FamilyCyperaceae Species name Cyperaceae Cyperaceae Cyperaceae Cyperaceae Euphorbiaceae Euphorbiaceae Euphorbiaceae Gentianaceae Gentianaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

Appendices 111 eld Co, UT (Welsh eld Co, UT (Jones 5663 on collections from outside the park. Huntington along Hwy 10, Emery Co, UT (Atwood & Atwood 24852 BRY) Laccolite, Garfi NY) UT (Jones s.n. POM) Penellen Pass, Garfi USFS: Region 4 & Moore 11740 BRY). Sensitive on collections from outside the park. locality is Frisco, Beaver Co, UT Type (Jones 180 MINN) below Kachina Bridge, San Juan Co, UT (Barneby 12778 CAS). USFWS: 3C UT (Jones 5355 NY) Range Comments Life form AnnF LocEn locality is Henry Mtns, Marvine Type ddleleaf AnnF Wide ower dragonhead ower PerF Wide eld milkvetch PerF RegEn Reports from CARE Herbarium based littlefi Fort Wingate milkvetch PerF RegEn Common name leafy fi Cottam’s phaceliaCottam’s AnnF LocEn locality is 4.5 miles N of Type Henry Mountains phacelia butternut Tree Intro Native to E North America Drummond’s rushDrummond’s PerG Wide marsh arrowgrass PerG Wide smallfl Eastwood’s milkvetchEastwood’s PerF RegEn Grand Co, locality is Westwater, Type Dana’s milkvetchDana’s PerF LocEn locality is The Horn near Type canyon kentrophytacobweb milkvetch PerF RegEn PerF RegEn Reports from CARE Herbarium based birds-nest milkvetch PerFSevier milkvetch LocEn locality is White Canyon, 2 miles Type PerF RegEn locality is Marysvale, Piute Co, Type Ellis’ milkvetch PerF Wide

& diphysus stocksii ora var. var. var. drummondii orus Synonyms/Taxonomic Synonyms/Taxonomic notes Var. not recognized in Var. Cronquist et al. (1984) subtriﬂ Includes vars. Moldavica parviﬂ A. argophyllus A. lentiginosus

var.

var. var.

var. rmed) taxa of Capitol Reef National Park, cont. orum orum orus coloradoensis araneosus laxifl Astragalus wingatanus Nama densum parvifl Phacelia cottamii Phacelia crenulata orbicularis Juglans cinerea Juncus drummondii Triglochin palustris Triglochin Dracocephalum parvifl Astragalus eastwoodiae Astragalus henrimontanensis Astragalus kentrophyta var. Astragalus lentiginosus var. Astragalus nidularius Astragalus piutensisAstragalus praelongus A. marianus ellisiae Astragalus preussii Leguminosae (Fabaceae) Appendix B. Potential (unconfi FamilyHydrophyllaceae Species name Hydrophyllaceae Hydrophyllaceae Juglandaceae Juncaceae Juncaginaceae Labiatae (Lamiaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

112 Annotated Checklist of Vascular Flora: Capitol Reef National Park 541 GH) eld Co, UT (Ward on collections from outside the park. locality is Courthouse Wash, Type Grand Co, UT (Eastwood s.n. CAS) on collections from outside the park. locality is W of Carmel, Kane Co, Type UT (Ferguson & Ottley 5613 NY) 14632 & Taylor Grand Co, UT (Welsh BRY) on collections from outside the park. Wasatch locality is Heber Valley, Type 234 GH) Co, UT (Watson Co, UT (Watson 234 US) Co, UT (Watson on collections from outside the park. locality is Kane Co, UT (Siler s.n. Type GH) Garﬁ Juan Co, UT (Eastwood s.n.) on collections from outside the park. ca 17 miles locality is Salt Wash, Type Co, UT WNW of Hanksville, Wayne UT BLM: Sensitive, 13348 BRY). (Welsh USFWS: former C2 candidate Range Comments Life form ax PerF RegEn locality is Willow Creek, San Type Common name creeping rush-pea PerF RegEn Reports from CARE Herbarium based Utah trefoilMoab lupine PerF RegEn Reports from CARE Herbarium based PerF RegEn Ranch, locality is Professor Valley Type King’s lupineKing’s AnnF Wide Reports from CARE Herbarium based stemless lupine PerF Wide Park, Summit locality is Parley’s Type white locoweed PerF Wide sink lupinemountain locoweed PerF PerF RegEndwarf clover Wide Reports from CARE Herbarium based locality is Aquarius Plateau, Type PerF Wide alsike clover PerF Intro Native to Europe Utah yellow-ﬂ slenderleaf globemallow PerF Wide scurfpea globemallow PerF LocEn Reports from CARE Herbarium based

var. utahensis caespitosus, L. Vars. not Vars. var. var. Synonyms/Taxonomic Synonyms/Taxonomic notes L. lepidus caespitosus L. barbiger. L. barbiger. recognized in Barneby (1989) gymnocarpon

var.

var. kingii L. kingii var. var. var. var. rmed) taxa of Capitol Reef National Park, cont. Caesalpinia repens repens Hoffmanseggia Lotus utahensis Lupinus argenteus moabensis Lupinus kingii Lupinus lepidus utahensis Oxytropis sericea sericea Lupinus sericeus barbiger gymnocarponTrifolium gymnocarpon T. Oxytropis oreophila oreophila Trifolium hybridum Trifolium Linum subteres Sphaeralcea leptophylla Sphaeralcea psoraloides Appendix B. Potential (unconﬁ FamilyLeguminosae (Fabaceae, Caesalpiniaceae) Species name Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Linaceae Malvaceae Malvaceae

Appendices 113 on collections from outside the park. on collections from outside the park. on collections from outside the park. locality is ca 2 miles S of US 506 Type Grand Co, UT on road to Westwater, (Reveal & Davidse 949 UTC) Co, UT (Ward 526 US) Co, UT (Ward Green River, Emery Co, UT (Ownbey & Green River, Ownbey 2146 ) (Jones s.n. POM) Range Comments Life form AnnF RegEn Reports from CARE Herbarium based PerF Wide PerF RegEn locality is Price, Carbon Co, UT Type AnnF Wide y wild buckwheat Shrub RegEn Common name butterﬂ smooth wild buckwheat AnnF wild Westwater Widebuckwheat Reports from CARE Herbarium based Wooton’s sand-verbenaWooton’s AnnF RegEn Reports from CARE Herbarium based desert olive Shrub Wide autumn willow-herb AnnF Wide Rocky Mountain willow- herb raceme groundsmoke AnnF Wide Jones’ evening-primrose PerF Wide Wayne locality is Rabbit Valley, Type yellow evening-primrose PerF Wide little grapefern Fern Sparse San Rafael prickly-poppy PerF RegEn locality is 37.9 miles SW of Type small collomia AnnF Wide scarlet gilia PerF Wide scarlet gilia PerF Wide Bateman’s wild Bateman’s buckwheat spreading wild buckwheat

& ssp.

var. var. compositum ava macrosiphon Synonyms/Taxonomic Synonyms/Taxonomic notes T. wootonii T. paniculatum Includes vars. tenebrosum I. tenuituba, Gilia aggregata var. Gilia aggregata maculata, I. aggregata formosissima E. batemanii Misspelled in Welsh et al. Misspelled in Welsh (2003)

var. var. var. avafl O. var. var. var. fl var. ava rmed) taxa of Capitol Reef National Park, cont. papiliunculi Tripterocalyx carneus Tripterocalyx wootonii Eriogonum salsuginosum Stenogonum salsuginosum Eriogonum scabrellum Forestiera pubescens neomexicana F. Epilobium brachycarpum E. paniculatum Epilobium saximontanum Gayophytum racemosum Oenothera caespitosa crinita Oenothera fl Botrychium simplex Argemone corymbosa arenicola Collomia linearis Ipomopsis aggregata macrosiphon Ipomopsis aggregata maculata Eriogonum batemanii batemanii Eriogonum divaricatum Eriogonum leptocladon var. Appendix B. Potential (unconfi FamilyNyctaginaceae Species name Polygonaceae Polygonaceae Oleaceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Ophioglossaceae Papaveraceae Polemoniaceae Polemoniaceae Polemoniaceae Polygonaceae Polygonaceae Polygonaceae

114 Annotated Checklist of Vascular Flora: Capitol Reef National Park rmed as present in the park. on collections from outside the park. on collections from outside the park. lacks location data and cannot be conﬁ Native to Eurasia Marysvale, Piute Co, UT (Rydberg & Carlton 7261 NY) Summit Co, UT (Watson 329 GH) Summit Co, UT (Watson Range Comments Life form Common name yellow paintbrush PerF Wide Reports from CARE Herbarium based cream wild buckwheat PerF Wide blue-eyed mary AnnF Wide sulfur wild buckwheat PerF Wide Douglas’ knotweed AnnF Wide Reports from CARE Herbarium based lady’s-thumb AnnF Intro Report from CARE Herbarium bushy knotweed AnnF Wide spiny cliff-brake Fern Wide Rocky Mountain woodsia Fern Wide prairie smoke PerF Wide modest cinquefoil PerF RegEn locality is Mt, Barrette near Type sheep cinquefoil PerF Wide locality is Uinta Mountains, Type milky kelloggia PerF Wide fringecup woodlandstar PerF Wide slender woodlandstar PerF Wide

& from

scopulina var. ora var. ssp. includes ava aureum in Holmgren et Combined with fl var. cum, P. ramosissimum cum, P. dichrocephalum ramosissimum ava Synonyms/Taxonomic Synonyms/Taxonomic notes E. umbellatum subalpinum, E. subalpinum; vars. desereticum Includes var. Includes var. Flora of North America (2005) P. douglasii P. prolifi var. Erythrocoma trifl P. concinna. P. rubricaulis P. al. (1997) L. bulbifera, glabra

var.

var. var. ora rmed) taxa of Capitol Reef National Park, cont. avafl C. orum majus umbellatum ramulosum Eriogonum umbellatum var. Collinsia parvifl Eriogonum umbellatum var. Polygonum douglasii douglasii Polygonum persicaria Persicaria maculosa Polygonum ramosissimum ramosissimum P. Pellaea truncata Woodsia scopulinaWoodsia scopulina W. Geum trifl ciliatum Potentilla concinna modesta Potentilla ovina decurrens Kelloggia galioides Lithophragma glabrum var. Lithophragma tenellum L. tenella Castilleja fl Appendix B. Potential (unconfi FamilyPolygonaceae Species name Scrophulariaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polypodiaceae (Adiantaceae, Sinopteridaceae) Polypodiaceae (Aspleniaceae, Woodsiaceae) Rosaceae Rosaceae Rosaceae Rubiaceae Saxifragaceae Saxifragaceae Scrophulariaceae

Appendices 115 Atlas of the on collections from outside the park. Summit Co, UT (Watson 458 GH) Summit Co, UT (Watson Summit Co, UT (Watson 454 GH) Summit Co, UT (Watson on collections from outside the park. ed here. Range Comments Life form ane Atwood of Brigham Young University in 2001, and the ane Atwood of Brigham Young eld, Emery, and Sevier counties, Utah, but have not previously been reported been reported and Sevier counties, Utah, but have not previously eld, Emery, Common name bird’s-beakKing’s AnnF RegEn butter-and-eggs PerF Intro Native to Eurasia Palmer’s penstemonPalmer’s PerF Wide Watson’s penstemonWatson’s purslane speedwell PerF Wide AnnF Reports from CARE Herbarium based Wide pale wolfberry Shrub Wide Torrey’s wolfberryTorrey’s Shrub Wide Fendler’s ground-cherryFendler’s PerF Wide pinnate angelica PerF Wide locality is Uinta Mountains, Type cutleaf water-parsnip PerF Wide sticky spring-parsley PerF RegEn cow parsnip PerF Wide fernleaf ligusticumPennsylvania pellitory PerF AnnFtobacco-root Wide Wide locality is Uinta Mountains, Type Reports from CARE Herbarium based PerF Wide western valerian PerF Wide

var. montanum fendleri lanatum, H. sphondylium Synonyms/Taxonomic Synonyms/Taxonomic notes P. palmeri P. cordifolia, P. hederaefolia cordifolia, P. var. P. fendleri, P. hederifolia fendleri, P. P. var. ssp.

var. var. var. var. incisa var. licinum rmed) taxa of Capitol Reef National Park, cont. Cordylanthus kingii kingii Linaria vulgaris Penstemon palmeri palmeri Penstemon watsonii peregrina Veronica xalapensis Lycium pallidum Lycium Lycium torreyi Lycium Physalis hederifolia fendleri Angelica pinnata Berula erecta Cymopterus newberryi Heracleum lanatum H. maximum, sphondylium Ligusticum fi Parietaria pensylvanica edulis Valeriana Valeriana occidentalis Valeriana (Albee et al. 1988). Some species from Appendix C could be considered as potentially occurring in the park, but are not includ as potentially occurring in the park, but are Appendix C could be considered (Albee et al. 1988). Some species from rmed for the park. Species on this list are derived from a checklist of potential species for NCPN parks prepared by Dr. N. Du by Dr. a checklist of potential species for NCPN parks prepared derived from rmed for the park. Species on this list are Appendix B. Potential (unconfi FamilyScrophulariaceae Species name Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Solanaceae Solanaceae Solanaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Urticaceae Valerianaceae Valerianaceae Note: The vascular plant species listed in this table are known from the vicinity of Capitol Reef National Park in Wayne, Garfi the vicinity of Capitol Reef National Park in Wayne, known from Note: The vascular plant species listed in this table are or confi Vascular Flora of Utah Vascular

116 Annotated Checklist of Vascular Flora: Capitol Reef National Park .

var. var. var. . C. is Salt . var. var. . . Also ora exus and utahensis A. retrofl C. crassisepala A. truncata C. graveolens C. crassisepala A. acanthicarpa. C. confertifl C. abata C. fontanum drummondii Lychnis A. canescens A. prostrata Type locality is St. George, Type . Native to Eurasia. . Type locality is Courthouse Wash, locality is Courthouse Wash, . Type Type locality of var. locality of var. Type ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of elachantha. avoculata misidentifi misidentifi misidentifi misidentifi Comments var. var. Co, UT (Parry 171 GH). Washington misidentifi Reported by Northern Colorado Plateau Network (in ed.). Native to Eurasia. neomexicanum misidentifi elachantha. misidentifi Type locality is Huntington Road, ca 15 mi W Type of US 506, Emery Co, UT (Higgins 1310 BRY). USFWS: 3C. misidentifi fl Grand Co, UT (Eastwood s.n. CAS). misidentifi misidentifi misidentifi canescens. Lake City, Salt Lake Co, UT (Jones 1760 POM). Lake City, misidentifi Park status Range Life form PerF LocEn FalsRep Reports from CARE herbarium are based on Powell’s amaranthPowell’s AnnFbearded cryptanth Sparse AnnF FalsRep Periph Reports from CARE Herbarium based on FalsRep Reports from CARE herbarium are based on tumbling orach AnnF Intro FalsRep Reports from CARE herbarium are based on Jerusalem-oak AnnF Intro FalsRep Reports from CARE herbarium are based on Common name beguiling cryptanth AnnF Periph FalsRep Reports from CARE herbarium are based on common ragweed AnnF Wide FalsRep Reports from CARE Herbarium based on Johnston’s Johnston’s cryptanth slender cryptanth PerF RegEn FalsRep Reports from CARE herbarium are based on nodding chickweed AnnFshy campion Sparse FalsRep Reports from CARE herbarium are based on PerF Periph FalsRep Reports from CARE Herbarium are based on basin saltbush Shrub Wide FalsRep Reports from CARE Herbarium are based on spear orach AnnF Sparse FalsRep Reports from CARE herbarium are based on

Flora var. elatior Synonyms/ notes Taxonomic Teloxys botrys, Teloxys Dysphania botrys A. artemisiifolia var. Vars. not Vars. recognized in of North America (2005) A. tridentata robusta andersonii utahensis barbigera Amaranthus powellii Atriplex rosea Cryptantha decipiens Chenopodium botrys Ambrosia artemisiifolia Cryptantha johnstonii Cryptantha tenuis Cerastium nutans Atriplex gardneri var. Silene verecunda ssp. Atriplex patula Boraginaceae Cryptantha FamilyAmaranthaceae Species name Chenopodiaceae Boraginaceae Chenopodiaceae Compositae (Asteraceae) Boraginaceae Boraginaceae Caryophyllaceae (Alsinaceae) Chenopodiaceae Caryophyllaceae Chenopodiaceae Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park. Appendix C. Rejected (falsely reported

Appendices 117

or Type Type var. ed var. var. H. nuttalllii. B. salicina. C. stevioides. C. nauseosus E. pumilus E. caespitosus G. marginata. G. wrightii. ed specimen of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of misidentifi specimen of misidentifi Comments misidentifi Co, UT locality is near St. George, Washington (Palmer 6 GH). misidentifi misidentifi misidentifi CARE Herbarium in 2005. junceus. Network (in ed.), but Capitol Reef NP is well outside the expected range of this species. Native to Europe. on specimens collected outside the park. Type Co, Wayne locality is 26.5 miles S of Antimony, 9388 BRY). & Welsh UT (Welsh misidentifi concinnus. Park status Range Life form AnnF Periph FalsRep Reports from CARE Herbarium are based on Shrub RegEn FalsRep Not known from Utah. No specimens found in ower PerF Wide FalsRep Reports from CARE herbarium are based on eabane AnnF Intro Ques? Reported by Northern Colorado Plateau little sunfl setose crustweed AnnF Periph FalsRep Reports from CARE herbarium based on sticky gumweed AnnF Sparse FalsRep Report from CARE based on a misidentifi Common name sticky seepwillow Shrub Wide FalsRep Reports from the CARE Herbarium are based on Fremont’s dusty- Fremont’s maiden Awapa daisyAwapa PerF LocEn FalsRep Reports from CARE Herbarium are based on shiny rabbitbrush Shrub RegEn FalsRep Reports from CARE herbarium based on annual fl daisyEngelmann’s PerF Wide FalsRep Reports from CARE herbarium are based on in

B. E. in var. var. (2006) by some included turbinatus Flora of North B. salicifolia C. nauseosus Included in salicina authors. Included in Synonyms/ notes Taxonomic in America G. viscosum, Pseudognaphalium viscosum, P. macounii Ericameria nauseosa latisquamea Cronquist (1994) Ericameria nauseosa abajoensis nitida; Included in in var. Cronquist (1994)

ssp. var. ora Helianthella uniﬂ Baccharis glutinosa Glyptopleura setulosa Gnaphalium macounii Chaenactis fremontii latisquameus Chrysothamnus nauseosus nitidus Chrysothamnus nauseosus Erigeron annuus Erigeron awapensis Erigeron engelmannii Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park, cont.

