1 How butterflies keep their cool: physical and ecological traits influence thermoregulatory ability 2 and population trends. 3 Andrew J. Bladon1*, Matthew Lewis1, Eleanor K. Bladon1, Sam J. Buckton1,2,3, Stuart Corbett4, Steven 4 R. Ewing5, Matthew P. Hayes1, Gwen E. Hitchcock2, Richard Knock2, Colin Lucas6, Adam McVeigh7, Rosa 5 Menéndez7, Jonah M. Walker1, Tom M. Fayle8, Edgar C. Turner1 6 1Department of Zoology, University of Cambridge, Downing Street, Cambridge, CB2 3EJ, UK. 7
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[email protected] 9 2The Wildlife Trust for Bedfordshire, Cambridgeshire & Northamptonshire, The Manor House, Broad 10 Street, Cambourne, Cambridge, CB23 6DH, UK.
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[email protected] 12 3Yorkshire Wildlife Trust, 1 St George’s Place, York, YO24 1GN, UK. 13 4RSPB Winterbourne Downs, RSPB Manor Farm, Newton Tony, Salisbury, Wiltshire, SP4 0HA, UK. 14
[email protected] 15 5RSPB Centre for Conservation Science, RSPB Scotland, 2 Lochside View, Edinburgh Park, Edinburgh, 16 EH12 9DH, UK.
[email protected] 17 649 Mill Road, Beccles, Suffolk, NR34 9UT, UK.
[email protected] 18 7Lancaster Environment Centre, Lancaster University, Lancaster, LA1 4YQ, UK.
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[email protected] 20 8Biology Centre of the Czech Academy of Sciences, Institute of Entomology, Branisovska 31, 37005, 21 Ceske Budejovice, Czech Republic.
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[email protected] 23 1 24 Abstract 25 1. Understanding which factors influence the ability of individuals to respond to changing 26 temperatures is fundamental to species conservation under climate change.