Studying Forest Layers

Total Page:16

File Type:pdf, Size:1020Kb

Studying Forest Layers FIELD ENHANCEMENT 2 Studying Forest Layers OBJECTIVES NUTSHELL In this lesson, students observe the structural Upon completion of this lesson, students will layers of a forest and draw a color-coded picture. be able to: They also embark on two exploration activities to • Identify the structural layers in a forest. discover which animals can be found in each of • Describe wildlife that can be found in each the forest layers. of the layers of a forest. BACKGROUND SUBJECT AREAS Forests vary in their composition (species Arts, Science present) and their structure (how species are arranged). Forests may LESSON/ACTIVITY TIME Forest Layers have any combination • Total Lesson Time: 75 minutes • Overstory of the following layers: • Time Breakdown: overstory, understory • Understory which includes trees, Introduction...........10 minutes - Trees shrubs, and forbs, and Activity 1 ...............30 minutes - Shrubs litter. A mixed forest, for Activity 2 ...............10 minutes - Forbs example, may contain Activity 3 ...............10 minutes • Litter all of the layers. A pine Conclusion............15 minutes plantation may only have TEACHING SITE an overstory, forb layer, and litter layer. Each layer contains its own set of plant species and A forested area that contains five layers provides food and shelter to wildlife. (overstory, understory, shrub, forb, litter) is preferable. The overstory is above all the layers and is made up of mature trees. Tree species CLASSROOM LESSON commonly found in the overstory include white CONNECTIONS and red pine, sugar and red maple, white ash, This lesson ties closely with Classroom red and white oak, and trembling aspen. Animals Lesson 2, What Makes a Forest? that use the overstory of a forest include blue jays, owls, orioles, hawks, woodpeckers, eagles, squirrels, crows, turkeys, and porcupines. The understory may contain a tree, shrub, and forb layer. Trees in this layer are usually immature. When a mature tree from the overstory dies, some of the immature trees may grow taller and become part of the overstory. The understory also contains smaller trees like choke or pin cherry, serviceberry, red cedar, ironwood, and musclewood. 170 Field Enhancement 2: Studying Forest Layers LEAF Guide • 5-6 UNIT VOCABULARY MATERIALS LIST FOR EACH STUDENT Forb Layer: Layer of the understory containing • Copy of Student Page 1, Forest Layers non-woody plants. • Five different colored pencils Litter Layer: Surface layer of the forest floor • Clipboard or other writing surface composed of leaves, twigs, needles, etc., with FOR EVERY 3 TO 4 STUDENTS minimal decomposition. • Bug boxes or magnifying lenses (optional) Overstory: The uppermost trees in a forest. • One white paper plate Shrub Layer: Layer of the understory FOR THE CLASS containing woody plants with multiple stems. • Copy of Student Page 2, Forest Layers Understory: Forest vegetation present Example Page under the overstory, which can include trees, shrubs, and forbs. TEACHER PREPARATION • Visit the teaching site in advance and choose boundaries for your students to work within. These trees are shade-tolerant. This means that they are able to grow well even when the • Choose an area on the site where your trees growing above them block most of the students can sit down and observe forest light. Animals in the tree layer of the understory layers independently. are similar to those in the overstory. • Choose an area where small groups of students can sit on the ground and study The shrub layer occupies the space between leaf litter. the understory trees and the forb layer. Woody • Choose a trail on which to take a short walk plants like raspberry, maple-leaved viburnum, to look for wildlife or evidence of wildlife. dogwood, and witch hazel are found here. This layer provides food and habitat for a SAFETY PRECAUTIONS variety of wildlife like squirrels and deer. Birds Visit the teaching site ahead of time to locate common to the shrub layer in Wisconsin forests any hazards such as hanging branches, include grouse, robins, chickadees, warblers, protruding tree roots, holes, poison ivy, sparrows, finches, cardinals, mourning doves, etc. Encourage students to walk, not run, and grosbeaks. at all times. The next layer of the understory is the forb layer. Small herbaceous (fleshy, not woody-stemmed) plants like bracken fern, wild columbine, pink The litter layer contains fallen leaves, branches, lady slipper, and common blue violet grow here. needles, and other organic matter. Items in this Chipmunks, snakes, toads, mice, moles, grouse, layer are easily recognizable. They are starting turkeys, rabbits, insects, turtles, and spiders all to break apart but have not begun to decompose. use this layer. Critters commonly found in this layer