118 Annotated Checklist of Vascular Flora: Capitol Reef National Park . . ed ed

. var. var. var. ora. Convolvulus T. aprica. T. S. velutina. Sonchus asper Prenanthella exigua M. canescens S. douglasii Vernonia marginata Vernonia sparsifl P. arizonica. var. H. acaulis T. spinosa. T. ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of . misidentifi on misidentifi arvensis misidentifi Comments specimens of misidentifi misidentifi misidentifi misidentifi specimens of longilobus. canescens. misidentifi misidentifi Park status Range PerF PeriphAnnF FalsRep Reports from CARE Herbarium are based on Intro FalsRep Reports from CARE Herbarium are based Life form Shrub Periph FalsRep Reports from CARE herbarium are based on ower Wyoming townsendia ivyleaf morning- glory stemless hymenoxys PerF RegEn FalsRep Reports from CARE are based on misidentifi Missouri goldenrod PerF Wide FalsRep Reports from CARE herbarium are based on Common name Canada lettucerush skeleton-plant PerF PerF Periph Periph FalsRep FalsRep Reports from CARE Herbarium are based on Reports from CARE Herbarium are based on bracteate aster PerF RegEn FalsRep Reports from CARE Herbarium based on two-leaf groundsel PerF RegEn FalsRep Reports from CARE Herbarium are based on Nuttall’s horsebrushNuttall’s Shrub RegEn FalsRep Reports from CARE herbarium are misidentifi arrowweedwhitestem paperfl Shrub Wide FalsRep Reports from CARE herbarium are based on

var. var.

Vars. Vars. var. M. var. by some (2006). according

epunctata Flora of North Tetraneuris acaulis Tetraneuris var. not recognized in America missouriensis. T. alpigena T. alpigena Ipomoea hederacea Synonyms/ notes Taxonomic authors Packera dimorphophylla var. dimorphophylla; UT material is var. paysonii to Cronquist (1994) Included in canescens canescens S. missouriensis

var. var. var.

Hymenoxys acaulis epunctata Townsendia Townsendia montana montana Ipomoea hirsutula Lactuca canadensis Lygodesmia juncea Tetradymia Tetradymia nuttallii dimorphophyllus Machaeranthera canescens monticola Senecio dimorphophyllus var. Solidago missouriensis Pluchea sericea sericea Tessaria Psilostrophe cooperi Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park, cont. FamilyCompositae (Asteraceae) Species name Compositae (Asteraceae) Convolvulaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Appendices 119 A. . . var. var. . ora. var. var. and . Also reported Fendlerella S. acutus Q. welshii. G. spinescens D. pinnata D. aurea A. stolonifera C. praegracilis E. quinquefl viridis. A. demissa var. var. C. stenophylla. E. viridis ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of of ed ed specimens of ed specimens of ed ed specimens of . Also reported by Northern Colorado misidentifi misidentifi on misidentifi selbyi. misidentifi misidentifi utahensis Plateau Network ( in ed.). microphyllum. misidentifi on misidentifi on misidentifi Type locality is Butler Wash, San Juan Co, UT locality is Butler Wash, Type (Eastwood 141 NY) misidentifi Native to Eurasia osmiarum. misidentifi Also reported by Northern Colorado Plateau Network (in ed.) misidentifi Comments misidentifi by Northern Colorado Plateau Network (in ed.) Network (in ed.), but Capitol Reef NP is well outside the expected range of this species. Park status Range AnnF Wide FalsRep Reports from CARE Herbarium are based on Life form spiny greasebush Shrub RegEn FalsRep Reports from CARE herbarium based on small-leaf rockcress PerF Wide FalsRep Reports from CARE herbarium based on Nevada greasebush Shrub Periph FalsRep Reports from CARE Herbarium are based western tansy- mustard sedgeHood’s PerG Wide FalsRep Reports from CARE Herbarium based on lanceleaf draba PerF Wide FalsRep Reports from CARE Herbarium are based on soft-stem bulrush PerG Wide FalsRep Reports from the CARE Herbarium are based Eastwood’s oakEastwood’s Shrub RegEn FalsRep Reports from CARE Herbarium are based colonial bentgrass PerG Intro FalsRep Reports from CARE Herbarium based on dryland sedge PerG Wide FalsRep Reports from CARE Herbarium are based on Nevada ephedra Shrub Wide FalsRep Reports from CARE herbarium are based on Common name beaked spikerush PerG Wide FalsRep Reports from CARE Herbarium are based on Mexican manzanita Shrub Wide Ques? Reported by Northern Colorado Plateau

Q. lipes ﬁ var. validus cana x var. ssp. var. D. incisa Forsellesia meionandra Boechera microphylla microphylla Forsellesia nevadensis breweri tabernaemontani, Schoenoplectus lacustris welshii Synonyms/ notes Taxonomic Hybrid between Q. gambelii

var.

var. var. var. x lipes Descurainia pinnata ﬁ Carex hoodii meionandrum Glossopetalon spinescens Arabis microphylla microphylla Agrostis capillaris A. tenuis Glossopetalon spinescens aridum Draba cana D. lanceolata, Carex tahoensis C. xerantica Scirpus validus Schoenoplectus Ephedra nevadensis Eleocharis rostellata eastwoodiae Arctostaphylos pungens Quercus Cruciferae (Brassicaceae) Cyperaceae Crossosomataceae Cruciferae (Brassicaceae) Gramineae (Poaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park, cont. FamilyCrossosomataceae Species name Cruciferae (Brassicaceae) Cyperaceae Ephedraceae Cyperaceae Ericaceae Fagaceae Cyperaceae

120 Annotated Checklist of Vascular Flora: Capitol Reef National Park var. var...... eldii eldii . . Not Festuca Calamagrostis A. aequalis Festuca sororia B. springfi Bromus inermis B. springfi B. gracilis E. trachycaulus B. gracilis B. japonicus ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of . Native to Eurasia. . misidentifi misidentifi stricta misidentifi Comments misidentifi misidentifi inermis misidentifi misidentifi Native to Eurasia. misidentifi misidentifi misidentifi on misidentifi arundinacea. included for UT in FNA 2003. Park status Range Life form PerG Intro FalsRep Reports from CARE Herbarium are based on reed canary grass PerG Wide FalsRep Reports from CARE Herbarium are based on Carolina foxtail AnnG Periph FalsRep Reports from CARE Herbarium based on Wasatch bluegrassWasatch PerG Wide FalsRep Reports from CARE herbarium are based on Common name cane bluestem PerG Wide FalsRep Reports from the CARE Herbarium are based on meadow fescue PerG Intro FalsRep Reports from CARE Herbarium based on silver bluestem PerG Periph FalsRep Reports from the CARE Herbarium are based on hairy grama PerG Periph FalsRep Reports from CARE Herbarium based on intermediate wheatgrass red grama PerG Periph FalsRep Reports from CARE Herbarium based on mountain brome PerG Wide FalsRep Reports from CARE Herbarium are based on blue wildrye PerG Wide FalsRep Reports from CARE Herbarium are based da

triﬁ var. var. da hirsuta hispidus, Phalaroides arundinacea Synonyms/ notes Taxonomic Schedonorus pratensis hirsutum, B. hirsuta ssp. var. Agropyron intermedium, Elytrigia intermedia, Thinopyrum intermedium B. polyanthus, Ceratochloa carinata glaucus

datriﬁ B. ssp. Alopecurus carolinianus Phalaris arundinacea Poa arnowiae Poa curta Bothriochloa barbinodis Bothriochloa laguroides torreyana Festuca pratensis Lolium pratense, Bouteloua hirsuta Chondrosum Bouteloua triﬁ Elymus hispidus E. hispidus Bromus carinatus B. marginatus, Elymus glaucus E. glaucus Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park, cont. FamilyGramineae (Poaceae) Species name Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

Appendices 121 . ed . . . and . var. var. var. var. S. pinetorum crenulata P. Glyceria striata N. retrorsum J. ensifolius Mentha spicata . A. consobrinus Diospyros virginiana . Also reported by parishii. S. demissum A. laccoliticus var. var. Type locality is Nipple Bench, Kane Type ed specimens of ed specimens of ed specimens of ed specimens of ed ed specimen of ed specimens of ed specimens of ed specimens of misidentifi misidentifi misidentifi S. coronata Northern Colorado Plateau Network (in ed.). Network (in ed.), but Capitol Reef NP is outside the known or expected range of this species. misidentifi corrugata. USFWS: 3C & Atwood 9809 BRY). Co, UT (Welsh Colorado Plateau Network (in ed.) are based on misidentifi misidentifi Native to E North America misidentifi Comments Colorado Plateau Network (in ed.) are based on misidentifi montanus. misidentifi specimens of Park status Range AnnF LocEn FalsRep Reports from CARE Herbarium are based on PerG Wide FalsRep Reports from CARE Herbarium are based on PerF Wide FalsRep Reports from CARE Herbarium and Northern Life form PerF Sparse FalsRep Reports from CARE Herbarium are based on Nipple Bench phacelia Nelson’s Nelson’s needlegrass weak alkaligrass PerG Wide FalsRep Reports from CARE Herbarium are based on purple-mathairy nama AnnF Wide AnnF Ques? Periph Reported by Northern Colorado Plateau Nevada blue-eyed FalsRepgrass Reports from CARE Herbarium are based on pecan Tree Intro FalsRep Reports from CARE Herbarium based on Nevada rush PerG Wide FalsRep Reports from CARE Herbarium are based on Common name chestnut milkvetch PerG Periph FalsRep Reports from CARE herbarium and Northern cicada milkvetch PerF RegEn FalsRep Reports from CARE are based on misidentifi American germander

var.

var. ora var. & ora, T. T. ora, ora holmii, chamaeleuce Cronquist et al. (1984) Achnatherum nelsonii nelsonii, S. columbiana crenulata nelsonii Included in Includes vars. badius var. P. paucifl P. Torreyochloa pallida paucifl paucifl Synonyms/ notes Taxonomic A. chamaeleuce var. nevadensis

var. var. ora nelsonii Stipa nelsonii Nama demissum Nama hispidum Phacelia mammillarensis Sisyrinchium halophilum Carya illinoensis Juncus nevadensis Puccinellia pauciﬂ Astragalus castaneiformis Astragalus chamaeleuce Teucrium Teucrium canadense occidentale Gramineae (Poaceae) Hydrophyllaceae Hydrophyllaceae Hydrophyllaceae Iridaceae Juglandaceae Juncaceae Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park, cont. FamilyGramineae (Poaceae) Species name Leguminosae (Fabaceae) Leguminosae (Fabaceae) Labiatae (Lamiaceae)

122 Annotated Checklist of Vascular Flora: Capitol Reef National Park . ed ed ed ed by the basalticus A. musiniensis A. tenellus. Type locality is 10 Type A. macropetalum Type locality is Salina Type Type locality is City Type zionis. avescens. zionis. fl ed specimen of boreale. intermontanus. var. var. var. var. ed specimens of attributed to var. attributed to var. var. ed specimens of var. var. sileranus Type locality is W of Loa Pass, 0.8 mi E Piute Type & Moore Isely, Co, UT (Welsh, Co line, Wayne 6473 BRY). on a misidentifi Center is based on a collection from Grand Staircase-Escalante National Monument. Type (near Alton), Kane Co, UT locality is Sink Valley (Jones s.n. POM). var. var. locality is 10 Type UT Conservation Data Center. mi S of Fremont Junction, Sevier Co, UT (Welsh, USFWS: former C2 & Moore 6447 BRY). Isely, candidate. Colorado Plateau Network (in ed.) based on misidentifi Canyon, Sevier Co, UT (Jones 5441l POM) L. pusillus on misidentifi H. boreale L. brachycalyx Also reported by Northern Colorado Plateau Network (in ed.). Comments mi E of Halls Crossing, San Juan Co, UT (Welsh mi E of Halls Crossing, San Juan Co, UT (Welsh UT BLM: Sensitive, USFWS: former C2. 5205 BRY). Reports from CARE Herbarium and UT Conservation Data Center are based on specimens of var. Reports from CARE Herbarium are misidentifi L. brachycalyx Creek Canyon, Salt Lake Co, UT (Leonard 101 NY). Park status Range PerF LocEn FalsRep Life form PerF RegEn FalsRep Reports from the CARE Herbarium are based Hole-in-the-Rock prairie-clover Welsh’s milkvetchWelsh’s PerF LocEn FalsRep Report by UT Conservation Data Center based Siler’s milkvetchSiler’s PerF RegEn FalsRep Report for CARE from UT Conservation Data basalt milkvetch PerF LocEn FalsRep Report from CARE based on BYU specimen of textile onion PerF Wide FalsRep Reports from CARE Herbarium and Northern shortstem lupine AnnF Wide FalsRep Reports from CARE herbarium are misidentifi western sweetvetch PerF Periph sweetpeaRydberg’s PerF FalsRep Reports from CARE Herbarium are misidentifi RegEnUtah sweetpea FalsRep PerF RegEn FalsRep Reports from CARE Herbarium are misidentifi Garrett’s weedy Garrett’s milkvetch Common name A. Var. not Var. by some D. epica. recognized by Barneby (1989) Included in loanus authors Vars. not Vars. recognized in Barneby (1989) Vars. not Vars. recognized in Barneby (1989) Synonyms/ notes Taxonomic

var. var. var. var. var. avescens orus epica tenuifolius Dalea ﬂ var. Astragalus welshii Astragalus subcinereus sileranus Allium textile Lupinus brevicaulis brachycalyx Lathyrus brachycalyx utahensis Astragalus miser var. Hedysarum occidentale occidentale Lathyrus pauciﬂ Astragalus subcinereus basalticus Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae (Alliaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park, cont. FamilyLeguminosae (Fabaceae) Species name Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

