Catalogue of the Heteroptera of the Palaearctic Region. Volume 5
Total Page:16
File Type:pdf, Size:1020Kb
See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/254899169 Catalogue of the Heteroptera of the Palaearctic Region. Volume 5. Pentatomomorpha II Book · July 2006 CITATIONS READS 105 164 2 authors: Berend Aukema Christian Rieger Naturalis Biodiversity Center 21 PUBLICATIONS 185 CITATIONS 157 PUBLICATIONS 1,265 CITATIONS SEE PROFILE SEE PROFILE Some of the authors of this publication are also working on these related projects: Heteroptera of the Canary Islands View project Faunistics Heteroptera of Belgium View project All content following this page was uploaded by Berend Aukema on 09 November 2018. The user has requested enhancement of the downloaded file. Catalogue of the Heteroptera of the Palaearctic Region edited by Berend Aukema & Christian Rieger Volume 5 Pentatomomorpha II CIP-DATA KONINKLIJKE BIBLIOTHEEK, DEN HAAG Aukema, B. & Chr. Rieger (editors) Catalogue of Heteroptera of the Palaearctic Region Catalogue. - With index, references ISBN-10: 90-71912-28-0 ISBN-13: 978-90-71912-28-3 Subject headings: Heteroptera / Palaearctic Region / checklist / type material/ distribution / references Published 18 July 2006 by The Netherlands Entomological Society c/o Plantage Middenlaan 64 NL-1018 DH Amsterdam The Netherlands Printed by Ponsen & Looijen, Wageningen, The Netherlands Cover: Aelia klugii Hahn (Pentatomidae), del. J. de Rond, Lelystad, The Netherlands ii PENTATOMOMORPHA II STENOCEPHALIDAE - W.R. DOLLING RHOPALIDAE - W.R. DOLLING ALYDIDAE - W.R. DOLLING COREIDAE - W.R. DOLLING UROSTYLIDIDAE - D.A. RIDER THAUMASTELLIDAE - J.A. LIS PARASTRACHIIDAE - J.A. LIS CYDNIDAE - J.A. LIS THYREOCORIDAE - J.A. LIS PLATASPIDAE - J. DAVIDOVÁ-VILÍMOVÁ ACANTHOSOMATIDAE - U. GÖLLNER-SCHEIDING TESSARATOMIDAE - D.A. RIDER SCUTELLERIDAE - U. GÖLLNER- SCHEIDING DINIDORIDAE - J.A. LIS PENTATOMIDAE - D.A. RIDER iii This volume is dedicated to the memory of the late Dr. N.M. Andersen, naturalist, colleague and co-author Nils Otto Møller Andersen was born 21 november 1940 near Saxkøbing on Lolland in the southeastern part of Denmark. Already as a boy he was interested in natural history, and this hobby was later shared with his schoolmate, Annemarie Børsen Pedersen, whom he later married. After serving his military duties he began to study biology at the University of Copenhagen in 1963. He was a very ambitious and productive young man, and when he graduated from the Zoological Museum in Copenhagen in 1970, on a study of wing polymorphism in two northern temperate water strider species, he already had a considerable publication record on systematics, morphology, ecology and faunistics of semiaquatic bugs (Gerromorpha). Soon after he was appointed as assistant professor, and later as associate professor at the Zoological Museum. Andersen became very interested in historical biogeography and ecological phylogenetics of Gerromorpha, both from a palaentological and a neontological viewpoint, and received a doctoral degree in 1982 on his book “The Semiaquatic Bugs (Hemiptera, Gerromorpha). Phylogeny, Adaptations, Biogeography and Classification”. This book was partly responsible for the dramatically increased interest for using water striders and their allies in studies of ecology, behaviour and evolution, which has resulted in numerous publications on the subject ever since. With the revolution in computer science Andersen became a pioneer in using computer based phylogenetic reconstructions of studying gerromorphan relationships, and with the upcoming DNA sequencing technology, he also became familiar with analyzing this kind of data. Andersen continued to describe more than 200 extant and fossil species of semiaquatic bugs, a large part of which resulted from his close collaboration with Tom Weir at the Australian National Insect Collection in Canberra. The studies resulted in numerous publications, and are largely summarized in their joint monography “Australian Water Bugs. Their Biology and Identification”, which was published shortly before his highly unexpected death on May 12, 2004. Up til his untimely death, he was working on a world catalogue of Gerromorpha, which will be published posthumous. iv CONTENTS Preface.................................................................................................................................vi Depositories (abbreviations)........................................................................................viii Distribution (abbreviations)............................................................................................x Literature............................................................................................................................xii