1

Version: 18.11.2008 CHECKLIST OF BRITISH AND IRISH SYMPHYTA

A. D. Liston & D. A. Sheppard

AL: Deutsches Entomologisches Institut, Eberswalder Str. 84, 15374 Müncheberg, Germany DS: 10 Stainfield Road, Hanthorpe, Bourne, Lincs., PE10 0RE

Introduction Fifty years have passed since the publication of the final part of Robert B. Benson’s three part identification key to the “Symphyta” of the British Isles (Benson 1951, 1952, 1958). The importance of these works can hardly be overstated. Whilst the first two parts were based substantially on the acute monography by Eduard Enslin (Enslin 19121918), Benson’s treatment of the represented an original and significant step forward in our knowledge of this subfamily. The results of subsequent research on the British and Irish fauna were collated by Fitton et al. (1978). Since then, renewed investigation of the of West Palaearctic Symphyta has led to a great number of taxonomic and nomenclatural changes, affecting many taxa occurring in the British Isles.

Of particular value in accessing the very extensive but widely scattered literature on , is the online database “Electronic World Catalog of Symphyta (ECatSym)” (Taeger & Blank 2008). Unfortunately, the only identification guide to the European “Symphyta” published since 1958 with a more than local geographical scope, by Zhelochovtsev (1988), suffers from its own particular weaknesses, such as the reliance placed on the opinions and illustrations of E. Lindqvist for the treatment of the Nematinae. Serious mistakes in the translation from Russian make the later English language editions of this work even more difficult to use.

Special problems are attached to the interpretation of records for some species not found in the British Isles since their inclusion in the works by Leach (1817) and Stephens (1835). Some of these are probably extinct, but their status as in the British Isles has in many cases never been unequivocally proved: see for example Benson (1943a). We therefore maintain the practice, as in Fitton et al. (1978), of marking names of such taxa with a preceding “?”. Further, the taxonomic status of some of the nominal taxa described from the British Isles by Hill, Stephens, Newman and Cameron is still unclear, particularly where no type material has been located.

Authorship and date of publication of taxa first described by Geoffroy, Panzer, Klug, Serville. Lepeletier, Zaddach and Brischke are adopted from the bibliographic researches of Taeger & Blank (1996, 2006) and Blank & Taeger (1998).

Arrangement of list

The listing of superfamilies is alphabetical, with families, subfamilies, tribes, genera, subgenera and species listed alphabetically within these.

Genera. The generic classification used here mainly follows the system of Benson (19511958). This has the advantages of being relatively simple and widely known. The present lack of phylogenetic analyses for most groups of Symphyta (but see Nyman et al. 2006 on the Nematinae), which might support alternative generic classifications, makes the adoption here of some of the recently proposed genera at least seem premature. Whilst it is probable that some of these will in the future come to be recognised as valid, they are listed here either as synonyms or as “subgenera” of the genera treated by Benson. Generally, the lack of detailed morphological and biological studies on the larvae of many European sawfly species, must be regarded as a major obstacle to our understanding of their phylogeny.

For each species name, the in which it was originally described follows the author’s name and date, where this differs from its current placement. Species names considered to be synonyms are treated in the same way as valid names, i. e. where necessary, the spelling is amended to agree with the gender of the genus in which it is at present placed. In some cases the most commonly used spelling in the literature therefore differs from that which appears here (e.g. astutus Cameron and (Lygaeotus) astutus (Cameron)). This should be borne in mind when searching the list for synonym names.

Synonymy. The genus and species group names are included selectively in the synonymy. For a complete current synonymy of all these, including important misidentifications, see ECatSym (Taeger & Blank 2006). Names have only been included below as synonyms when they fulfil one or both of the following criteria:

has at some time been used for a taxon occurring in the British Isles. 2

used in widely consulted works on taxonomy or distribution, or in original accounts of biological characters / descriptions of immature stages.

The use of a question mark (?) preceding names in the list has two different meanings: 1) in combination with a taxon listed as valid, doubt is indicated as to validity of previous British records 2) in combination with a synonym, doubt is indicated on the taxonomic status of that taxon.