Appendices 123

. ed . var. . Type . Type ana . Type . Type . . orum. (stonewort G. imperialis M. marginata S. romanzoffi E. sparsifl S. parvifolia S. grossulariifolia. Chara C. eastwoodiae C. walkeri G. ramosissimum O. caespitosa O. albicaulis. ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of diluvialis. misidentifi misidentifi misidentifi var. misidentifi locality is valley of the Virgin River near St. Co, UT (Parry 188 GH). George, Washington Comments locality is Rockville, Washington Co, UT (Jones locality is Rockville, Washington 6082c) misidentifi specimens of misidentifi misidentifi Type locality is E side of Last Chance Bay, Lake locality is E side of Last Chance Bay, Type & Atwood 11597 Powell, Kane Co, UT (Welsh BRY) algae). misidentifi misidentifi misidentifi misidentifi marginata. Park status Range Life form PerF RegEn FalsRep Reports from CARE are based on misidentifi PerF Wide FalsRep Reports from CARE Herbarium are based on PerF Wide FalsRep Reports from CARE Herbarium are based on PerF LocEn FalsRep Reports from CARE Herbarium are based on AnnF Periph FalsRep Reports from CARE Herbarium are based on PerF Periph FalsRep Reports from CARE Herbarium are based on spiny gilia AnnF RegEn FalsRep Reports from CARE Herbarium are based on Virgin stickleaf PerF Wide FalsRep Reports from CARE herbarium based on primrose creamy ladies’- tresses spreading eriastrumspreading AnnF Wide FalsRep Reports from CARE herbarium is based on Common name Mohave globemallow Moore’s Moore’s globemallow southern waternymph showy camissonia AnnF Periph FalsRep Reports from CARE Herbarium are based on plains evening- primrose clubpod camissonia AnnF RegEnlow groundsmoke FalsRep AnnF Reports from CARE Herbarium are based on Periph FalsRep Reports from CARE Herbarium are based on pale evening-

var. ana var. ora guadalupensis porrifolia moorei S. romanzofﬁ var. latifolia, Aliciella latifolia Synonyms/ notes Taxonomic integra S. grossulariifolia var. N. guadalupensis ssp.

var.

var. var. var. Mentzelia integra M. multiﬂ Spiranthes porrifolia Eriastrum diffusum Gilia latifolia G. latifolia ambigua Sphaeralcea ambigua Sphaeralcea moorei Najas guadalupensis Camissonia brevipes Oenothera caespitosa macroglottis Camissonia claviformis claviformis Gayophytum humile Oenothera pallida trichocalyx Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park, cont. FamilyLoasaceae Species name Orchidaceae Polemoniaceae Polemoniaceae Malvaceae Malvaceae Najadaceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae

124 Annotated Checklist of Vascular Flora: Capitol Reef National Park

. var. var. var. var. var. var. eld Co, UT eld Co, UT Leptodactylon E. palmerianum P. austromontana P. Leptodactylon E. microthecum E. corymbosum E. microthecum Type locality is along the Type . Also reported by Northern ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of USFWS: 3C. . Also reported by Northern Colorado (newly described in 2005). Type locality is (newly described in 2005). Type austromontana. misidentifi misidentifi Network, but Capitol Reef NP is outside the expected range of this species. pungens Plateau Network (in ed.). misidentifi misidentifi misidentifi simpsonii. Colorado Plateau Network (in ed.) based on misidentifi near milepost 26 S of Antimony, Garfi near milepost 26 S of Antimony, USFWS: 3C. 9389 BRY). & Welsh (Welsh Network (in ed.), but Capitol Reef NP is well outside the expected range of this species. misidentifi simpsonii. var. var. margin of Cataract Canyon, Garfi & Chatterley 21972 BRY). Welsh, (Welsh, caespitosum Colorado Plateau Network (in ed.). heilii Comments Park status Range AnnF Periph FalsRep Reports from CARE Herbarium are based on Shrub RegEn FalsRepShrub Reports from CARE Herbarium are based on LocEn FalsRep Reports from CARE herbarium and Northern Life form AnnF Wide Ques? Reported by Northern Colorado Plateau Shrub Wide FalsRep Reports from CARE Herbarium are based on Nuttall’s linanthusNuttall’s PerF Wide FalsRep Reports from CARE Herbarium are based on nodding wild buckwheat ball-head gilia PerF Wide Ques? Reported by Northern Colorado Plateau yellowish phlox PerF LocEn FalsRep Reports from CARE Herbarium are based on moss phlox PerF Wide FalsRep Reports from CARE Herbarium are based on Comb Wash wild Comb Wash buckwheat wild Reveal’s buckwheat spotted wild buckwheat slender wild buckwheat Common name

var. Flora

ssp. var. ssp. lutescens Linanthastrum nuttallii nuttallii, Leptosiphon nuttallii, Linanthus nuttallii nuttallii var. Vars. not Vars. recognized in of North America (2005) Gilia congesta congesta muscoides, P. muscoides, P. bryoides Synonyms/ notes Taxonomic

var. var.

var. var. var. orum Linanthastrum nuttallii Phlox lutescens austromontana P. viminale Eriogonum corymbosum revealianum Eriogonum clavellatum Eriogonum maculatum Eriogonum microthecum laxiﬂ Ipomopsis congesta congesta Phlox muscoides hoodii P. Eriogonum cernuum Polemoniaceae Polemoniaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae FamilyPolemoniaceae Species name Polemoniaceae Polygonaceae Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park, cont.

Appendices 125

var. var. var. var. . Native . Native var. var. var. var. var. orum Philadelphus liformis liformis P. gracilis P. G. multifl E. palmerianum douglasii P. P. fi P. fi M. sylvestris P. domestica P. S. eriocephala F. virginiana. F. ed specimens of ed specimens of . . Also reported by Northern . ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of orum. misidentifi misidentifi misidentifi Comments on misidentifi multifl misidentifi microphyllus misidentifi misidentifi Type locality is Virgin River Valley, Washington Washington locality is Virgin River Valley, Type Co, UT (Parry 237 ISC). douglasii. misidentifi ligulifolia. occidentalis pulcherrima Colorado Plateau Network (in ed.). misidentifi occidentalis. Network (in ed.), but Capitol Reef NP is outside the known range of this species. Report probably based on to C North America. on misidentifi to C North America. Park status Range Life form AnnF RegEn FalsRep Reports from CARE Herbarium are based on PerF Wide Ques? Reported by Northern Colorado Plateau PerF Wide FalsRep Reports from CARE Herbarium are based small bedstraw PerF Wide FalsRep Reports from CARE Herbarium are based on kidney-shaped wild buckwheat Kellogg’s knotweedKellogg’s AnnF Wide FalsRep Reports from CARE Herbarium are based on Common name Fendlerbush Shrub Wide FalsRep Reports from CARE Herbarium are based Geyer’s willowGeyer’s Shrub Wide FalsRep Reports from CARE Herbarium based on dwarf pondweed PerF Wide FalsRep Reports from CARE Herbarium are based on Iowa crabapple Tree Intro FalsRep Reports from CARE Herbarium are based on leafy pondweed PerF Wide FalsRep Reports from CARE Herbarium are based on American plum Shrub Intro FalsRep Reports from CARE Herbarium based on woodland strawberry wedgeleaf cinquefoil

var.

var. var. var. & orum kelloggii, dum Synonyms/ notes Taxonomic subbiﬂ Includes ssp. columbianum, triﬁ P. kelloggii P. P. polygaloides P. ssp. kelloggii rupicola, F. falcata rupicola, F. P. pusillus P. pusillus

var. dum var. geyeriana bracteata Eriogonum subreniforme Salix geyeriana var. Malus ioensis Polygonum kelloggii Potamogeton foliosus foliosus Fendlera rupicola rupicola F. Potamogeton pusillus Prunus americana Fragaria vesca var. Potentilla diversifolia diversifolia Galium triﬁ Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park, cont. FamilyPolygonaceae Species name Salicaceae Rosaceae Polygonaceae Potamogetonaceae Saxifragaceae (Hydrangeaceae) Potamogetonaceae Rosaceae Rosaceae Rosaceae Rubiaceae

126 Annotated Checklist of Vascular Flora: Capitol Reef National Park

var. var.

Type Type var. . Not var. var. var. . This var. var. Cordylanthus P. caespitosus P. scariosus P. scariosus P. P. ophianthus P. P. pachyphyllus P. P. ophianthus. P. P. scariosus P. L. junceum P. palustris P. is not otherwise eld Co, UT (Jones ora P. parviﬂ P.

Type locality is near Castle Gate, Type Type locality is Cottonwood Canyon, Type ed specimen of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of ed specimens of Type locality is Mammoth Creek, Type Also reported by Northern Colorado orus. misidentifi misidentifi desertipicti. Carbon Co, UT (Pennell 6138 US). misidentifi montanensis. known from Utah. parvifl misidentifi scariosus. misidentifi scariosus. Markagunt Plateau, Garfi 6026b US). misidentifi Comments congestus. 787 GH). Salt Lake Co, UT (Watson misidentifi scariosus. Plateau Network (in ed.). misidentifi misidentifi locality is near Price, Carbon Co, UT (Jones s.n. POM). misidentifi native to Utah. species is not known from Utah, but native to Wyoming and Nebraska. Park status Range PerF Wide FalsRep Report from Meyer (1982) based on Life form PerF RegEn FalsRep Reports from CARE Herbarium are based on PerF RegEn FalsRep Reports from CARE herbarium based on PerF LocEn FalsRep Reports from CARE Herbarium are based on PerF LocEn FalsRep Reports from CARE Herbarium are based on PerF RegEn FalsRepPerF Reports from CARE Herbarium are based on RegEn FalsRep Reports from CARE Herbarium are based on PerF RegEn FalsRep Reports from CARE Herbarium based on PerF LocEn FalsRep Reports from CARE Herbarium are based on marsh grass-of- Parnassus violet owl-clover AnnF RegEn FalsRep Reports from CARE Herbarium based on bluestem penstemon shortleaf penstemon Markagunt penstemon Marcus Jones’ penstemon Common name Natural Bridges penstemon upright blue penstemon Nuttall’s desert Nuttall’s parsley broadleaf penstemon

var. ora P. palustris P. parviﬂ O. purpureo-albus Synonyms/ notes Taxonomic L. megarrhizum, Aletes nuttallii

var. var. ora Penstemon cyanocaulis Parnassia parviﬂ Orthocarpus purpureoalbus Penstemon caespitosus perbrevis Penstemon leiophyllus leiophyllus Penstemon marcusii Penstemon strictiformis Penstemon virgatus Penstemon platyphyllus Lomatium nuttallii Scrophulariaceae Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park, cont. FamilySaxifragaceae (Parnassiaceae) Species name Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Umbelliferae (Apiaceae)

Appendices 127 . Also Type Type L. parryi. O. depauperata adunca. V. ed specimens of ed specimens of ed specimens of misidentifi misidentifi misidentifi locality is Frisco, Beaver Co, UT (Jones 1864 US). reported by Northern Colorado Plateau Network (in ed.). Comments Park status ed literature (Heil et al. 1993, Meyer 1982, Northern ed literature Colorado Plateau Network, Range PerF Wide FalsRep Reports from CARE Herbarium based on Life form cations (False Reports or FalsRep) because the park is well outside known expected range of rough lomatium PerF RegEn FalsRep Reports from CARE Herbarium are based on Chilean sweet-cicelyChilean PerF Wide FalsRep Reports from CARE Herbarium are based on Canadian white violet Common name rugulosa, V. rugulosa, V. O. berteroi var. scopulorum, V. scopulorum, V. rydbergii Synonyms/ notes Taxonomic

var. scabrum Lomatium scabrum Osmorhiza chilensis Viola canadensis canadensis V. Appendix C. Rejected (falsely reported or questionable) taxa of Capitol Reef National Park, cont. FamilyUmbelliferae (Apiaceae) Species name Umbelliferae (Apiaceae) Violaceae The vascular plant species listed in this report have been reported for Capitol Reef National Park from published and unpublish for Capitol Reef National Park from have been reported The vascular plant species listed in this report because of misidentiﬁ now believed to be erroneous herbarium specimens, but are in ed.) or from species (Questionable or Ques?).

128 Annotated Checklist of Vascular Flora: Capitol Reef National Park areas Wet Comments con Mix Sage PJ des Salt Flower time Park status r Native Pres NA x box elder Native Pres Apr–Jun x water birch Native Pres Apr–May x European white birch Intro Pres Apr–May Cultivated western hop-hornbeam Native Pres Jul x showy catalpa Intro Pres May–Jun Cultivated Rocky Mountain juniper Native Pres NA x x x white cedar Intro Pres NA Cultivated common persimmon Intro Pres May–Jun Cultivated Russian oliveGambel oak Intro Pres Native Pres May–Jul Apr–Jun x x x x x Disturbed black walnut Intro Pres Apr–May Cultivated English walnut Intro Pres Apr–May Cultivated honey locust Intro Pres Apr–May Cultivated black locust Intro Pres May–Jun x Cultivated Osage orange Intro Pres May Cultivated white mulberry Intro Pres Apr–Jun Cultivated white ash Intro Pres May–Jun Cultivated green ash Intro Pres Jun Cultivated Engelmann spruce Native Pres NA x blue spruce Native Pres NA x two-needle pinyon Native Pres NA x x x limber pine Native Rep NA x x western bristlecone pine Native Pres NA x ponderosa pine Native Pres NA x Douglas-ﬁ quinceshowy crabapple Intro Pres Intro May Pres May Cultivated Cultivated

var. gambelii

var. var. oribunda exilis Acer negundo Betula occidentalis Betula pendula Ostrya knowltonii Catalpa speciosa Juniperus scopulorum Thuja occidentalis Diospyros virginiana Quercus gambelii Elaeagnus angustifolia Juglans nigra Juglans regia Gleditsia triacanthos Robinia pseudoacacia Maclura pomifera Morus alba Fraxinus americana Fraxinus pennsylvanica Picea engelmannii Picea pungens Pinus edulis Pinus fl Pinus longaeva scopulorum Pinus ponderosa Pseudotsuga menziesii glauca Cydonia oblonga Malus fl taxa of Capitol Reef National Park, organized by life form with ecological notes. rmed, historical, and reported FamilyTrees Aceraceae Species name Common name Nativity Betulaceae Betulaceae Betulaceae Bignoniaceae Cupressaceae Cupressaceae Ebenaceae Fagaceae Elaegnaceae Juglandaceae Juglandaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Moraceae Moraceae Oleaceae Oleaceae Pinaceae Pinaceae Pinaceae Pinaceae Pinaceae Pinaceae Pinaceae Rosaceae Rosaceae Appendix D. Confi

Appendices 129 areas Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres Apr–Jun x owering plum owering Intro Pres Mar–Apr Cultivated common hackberrynetleaf hackberry IntroChinese elm PresSiberian elm Native Apr–May PresRocky Mountain maplenarrowleaved yucca Apr–May Native Intro Intro x Pres Native Pres Rep x Apr–Jun Pres Mar–Apr Mar–Apr May–Jul x x x x Cultivated x x x x Cultivated Sandy soils common applecrabappleapricotsweet cherry Introﬂ sour cherry Prescommon plum Introalmond May Intropeach Pres Introcommon pear Pres May Pres Intro Intronarrow-leaved cottonwood Apr–May Mar–Apr Pres PresFremont cottonwood Intro Intro Mar–Apr Mayblack poplar Native Introquaking aspen Pres Pres Preslance-leaf cottonwood Pres Apr–Maypeach-leaf willow Apr–May Mar–Jun Nativeweeping willow Apr–May Intro Native Pres black willowGoodding’s Cultivated Native Native willowMatsuda’s Pres Pres Apr–Juntree-of-heaven Pres Pres Cultivated Intro Cultivated Mar–Jun Apr–Jun Cultivated Apr–Jun Intro Apr–May Pres Cultivated Intro Pres May–Jun Cultivated x Pres Apr–May Cultivated Cultivated Jun–Jul Cultivated x x x x x Cultivated Cultivated Disturbed

var. glabrum var. intercurrens x Celtis occidentalis Celtis reticulata Ulmus parvifolia Ulmus pumila Acer glabrum angustissima Yucca Malus pumila Prunus cerasus Populus fremontii fremontii Malus sylvestris Prunus armeniaca Prunus avium Prunus cerasifera Prunus domestica Prunus dulcis Prunus persica Pyrus communis Populus angustifolia Populus nigra Populus tremuloides Populus Salix amygdaloides Salix babylonica Salix gooddingii Salix matsudana Ailanthus altissima rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Ulmaceae (Celtidaceae) Ulmaceae Ulmaceae Shrubs Aceraceae Agavaceae Ulmaceae (Celtidaceae) Family cont. Trees, Rosaceae Species name Common name Nativity Rosaceae Salicaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Simaroubaceae Appendix D. Conﬁ

130 Annotated Checklist of Vascular Flora: Capitol Reef National Park Tropic, Tropic, Morrison, & Duchesne River F. gravelly soils Wet Comments con Mix Sage PJ des Salt Flower time Park status ower snowberry ower Native Pres May–Jul x x x x x Castle Valley saltbushCastle Valley Native Pres May–Sep x x Saline clays mat saltbush Nativespiny hopsage Pres Apr–Sep Native x Rep Apr–Jul x x x Mancos, coralberrymountain snowberrysnowball-bush Native Presmountain-lover Introiodine bush Jun–Augfourwing saltbush Intro Pres Nativeshadscale Pres May–Jul Native Pres x Jun–Jul Native Pres May–Aug Pres May–Sep x Jul–Sep Native x x Pres x x x May–Sep x x x Cultivated x Cultivated x Sandy soils x Clayey to Harriman’s yuccaHarriman’s squawbush Nativesquawbush Preswestern poison ivy barberryThunberg’s Apr–Maycommon barberry Native Native mahoniaFremont’s x Intro Pres NativeOregon-grape Pres x Intro Presthinleaf alder May–Jul Pres Native May–JunUtah honeysuckle x Pres May Pres x May–Julblue elderberry Native May–Jun x Apr–Jul Native Native Pres x Rep x Pres Native Apr–Jun Pres x Jun–Jul Apr–May x May–Jul x x x x x x Cultivated Cultivated x x x x x x long-ﬂ

var.