are centipedes, millipedes, ants, beetles, worms, snails, slugs, maggots, spiders, and pill bugs. LEAF Guide • 5-6 UNIT Field Enhancement 2: Studying Forest Layers 171 Although the plant species in each forest layer 5. The next layer is composed of the understory are more clearly defined, wildlife frequently use trees. Ask your students to look around for more than one or all the layers of a forest. For trees whose crowns fit underneath the crowns example, we commonly find squirrels leaping of larger trees. Explain that these trees can along the forest floor, as well as from branch to tolerate a lot of shade. They are either young branch up in the treetops. In fact, squirrels make trees or species that do not get as big as the their nests in the branches of trees. Wild turkeys other trees around them. Tell your students eat nuts and insects from the leaf litter; they that these trees make up part of the understory. also eat food found in the forb and shrub layer. The shrub layer and forb layer are also part of In addition, they roost on branches in the the understory with the trees. understory or overstory. 6. Finally, ask your students to look directly PROCEDURE overhead at the tallest trees in the forest. Explain that this layer is called the overstory. INTRODUCTION These trees dominate the forest. Very little 1. Take students to the forested area you have light passes through their branches to the chosen for exploration. forest floor. Tell your students that when one 2. Have students sit in a circle on the ground in of the trees in the overstory dies, a tree in the forest. Explain that the forest is comprised the understory may have an opportunity to of different layers. Ask them to pick up a become a dominant tree. handful of debris from the ground. Ask what NOTE: If your forested area does not contain they see in this debris. (Leaves, twigs, all five layers, explain all the layers but modify needles, nuts, moss, and other decaying the activity to accommodate your site. plant material.) Explain that this layer, called leaf litter, is where things in the forest ACTIVITY 1 are recycled. Decomposers like fungi, worms, 1. Hand each student a copy of Student Page beetles, and ants break down dead plant 1, Forest Layers, a clipboard, and a set of material and return the nutrients to the soil colored pencils. Explain that they will have so that living plants can use them. time to sit alone and observe the different layers in the forest. Tell them they need to 3. Next, ask someone to point to a non-woody make a drawing of what they observe. The plant that is growing close to the ground. drawing should contain the plants found in Explain that small herbaceous (fleshy, not each of the layers. Ask them to choose a woody-stemmed) plants like these make up different color to draw each of the layers. For the forb layer of the forest. This layer provides example, the forb layer may be purple, the shelter to animals like snakes, toads, rabbits, shrub layer orange, and so on. Tell them that insects, turtles, and spiders. after their drawings are complete, they should 4. Ask your students to look around and spot a fill in the name of each layer on the blank woody-stemmed bush or a sapling. Explain provided. It is important that the drawings that these types of plants make up the shrub show detail and are complete. You can show layer of the forest. Squirrels, deer, and many Student Page 2, Forest Layers Example types of birds benefit from the cover and food Page, as an example. these plants offer. 172 Field Enhancement 2: Studying Forest Layers LEAF Guide • 5-6 UNIT 2. After you have given the directions, explain them to get a closer look. Show your students the boundaries you have chosen for this where to make note of any animals they find lesson. Then ask your students to spread on their worksheet. (There are examples of out within the boundaries and find their own wildlife for all the layers listed along the edge place to sit and work. Give your students time of the student page.) to draw, then bring them back together to sit in a circle. 2. After you have explained the activity, ask the groups to spread out and begin working. As 3. During Activities 2 and 3, your students will they work, walk around and check on their add more information to their worksheets. progress. If anyone has trouble with finding Before continuing, quickly check to be sure critters in the leaf litter, have them put their that everyone drew five layers. Ask your group sample back, move to another location, and to put their drawings on the ground in front of try again. them. As you walk around and check their work, ask the group to point to the overstory 3. When all the groups have had time to work, layer as well as the understory trees, shrub, remind them to put the leaf litter and critters forb, and leaf litter layers.