Distribution. Area covered is the British Isles north to and including Shetland, but excluding the Channel Islands: see Sheppard (1990) on the sawfly fauna of the latter. Distribution of species within the British Isles is roughly indicated by the letter(s) following the valid species name, representing the countries from which it is recorded: England (E), Wales (W), Scotland (S) and Ireland (I) . The Isle of Man is not considered separately, because of a lack of data. It should be noted that several species were mistakenly listed as occurring in Scotland by Liston (1995). A reference is given for species first recorded in the British Isles since the last checklist by Fitton et al. (1978).

Acknowledgements We are most grateful to Dr. M. G. Fitton for his work on an earlier version of this checklist. Guy Knight supplied much new information on the fauna of Wales, based mostly on his original research. Our warmest thanks for many further useful additions, corrections and suggestions are due to Dr. Alison Barker, Dr. Keith Bland, Dr. Stephan Blank, Jerry Bowdrey, Laurence Clemons, Rob Edmunds, K. John Grearson, Andrew Halstead, Dr. Charles Hussey, Dr. Mikk Heidemaa, Keith Palmer, Dr. Andreas Taeger and Dr. Veli Vikberg. 3

Superfamily CEPHOIDEA Family CEPHIDAE Subfamily CEPHINAE Tribe CEPHINI CALAMEUTA Konow, 1896 MONOPLOPUS Konow, 1896 HAPLOCEPHUS Benson, 1935 filiformis (Eversmann, 1847, Cephus ) E W elongatus (Vollenhoven, 1858, Cephus ) Cephus arundinis Giraud, 1863 Cephus analis (Klug, 1803): Stephens, 1835 misidentification pallipes (Klug, 1803, Astatus ) E W S I phthisica (Fabricius, 1805, Cephus ) misspelling pusilla (Stephens 1835, Cephus ) immaculata (Stephens, 1835, Cephus ) cultrarius (Hartig, 1837, Cephus ) pallidipes (Dalla Torre, 1894, Cephus )

CEPHUS Latreille, 1802 spinipes (Panzer, [1800], Banchus ) E W pilosulus Thomson, 1871 pusillus : Stephens, 1835 misidentification cultratus : Eversmann, 1847 misidentification and misspelling of cultrarius Hartig nigrinus Thomson, 1871 E pallipes (Klug, 1803): Stephens, 1835 misidentification pygmeus (Linnaeus, 1767, Sirex ) E W floralis (Klug, 1803, Astatus ) atripes Stephens, 1835 pygmaeus : misspelling

TRACHELUS Jurine, 1807 CEPHA Billberg, 1820 ATEUCHOPUS Konow, 1896 EUMETABOLUS Schulz, 1906 TRACHELASTATUS Morice & Durrant, 1915 NEATEUCHOPUS Benson, 1935 MICROCEPHUS Benson, 1935 tabidus (Fabricius, 1775, Sirex ) E W macilentus (Fabricius, 1793, Sirex ) mandibularis (Serville, 1823, Cephus ) nigritus (Serville, 1823, Cephus ) troglodyta (Fabricius, 1787, Sirex ) E extinct niger (Harris, 1776): Cameron, 1890 misidentification

Tribe HARTIGIINI HARTIGIA Schiødte, 1839 MACROCEPHUS Schlechtendal, 1878 COPIOSOMA W. F. Kirby, 1882 ADIRUS Konow, 1899 linearis (Schrank, 1781, ) E W quinquefasciata (Stephens, 1835, Cephus ) major (Eversmann, 1847, Cephus ) agrimoniae (Goury, 1912, Macrocephus ) nigra (Harris, 1776, Sirex ) E W satyrus (Panzer, 1801, Astatus ) brachyptera (Damianitsch, 1866, Cephus ) xanthostoma (Eversmann, 1847, Cephus ) E W ulmariae (Schlechtendal, 1878, Macrocephus )

JANUS Stephens, 1829 PHYLLOECUS Newman, 1838 4

EPHIPPIONOTUS Costa, 1860 femoratus (Curtis, 1830, Cephus ) E faunus (Newman, 1838, Phylloecus ) niger (Brischke, 1892, Cephus ) cynosbati (Linnaeus, 1758): misidentification luteipes (Lepeletier, 1823, Cephus ) E W connectens Stephens, 1829 5