orus cuneata

trilobata var. var. var. occidentalis var. var. var. Atriplex gardneri Grayia spinosa Atriplex corrugata Symphoricarpos orbiculatus Symphoricarpos oreophilus utahensis Viburnum opulus Pachystima myrsinites Allenrolfea occidentalis Atriplex canescens canescens Atriplex confertifolia Yucca harrimaniae Yucca harimaniae Rhus aromatica simplicifolia Rhus aromatica rydbergii Toxicodendron Berberis thunbergii Berberis vulgaris Mahonia fremontii Mahonia repens Alnus incana Lonicera utahensis Sambucus caerulea Symphoricarpos longiﬂ rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Chenopodiaceae Chenopodiaceae Chenopodiaceae Caprifoliaceae Caprifoliaceae (Adoxaceae) Celastraceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Caprifoliaceae Appendix D. Conﬁ FamilyShrubs, cont. Agavaceae Species name Common name Nativity Anacardiaceae Anacardiaceae Anacardiaceae Berberidaceae Berberidaceae Berberidaceae Berberidaceae Betulaceae Caprifoliaceae Caprifoliaceae (Adoxaceae) Caprifoliaceae

Appendices 131 Morrison, & Duchesne F seleniferous soils gravelly saline soils Wet Comments con Mix Sage PJ des Salt Flower time Park status winterfat Native Pres May–Jul x x x greasewood Native Pres May–Aug x Arizona siltbush Native Pres Jun–Aug x Entrada, Brandegee’s siltbushBrandegee’s Native siltbushPlummer’s Pres Jun–Aug Native Pres x Jun–Aug x Saline or Bigelow’s sagebrushBigelow’s Native Pres Aug–Oct x x Rocky areas sand sagebrush Native Pres Aug–Oct x x x fringed sagebrush Native Pres Jul–Sep x x x x black sagebrush Native Pres Sep–Oct x x x pygmy sagebrush Native Pres Aug–Sep x x x x budsagebasin big sagebrush Native Native Pres Pres Jul–Sep Apr–Jun x x x x x Clayey to mountain big sagebrush Native Rep Jul–Sep x x x Wyoming big sagebrush Native Rep Jul–Sep x Rio Grande seepwillow Native Pres May–Nov x spiny brickellbush Native Pres Apr–May x Rock crevices

var.

arizonica var. var. var. var. nova var. var. var. var. lifolia Krascheninnikovia lanata lanata Sarcobatus vermiculatus Zuckia brandegei Zuckia brandegei brandegei Zuckia brandegei plummeri Artemisia bigelovii Artemisia ﬁ Artemisia frigida Artemisia nova Artemisia pygmaea Artemisia spinescens Artemisia tridentata tridentata Artemisia tridentata vaseyana Artemisia tridentata wyomingensis Baccharis salicina Brickellia atractyloides rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyShrubs, cont. Chenopodiaceae Species name Common name Nativity Chenopodiaceae (Sarcobataceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

132 Annotated Checklist of Vascular Flora: Capitol Reef National Park soils Wet Comments con Mix Sage PJ des Salt Flower time Park status lanceleaf rabbitbrush Native Pres Jul–Sep x x x California brickellbush Native Pres Aug–Sep Rock crevices longleaf brickellbush Native Pres Aug–Sep x rough brickellbush Native Pres Aug–Oct x x x Sandstone Mohave brickellbush Native Pres Jun–Aug x x x x dwarf rabbitbrush Native Pres Jul–Sep x x x x Greene’s rabbitbrushGreene’s Native Pres Jul–Sep x x x Bigelow’s rabbitbrushBigelow’s Native Rep Aug–Oct x x Rocky areas spear-leaf rabbitbrushspear-leaf Native Pres Jul–Sep x Sandy to saline greenish rabbitbrush Native Pres Aug–Oct x x x x x graystem rabbitbrush Native Pres Aug–Oct x x x glabrate rabbitbrush Native Pres Aug–Oct x x x x rush rabbitbrush Native Pres Aug–Oct x x x smoothseed rabbitbrush Native Pres Aug–Oct x x x x whitestem rabbitbrush Native Rep Aug–Oct x x x dune rabbitbrush Native Pres Aug–Oct x x x Nevada rabbitbrush Native Pres Jul–Sep x x x

var. var. var. var. var. var. var. var.

orus var. var. var. lanceolatus Chrysothamnus viscidiﬂ var. Brickellia californica Brickellia longifolia Brickellia microphylla scabra Brickellia oblongifolia linifolia Chrysothamnus depressus Chrysothamnus greenei Chrysothamnus nauseosus bigelovii Chrysothamnus linifolius Chrysothamnus nauseosus consimilis Chrysothamnus nauseosus gnaphalodes Chrysothamnus nauseosus graveolens Chrysothamnus nauseosus junceus Chrysothamnus nauseosus leiospermus Chrysothamnus nauseosus speciosus Chrysothamnus nauseosus turbinatus Chrysothamnus parryi nevadensis rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyShrubs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Appendices 133 Wet Comments con Mix Sage PJ des Salt Flower time Park status puberulent rabbitbrush Native Pres Jul–Sep x x x x slenderleaf rabbitbrush Native Pres Jul–Sep x x x green rabbitbrush Native Pres Jul–Sep x x x x thread snakeweed Native Pres Jul–Oct x x x broom snakeweed Native Pres Jul–Oct x x x Rusby’s goldenbushRusby’s Native Pres Aug–Sep x x copperweed Native Pres Jun–Oct x Saline soils spineless horsebrush Native Pres Jun–Sep x x littleleaf horsebrush Native Pres May–Jul x x x shortspine horsebrush Native Pres Apr–Jun x x x resinbush Native Pres Jul–Sep x x x red-osier dogwood Native Pres May–Jul x small-leaved greasebush Native Pres Mar–Jun x x x common juniper Native Pres NA x Utah juniper Native Pres NA x x x x x silver buffaloberry Native Pres Apr–May x Canada buffaloberry Native Pres May–Jul x roundleaf buffaloberry Native Pres Mar–May x x x x Torrey’s ephedraTorrey’s Native Pres NA x x green ephedra Native Pres NA x x x Cutler’s ephedraCutler’s Native Pres NA x x x Sandy soil

var.

orus orus orus var. viridis viscida sericea var. var. var. orus puberulus stenophyllus viscidifl Chrysothamnus viscidifl var. Chrysothamnus viscidifl var. Chrysothamnus viscidifl var. Gutierrezia microcephala Gutierrezia sarothrae Haplopappus rusbyi Oxytenia acerosa Tetradymia canescens Tetradymia Tetradymia glabrata Tetradymia Tetradymia spinosa Tetradymia Vanclevea stylosa Vanclevea Cornus sericea Glossopetalon spinescens microphyllum depressa Juniperus communis Juniperus osteosperma Shepherdia argentea Shepherdia canadensis Shepherdia rotundifolia Ephedra torreyana Ephedra viridis Ephedra viridis rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyShrubs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cornaceae Crossosomataceae Cupressaceae Cupressaceae Elaegnaceae Elaegnaceae Elaegnaceae Ephedraceae Ephedraceae Ephedraceae

134 Annotated Checklist of Vascular Flora: Capitol Reef National Park boulders Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres Jul–Nov x Native Pres Apr–Jun x x x Rock crevices Native Pres May–Jun x x Rocky sites greenleaf manzanita Native Pres Mar–Jul x x x shinnery oak Native Pres Apr–Jun x x x Sandy soils purple sage Native Pres May–Jun x x x Fremont’s indigo-bushFremont’s Native Pres Apr–Jun x x rose-of-sharon Intro Pres Jul–Sep Cultivated Thompson’s indigo-bushThompson’s Native Rep May–Aug x x Sandy soils & singleleaf ash Native Pres Apr–May x x x common lilac Intro Pres May Cultivated prickly-phlox Native Pres May–Jul x x x x golden wild buckwheat Native Pres Aug–Oct x x x x crisp-leaf wild buckwheat Native Pres Jul–Oct x x x Heil’s wild buckwheatHeil’s Native Pres Jul–Aug x rimrock wild buckwheat Native Pres Aug–Oct x x x Sandstone sand wild buckwheat Native Pres Jun–Oct x x x San Juan wild buckwheat Native Pres Jun–Oct x x x slenderleaf wild buckwheat Fendler’s mountain-lilacFendler’s Native Pres Jun–Aug x x birchleaf buckthorn Native Pres May–Jun x Rock crevices Saskatoon serviceberry Native Pres May–Jul x x x Utah serviceberry Native Pres Apr–Jun x x x x dwarf mountain mahogany curl-leaf mountain mahogany

var. var. var. var. var. var. var. thompsoniae Arctostaphylos patula Quercus welshii Poliomintha incana Psorothamnus fremontii Hibiscus syriacus Psorothamnus thompsoniae var. Fraxinus anomala Syringa vulgaris Leptodactylon pungens Eriogonum corymbosum aureum Eriogonum corymbosum corymbosum Eriogonum corymbosum heilii Eriogonum corymbosum orbiculatum leptocladon Eriogonum leptocladon ramosissimum Eriogonum leptocladon Eriogonum leptophyllum Ceanothus fendleri Rhamnus betulifolia Amelanchier alnifolia Amelanchier utahensis Cercocarpus intricatus Cercocarpus ledifolius intermontanus rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyShrubs, cont. Ericaceae Species name Common name Nativity Fagaceae Labiatae (Lamiaceae) Leguminosae (Fabaceae) Malvaceae Leguminosae (Fabaceae) Oleaceae Oleaceae Polemoniaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Rhamnaceae Rhamnaceae Rosaceae Rosaceae Rosaceae Rosaceae

Appendices 135 to clay soil Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres May–Jun x x x ve-stamen tamarisk ve-stamen Intro Pres Apr–Sep x alder-leaf mountain alder-leaf mahogany blackbrushApache plume Native Native Pres Pres Apr–Jun Apr–Aug x x x x x Shallow sandy bush oceanspray Native Pres Jun–Aug x x x Rock crevices dwarf ninebark Native Pres May–Aug x x x Rock crevices cliff-rose Native Pres May–Jun x x x x antelope bitterbrush Native Pres Apr–Jul x x x Woods’ roseWoods’ Native Pres May–Jul x x Booth’s willowBooth’s Rep May–Jun x strapleaf willow Native Pres Apr–Jun x yellow willow Native Pres Apr–Jun x coyote willow Native Pres Apr–Aug x arroyo willow Native Pres Apr–May x whiplash willow Native Pres Apr–Jun x Utah fendlerella Native Pres Apr–Sep x x x littleleaf mock-orange Native Pres May–Sep x x golden currant Native Pres Mar–Jul x x x x x wax currant Native Pres Apr–Aug x x x whitestem gooseberry Native Pres Apr–Jul x x x Anderson’s wolfberryAnderson’s Native Pres Feb–May x x x ﬁ thicket-creeper Native Pres May–Jul x northern fox grape Intro Pres May–Jun Cultivated northern arrowhead Native Pres Jun–Sep x

ligulifolia watsonii var. var. var. inebrians stenophylla caudata var. var. ssp. Cercocarpus montanus Coleogyne ramosissima Fallugia paradoxa Holodiscus dumosus Physocarpus alternans stansburyana Purshia mexicana Purshia tridentata Rosa woodsii Salix boothii Salix eriocephala Salix eriocephala Salix exigua Salix lasiolepis Salix lucida Fendlerella utahensis Philadelphus microphyllus Ribes aureum Ribes cereum Ribes inerme Lycium andersonii Lycium Tamarix chinensis Tamarix Parthenocissus vitacea Vitis labrusca Sagittaria cuneata rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyShrubs, cont. Rosaceae Species name Common name Nativity Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Saxifragaceae (Hydrangeaceae) Saxifragaceae (Hydrangeaceae) Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Solanaceae Tamaricaceae Vitaceae Vitaceae Perennial Forbs Perennial Alismataceae

136 Annotated Checklist of Vascular Flora: Capitol Reef National Park Summerville F. areas Wet Comments con Mix Sage PJ des Salt Flower time Park status Jones’ amsonia Native Pres Apr–Jun x x x showy stoneseed Native Pres Apr–Jul x x x tomentose amsonia Native Pres Apr–May x x common dogbane Native Pres Jun–Aug x Jones’ cycladeniaspider milkweed Native Pres Native May–Jun Pres x May–Jul x x x x Chinle, Cutler, pallid milkweed Native Pres Apr–Jun x x x Circle Cliffs milkweedCircle Cliffs Native Pres May–Aug x x x broadleaf milkweed Native Pres Jun–Aug x x x x bigseed milkweed Native Pres Apr–Jul x x x Ruth’s milkweedRuth’s Native Pres Apr–Jun x x x showy milkweed Native Pres May–Aug x whorled milkweedlow cryptanth Native Rep May–Sep Native Pres x May–Jun x x x x Disturbed x head cryptanth Native Pres May–Jun x x ashy cryptanth Native Pres May–Aug x x x James’ cryptanth Native Pres May–Aug x x x x San Juan cryptanth Native Pres May–Aug x x x basin yellow cryptanth Native Pres Apr–Jun x x x x plateau yellow cryptanth Native Pres Apr–Jun x x x yellow-eye cryptanth Native Pres Apr–Jun x x x plateau cryptanth Native Pres May–Jun x x x Mancos Shale dwarf cryptanth Native Pres Apr–Jun x x x carbon cryptanth Native Pres May–Jun x x x x Osterhout’s cryptanthOsterhout’s Native Pres May–Jun x x Rollins’ cryptanth Native Pres May–Jun x x x bristly cryptanth Native Pres May–Aug x x x

cinerea jamesii jonesii asperula var. var. ora var. var. var. var. var. ava avoculata Amsonia jonesii Lithospermum incisum Amsonia tomentosa stenophylla Apocynum cannabinum Cycladenia humilis Asclepias asperula Asclepias cryptoceras cryptoceras Asclepias labriformis Asclepias latifolia Asclepias macrosperma Asclepias ruthiae Asclepias speciosa Asclepias subverticillata Cryptantha abata Cryptantha capitata Cryptantha cinerea Cryptantha cinerea Cryptantha cinerea pustulosa Cryptantha confertifl Cryptantha fl Cryptantha fl Cryptantha fulvocanescens Cryptantha humilis Cryptantha mensana Cryptantha osterhoutii Cryptantha rollinsii Cryptantha setosissima rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Apocynaceae Species name Common name Nativity Boraginaceae Apocynaceae Apocynaceae Apocynaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae

Appendices 137 Curtis F. Dakota, Morrison, & Entrada F. Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres May–Jul x x x x Native Pres Apr–May x x x x Native Pres Mar–May x Dakota & shhook shhook cactus shhook Native Pres Apr–May x x x Mancos, ower Native Pres Aug–Oct x ower fi ower basin sandwort Native Rep May–Aug x x x x southwestern stoneseed Native Pres Jun–Aug x x x matted tiquilia Native Pres May–Aug x x x Arizona pincushion Native Pres Apr–Aug x x x claretcup Native Pres Apr–Jun x x x x Heil’s beavertailHeil’s Native Pres Mar–Apr x brittle pricklypear Native Pres Jun–Jul x x x x large pricklypear Native Pres Apr–Jul x x plains pricklypear Native Pres May–Jun x x x x rufous-spine pricklypear Native Pres May–Jun x x x Simpson’s hedgehog Simpson’s cactus cactus Winkler’s pincushion Winkler’s cactus small-fl fi Wright’s harebellParry’s Native Pres Jun–Sep x cardinal-fl common hops Native Pres Jul–Aug x Eastwood’s sandwortEastwood’s Native Pres May–Sep x x x x

var. var. var. var. var.