Recommended publications
  • The Story About Understory Why Everyone on Bald Head Island Should Care
    Bald Head Association 910-457-4676 • www.BaldHeadAssociation.com 111 Lighthouse Wynd • PO Box 3030, Bald Head Island, NC 28461 The Story About Understory Why Everyone on Bald Head Island Should Care Bald Head Island is truly unique. As a barrier island, it at ground level. Bald Head Island’s latitudinal position is bordered by the Atlantic Ocean and Cape Fear River. BHI has supports both northern and southern species of plants.” 14 miles of pristine beaches; 12,400 acres, of which 10,000 acres Important understory plants include vines, small plants and are protected; over 244 species of birds, including the bald eagle; trees, mosses, lichens and even weeds. All are needed to have and the North Carolina Forest Preserve of nearly 200 acres. a healthy forest habitat. Many extreme forces of nature that have helped shape this The BHIC explains, “Vines play an important role. The very special island include hurricane-force winds, salt water and vines and herbaceous plants intertwine, further developing the salt spray, flooding and drought. There is another force of nature structural integrity of the forest and forming pockets of vegetation that impacts BHI — humans — and with us, development. that provide a base for songbirds to build nests. The vines twist The Bald Head Association is mandated by its Covenants to around the canopy and are the secret to wind protection. These help sustain BHI by managing the buildout of homes and by vines actually weave together the canopy so blowing winds don’t managing vegetation trimming/removal, to help protect its penetrate through the top layer and keep homes, plants, and members’ property values.
    [Show full text]
  • Old-Growth Forests
    Pacific Northwest Research Station NEW FINDINGS ABOUT OLD-GROWTH FORESTS I N S U M M A R Y ot all forests with old trees are scientifically defined for many centuries. Today’s old-growth forests developed as old growth. Among those that are, the variations along multiple pathways with many low-severity and some Nare so striking that multiple definitions of old-growth high-severity disturbances along the way. And, scientists forests are needed, even when the discussion is restricted to are learning, the journey matters—old-growth ecosystems Pacific coast old-growth forests from southwestern Oregon contribute to ecological diversity through every stage of to southwestern British Columbia. forest development. Heterogeneity in the pathways to old- growth forests accounts for many of the differences among Scientists understand the basic structural features of old- old-growth forests. growth forests and have learned much about habitat use of forests by spotted owls and other species. Less known, Complexity does not mean chaos or a lack of pattern. Sci- however, are the character and development of the live and entists from the Pacific Northwest (PNW) Research Station, dead trees and other plants. We are learning much about along with scientists and students from universities, see the structural complexity of these forests and how it leads to some common elements and themes in the many pathways. ecological complexity—which makes possible their famous The new findings suggest we may need to change our strat- biodiversity. For example, we are gaining new insights into egies for conserving and restoring old-growth ecosystems. canopy complexity in old-growth forests.