Unplaced species of Cephidae: Janus bifrons Newman, 1838 6

Superfamily ORUSSOIDEA Family ORUSSIDAE ORUSSUS Latreille, 1797 ORYSSUS Fabricius, 1798 ? abietinus (Scopoli, 1763, Sphex ) E [note] 7

Superfamily PAMPHILIOIDEA MEGALODONTOIDEA Family homonym MEGALODONTES Latreille, 1802 [note] TARPA Fabricius, 1804 ? cephalotes (Fabricius, 1781, Tenthredo ) E klugii (Leach, 1817, Tarpa ) spissicornis (Klug, 1824, Tarpa ) klugi : misspelling

Family PAMPHILIIDAE Subfamily CEPHALCIINAE ACANTHOLYDA Costa, 1894 Subgenus ACANTHOLYDA Costa, 1894 erythrocephala (Linnaeus, 1758, Tenthredo ) S

Subgenus ITYCORSIA Konow, 1897 posticalis Matsumura, 1912 E S stellata (Christ, 1791, Tenthredo ) homonym pinivora Enslin, 1918 nemoralis (Linnaeus, 1758): Thomson, 1871 misidentification

CEPHALCIA Panzer, 1805 CEPHALCIA Jurine, 1801 suppressed CEPHALEIA Jurine, 1807 lariciphila (Wachtl, 1898, Cephaleia ) E W S alpina (Klug, 1808): misidentification falleni (Dalman, 1823): misidentification annulata (Hartig, 1837): misidentification

Subfamily PAMPHILIINAE NEUROTOMA Konow, 1897 GONGYLOCORSIA Konow, 1897 GONGYLOCORISA : Ashmead, 1898 misspelling mandibularis (Zaddach, 1866, Lyda ) E saltuum (Linnaeus, 1758, Tenthredo ) E W flaviventris (Retzius, 1783, Tenthredo ) fasciata (Curtis, 1831, Lyda )

PAMPHILIUS Latreille, 1802 LYDA Fabricius, 1804 ANOPLOLYDA Costa, 1894 BACTROCERUS Konow, 1897 albopictus (Thomson, 1871, Lyda ) S added by Shinohara (1998) balteatus (Fallén, 1808, Lyda ) E W S I cingulatus Latreille, 1812 betulae (Linnaeus, 1758, Tenthredo ) E fumipennis (Curtis, 1831, Lyda ) E W gyllenhali (Dahlbom, 1835, Lyda ) E W S I [note] histrio (Latreille, 1812, Lyda ) E W S flaviventris (Retzius, 1783): Stephens, 1835 misidentification hortorum (Klug, 1808, Lyda ) E W S I hortorum ssp. bicinctus Benson, 1945 inanitus (Villers, 1789, Tenthredo ) E W S I latifrons (Fallén, 1808, Lyda ) E [note] maculosus (Zaddach, 1866, Lyda ) pallipes (Zetterstedt, 1838, Lyda ) E W S I stramineipes (Hartig, 1837, Lyda ) S arbustorum (Fabricius, 1793): Cameron, 1885 misidentification sylvarum (Stephens, 1835, Lyda ) E W 8

nigricornis (Vollenhoven, 1858, Lyda ) sylvaticus (Linnaeus, 1758, Tenthredo ) E W S I vafer (Linnaeus, 1767, Tenthredo ) E W S I depressus (Schrank, 1781, Tenthredo ) varius (Serville, 1823, Lyda ) E W S I vafer (Linnaeus, 1767): misidentification

Unplaced species of Pamphiliidae: Tenthredo variegata Hill, 1773 S 9

Superfamily SIRICOIDEA Family SIRICIDAE SIREX Linnaeus, 1761 PAURURUS Konow, 1896 areolatus (Cresson, 1867, Urocerus ) E introduced, not established cyaneus Fabricius, 1781 E W S I torvus Harris, 1782 duplex Shuckard, 1837 juvencus (Linnaeus, 1758, Ichneumon ) E W S nigricornis Acerbi, 1802 homonym dubia W. F. Kirby, 1882 ? atricornis Kjellander, 1945 noctilio Fabricius, 1793, Sirex E W S I melanocerus Thomson, 1871