orum heilii

ssp. var. var. var. var. melanacanthus spina glabrescens Lithospermum multifl Tiquilia latior Tiquilia Coryphantha vivipara arizonica Echinocereus triglochidiatus var. Opuntia basilaris Opuntia fragilis major Opuntia phaeacantha Opuntia polyacantha polyacantha Opuntia polyacantha rufi Pediocactus simpsonii Pediocactus winkleri Sclerocactus whipplei roseus Sclerocactus wrightiae Campanula parryi graminea Lobelia cardinalis Humulus lupulus neomexicanus Arenaria fendleri eastwoodiae Arenaria fendleri rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Boraginaceae Species name Common name Nativity Boraginaceae (Ehretiaceae) Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Campanulaceae Campanulaceae Cannabaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae)

138 Annotated Checklist of Vascular Flora: Capitol Reef National Park areas areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status y Native Pres May–Sep x x x c aster c Native Pres Jul–Oct x x x x Drummond’s catchﬂ Drummond’s creeping nailwort Native Pres May–Aug x x x mouse-ear chickweed Intro Pres Apr–Sep x Disturbed long-stalk starwort Native Pres May–Aug x x bouncing-bet Intro Pres May–Sep Disturbed greenmolly Native Pres Aug–Oct x x western spiderwortcommon yarrow Native Pres Native May–Aug Pres May–Sep x x x x x x x cutleaf agoseris Native Pres May–Sep x x x small-leaf pussyoes Native Pres Jun–Aug x x x common pussytoes Native Pres May–Jul x x x breaks pussytoes Native Rep Jun–Aug x x burdock Intro Pres Jul–Sep x Disturbed sand wormwood Native Pres Jul–Sep x x x Sandy soils tarragon Native Pres Jul–Oct x x x x x Louisiana wormwood Native Pres Jul–Oct x x Louisiana wormwood Native Pres Jul–Oct x x x Paciﬁ Parry’s asterParry’s Native Pres Jul–Sep x

var. var. var. longipes var. var. laciniata parryi ora var. var. var. cinalis Lychnis drummondii Lychnis Paronychia sessilifl Cerastium fontanum Stellaria longipes Saponaria offi Bassia americana Tradescantia occidentalis Tradescantia lanulosa Achillea millefolium Agoseris glauca Antennaria microphylla Antennaria parvifolia Antennaria rosulata Arctium minus Artemisia campestris scouleriana Artemisia dracunculus glauca Artemisia ludoviciana albula Artemisia ludoviciana mexicana Aster ascendens Aster foliaceus rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Caryophyllaceae Caryophyllaceae (Alsinaceae) Appendix D. Confi FamilyPerennial Forbs, cont. Caryophyllaceae (Alsinaceae) Species name Common name Nativity Caryophyllaceae (Alsinaceae) Caryophyllaceae Chenopodiaceae Commelinaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Appendices 139 areas Wet Comments con Mix Sage PJ des Salt Flower time Park status ower brickellbush ower Native Pres Jul–Oct x x blueleaf aster Native Pres Jul–Sep x x x x alkali aster Native Pres May–Sep x western aster Native Pres Jul–Sep x x Welsh’s asterWelsh’s Native Pres Aug–Oct x cutleaf Native Rep Jul–Sep x x x nodding bur-marigold Native Pres Jul–Sep x tassel-ﬂ Russian knapweedhoary dusty-maiden Intro Native Pres Pres Jun–Sep May–Sep x x x x x Disturbed rose-heath Native Pres May–Sep x x x x Eastwood-plant Native Pres May–Jun x x x x hispid golden-aster Native Pres May–Sep x x x x wingless thistle Native Pres Jun–Sep x x Caineville thistle Native Pres Jun–Sep x pretty thistle Native Pres Jun–Sep x x x x New Mexico thistle Native Pres May–Jun x x x Utah thistle Native Pres May–Jun x x x

var. var.

minor var. var. var. var. ora orus Aster glaucodes Aster paucifl Aster spathulatus Aster welshii Bahia dissecta Bidens cernua Brickellia grandifl Centaurea repens Chaenactis douglasii Chaetopappa ericoides Chamaechaenactis scaposa Chrysopsis villosa Cirsium calcareum bipinnatum Cirsium calcareum calcareum Cirsium calcareum pulchellum Cirsium neomexicanum neomexicanum Cirsium neomexicanum utahense rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

140 Annotated Checklist of Vascular Flora: Capitol Reef National Park Morrison, Summerville F Navajo F Wet Comments con Mix Sage PJ des Salt Flower time Jun–Aug x Park status Rep Native eabane Native Pres May–Aug x x x x eabane Native Pres Jun–Aug x x x elk thistle wavy-leaved thistle Native Pres Jun–Sep bull thistle Intro Pres Jul–Aug glaucous hawksbeard Native Pres May–Aug x nakedstem Native Pres May–Jun x x x Abajo daisy Native Pres Jul–Sep x x congested hairy daisy Native Pres May–Jul x x x silver daisy Native Pres May–Jun x x x tufted ﬂ San Rafael daisyspreading daisy Native Pres Native May–Jun Pres x May–Sep x x Moenkopi, x Eaton’s daisyEaton’s Native Pres May–Jul x x x trailing daisy Native Pres May–Aug x x x longleaf daisy Native Pres Jun–Sep x Maguire’s daisyMaguire’s basin daisy Native Pres Jun–Jul Native Pres May–Jun x x x Wingate & Navajo ﬂ

var.

var. var. glauca var. var. var. var. var. agellaris scariosum Cirsium scariosum Cirsium undulatum undulatum Cirsium vulgare Crepis runcinata Enceliopsis nudicaulis nudicaulis Erigeron abajoensis Erigeron aphanactis congestus Erigeron argentatus Erigeron caespitosus Erigeron compactus consimilis Erigeron divergens cinereus Erigeron eatonii Erigeron ﬂ Erigeron lonchophyllus Erigeron maguirei Erigeron pulcherrimus concinnus Erigeron pumilus rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Appendices 141 areas Wet Comments con Mix Sage PJ des Salt Flower time Park status ower Native Pres Jul–Sep x x x ower Native Pres May–Oct x ower Native Pres Apr–Aug x x ower Native Pres Jul–Sep x lanate hymenoxys Native Rep May–Jul x x vernal daisy Native Pres May–Aug x x x Canyonlands hymenoxys Native Pres Apr–Jun x x x aspen daisy Native Pres Jun–Aug x x slenderleaf daisy Native Pres Jun–Sep x x Sandstone Utah daisy Native Pres Apr–Jun x x western goldenrod Native Pres Jul–Oct x Hopi blanketfl basin blanketfl Wright’s cudweedWright’s Native Pres Aug–Oct x curly-cup gumweedstemless goldenweed Native Native Pres Pres Jul–Sep May–Jul x x x x Disturbed thrift goldenweed Native Pres May–Jun x x x x purpledisk sunfl Nuttall’s sunfl Nuttall’s common hyalineherb Native Pres May–Oct x x x x Arizona hymenoxys Native Pres Apr–Jun x x x x

var. var.

ivesiana

var. var. var. var. var. var. var. var. da var. lifolius orus caespitosa condensatus Erigeron pumilus Hymenoxys acaulis Erigeron speciosus macranthus Erigeron utahensis sparsifl Erigeron utahensis utahensis Euthamia occidentalis Gaillardia pinnatifi Gaillardia spathulata Gnaphalium wrightii quasiperennis Haplopappus acaulis acaulis Grindelia squarrosa Haplopappus armerioides armerioides Helianthella microcephala Helianthus nuttallii Hymenopappus fi cinereus Hymenoxys acaulis arizonica Hymenoxys acaulis rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

142 Annotated Checklist of Vascular Flora: Capitol Reef National Park Wet Comments con Mix Sage PJ des Salt Flower time Park status ower Native Pres May–Sep x x x many-headed groundsel Native Pres Jun–Oct x x x Saline soils low hymenoxys Native Pres May–Jun x x x Colorado rubber-plant Native Pres May–Aug x x x x broom groundsel Native Pres Jun–Oct x x x x Sandy soils poverty-weed Native Pres May–Sep x x x x Arizona rush-pink Native Pres May–Jun x x x western rush-pink Native Rep May–Jun x x showy rush-pink Native Pres May–Jun x x x x aristate aster Native Pres Jul–Oct x x x x hoary aster Native Pres May–Sep x x x x gumweed aster Native Pres May–Jul x x x low feverfew Native Pres Apr–Jun x x x rock-goldenrod Native Pres Jul–Sep x x x x desert bahia Native Pres May–Jun x x greenstem paperﬂ Douglas’ groundsel Native Pres May–Oct x x x basin groundsel Native Pres Apr–Aug x x x x

var. var. var. var. var. var. var.

nana pumila

var. var. ora ora ora var. ora var. var. ora grindelioides oribunda multicapitatus Hymenoxys acaulis fl Senecio spartioides spartioides Hymenoxys richardsonii Iva axillaris Lygodesmia grandifl Lygodesmia arizonica Lygodesmia grandifl Lygodesmia dianthopsis Lygodesmia grandifl Lygodesmia grandifl Machaeranthera canescens aristata Machaeranthera canescens canescens Machaeranthera grindelioides var. Parthenium ligulatum Petradoria pumila Platyschkuhria integrifolia desertorum Psilostrophe sparsifl Senecio douglasii longilobus Senecio multilobatus Senecio spartioides rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Appendices 143 areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Rep Jun–Aug x x Rocky Mountain groundsel Canadian goldenrod Native Pres Aug–Oct x coast goldenrod Native Pres Jul–Sep x three-nerve goldenrod Native Pres Aug–Oct x x x desert wirelettuce Native Pres Jul–Aug x x slender wirelettucecommon dandelion Nativealpine greenthread Pres Intro Jun–Sep Pres Native x Pres Apr–Dec Jun–Jul x x x x x x x x x Disturbed scapose greenthread Native Pres May–Aug x x x scale glandweed Native Pres Aug–Oct x Last Chance townsendia Native Pres Apr–May x x Mancos Shale stemless townsendia Native Pres May–Jun x hoary townsendia Native Pres Apr–Jul x x x yellow salsifyplains ironweed Intro Intro Pres Pres May–Jul Sep x x Disturbed x showy goldeneye Native Pres Jun–Sep x x x x rough mule’s-ears Native Pres May–Aug x x x x Sandy soils

var. var. var. var. var.

var. var. ora cinale ora Senecio streptanthifolius streptanthifolius Solidago canadensis salebrosa Solidago simplex Solidago velutina Stephanomeria runcinata Stephanomeria tenuifolia tenuifolia Thelesperma subnudum alpinum Taraxacum offi Taraxacum Thelesperma subnudum subnudum belenidium Thymophylla pentachaeta Townsendia aprica Townsendia Townsendia exscapa Townsendia Townsendia incana Townsendia Tragopogon dubius Tragopogon Vernonia marginata Vernonia Viguiera multifl multifl Wyethia scabra rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

144 Annotated Checklist of Vascular Flora: Capitol Reef National Park Morrison, Moenkopi F seleniferous soils areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status eld bindweed eld Intro Pres Jun–Aug Disturbed hurtleaf woodyaster Native Pres Apr–Jun x x x ﬁ evolvulus Native Pres May–Jun x x Henrieville woodyaster Native PresCisco woodyaster May–Jul x Native Pres x Apr–Jun x Kaiparowits, Saline & lance-leaved stonecrop Native Pres Jun–Aug x x x nodding rock cress Native Pres May–Aug x x x Drummond’s rockcressDrummond’s Native Pres May–Sep x x hairy rockcress Native Pres Jun–Aug x x x Fendler’s rockcressFendler’s Native Pres Apr–Aug x x Holboell’s rockcressHolboell’s Native Pres May–Jul x x x x Holboell’s rockcressHolboell’s Native Pres May–Jul x x x perennial rockcress Native Pres Mar–Jul x x x x desert rockcress Native Pres Mar–Jun x x x Selby’s rockcressSelby’s heartleaf bittercress Native Native Pres Rep May–Aug Jun–Sep x x x x x whitetop Intro Pres May–Aug Disturbed

var.

fendleri secunda var. pallens var. var. var. var. var. imberbis Xylorhiza tortifolia Convolvulus arvensis Evolvulus nuttallianus Xylorhiza confertifolia Xylorhiza venusta Sedum lanceolatum Arabis demissa Arabis drummondii Arabis hirsuta Arabis holboellii Arabis holboellii pinetorum Arabis holboellii Arabis perennans perennans Arabis pulchra Arabis selbyi cordifolia Cardamine cordifolia Cardaria draba rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Convolvulaceae Convolvulaceae Appendix D. Conﬁ FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species nameCompositae (Asteraceae) Common name Nativity Crassulaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

Appendices 145 Formation Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres Apr–Aug x x x ower Native Pres Apr–Aug x x x x ower Native Pres Apr–Jul x x smooth spindlestem Native Pres May–Jul x x x California tansy-mustard Native Pres Jun–Sep x x x golden draba Native Pres Jun–Aug x x slender whitlow-grass Native Pres Jun–Aug x sand dune wallﬂ shy wallﬂ Jones’ pepperwort Native Pres Apr–Sep x x x watercress Intro Pres May–Sep x sharp-leaf twinpod Native Pres May–Jul x x x x purple twinpod Native Pres May–Jul x Watson’s bladderpodWatson’s Native Pres May–Aug x x x silver bladderpod Native Pres May–Jun x x x Newberry’s twinpodNewberry’s Native Pres Mar–Jun x x x Colorado Plateau bladderpod Ward’s bladderpodWard’s Native Pres Jun–Jul x x x smooth yellowcress Native Rep Jun–Sep x Barneby’s reed-mustardBarneby’s Native Pres May x Chinle

var. purshii

var. glabra nana var. var. var. var. var. cinale Caulanthus crassicaulis glaber Descurainia californica Draba aurea Draba stenoloba Erysimum asperum Erysimum inconspicuum Lepidium montanum jonesii Nasturtium ofﬁ Physaria acutifolia acutifolia Physaria acutifolia purpurea Physaria intermedia Physaria ludoviciana Physaria newberryi Physaria rectipes Physaria wardii Rorippa islandica Schoenocrambe barnebyi rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyPerennial Forbs, cont. Cruciferae (Brassicaceae) Species name Common name Nativity Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

146 Annotated Checklist of Vascular Flora: Capitol Reef National Park seleniferous soils Tropic, Dakota, Tropic, Morrison, Chinle, & Moenkopi F Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres Apr–Sep x x x x ower Native Pres Apr–Jul x x x x ax-leaved plainsmustard ax-leaved Native Pres Apr–Jul x x x x ﬂ smoothleaf prince’s- plume green prince’s-plume Native Pres May–Aug x x x prince’s-plume Native Pres Apr–Sep x x x Often on heart-leaf jewelﬂ candelabra thelypody Native Pres Apr–Jun x x Mancos, entire-leaf thelypody Native Pres Jun–Sep x Fendler’s spurgeFendler’s Native Pres May–Sep x x x x shorthorn spurge Native Pres May–Jun x x x x noseburn Native Rep May–Aug x x x golden corydalis Native Pres Apr–Sep x x x x Great Basin centaury Native Pres Jun–Sep x Rocky Mountain gentian Native Pres Jul–Sep x x Engelmann’s gentianEngelmann’s Native Pres Jun–Aug x x white-margined swertia Native Pres May–Jul x x x elkweed Native Pres Jun–Aug x x small-leaf geranium Native Pres Jun–Aug x x x Rchardson’s crane’s-billRchardson’s Native Rep Jun–Aug x common mare’s-tail Native Pres Jun–Sep x

var.

var. nis afﬁ

var. var. ora nis Schoenocrambe linifolia Stanleya integrifolia Stanleya pinnata Stanleya viridifl Streptanthus cordatus Thelypodiopsis divaricata Thelypodium integrifolium integrifolium Chamaesyce fendleri Euphorbia brachycera Tragia ramosa Tragia Corydalis aurea occidentalis Centaurium exaltatum Gentiana affi Gentianella heterosepala Swertia albomarginata Swertia radiata Geranium caespitosum marginale Geranium richardsonii Hippuris vulgaris rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Cruciferae (Brassicaceae) Species name Common name Nativity Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Euphorbiaceae Euphorbiaceae Euphorbiaceae Fumariaceae Gentianaceae Gentianaceae Gentianaceae Gentianaceae Gentianaceae Geraniaceae Geraniaceae Hippuridaceae