    [Show full text]
  • Understory Forest Monitoring: a Guide for Small Forest Managers
    Understory Forest Monitoring: A Guide for Small Forest Managers With funding provided by BEFORE: A dense monoculture of invasive periwinkle (Vinca species). AFTER: A diverse mix of native vegetation beginning to establish. It will mature into a lush forest Á oor of native wildÁ owers. Managing a healthy forest is not just about having healthy trees. Foresters, scientists, and private woodland owners have an interest in what is growing on the forest Á oor. A healthy understory can offer: • Flowers for native pollinators • Food for wildlife • Resiliency against invasion by forest weeds • Organic material to build healthier soil • Stable soil that doesn’t erode into nearby streams • Beautiful views while recreating in the forest Just like taking inventory of the forest, measuring tree diameters and spacing, it is also important to monitor the changes over time on the forest Á oor. Herbaceous forest plants tend to reach their peak bloom and cover in mid to late spring (mid-May to mid-June in western Oregon). It’s best to monitor forest understory consistently at this time of year. There are dozens of ways to monitor the understory plants in a forest. Two effective methods are described here. Point intercept method This method is performed by laying out a straight line transect in the forest and documenting what is growing on the forest Á oor at one- foot increments along the transect. West Multnomah Soil & Water Conservation District often sets up multiple transects that are 33’ in length giving us 33 data points per transect. The data gathered from the plots can be used to À nd the average cover of different vegetation types in your forest.
    [Show full text]
  • Accelerating the Development of Old-Growth Characteristics in Second-Growth Northern Hardwoods
    United States Department of Agriculture Accelerating the Development of Old-growth Characteristics in Second-growth Northern Hardwoods Karin S. Fassnacht, Dustin R. Bronson, Brian J. Palik, Anthony W. D’Amato, Craig G. Lorimer, Karl J. Martin Forest Northern General Technical Service Research Station Report NRS-144 February 2015 Abstract Active management techniques that emulate natural forest disturbance and stand development processes have the potential to enhance species diversity, structural complexity, and spatial heterogeneity in managed forests, helping to meet goals related to biodiversity, ecosystem health, and forest resilience in the face of uncertain future conditions. There are a number of steps to complete before, during, and after deciding to use active management for this purpose. These steps include specifying objectives and identifying initial targets, recognizing and addressing contemporary stressors that may hinder the ability to meet those objectives and targets, conducting a pretreatment evaluation, developing and implementing treatments, and evaluating treatments for success of implementation and for effectiveness after application. In this report we discuss these steps as they may be applied to second-growth northern hardwood forests in the northern Lake States region, using our experience with the ongoing managed old-growth silvicultural study (MOSS) as an example. We provide additional examples from other applicable studies across the region. Quality Assurance This publication conforms to the Northern Research Station’s Quality Assurance Implementation Plan which requires technical and policy review for all scientific publications produced or funded by the Station. The process included a blind technical review by at least two reviewers, who were selected by the Assistant Director for Research and unknown to the author.
    [Show full text]
  • E666134Z Enhance Development of the Forest Understory to Create
    CONSERVATION ENHANCEMENT ACTIVITY E666134Z Enhance development of the forest understory to create conditions resistant to pests Conservation Practice 666: Forest Stand Improvement APPLICABLE LAND USE: Forest RESOURCE CONCERN ADDRESSED: Degraded Plant Condition PRACTICE LIFE SPAN: 10 Years Enhancement Description: Forest stand improvement that manages the structure and composition of overstory and understory vegetation to reduce vulnerability to damage by insects and diseases of forest trees. Managing the understory vegetation will also reduce the risk of wildfire, and promote development of herbaceous plants that benefit wildlife. This enhancement provides for management of the understory vegetation in a forested area, using mechanical, chemical or manual methods to improve the plant species mix and the health of the residual vegetation. These practices encourage the development of a less dense forested ecosystem. The canopy gaps and open understory allow for air circulation that reduces the incidence of disease, and the improved health of the residual trees increases their ability to withstand insect attacks. Forest stand improvement (FSI) activities are used to remove trees of undesirable species, form, quality, condition, or growth rate. The quantity and quality of forest for wildlife and/or timber production will be increased by manipulating stand density and structure. These treatments can also reduce wildfire hazards, improve forest health, restore natural plant communities, and achieve or maintain a desired native understory plant community for soil health, wildlife, grazing, and/or browsing. Criteria: States will apply general criteria from the NRCS National Conservation Practice Standard (CPS) 666 as listed below, and additional criteria as required by the NRCS State Office. The enhancement will be applied to sites which have an uncharacteristically dense understory of shrubs and small trees that limit development of ground cover.