TREMEX Jurine, 1807 XYLOTERUS Hartig, 1837 XYLOECEMATIUM Heyden, 1868 columba (Linnaeus, 1763, Sirex ) E S introduced, not established

UROCERUS Geoffroy, 1785 UROCERUS Geoffroy, 1762 suppressed XANTHOSIREX SemenovTianShanskij, 1921 albicornis (Fabricius, 1781, Sirex ) introduced, not established stephensii (W. F. Kirby, 1882, Sirex ) augur (Klug, 1803, Sirex ) E S I, established in Ireland [note] cedrorum (Smith, 1860, Sirex ) fantoma (Fabricius, 1781): Benson, 1958 misidentification californicus Norton, 1869, introduced, not established fantoma (Fabricius, 1781, Sirex ) introduced, not established (?) [note] tardigradus Cederhjelm, 1798 phantoma : misspelling gigas (Linnaeus, 1758, Ichneumon ) E W S I bizonatus (Stephens, 1835, Sirex ) [note] taiganus Benson, 1943

XERIS Costa, 1894 spectrum (Linnaeus, 1758, Ichneumon ) E W I

Unplaced species of Siricidae Sirex bimaculatus Donovan, 1808 [note]

Superfamily Family Subfamily Schrank, 1802 CRYPTUS Jurine, 1801 suppressed HYLOTOMA Latreille, 1802 berberidis Schrank, 1802 E, added by Halstead (2004) ciliaris (Linnaeus, 1767, Tenthredo ) E S I coerulea (Klug, 1814, Hylotoma ) corusca (Zaddach, 1859, Hylotoma ) cyanocrocea (Forster, 1771, Tenthredo ) E W I coerulescens (Fabricius, 1775, Tenthredo ) bicolor (Schrank, 1776, Tenthredo ) coerulea (Latreille, 1805, Hylotoma ) ? syriaca (Mocsáry, 1880, Hylotoma ), possibly a distinct species (Pesarini 2002) enodis (Linnaeus, 1767, Tenthredo ) E coeruleipennis (Retzius, 1783, Tenthredo ) 10 expansa (Klug, 1834, Hylotoma ) S, I clavicornis (Fabricius, 1781): misidentification. fuscipes (Fallén, 1808, Hylotoma ) E W S I clavicornis (Fabricius, 1781): misidentification. violacea (Klug, 1814, Hylotoma ) atrocoerulea (Serville, 1823, Hylotoma ) fuscinervis Lindqvist, 1974 gracilicornis (Klug, 1814, Hylotoma ) E W S I coerulescens (Geoffroy, 1785, Tenthredo ) homonym incolorata (Christ, 1791, Tenthredo ) nomen oblitum pilicornis (Leach, 1817, Hylotoma ) cyanella (Klug, 1834, Hylotoma ) crassa (Konow, 1884, Hylotoma ) melanochra (Gmelin, 1790, Tenthredo ) E melanochroa : misspelling femoralis (Klug, 1814, Hylotoma ) dimidiata (Serville, 1823, Hylotoma ) homonym nigritarsis (Klug, 1834, Hylotoma ) bicolor (Gimmerthal, 1846, Hylotoma ) fuliginata Konow, 1907 metallica (Klug, 1834, Hylotoma ) I nigripes (Retzius, 1783, Tenthredo ) E W S enodis (Linnaeus, 1767): misidentification anglica (Leach, 1817, Hylotoma ) ochropus (Gmelin, 1790, Tenthredo ) E ochropa : misspelling rosincola Schrank, 1802 rosarum (Klug, 1814, Hylotoma ) pyrenaica (André, 1879, Hylotoma ) rosae (Linnaeus, 1758): misidentification pagana (Panzer, 1798, Tenthredo ) E W ?stephensii (Leach, 1817, Hylotoma ) [note] rustica (Linnaeus, 1758, Tenthredo ) E atrata (Forster, 1771, Tenthredo ) segmentaria (Panzer, 1804, Cryptus ) klugii (Leach, 1817, Hylotoma ) leachii (Stephens, 1835, Hylotoma ) ustulata (Linnaeus, 1758, Tenthredo ) E W S I