Acknowledgements...........................................................................................................xiii Family Stenocephalidae..................................................................................................2 Family Rhopalidae..........................................................................................................8 Family Alydidae ...........................................................................................................28 Family Coreidae ...........................................................................................................43 Family Urostylididae ..................................................................................................102 Family Thaumastellidae..............................................................................................117 Family Parastrachiidae................................................................................................118 Family Cydnidae ........................................................................................................119 Family Thyreocoridae ................................................................................................148 Family Plataspidae......................................................................................................150 Family Acanthosomatidae ..........................................................................................166 Family Tessaratomidae ...............................................................................................182 Family Scutelleridae ...................................................................................................190 Family Dinidoridae.....................................................................................................228 Family Pentatomidae ..................................................................................................233 References........................................................................................................................415 Index.................................................................................................................................495 v PREFACE The fifth volume of the Catalogue of Palaearctic Heteroptera (Pentatomomorpha II) contains the following families: Stenocephalidae, Rhopalidae, Alydidae, Urostylididae, Thaumastellidae, Parastrachiidae, Cydnidae, Thyreocoridae, Plataspidae, Acanthosomatidae, Tessaratomidae, Scutelleridae, Dinidoridae and Pentatomidae. The included families are represented by altogether 2133 species and 43 subspecies of 465 genera (table 1). Numbers of Palaearctic Heteroptera Genera Species Subspecies Stenocephalidae 1 18 - Rhopalidae 14 69 6 Alydidae 26 69 - Coreidae 84 306 5 Urostylididae 8 131 - Thaumastellidae 1 1 - Parastrachiidae 1 2 - Cydnidae 37 167 2 Thyrocoridae 1 4 - Plataspidae 10 104 2 Acanthosomatidae 9 107 4 Tessaratomidae 12 30 - Scutelleridae 38 158 5 Dinidoridae 4 19 - Pentatomidae 219 841 19 Incertae sedis 107 ____ ____ ____ Totals 465 2133 43 Table 1. Numbers of Palaearctic Heteroptera listed in volume 5. The following new names and synonymies are included. Stenocephalidae: Dicranocephalus medius (Mulsant & Rey) = Cimex gonymelas Donovan. Rhopalidae: Leptocoris amictus Germar = Leptocoris amicta rubra Göllner-Scheiding. Alydidae: Marcius Stål = Rhodoclia Distant; Alydinae Amyot & Serville = Daclerini Ahmad, Abbas, Shadab & Khan. Coreidae: Centrocoris spiniger (Fabricius) = Cimex branderi Linnaeus; Coreus marginatus marginatus (Linnaeus) = Copium serratum Thunberg; Gonocerus acuteangulatus (Goeze) = Cimex ictericus Villers = Syromastes cinnameus Millet de Turtaudière; Plinachtus Stål = Metacletus Matsumura; Plinachtus basalis (Westwood) = Metacletus bivittatus Matsumura; Plinachtus imitator (Reuter) = Plinachtus falcatus Distant; Anacanthocoris Uhler = Muzafferiana Ahmad & Perveen. Pentatomidae: Eysarcoris Hahn = Gobisa Gorski. New combinations. Alydidae: Melanacanthus marginatus (Hsiao) = Mirperus marginatus Hsiao; Nariscus fisheri (Distant) = Akbaratus fisheri Distant. Coreidae: Plinachtus imitator (Reuter) = Gonocerus imitator Reuter; Prismatocerus borealis (Distant) = Homoeocerus borealis Distant; Prismatocerus inornatus (Stål) = Homoeocerus inornatus Stål; Prismatocerus kiritshenkoi (Bergevin) = Homoeocerus kiritshenkoi Bergevin; Derepteryx vi yunnana (Ren) = Pterygomia yunnana Ren; Tongorma latreillii (Guérin-Méneville) = Phyllomorpha latreillii Guérin-Méneville. Type species are designated for the following taxa. Coreidae: Agonotomus Signoret: Gonocerus amyoti Montrouzier; Tliponius Stål: Homoeocerus puncticornis Burmeister; Pephricus Amyot & Serville,: Cimex paradoxus Sparrmann. Pentatomidae: Cimex perlatus Fabricius (= Cimex aeneus Scopoli). The arrangement