    [Show full text]
  • Forest Biomes DAY ONE Biomes Section 2
    Biomes Section 2 Chapter 6: Biomes Section 2: Forest Biomes DAY ONE Biomes Section 2 Forest Biomes • Of all the biomes in the world, forest biomes are the most widespread and the most diverse. • The large trees of forests need a lot of water, so forests can be found where temperatures are mild to hot and where rainfall is plenty. • There are three main forest biomes of the world: tropical, temperate, and coniferous. Biomes Section 2 Tropical Rainforests • Tropical rain forests are forests or jungles near the equator. • They are characterized by large amounts of rain and little variation in temperature and contain the greatest known diversity of organisms on Earth. • They help regulate world climate an play vital roles in the nitrogen, oxygen, and carbon cycles. • They are humid, warm, and get strong sunlight which allows them to maintain a fairly constant temperature that is ideal for a wide variety of plants and animals. Biomes Section 2 Tropical Rainforests Biomes Section 2 Nutrients in Tropical Rainforests • Most nutrients are within the plants, not the soil. • Decomposers on the rainforest floor break down dead organisms and return the nutrients to the soil, but plants quickly absorb the nutrients. • Some trees in the tropical rain forest support fungi that feed on dead organic matter on the rainforest floor. • In this relationship, the fungi transfer the nutrients from the dead matter directly to the tree. Biomes Section 2 Nutrients in Tropical Rainforests • Nutrients from dead organic matter are removed so efficiently that runoff from rain forests is often as pure as distilled water.
    [Show full text]
  • The Formation of Dense Understory Layers in Forests Worldwide: Consequences and Implications for Forest Dynamics, Biodiversity, and Succession
    Previous Advances in Threat Assessment and Their Application to Forest and Rangeland Management The Formation of Dense Understory Layers in Forests Worldwide: Consequences and Implications for Forest Dynamics, Biodiversity, and Succession Alejandro A. Royo and Walter P. Carson by land ownership and administrative boundaries. In many cases, the risk to forest understories was particularly acute Alejandro A. Royo, research ecologist, Forestry Sciences if the effects of multiple stressors occurred in a stand, either Laboratory, USDA Forest Service, Northern Research in tandem or within a short period of time. Specifically, the Station, Irvine, PA 16329; and Walter P. Carson, associate synergy between overstory disturbance and uncharacteristic professor, Department of Biological Sciences, University of fire regimes or increased herbivore strongly controls species Pittsburgh, Pittsburgh, PA 15260. richness and leads to depauperate understories dominated Abstract by one or a few species. We suggest that aggressive expansion by native Alterations to natural herbivore and disturbance regimes understory plant species can be explained by considering often allow a select suite of forest understory plant species their ecological requirements in addition to their environ- to dramatically spread and form persistent, mono-dominant mental context. Some plant species are particularly invasive thickets. Following their expansion, this newly established by virtue of having life-history attributes that match one or understory canopy can alter tree seedling recruitment rates more of the opportunities afforded by multiple disturbances. and exert considerable control over the rate and direction Increased overstory disturbance selects for shade-intolerant of secondary forest succession. No matter where these species with rapid rates of vegetative spread over slower native plant invasions occur, they are characterized by one growing, shade-tolerant herbs and shrubs.