Subfamily STERICTIPHORINAE APROSTHEMA Konow, 1899 COPIDOCEROS Forsius, 1921 fusicorne (Thomson, 1871, Schizocera ) E, added by Vikberg (2004) melanurum (Klug, 1814, Hylotoma ) E friesei (Konow, 1895, Schizocera ) alfkeni (Konow, 1895, Schizocera ) melanura : misspelling tardum (Klug, 1814, Hylotoma ) E, added by Vikberg (2004)

STERICTIPHORA Billberg, 1820 SCHIZOCERUS Berthold, 1827 SCHIZOCERA Lepeletier & Serville, 1828 CYPHONA Dahlbom, 1835 SCHIZOCEROS Konow, 1899 STERICTOPHORA Benson,195l: misspelling angelicae (Panzer, 1799, Tenthredo ) E W villersii (Leach, 1817, Cryptus ) melanocephala (Fabricius, 1798): misidentification furcata (Villers,1789): misidentification geminata (Gmelin, 1790, Tenthredo ) E W S I pallipes (Leach, 1817, Cryptus ) 11

Family BLASTICOTOMIDAE BLASTICOTOMA Klug, 1834 filiceti Klug, 1834 E W I

Family Subfamily ABIINAE [note] ABIA Leach, 1817 candens Konow, 1887 E W S I sericea (Linnaeus, 1767, Tenthredo ) E W S I

ZARAEA Leach, 1817 AENOABIA Kangas, 1946 AUROABIA Kangas, 1946 aenea (Klug, 1820, Cimbex ) E bifida (Thomson, 1871, Abia ) bigens (Kangas, 1946, Abia ) lonicerae (Linnaeus, 1758): Enslin, 1917 misidentification fasciata (Linnaeus, 1758, Tenthredo ) E W S I lonicerae (Linnaeus, 1758, Tenthredo ) E W I nigricornis Leach, 1817

Subfamily CIMBICINAE Tribe CIMBICINI CIMBEX Olivier, 1790 CRABRO Geoffroy, 1762 suppressed CLAVELLARIUS Olivier, 1789 suppressed CLAVELLARIA Lamarck, 1801 PALAEOCIMBEX Semenov TianShanskij, 1935 ALLOCIMBEX Zirngiebl, 1953 connatus (Schrank, 1776, Tenthredo ) E W I femoratus (Linnaeus, 1758, Tenthredo ) E W S I europaea Leach, 1817 sylvarum (Fabricius, 1793, Tenthredo ) varians Leach, 1817 luteus (Linnaeus, 1758, Tenthredo ) E W S griffinii Leach, 1817 annulata (Geoffroy, 1785): Leach, 1817 misidentification quadrimaculatus (Müller, 1766, Tenthredo ) E extinct [note] humeralis (Geoffroy, 1785, Crabro ) axillaris (Panzer, 1801, Tenthredo )

Tribe TRICHIOSOMINI PSEUDOCLAVELLARIA Schulz, 1906 CLAVELLARIA Lamarck, 1801: misidentification amerinae (Linnaeus, 1758, Tenthredo ) E extinct [note] marginata (Linnaeus, 1767, Tenthredo )

TRICHIOSOMA Leach, 1817 [note] ASITRICHIOSOMA Malaise, 1937 laterale Leach, 1817, E S I [note] ?marginale Leach, 1817 latreillei Leach, 1817 lucorum (Linnaeus, 1758, Tenthredo ) E W S I pusillum Stephens, 1835 E [note] nigripes Gussakovskij, 1947 scalesii Leach, 1817 E S unidentatum Leach, 1817 silvaticum Leach, 1817 biverrucatum Stephens, 1835 12

betuleti (Klug, 1834): Cameron, 1875 misidentification sorbi Hartig, 1840 E W S I scalesii Leach, 1817: Cameron, 1885 misidentification tibiale Stephens, 1835 E W S I crataegi (Zaddach, 1863, Cimbex ) vitellina (Linnaeus, 1761, Tenthredo ) E S I boreale Gussakovskij, 1947 vitellinae : misspelling

Subfamily CORYNINAE CORYNIS Thunberg, 1789 AMASIS Leach, 1817 ? crassicornis (Rossi, 1790, Tenthredo ) E [note] laeta (Fabricius, 1798, Tenthredo ) ? obscura (Fabricius, 1775, Tenthredo ) E [note]