Appendices 147 Summerville, Moenkopi F areas Wet Comments con Mix Sage PJ des Salt Flower time Park status eld mint eld Native Pres Jun–Sep x eld milkvetch eld Native Pres May–Aug x x Rocky Mountain iris Native Pres May–Jul x San Rafael phacelia Native Pres Apr–Jun x x Carmel, Chinle, blue-eyed grass Native Pres Apr–Aug x mock pennyroyal Native Pres May–Aug x x rough bugleweed Native Pres Jun–Aug x ﬁ common horehound Intro Pres May–Oct Disturbed peppermint Intro Pres Jul–Sep x spearmint Intro Pres Jun–Aug x Cultivated stinking horsemint Native Rep May–Aug x x x catnip Intro Pres Jun–Sep x x annual sage Native Pres Jun–Sep x x x x little skullcap Native Pres Jun–Aug x hairy hedge-nettle Native Pres Jun–Aug x ﬁ crescent milkvetch Native Pres Mar–Jun x x Sheldon’s milkvetchSheldon’s Native Pres Mar–Jun x x x milkweed milkvetch Native Pres May–Jun x Barneby’s milkvetchBarneby’s Native Pres Apr–Jun x x Brandegee’s milkvetchBrandegee’s Native Pres May–Sep x x x Torrey’s milkvetchTorrey’s Native Pres Apr–Jul x x x x painted milkvetch Native Pres Apr–Jun x x x x Sandy soils

var. var. var. var. glabrata pilosa var. var. exa Iris missouriensis Phacelia rafaelensis Sisyrinchium demissum Hedeoma drummondii Lycopus asper Lycopus Marrubium vulgare Mentha arvensis Mentha piperita Mentha spicata Monardella odoratissima Nepeta cataria Salvia reﬂ Scutellaria galericulata Stachys palustris Astragalus agrestis Astragalus amphioxys amphioxys Astragalus amphioxys vespertinus Astragalus asclepiadoides Astragalus barnebyi Astragalus brandegei Astragalus calycosus calycosus Astragalus ceramicus ceramicus rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Iridaceae Appendix D. Conﬁ FamilyPerennial Forbs, cont. Hydrophyllaceae Species name Common name Nativity Iridaceae Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

148 Annotated Checklist of Vascular Flora: Capitol Reef National Park Wingate, & Chinle F Wet Comments con Mix Sage PJ des Salt Flower time Park status Colton’s milkvetchColton’s Native Pres Apr–Jun x x x Bicknell milkvetch Native Pres May–Jul x x lesser rushy milkvetch Native Pres May–Aug x x x x rockloving milkvetch Native Pres May–Jul x x x bishop’s milkvetchbishop’s Native Pres May–Jun x x clay milkvetch Native Pres Apr–Jul x yellow milkvetch Native Pres Apr–Jul x x Harrison’s milkvetchHarrison’s Native Pres Apr–Jun x Kayenta, tall kentrophyta Native Pres Jun–Sep x x x x laccolite milkvetch Native Pres Apr–Jun x x x straggling milkvetch Native Pres Apr–Jun x x x great rushy milkvetch Native Pres May–Jul x x Kaiparowits milkvetch Native Pres Apr–Jun x x x Moenkopi milkvetch Native Pres May–Jul x x x woolly milkvetch Native Pres Feb–Jun x x x Ferron milkvetch Native Pres Apr–Jun x x x panther milkvetch Native Pres Apr–Jun x x

var. var. var. var. coltonii var. avus argillosus fl var. var. var. avus avus Astragalus coltonii Astragalus consobrinus Astragalus convallarius convallarius Astragalus desperatus petrophilus Astragalus episcopus Astragalus fl Astragalus fl Astragalus harrisonii Astragalus kentrophyta elatus Astragalus laccoliticus Astragalus lentiginosus palans Astragalus lonchocarpus Astragalus malacoides Astragalus moencoppensis Astragalus mollissimus thompsoniae Astragalus musiniensis Astragalus pardalinus rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Species name Common name Nativity Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

Appendices 149 areas Wet Comments con Mix Sage PJ des Salt Flower time Park status Patterson’s milkvetchPatterson’s Native Pres May–Jul x x x stinking milkvetch Native Pres Apr–Jul x x sandstone milkvetch Native Pres May–Aug x x x pulse milkvetch Native Pres May–Aug x x x Ward’s milkvetchWard’s Native Rep May–Sep x x x x x Woodruff’s milkvetchWoodruff’s Native Pres May–Jun x Entrada Zion milkvetch Native Pres Apr–Jun x x x x Kanab prairie-clover Native Pres May–Jul x x x Sandy soils white prairie-clover Native Pres May–Sep x x x x Sandy soils Nuttall’s licoriceNuttall’s Native Pres May–Jul x northern sweetvetch Native Pres May–Aug x x x Zion sweetpea Native Pres Apr–Jun x x x Sandy soils silvery lupine Native Pres May–Aug x x x x subalpine lupine Native Pres May–Aug x x Utah lupine Native Pres May–Aug x x x silky lupine Native Pres May–Aug x x x alfalfa Intro Pres May–Sep Disturbed

zionis

var.

avescens utahensis var. zionis i fl sericeus var. var. var. orus var. var. var. var. avescens Astragalus pattersonii Astragalus praelongus praelongus Astragalus sesquifl Astragalus tenellus Astragalus wardii Astragalus woodruffi Astragalus zionis Dalea fl Dalea oligophylla Glycyrrhiza lepidota Hedysarum boreale boreale Lathyrus brachycalyx argenteus Lupinus argenteus rubricaulis Lupinus argenteus Lupinus caudatus Lupinus sericeus Medicago sativa rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Species name Common name Nativity Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

150 Annotated Checklist of Vascular Flora: Capitol Reef National Park areas areas areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status southern locoweed Native Pres Jun–Aug x x x yellow sweet-clover Intro Pres May–Oct Disturbed nodding locoweed Native Pres Jun–Aug x x x Lambert’s locoweedLambert’s Native Pres Apr–Sep x x Bicknell locoweed Native Pres May–Aug x x Thompson’s peteriaThompson’s Native Pres Apr–Jun x x x rush scurfpea Native Pres May–Jul x x lemon scurf-pea Native Pres Apr–Aug x slenderleaf scurfpea Native Pres Apr–Aug x Sandy soils montane golden pea Native Pres May–Aug x x strawberry cloverred clover Introwhite clover PresAmerican vetch May–Sep Intro Intro Pres Native Pres Pres May–Oct Apr–Sep May–Aug x x Disturbed x x x Disturbed x Disturbed Chinese wisteria Intro Pres May Cultivated common bladderwort Native Pres Jul–Aug x nodding onion Native Pres Jun–Jul x x San Juan onion Native Pres May–Jun x x Nevada onion Native Pres Apr–Jul x x x

var. var.

australis sericea var. americana var. var. var. var. exa cinalis Oxytropis borealis Melilotus offi Oxytropis defl Oxytropis lambertii bigelovii Oxytropis oreophila juniperina Peteria thompsoniae Psoralidium junceum Psoralidium lanceolatum lanceolatum Psoralidium lanceolatum stenophyllum Thermopsis montana Trifolium fragiferum Trifolium Trifolium pratense Trifolium repens Trifolium Vicia americana Wisteria sinensis Utricularia vulgaris Allium cernuum Allium macropetalum Allium nevadense rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Leguminosae (Fabaceae) Appendix D. Confi FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Species name Common name Nativity Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Lentibulariaceae Liliaceae (Alliaceae) Liliaceae (Alliaceae) Liliaceae (Alliaceae)

Appendices 151 seleniferous soils areas Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres Apr–Jun x x x x Native Pres Mar–Oct x x x x Native Pres Jun–Aug x x x ax Native Pres Apr–Aug x x x pink funnel-lilyasparagus Nativegolden mariposa Pres Apr–May Native Intro x Pres Pres x May–Jun x Jun–Jul x x x Saline to x Disturbed sego-lily Native Pres Jun–Jul x x x x sand lily Native Pres Apr–Jun x x leopard-lily Native Pres Apr–Jul x x x x starry false Solomon’s- seal mountain death camas Native Pres Jun–Aug x x blue fl Colorado stickleaf Native Pres Apr–Oct x x x desert stickleaf Native Pres Apr–Sep x x wing-seed stickleaf Native Pres Mar–Aug x x x scarlet globemallow Native Pres Apr–Oct x x x x gooseberry-leaf globemallow small-leaf globemallow Native Pres Mar–Oct x x x woodland pinedrops Native Pres Jun–Sep x x fragrant sand-verbena Native Pres Apr–Sep x x x x low sand-verbena Native Pres Apr–Sep x x x trailing four-o’clock Native Pres Apr–Sep x x New Mexico umbrellawort narrow-leaf four-o’clock Native Pres May–Aug x x x x x orum decipiens linearis lewisii var. var. ora ssp. cinalis Androstephium brevifl Asparagus offi Calochortus aureus Calochortus nuttallii Eremocrinum albomarginatum Fritillaria atropurpurea Smilacina stellata Zigadenus elegans Linum perenne Mentzelia marginata Mentzelia multifl Mentzelia pterosperma Sphaeralcea coccinea Sphaeralcea grossulariifolia Sphaeralcea parvifolia Pterospora andromedea Abronia fragrans Abronia nana Allionia incarnata Mirabilis linearis Mirabilis linearis rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Liliaceae (Themidaceae) Species name Common name Nativity Liliaceae (Asparagaceae) Liliaceae (Calochortaceae) Liliaceae (Calochortaceae) Liliaceae (Anthericaceae) Liliaceae Liliaceae (Convallariaceae or Ruscaceae) Liliaceae (Melanthiaceae) Linaceae Loasaceae Loasaceae Loasaceae Malvaceae Malvaceae Malvaceae Monotropaceae (Ericaceae) Nyctaginaceae Nyctaginaceae Nyctaginaceae Nyctaginaceae Nyctaginaceae

152 Annotated Checklist of Vascular Flora: Capitol Reef National Park Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres Apr–Jun x x x Native Pres Jun–Aug x x x reweed Native Rep Jun–Sep x x armed prickly-poppy Native Rep May–Sep x x x showy four-o’clock Native Pres Apr–Jul x x English plantain Intro Pres May–Aug x spreading four-o’clock Native Pres May–Sep x x lavender-leaf sundropslavender-leaf Native Pres May–Aug x x x x ﬁ northern willow-herb Native Pres Jun–Aug x Hornemann’s willow-herbHornemann’s Native Pres Jun–Aug x long-tube evening- primrose Paria evening-primrose Native Pres Apr–Jun x x x hairy-throat evening- primrose western evening-primrose Native Rep Jun–Aug x x x x bronze evening-primrose Native Pres May–Jul x x x x Bridges evening-primrose Native Pres Jul–Sep x pale evening-primrose Native Pres May–Jul x x x x spotted coral-root Native Rep Jun–Jul x giant helleborine Native Pres Apr–Aug x northern bog orchid Native Pres Jun–Aug x slender bog orchid Native Pres Jun–Jul x alcove bog-orchid Native Pres Jul–Aug x Ute ladies’-tresses Native Ext Jul–Sep x clustered broomrape Native Pres Jun–Jul x x x x Cooper’s broomrapeCooper’s Native Pres May–Jul x x x

var.

var. var. var.

pallida var. var. var. ana var. var. ora ora ora Mirabilis multifl Plantago lanceolata Mirabilis oxybaphoides Calylophus lavandulifolius Epilobium angustifolium canescens Epilobium ciliatum hornemannii Epilobium hornemannii Oenothera caespitosa marginata Oenothera caespitosa navajoensis Oenothera coronopifolia Oenothera elata Oenothera howardii Oenothera longissima Oenothera pallida Corallorhiza maculata Epipactis gigantea Habenaria hyperborea Habenaria sparsifl sparsifl Habenaria zothecina Spiranthes romanzoffi diluvialis Orobanche fasciculata Orobanche ludoviciana cooperi Argemone munita rotundata rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Nyctaginaceae Species name Common name Nativity Plantaginaceae Nyctaginaceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Orchidaceae Orchidaceae Orchidaceae Orchidaceae Orchidaceae Orchidaceae Orobanchaceae Orobanchaceae Papaveraceae

Appendices 153 areas Sandstone & Carmel F gravelly soils Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres May–Aug x x x Native Pres May–Jul x x owered gilia owered Native Pres May–Aug x x x Rabbit Valley giliaRabbit Valley carmine gilia Native Pres Jun–Jul Native Pres May–Jun x x x x x Navajo common plantain Intro Pres May–Aug x Disturbed Mussentuchit gilia Native Pres May–Jun x x x scarlet gilia Native Pres May–Aug x x x x shrubby gilia Native Pres Jun–Sep x x x x Sandy soil long-ﬂ San Rafael gilia Native Pres May–Jun x x x matted prickly-phlox Native Pres May–Jul x x x Watson’s prickly-phloxWatson’s Native Pres May–Aug x x x x desert phlox Native Pres Apr–Jun x x x x carpet phlox Native Pres Apr–Jun x x x x long-leaf phlox Native Pres Apr–Jul x x x x cushion phlox Native Rep Jun–Aug x x cushion milkwort Native Pres May–Jun x x x x winged wild buckwheat Native Pres Jun–Oct x x Sandy soils pretty wild buckwheat Native Pres Apr–Jul x x x desert trumpetslender wild buckwheat Native Native Pres Pres Jun–Oct Mar–Oct x x x x x x Sandy or cushion wild buckwheat Native Pres Apr–Aug x x x redroot wild buckwheat Native Pres Jun–Oct x x x Shockley’s wild Shockley’s buckwheat Woodside wild Woodside buckwheat

var.

var. var. var. var. canescens ora atum var. atum Gilia caespitosa Gilia subnuda Plantago major Gilia tenuis Ipomopsis aggregata aggregata Ipomopsis congesta frutescens Ipomopsis longifl Ipomopsis roseata Leptodactylon caespitosum Leptodactylon watsonii Phlox austromontana austromontana Phlox hoodii Phlox longifolia Phlox pulvinata Polygala subspinosa Eriogonum alatum Eriogonum bicolor Eriogonum infl Eriogonum microthecum simpsonii infl Eriogonum ovalifolium Eriogonum racemosum racemosum Eriogonum shockleyi Eriogonum tumulosum rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Polemoniaceae Polemoniaceae Appendix D. Confi FamilyPerennial Forbs, cont. Plantaginaceae Species name Common name Nativity Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polygalaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae

154 Annotated Checklist of Vascular Flora: Capitol Reef National Park areas areas Cedar Mesa sandstone Wet Comments con Mix Sage PJ des Salt Flower time Park status ower meadow-rue ower Native Rep Jun–Aug x x arid wild buckwheat Native Pres Jun–Oct x x x curly dock Intro Pres May–Sep x Disturbed canaigre dock Native Pres Apr–Jun x x sausageleaf talinum Nativeslenderleaf pondweed Pres Native Jun–Oct Pres Jul–Sep x Navajo & x willow dock Native Pres May–Sep x x x x Disturbed sea-milkwort Native Pres May–Jul x cave primrose Native Pres Feb–Jul x Columbian monkshood Native Rep Jun–Aug x x red baneberry Native Rep May–Aug x x Colorado columbine Native Pres Jun–Aug x x x alcove columbine Native Pres May–Jun x Columbian virgin’s-bower Native Pres May–Sep x x white virgin’s-bower Native Pres Jul–Aug x pale larkspur Native Pres Apr–Jun x x x marsh buttercup Native Pres May–Sep x Fendler’s meadow-rueFendler’s Native Pres Jun–Aug x x few-ﬂ Virginia strawberry Native Rep Jun–Jul x x rock spiraea Native Pres Aug–Sep x x x Rock crevices

var. var. var. var.

ssp. orum liformis Eriogonum umbellatum subaridum Rumex crispus Rumex hymenosepalus Talinum brevifolium Talinum Potamogeton fi occidentalis Rumex salicifolius triangulivalvis Glaux maritima Primula specuicola Aconitum columbianum columbianum Actaea rubra Aquilegia corulea Aquilegia micrantha Clematis columbiana Clematis ligusticifolia Delphinium andersonii scaposum Ranunculus cymbalaria Thalictrum fendleri Thalictrum sparsifl Fragaria virginiana Petrophytum caespitosum rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Polygonaceae Species name Common name Nativity Polygonaceae Polygonaceae Portulacaceae Potamogetonaceae Polygonaceae Primulaceae (Myrsinaceae) Primulaceae Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae (Thalictraceae) Ranunculaceae (Thalictraceae) Rosaceae Rosaceae