    [Show full text]
  • Manual Herbicide Application Methods for Managing Vegetation in Appalachian Hardwood Forests
    United States Department of Manual Herbicide Agriculture Application Methods for Forest Service Managing Vegetation in Northern Appalachian Hardwood Research Station Forests General Technical Report NRS-96 Jeffrey D. Kochenderfer James N. Kochenderfer Gary W. Miller The Authors JEFFREY D. KOCHENDERFER is a forester with the U.S. Forest Service’s Monongahela National Forest at Petersburg, WV. JAMES N. KOCHENDERFER is a retired research forester with the U.S. Forest Service’s Northern Research Station at Parsons, WV. GARY W. MILLER is a research forester with the U.S. Forest Service’s Northern Research Station at Morgantown, WV. Cover photos Front cover: Stem injection on a cull tree using the hack-and-squirt method. Photo by James N. Kochenderfer, U.S. Forest Service (ret.). Back cover: Four primary application methods used in the central Appalachians. Clockwise from upper left: stem injection, foliar spray, cut-stump treatment, and basal spray. Upper left, upper right, and lower right photos by James N. Kochenderfer, U.S. Forest Service (ret.). Lower left photo by Jeffrey D. Kochenderfer, U.S. Forest Service. This publication/database reports research involving pesticides. It does not contain recommendations for their use, nor does it imply that the uses discussed here have been registered. All uses of pesticides must be registered by appropriate State and/or Federal agencies before they can be recommended. CAUTION: Pesticides can be injurious to humans, domestic animals, desirable plants, and fi sh or other wildlife—if they are not handled or applied properly. Use all pesticides selectively and carefully. Follow recommended practices for the disposal of surplus pesticides and pesticide containers.
    [Show full text]
  • Factors Influencing Growth and Flowering of Understory Plants in Conifer Stands in Western Washington
    Proceedings from the Wood Compatibility Initiative Workshop, number 17. FACTORS INFLUENCING GROWTH AND FLOWERING OF UNDERSTORY PLANTS IN CONIFER STANDS IN WESTERN WASHINGTON Constance A. Harrington1, Karl R. Buermeyer2, Leslie Chandler Brodie3, and Bryan W. Wender4 ABSTRACT Managing forest stands to produce wood while also producing other values, such as wildlife habitat, recreation, or spe­ cial forest products, requires information on forest components not always measured in traditional silviculture studies. For example, some managers may want to predict potential growth rates of herbs, shrubs, and trees in the understory—as these components will determine future forest composition and structure—or may be interested in flowering of plants in forest understories, since berry and seed production are important factors in determining species habitat suitability. We conducted several studies in conifer stands in western Washington to determine the effects of stand and site conditions on (1) response of understory vegetation to changes in overstory density, (2) flowering of selected shrub species, and (3) tree growth in for­ est understories. Responses of most factors were quite variable and indicate that managers may have to accept a lower level of certainty or invest more in research and monitoring when managing for components of forest stands other than overstory trees. The general results and management implications from the three studies are discussed and a reading list presented. KEY WORDS: Plant ecology, silviculture, flowering, shrubs, understory, tree growth. INTRODUCTION new information needs: development of understory vegeta­ tion after thinning, flowering of shrubs in forest under- For many forest lands in the Pacific Northwest, fiber stories, and height growth of understory trees.
    [Show full text]
  • FOREST MANAGEMENT 101 a Handbook to Forest Management in the North Central Region
    FOREST MANAGEMENT 101 A handbook to forest management in the North Central Region This guide is also available online at: http://ncrs.fs.fed.us/fmg/nfgm A cooperative project of: North Central Research Station Northeastern Area State & Private Forestry Department of Forest Resources, University of Minnesota Silviculture 21 Silviculture Silviculture The key to effective forest management planning is determining a silvicultural system. A silvicultural system is the collection of treatments to be applied over the life of a stand. These systems are typically described by the method of harvest and regeneration employed. In general these systems are: Clearcutting - The entire stand is cut at one time and naturally or artificially regenerate. Clearcutting Seed-tree - Like clearcutting, but with some larger or mature trees left to provide seed for establishing a new stand. Seed trees may be removed at a later date. Seed-tree 22 Silviculture Shelterwood - Partial harvesting that allows new stems to grow up under an overstory of maturing trees. The shelterwood may be removed at a later date (e.g., 5 to 10 years). Shelterwood Selection - Individual or groups of trees are harvested to make space for natural regeneration. Single tree selection 23 Silviculture Group tree selection Clearcutting, seed-tree, and shelterwood systems produce forests of primarily one age class (assuming seed trees and shelterwood trees are eventually removed) and are commonly referred to as even-aged management. Selection systems produce forests of several to many age classes and are commonly referred to as uneven-aged management. However, in reality, the entire silvicultural system may be more complicated and involve a number of silvicultural treatments such as site preparation, weeding and cleaning, pre-commercial thinning, commercial thinning, pruning, etc.