Appendices 155 Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres May–Sep x Sandstone ower Native Pres Jun–Aug x ower Native Pres Jun–Aug x ax Native Pres Apr–Aug x x x x eshy penstemon eshy Native Pres May–Jul x x x ower beardlip penstemon Native Pres Jun–Jul x Sandy soils silverweed Native Rep May–Aug x painted penstemon Native Pres May–Jul x x x Claron F bicrenate cinquefoil Native Pres May–Jun x x x fl hair-tuft cinquefoilhair-tuft Native Pres Jul–Aug x x x beautiful cinquefoil Native Pres Jun–Aug x x woolly cinquefoil Native Rep Jul–Aug x Pennsylvania cinquefoil Native Pres Jul–Aug x x shrubby bedstraw Native Pres May–Aug x x x x x bastard toadfl littleleaf alumroot Native Pres May–Sep x x x red alumroot Native Rep Apr–Sep x x x marsh grass-of-Parnassus Native Pres May–Oct x desert paintbrush Native Pres Apr–Jun x x x Wyoming paintbrush Native Pres Jul–Aug x x x Eastwood’s paintbrushEastwood’s Native Pres May–Jul x x x x Sandstone Eastwood’s monkey- Eastwood’s fl glabrous monkeyfl yellow monkeyfl dwarf lousewort Native Pres Apr–Jun x x bush penstemon Native Pres May–Aug x x x x veined penstemon Native Pres May–Jun x x x

var.

var. scabrida var.

var. var. var. var. var. var. var. orum orum Potentilla anserina Penstemon caespitosus desertipicti Potentilla concinna bicrenata Penstemon carnosus Potentilla crinita Potentilla gracilis pulcherrima Potentilla hippiana Potentilla pensylvanica Galium multifl multifl pallida Comandra umbellata Heuchera parvifolia Heuchera rubescens Parnassia palustris montanensis Castilleja chromosa Castilleja linariifolia Castilleja scabrida Mimulus eastwoodiae Mimulus glabratus utahensis Mimulus guttatus Pedicularis centranthera Penstemon ambiguus laevissimus Penstemon angustifolius venosus barbatus Penstemon barbatus rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Forbs, cont. Rosaceae Species name Common name Nativity Scrophulariaceae Rosaceae Scrophulariaceae Rosaceae Rosaceae Rosaceae Rosaceae Rubiaceae Santalaceae Saxifragaceae Saxifragaceae Saxifragaceae (Parnassiaceae) Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae

156 Annotated Checklist of Vascular Flora: Capitol Reef National Park areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres Jun–Aug x x x recracker Native Pres May–Jul x x x x dusty penstemon Native Pres Jun–Aug x x x Eaton’s ﬁ Eaton’s undulate penstemon Native Pres May–Jul x x x x Moffatt penstemonMoffatt Native Pres May–Jun x x x Loa penstemon Native Pres May–Jun x x x Rockville penstemon Native Pres May–Jul x x x x Palmer’s penstemonPalmer’s Native Pres May–Aug x x x beaked penstemon Native Pres Jun–Aug x x x x x plateau penstemon Native Pres May–Jul x x x Rocky Mountain penstemon Utah penstemon Native Pres Apr–Jun x x x woolly mullein Intro Rep Jun–Aug x x x Disturbed American brooklime Native Pres Jul–Aug x water speedwell Intro Pres Jun–Aug x long-leaf ground-cherry Native Pres Apr–Aug x x x silverleaf nightshade Intro Pres Apr–Aug x x Disturbed water-hemlock Native Pres Jun–Sep x Fendler’s cymopterusFendler’s Native Pres Apr–May x x x Sandy soils Beck’s spring-parsleyBeck’s Native Pres Apr–Jun x x onion spring-parsley Native Pres Apr–May x x x Lemmon’s spring-parsleyLemmon’s Native Pres Jul–Aug x

var.

eatonii var.

var. var. var. var. orus var. Penstemon comarrhenus Penstemon eatonii Penstemon eatonii undosus Penstemon moffatii Penstemon ophianthus Penstemon pachyphyllus congestus eglandulosus Penstemon palmeri Penstemon rostriﬂ Penstemon scariosus scariosus Penstemon strictus Penstemon utahensis Verbascum thapsus Verbascum Veronica americana Veronica Veronica anagallis-aquatica Veronica Physalis longifolia Solanum elaeagnifolium Cicuta maculata angustifolia fendleri Cymopterus acaulis Cymopterus beckii Cymopterus bulbosus Cymopterus lemmonii rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyPerennial Forbs, cont. Scrophulariaceae Species name Common name Nativity Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Solanaceae Solanaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae)

Appendices 157 clay soils clay soils areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres Apr–Jun x x x x Sandy to heavy Native Pres Mar–May x x x x Sandy to heavy Native Pres Mar–May x Native Pres May–Jun x r dwarf- r at-seed lomatium at-seed Native Pres May–Jul x x x Colorado Plateau spring- parsley southern ligusticum Native Rep Jun–Aug x x basin white-cup spring- parsley rush lomatium Native Pres May x x x x Parry’s lomatiumParry’s ﬂ Native Pres Apr–Jun x x x x blunt sweet-cicely Native Rep Jun–Jul x x common yampah Native Rep Jul–Aug x x x water parsnip Native Rep Jul–Aug x stinging nettle Native Pres Jul–Aug x x x prostrate vervain Native Pres May–Sep x x x Disturbed hook-spur violet Native Pres Mar–Aug x northern bog violet Native Pres Apr–Jul x pinyon dwarf-mistletoe Native Pres Aug–Sep x Douglas-ﬁ mistletoe southwestern dwarf- mistletoe juniper mistletoe Native Pres Jul–Sep x horned pondweed Native Pres Mar–Nov x tumble pigweed Intro Pres Jun–Oct Disturbed

var.

var. ssp. var. occidentalis var. Cymopterus purpureus purpureus Ligusticum porteri Cymopterus purpurascens Lomatium junceum Lomatium parryi platycarpum Lomatium triternatum Osmorhiza depauperata borealis Perideridia montana Sium suave Urtica dioica Verbena bracteata Verbena Viola adunca Viola nephrophylla Arceuthobium divaricatum Arceuthobium douglasii Arceuthobium vaginatum cryptopodum Phoradendron juniperinum Zannichellia palustris Amaranthus albus rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Umbelliferae (Apiaceae) Species name Common name Nativity Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Urticaceae Verbenaceae Violaceae Violaceae Viscaceae Viscaceae Viscaceae Viscaceae Zannichelliaceae Annual Forbs Amaranthaceae

158 Annotated Checklist of Vascular Flora: Capitol Reef National Park areas areas areas areas Morrison, & F Tropic Tropic F. Tropic Morrison F areas Wet Comments con Mix Sage PJ des Salt Flower time Park status y Native Pres May–Aug x x ve-horn smotherweed ve-horn Intro Pres Aug–Oct x x Disturbed redroot pigweedcushion cryptanth Intro Native Pres Pres Jun–Sep Apr–Jul x x x Disturbed prostrate pigweed Native Pres Jun–Sep Disturbed thick-sepaled cryptanth Native Pres Apr–Jun x x x sand dune cryptanth Native Pres May–Jul x x x slender cryptanth Native Pres Apr–Jul x x x redroot cryptanth Native Pres Mar–May x x x wing-nut cryptanth Native Pres Apr–Jul x x x recurved cryptanth Native Pres Apr–Jun x x x bindweed heliotrope Native Pres May–Sep x x Sandy soils yellow beeplant Native Pres Apr–Sep x x x x cupseed stickseed Native Pres Apr–Jun x x x x Disturbed sleepy catchﬂ Rocky Mountain beeplant Native stinkweedPalmer’s Pres May–Oct Native Pres Apr–Oct x x Disturbed Mancos, silver orach Native Rep Jun–Sep x Blue Valley orachBlue Valley orachPowell’s Native Pres Native Jun–Aug Pres x Jun–Sep x x x x Mancos & thinleaf orach Native Pres Jun–Sep x stalked orachwedge orach Native Pres Native Aug–Sep Pres x Jun–Sep x x Mancos & x ﬁ

var. var. var. argentea var. powellii saccaria exus var. var. var. ora Amaranthus retrofl Cryptantha circumscissa Amaranthus blitoides Cryptantha crassisepala elachantha Cryptantha fendleri Cryptantha gracilis Cryptantha micrantha pterocarya Cryptantha pterocarya Cryptantha recurvata Heliotropium convolvulaceum Cleome lutea Lappula occidentalis cupulata Cleomella palmeriana Silene antirrhina Cleome serrulata palmeriana Atriplex argentea Atriplex gracilifl Atriplex powellii Atriplex prostrata Atriplex saccaria Atriplex truncata Bassia hyssopifolia rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Amaranthaceae Boraginaceae Appendix D. Confi FamilyAnnual Forbs, cont. Amaranthaceae Species name Common name Nativity Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Capparaceae (Cleomaceae) Boraginaceae Capparaceae (Cleomaceae) Caryophyllaceae Capparaceae (Cleomaceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae

Appendices 159 areas areas areas areas areas areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status lambsquartermountain goosefoot Native Intro Pres Pres Jul–Sep Jun–Sep x x x Disturbed summer-cypress Intro Pres Aug–Oct x x x Disturbed desert goosefoot Native Pres Jun–Sep x x x x silvery goosefoot Native Pres Jun–Aug x x x oakleaf goosefoot Native Pres Aug–Sep x New Mexico goosefoot Nativenarrow-leaf goosefoot Pres Native Jul–Sep Pres Jul–Sep x x x x Disturbed American bugseed Native Pres Aug–Oct x x Sandy soils halogeton poverty-weedNuttall’s Native Pres Intro Apr–Jul Pres x Jul–Aug x x x x x Disturbed barbwire Russian thistle Intro Pres Jun–Sep Disturbed Russian thistlebroom seepweed Intro Native Pres Pres Jun–Sep Jul–Sep x Disturbed Torrey’s seepweedTorrey’s Native Pres Jun–Sep x x bur ragweed Native Pres Jul–Oct x x x x Disturbed rayless alkali aster Native Pres Aug–Sep x leafy aster Native Pres Aug–Oct x stevia dusty-maiden Native Pres Apr–Jul x x x Canadian horseweed Native Pres Jul–Sep x

var.

var. var.

var. var. var. Chenopodium album album Chenopodium atrovirens Bassia scoparia Chenopodium desiccatum incanum Chenopodium fremontii salinum Chenopodium glaucum Chenopodium graveolens neomexicanum Chenopodium leptophyllum Corispermum americanum Halogeton glomeratus Monolepis nuttalliana Salsola paulsenii Salsola tragus Suaeda calceoliformis Suaeda torreyana torreyana Ambrosia acanthicarpa Aster brachyactis Aster frondosus Chaenactis stevioides Conyza canadensis glabrata rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Chenopodiaceae Chenopodiaceae Appendix D. Conﬁ FamilyAnnual Forbs, cont. Chenopodiaceae Species name Common name Nativity Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

160 Annotated Checklist of Vascular Flora: Capitol Reef National Park areas areas areas areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres Apr–Jun x x x ower Native Pres Jun–Sep x x Disturbed ower Native Pres Jun–Sep x x x x ower Native Rep May–Sep x x x liform desert-dandelion liform Native Pres Apr–Jul x x x Brandegee’s sandplantBrandegee’s Native Pres Aug–Oct x pappose glandweed Native Pres Aug–Sep Sandy areas pretty daisy Native Pres May–Aug x x x crustweed Native Pres May–Jun x x sand sunfl common sunfl prairie sunfl marsh-elderprickly lettucethorny wirelettuce Native Intro Pres Native Pres Pres Aug–Oct Jul–Sep Jul–Sep x x x x x x x Disturbed Disturbed tansyleaf aster Native Pres May–Sep x x x fi sowthistle desert- dandelion spiny-leaf sow-thistlewhite-plume wirelettuce Intro Native Pres Pres Jul–Oct May–Sep x x x x Disturbed annual townsendia Native Pres Apr–Jun x x rough cocklebur Native Pres Apr–Oct x Disturbed

fallax

var. ssp. var. Dicoria brandegei Dyssodia papposa Erigeron bellidiastrum Glyptopleura marginata Helianthus anomalus Helianthus annuus lenticularis Helianthus petiolaris Iva xanthifolia Lactuca serriola spinosa Lygodesmia Machaeranthera tanacetifolia Malacothrix glabrata Malacothrix sonchoides Sonchus asper Stephanomeria exigua Townsendia annua Townsendia Xanthium strumarium canadense rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyAnnual Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Appendices 161 areas areas areas areas areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status eld dodder eld Native Pres Jul–Sep x x x ixweed Intro Pres Mar–Aug x x x Disturbed ﬁ European madwort Intro Pres Apr–Jul x x shepherd’s-purse Intro Pres Apr–Sep Disturbed blue mustardmountain tansy-mustard Native Pres Intro Apr–Jul Pres Apr–Jun x x Disturbed western tansy-mustard Native Pres Mar–Aug x x western tansy-mustard Native Pres Mar–Aug x x x x spectacle-pod Native Pres Apr–Sep x x x x Sandy soils ﬂ wedgeleaf draba Native Pres Feb–Jul x x x x dwarf draba Native Rep Apr–Jul x x x ovalpurse Native Pres Apr–Jun x prairie pepperwort Native Pres May–Jul x x x x prairie pepperwort Native Pres May–Jul x x x x African mustardlow yellowcress Intro Pres Native Pres Apr–Jul Jun–Sep x Disturbed clasping pepperwort Intro Pres May–Jul x x x Disturbed tumble mustard Intro Pres May–Aug Disturbed

sonnei var. var. cuneifolia var. var. var. micranthum var. orum orum var. orum Cuscuta pentagona Alyssum minus Capsella bursa-pastoris Chorispora tenella Descurainia incana Descurainia pinnata intermedia Descurainia pinnata osmiarum Dithyrea wislizenii Descurainia sophia Draba cuneifolia Draba reptans Hutchinsia procumbens Lepidium densifl densifl Lepidium densifl ramosum Malcolmia africana Rorippa tenerrima Lepidium perfoliatum Sisymbrium altissimum rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyAnnual Forbs, cont. Convolvulaceae Species name Common name Nativity Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

162 Annotated Checklist of Vascular Flora: Capitol Reef National Park clayey soils areas areas Mancos, Morrison, & F. Tropic areas areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status ddle-mustard Native Pres Mar–Jun x x xor Sandy ax Native Pres Apr–Jun x x x xsoils Sandy ax Native Pres May–Jun x x owered milkvetch owered Native Pres Apr–Jun x x x ower sandpuffs ower Native Pres Apr–Sep x x long-beak fi ridge-seeded spurge spurgeParry’s Native Pres Jun–Aug Native x Pres x May–Sep x x x x x Disturbed stork’s-bill Intro Pres Feb–Oct x x x Disturbed Howell’s namaHowell’s Native Rep May–Aug x x corrugate phacelia Native Pres May–Jun x x x Ives’ phacelia Native Pres Apr–Jun x x x brittle phacelia Native Pres May–Jun x x xDakota, Chinle, small-fl gravel milkvetch Native Pres Feb–Jul x x x Great Basin lupine Native Pres Apr–Jul x x x dwarf lupine Native Pres Apr–Jul x x x Sandy soils black medickwhite sweet-cloverbroom-fl Intro Intro Pres Pres May–Oct May–Oct x x Disturbed Disturbed downy fl white-stem blazingstar Native Pres Apr–Aug x x x x common mallow Intro Pres Apr–Oct Disturbed small-fl camissoniaEastwood’s Native Pres Apr–Jun x x

var.

demissa

pusillus var. var. var. var. Streptanthella longirostris Chamaesyce glyptosperma Chamaesyce parryi Erodium cicutarium Nama retrorsum corrugata Phacelia crenulata Phacelia ivesiana Phacelia demissa Astragalus sabulonum micranthiformis Astragalus nuttallianus Lupinus pusillus intermontanus Lupinus pusillus Medicago lupulina Melilotus alba Linum aristatum Linum puberulum Mentzelia albicaulis Malva neglecta Camissonia eastwoodiae Tripterocalyx micranthus Tripterocalyx rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyAnnual Forbs, cont. Cruciferae (Brassicaceae) Species name Common name Nativity Euphorbiaceae Euphorbiaceae Geraniaceae Hydrophyllaceae Hydrophyllaceae Hydrophyllaceae Hydrophyllaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Linaceae Linaceae Loasaceae Malvaceae Onagraceae Nyctaginaceae

Appendices 163 areas Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres Mar–Jun x x Native Pres Apr–Oct x x Native Pres Apr–Jul x Clay soils Native Pres Apr–Oct x x owered gaura owered Native Pres May–Aug x ower eriastrum ower Native Pres May–Aug x x x Paiute suncup Native Pres May–Jun x x x Walker’s camissoniaWalker’s Native Pres Apr–Jun x x x small-ﬂ branching groundsmoke Native Pres May–Aug x x x white-stem evening- primrose woolly plantain Native Pres Mar–Aug x x x few-ﬂ cataract gilia Native Pres Jun–Sep x shy gilia Native Pres Apr–Jul x x x common gilia Native Pres May–Jun x x x x common gilia Native Pres May–Jun x x x Gunnison’s giliaGunnison’s Native Pres May–Jun x x x Sandy soils spreading gilia Native Pres Apr–Jun x x x dwarf gilia Native Pres Apr–Jun x x x nodding wild buckwheat Native Pres Apr–Oct x x x x skeletonweed wild buckwheat bent wild buckwheat Native Pres Apr–Sep x x Gordon’s wild buckwheatGordon’s Native Pres May–Oct x x Hooker’s wild buckwheatHooker’s Native Pres Jun–Oct x x x x Grand Valley desert Grand Valley trumpet Palmer’s wild buckwheatPalmer’s Native Pres Mar–Oct x x x Wetherill’s wild Wetherill’s buckwheat yard knotweed Intro Pres May–Oct Disturbed

var. var. var.

var. var. orum exum atum exum ora scapoidea Camissonia scapoidea Camissonia walkeri Gaura parvifl Gayophytum ramosissimum Oenothera albicaulis Plantago patagonica Eriastrum sparsifl Gilia imperialis Gilia inconspicua Gilia leptomeria leptomeria Gilia leptomeria micromeria Ipomopsis gunnisonii Ipomopsis polycladon Ipomopsis pumila cernuum Eriogonum cernuum Eriogonum defl Eriogonum fl Eriogonum gordonii Eriogonum hookeri Eriogonum infl fusiforme Eriogonum palmerianum Eriogonum wetherillii Polygonum aviculare rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyAnnual Forbs, cont. Onagraceae Onagraceae Species name Common name Nativity Onagraceae Onagraceae Onagraceae Plantaginaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae

164 Annotated Checklist of Vascular Flora: Capitol Reef National Park areas areas areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status Native Pres May–Aug x x x ower rock- ower ower bird’s-beak ower Native Pres Aug–Sep x x x x Sawatch knotweed Native Pres Jun–Aug x x x willow-weedgolden dock Intro Native Pres Pres Jul–Nov May–Sep x x common purslanepygmy-ﬂ jasmine Intro Pres Jun–Sep Disturbed blister buttercup Native Pres May–Sep x bur buttercup Intro Pres Mar–Jun x x x Disturbed cleavers Native Pres Apr–Jun x x x x x annual paintbrush Native Pres Jun–Aug x smallﬂ bird’s-beakWright’s Native Pres Jul–Sep x x x x Sandy soils angel’s trumpetangel’s Native Pres Apr–Oct x x x coyote tobacco Native Pres Jun–Sep x x x desert tobacco Native Pres Mar–Oct x Rock crevices ground-cherry nightshade Introcut-leaf nightshade Pres May–Oct Native Pres May–Sep x x x Disturbed dill Intro Pres Jul–Aug puncture vine sedgeslender-beaked Intro Native Pres Pres May–Sep May–Aug x x Disturbed golden sedge Native Pres Apr–Aug x Douglas’ sedge Native Pres Apr–Aug x x x narrowleaf sedge Native Rep Jun–Jul x x threadleaf sedge Native Pres Apr–Jul x x x

var. var. fueginus orus

var. var. orum lifolia dus Polygonum douglasii johnstonii Polygonum lapathifolium Rumex maritimus Portulaca oleracea Androsace septentrionalis Ranunculus sceleratus multifi Ranunculus testiculatus Galium aparine echinospermon Castilleja exilis Cordylanthus parvifl Cordylanthus wrightii Datura wrightii Nicotiana attenuata Nicotiana trigonophylla Solanum sarrachoides Solanum trifl Anethum graveolens Tribulus terrestris Tribulus Carex athrostachya Carex aurea Carex douglasii Carex duriuscula Carex fi rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyAnnual Forbs, cont. Polygonaceae Species name Common name Nativity Polygonaceae Polygonaceae Portulacaceae Primulaceae Ranunculaceae Ranunculaceae Rubiaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Solanaceae Solanaceae Solanaceae Solanaceae Solanaceae Umbelliferae (Apiaceae) Zygophyllaceae Perennial Graminoids Perennial Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae

Appendices 165 areas Wet Comments con Mix Sage PJ des Salt Flower time Park status eld’s bluestem eld’s Native Pres Jun–Oct x x x ower spikerush ower Native Pres Jun–Aug x White Mountain sedge Native Pres May–Aug x x bottlebrush sedge Native Pres May–Jun x small-wing sedge Native Pres Jun–Aug x Nebraska sedge Native Pres May–Jul x western sedge Native Pres Jun–Aug x woolly sedge Native Pres May–Sep x blackcreeper sedge Native Pres May–Aug x Ross’ sedge Native Pres May–Aug x x x valley sedge Native Rep May–Jul x x slender spikerush Native Pres Jun–Sep x common spikerush Native Pres May–Aug x few-ﬂ hardstem bulrush Native Pres Jun–Aug x American bulrush Native Pres Jun–Sep x alkali bulrush Native Pres Jun–Sep x panicled bulrush Native Rep Jun–Aug x three-square bulrush Native Pres May–Aug x crested wheatgrass Intro Pres Jun–Aug x x x x Disturbed spike bentgrass Native Pres Jul–Aug x redtop Intro Pres Jun–Sep x shortawn foxtail Native Pres Jun–Aug x Hall’s bluestemHall’s Native Pres Jul–Sep x x x x big bluestem Native Pres Jul–Sep x x x bushy bluestem Native Pres Apr–Oct x purple three-awn Native Pres Mar–Sep x x x x Springﬁ sideoats grama Native Pres Jul–Sep x x x x

var. var. var. eldii

ora var. Carex geophila Carex hystericina Carex microptera Carex nebrascensis Carex occidentalis Carex pellita Carex praegracilis Carex rossii Carex vallicola Eleocharis acicularis Eleocharis palustris Eleocharis quinquefl Scirpus acutus Scirpus americanus Scirpus maritimus Scirpus microcarpus Scirpus pungens longispicatus Agropyron cristatum Agrostis exarata Agrostis stolonifera Alopecurus aequalis Andropogon gerardii chrysocomus Andropogon gerardii gerardii Andropogon glomeratus scabriglumis Aristida purpurea Bothriochloa springfi Bouteloua curtipendula rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Graminoids, cont. Cyperaceae Species name Common name Nativity Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

166 Annotated Checklist of Vascular Flora: Capitol Reef National Park areas areas areas areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status uffgrass Native Pres Apr–Aug x x x black grama Native Pres Jun–Aug x x x blue grama Native Pres Jul–Sep x x x x nodding brome Native Rep Jun–Aug x x x fringed brome Rep Jul–Sep x Jones’ reedgrass Native Pres Jul–Aug x x smooth brome Intro Pres Jun–Aug x x x Disturbed slimstem reedgrass Native Pres Jul–Aug x orchard grassdesert saltgrass Intro Native Pres Pres Jun–Aug Jun–Sep x x x x Disturbed Canada wildrye Native Pres Jul–Aug x squirreltail Native Pres May–Aug x x x x x thickspike wheatgrass Native Pres Jun–Aug x x x Russian wildrye Intro Pres Jun–Jul x x x x Disturbed Salina wildrye Native Pres Jun–Jul x x x quackgrass Intro Pres Jun–Aug x x Disturbed western wheatgrass Native Pres Jun–Aug x x x bluebunch wheatgrass Native Pres Jun–Aug x x x slender wheatgrass Native Pres Jun–Aug x x x Saunders’ wheatgrass Native Pres Jul–Aug x spike pappusgrass Native Pres Aug–Sep x x Slickrock hairy tridens Native Pres May–Jul x x x ﬂ tall fescueRocky Mountain fescue Native Pres Intro Jul–Aug Pres May–Jul x x x Disturbed ravine fescue Native Pres Aug–Sep x fowl mannagrass Native Pres Jul–Aug x galleta Native Pres May–Aug x x x inermis saximontana var. var. saundersii x Bouteloua eriopoda Bouteloua gracilis Bromus anomalus Bromus ciliatus Calamagrostis scopulorum Bromus inermis Calamagrostis stricta Dactylis glomerata Distichlis spicata Elymus canadensis Elymus elymoides Elymus lanceolatus Elymus junceus Elymus salinus Elymus repens Elymus smithii Elymus spicatus Elymus trachycaulus Elymus Enneapogon desvauxii Erioneuron pilosum Erioneuron pulchellum Festuca arundinacea Festuca ovina Festuca sororia Glyceria striata Hilaria jamesii rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

Appendices 167 areas areas areas areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status junegrass Native Pres Jun–Aug x x x foxtail barley Native Pres Jun–Aug x x x x x Disturbed wolftail Native Pres Jun–Aug x x perennial ryegrass Intro Pres Jun–Aug Disturbed foxtail muhly Native Pres Jul–Sep x scratchgrass Native Pres Jul–Aug x New Mexican muhly Native Pres Aug–Sep x x x x sandhill muhly Native Pres Jul–Sep x x green muhly Native Pres Jul–Sep x Thurber’s muhlyThurber’s Native Pres Jul–Sep x x x x littleseed ricegrass Native Pres Jun–Aug x x x bundle panicgrass Native Pres Jul–Sep x Sandy soils Scribner panicgrass Native Pres Jun–Sep x x switchgrass Native Pres Jun–Sep x x timothycommon reed Intro Native Pres Pres Jul–Sep Jun–Sep x x x x x Disturbed Canada bluegrass Intro Pres Jun–Aug x x x x muttongrass Native Pres May–Aug x x x x fowl bluegrass Native Pres May–Aug x Kentucky bluegrass bluegrassSandberg’s Intro Native Pres Pres May–Aug Apr–Sep x x x x x x x x x Disturbed x water polypogon Intro Pres Jul–Aug x weeping alkaligrass alkaligrassNuttall’s Intro Pres Native Pres Jun–Aug May–Aug x x x x x Disturbed tumble grass Native Pres Jun–Aug false melic Native Pres Jun–Aug x x

var. ora Koeleria macrantha Hordeum jubatum Lycurus phleoides Lycurus Lolium perenne Muhlenbergia andina Muhlenbergia asperifolia Muhlenbergia pauciﬂ Muhlenbergia pungens Muhlenbergia racemosa Muhlenbergia thurberi Oryzopsis micrantha Panicum acuminatum Panicum oligosanthes scribnerianum Panicum virgatum Phleum pratense Phragmites australis Poa compressa Poa fendleriana Poa palustris Poa pratensis Poa secunda Polypogon semiverticillatus Puccinellia distans Puccinellia nuttalliana Schedonnardus paniculatus Schizachne purpurascens rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Gramineae (Poaceae) Appendix D. Conﬁ FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

168 Annotated Checklist of Vascular Flora: Capitol Reef National Park Wet Comments con Mix Sage PJ des Salt Flower time Park status southern cattail Native Pres Jul–Aug x little bluestem Native Pres Jul–Sep x x x x alkali cordgrass Native Pres Jun–Sep x prairie wedgegrass Native Pres Jul–Sep x x alkali sacaton Native Pres Jun–Aug x x x x spike dropseed Native Pres Jun–Sep x x sand dropseed Native Pres Jun–Aug x x mesa dropseed Native Pres Jun–Sep x x x giant dropseed Native Pres Jun–Sep x Mormon needlegrass Native Pres May–Jul x x x needle-and-thread Native Pres May–Jul x x x x crested needlegrass Native Pres Jun–Jul x x x Indian ricegrass Native Pres May–Jul x x x x Letterman’s needlegrassLetterman’s Native Rep Jul–Sep x x x New Mexico feathergrass Native Pres Jun–Jul x x x pine needlegrass Native Pres Jun–Aug x x x Scribner needlegrass Native Pres Jun–Aug x desert needlegrass Native Pres Apr–Jun x x x Bloomer’s ricegrassBloomer’s Native Pres Jun–Aug x x x northern rush Native Pres Jul x Baltic rush Native Pres May–Jul x Tracy’s rushTracy’s Native Pres Jun–Aug x mountain rush Native Pres Jun–Aug x longstyle rush Native Pres Jun–Aug x poverty rush Native Pres Jun–Aug x Torrey’s rushTorrey’s Native Pres Jun–Aug x low arrowgrass Native Pres May–Aug x seaside arrowgrass Native Pres May–Aug x

var.

montanus var. parishii var. var. comata var. var. exuosus bloomeri x Typha domingensis Typha Schizachyrium scoparium Spartina gracilis Sphenopholis obtusata Sporobolus airoides airoides Sporobolus contractus Sporobolus cryptandrus Sporobolus ﬂ Sporobolus giganteus Stipa arida Stipa comata Stipa coronata Stipa hymenoides Stipa lettermanii Stipa neomexicana Stipa pinetorum Stipa scribneri Stipa speciosa Stipa Juncus alpinus Juncus arcticus brunnescens Juncus ensifolius Juncus ensifolius Juncus longistylus Juncus tenuis Juncus torreyi Triglochin concinnum Triglochin debilis Triglochin maritima Triglochin rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Conﬁ FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae Juncaginaceae Typhaceae

Appendices 169 areas areas areas areas areas areas areas areas areas areas Wet Comments con Mix Sage PJ des Salt Flower time Park status at-sedge Native Pres Jun–Aug x eld sandbur eld Native Pres Jul–Sep Disturbed broad-leaved cattail Native Pres Jun–Aug x bearded fl sixweeks three-awn Native Pres Mar–Sep x oatssixweeks grama Native Pres Intro Aug–Sep Rep x Jun–Aug x Disturbed Japanese chess Intro Pres Jun–Aug x x x x rescue grass Intro Pres Mar–Jun x Disturbed red brome Intro Pres Mar–Jun x cheatgrassfi barnyard-grass Introtufted lovegrass Pressix-weeks fescue Intro May–Jul Native Pres Pres x Native Aug–Sep Pres Jul–Aug x x Apr–Jun x x Disturbed x x x Disturbed x Disturbed rabbit barley Intro Pres May–Jul x Disturbed little barley Native Pres May–Aug x x x false buffalograss Native Pres Jun–Sep x x x witchgrass Native Pres Jun–Sep x x x broomcorn panicgrass Intro Pres Jul–Aug Disturbed Bigelow’s bluegrassBigelow’s Native Pres Mar–May x x x rabbitfoot-grass Intro Pres Jun–Aug x yellow bristlegrassgreen bristlegrass Intro Pres Intro Jul–Sep Pres Jun–Sep Disturbed Disturbed sativa ora var. Typha latifolia Typha Cyperus squarrosus Aristida adscensionis Avena fatua Avena Bouteloua barbata Bromus japonicus Bromus catharticus Bromus rubens Bromus tectorum Cenchrus longispinus Echinochloa crus-galli Eragrostis pectinacea Festuca octofl Hordeum murinum Hordeum pusillum Munroa squarrosa Panicum capillare Panicum miliaceum Poa bigelovii Polypogon monspeliensis Setaria glauca Setaria viridis rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Appendix D. Confi FamilyPerennial Graminoids, cont. Typhaceae Species name Common name Nativity Annual Graminoids Cyperaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

170 Annotated Checklist of Vascular Flora: Capitol Reef National Park areas Wet Comments con Mix Sage PJ des Salt Flower time Park status eld horsetail eld Native Pres NA x x x x toad rush Native Pres Jun–Aug x maize Intro Pres Jul–Aug Disturbed fi tall scouring-rush Native Pres NA x smooth scouring-rush Native Pres NA x southern maidenhair fern Native Pres NA x slender lip-fern Native Pres NA x x x Rock crevices border cloak-fern Native Pres NA x Rock crevices brittle bladder fern Native Pres NA x x male fern Native Pres NA x Rock crevices Oregon woodsia Native Pres NA x x x blunt-leaf spike-moss Native Pres NA x x x ne affi var. lix-mas Juncus bufonius Zea mays Equisetum arvense Equisetum hyemale Equisetum laevigatum Adiantum capillus-veneris Cheilanthes feei Pellaea limitanea Cystopteris fragilis Dryopteris fi Woodsia oregana Woodsia Selaginella mutica rmed, historical, and reported taxa of Capitol Reef National Park, organized by life form with ecological notes, cont. Juncaceae Appendix D. Confi FamilyAnnual Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Ferns and Allies Equisetaceae Equisetaceae Equisetaceae Polypodiaceae (Adiantaceae) Polypodiaceae (Adiantaceae) Polypodiaceae (Adiantaceae, Sinopteridaceae) Polypodiaceae (Aspleniaceae) Polypodiaceae (Aspleniaceae) Polypodiaceae (Aspleniaceae, Woodsiaceae) Selaginellaceae

Appendices 171 172 Annotated Checklist of Vascular Flora: Capitol Reef National Park The Department of the Interior protects and manages the nation’s natural resources and cultural heritage; provides scientiﬁ c and other information about those resources; and honors its special responsibilities to American Indians, Alaska Natives, and afﬁ liated Island Communities.

NPS D-243, January 2009 National Park Service U.S. Department of the Interior

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