    [Show full text]
  • Understory Structure in a 23-Year-Old Acacia Koa Forest and 2-Year Growth Responses to Silvicultural Treatments Paul G
    Available online at www.sciencedirect.com Forest Ecology and Management 255 (2008) 1604–1617 www.elsevier.com/locate/foreco Understory structure in a 23-year-old Acacia koa forest and 2-year growth responses to silvicultural treatments Paul G. Scowcroft *, Janis E. Haraguchi, David M. Fujii Institute of Pacific Islands Forestry, Pacific Southwest Research Station, USDA Forest Service, 60 Nowelo Street, Hilo, HI 96720, United States Received 19 July 2007; received in revised form 17 November 2007; accepted 21 November 2007 Abstract Restoration of degraded Acacia koa forests in Hawaii often involves mechanical scarification to stimulate germination of seed buried in the soil and to suppress vegetation that competes with shade intolerant A. koa. Resulting even-age stands are gradually colonized by other plant species, but understory structure is poorly quantified, and the effects of management on understory vegetation are unknown. We examined the structure of the dominant understory species 23 years after stand initiation and determined their subsequent 2-year growth responses to silvicultural treatments prescribed to improve growth of koa. Release thinning, chemical control of introduced grasses, and phosphorus fertilization were applied in a split- plot experimental design. Results for DBH and height class distributions indicated that recruitment for most tree species began within a few years of stand initiation. By 23 years understory trees made up only 14% of total stand basal area, but they added greatly to native plant biodiversity. Alien grasses covered 92% of the forest floor and averaged 1.35 Mg haÀ1. Of the four most abundant native understory tree species, only shade intolerant Myoporum sandwicense grew faster in response to the treatments.
    [Show full text]
  • Understory Plants in Temperate Forests - J
    FORESTS AND FOREST PLANTS – Vol. I - Understory Plants in Temperate Forests - J. A. Antos UNDERSTORY PLANTS IN TEMPERATE FORESTS J. A. Antos Department of Biology, University of Victoria, Canada Keywords: Clonal growth, disturbance, growth forms, herbaceous plants, seedling bank, seedling establishment, shade tolerance, shrubs, species diversity, succession Contents 1. Introduction 2. The Understory Environment 3. Structure of Forest Understory Vegetation 4. Diversity and Composition of Forest Understory Plants 4.1 Species Diversity 4.2 Growth Forms 4.3 Phenology 5. Physiology of Forest Understory Plants 6. Reproduction and Clonal Growth 6.1 Seed Production 6.2 Seedling Establishment 6.3 Spread of Clonal Species 7. Effects of Canopy Variation 8.Responses to Disturbance 9. Changes During Succession 10. Dynamics of the Understory 11. Effects of Understory Plants on Tree Seedling Establishment 12. The Tree Seedling Bank 13. Functional Aspects of the Understory 14. Conservation and Management of Forest Understory Plants 15. Conclusions Acknowledgements Glossary Bibliography Biographical Sketch SummaryUNESCO – EOLSS Much of the vascularSAMPLE plant diversity in temperate CHAPTERS forests occurs in the understory, and the species present there represent a wide variety of growth forms and functional groups. Understory plants comprise the habitat for many kinds of animals, influence ecosystem functions such as nutrient cycling, and can have important effects on the canopy. Many canopy trees established as post-disturbance cohorts, but many others started in the understory, especially in old forests, and understory species can have a major influence on which tree species are successful. Thus the understory is an important structural and functional component of temperate forests. Although many species grow in the forest understory, this environment is not ©Encyclopedia of Life Support Systems (EOLSS) FORESTS AND FOREST PLANTS – Vol.
    [Show full text]