Family DIPRIONIDAE Subfamily DIPRIONINAE DIPRION Schrank, 1802 PTERONUS Jurine, 1801 suppressed LOPHYRUS Latreille, 1802 suppressed pini (Linnaeus, 1758, Tenthredo ) E W S I similis (Hartig, 1836, Lophyrus ) E W S eremita (Thomson, 1871, Lophyrus )

GILPINIA Benson, 1939 frutetorum (Fabricius, 1793, Tenthredo ) S thomsoni (Konow, 1884, Lophyrus ) Lophyrus variegatus Hartig, 1834: Cameron, 1890 misidentification hercyniae (Hartig, 1837, Lophyrus ) E W polytoma (Hartig, 1834): misidentification pallida (Klug, 1812, Lophyrus ) E S Lophyrus virens (Klug, 1812): Cameron, 1895 misidentification Lophyrus dorsatus (Fabricius, 1781): Cameron, 1890 misidentification virens (Klug, 1812, Lophyrus ) E

MICRODIPRION Enslin, 1917 pallipes (Fallén, 1808, Hylotoma ) E W S

NEODIPRION Rohwer, 1918 sertifer (Geoffroy, 1785, Tenthredo ) E W S I rufa (Retzius, 1783): Latreille, 1807 misidentification

Subfamily MONOCTENINAE MONOCTENUS Dahlbom, 1835 juniperi (Linnaeus, 1758, Tenthredo ) S I

Family Subfamily ALLANTINAE Tribe ALLANTINI EMPHYTINI ALLANTUS Panzer, 1801 [note] Subgenus ALLANTUS Panzer, 1801 togatus (Panzer, 1801, Tenthredo ) E W S I [note]

Subgenus EMPHYTUS Klug, 1815 basalis (Klug, 1818, Tenthredo ) E S ? basalis ssp. caledonicus (Benson, 1945, Emphytus ) [note] calceatus (Klug, 1818, Tenthredo ) E W S I cinctus (Linnaeus, 1758, Tenthredo ) E W S I togatus Zetterstedt, 1838 homonym 13

cingulatus Blanchard, 1840 homonym neglectus (Zaddach, 1859, Emphytus ) cinctipes (Norton, 1867, Emphytus ) cingulatus (Scopoli, 1763, Tenthredo ) E W I coryli (Stritt, 1937, Emphytus ) E laticinctus (Serville, 1823, Dolerus ) W, added by Knight (2006) balteatus (Klug, 1818, Tenthredo ) homonym melanarius (Klug, 1818, Tenthredo ) E rufocinctus (Retzius, 1783, Tenthredo ) E W S I togatus (Panzer, 1801, Tenthredo ) E W S I [note] truncatus (Klug, 1818, Tenthredo ) E W S cingillum (Klug, 1818): Morice, 1909 misidentification melanarius (Klug, 1818): Morice, 1909 misidentification

APETHYMUS Benson, 1939 filiformis (Klug, 1818, Tenthredo ) E W S I serotinus (Klug, 1818, Tenthredo ) homonym abdominalis (Serville, 1823, Dolerus ) autumnalis (Forsius, 1933, Emphytus ) serotinus (O. F. Müller, 1776, Tenthredo ) E W S I braccatus (Gmelin, 1790, Tenthredo ) tibialis (Panzer, 1799, Tenthredo ) panzeri (W. F. Kirby, 1882, Emphytus )

TAXONUS Hartig, 1837 agrorum (Fallén, 1808, Tenthredo ) E S I [note]

Tribe ATHALIINI Leach, 1817 DENTATHALIA Benson, 1931 ancilla Serville, 1823 E W S [note] glabricollis Thomson, 1870 bicolor Serville, 1823 E W annulata (Fabricius, 1787, Tenthredo ) homonym richardi Serville, 1823 circularis (Klug, 1815, Tenthredo ) E W S I [note] lineolata Serville, 1823 bolivari Dusmet, 1896 cordatoides Kontuniemi, 1951 longifoliae Kontuniemi, 1951 cordata Serville, 1823 E W S I cornubiae Benson, 1931 E liberta (Klug, 1815, Tenthredo ) E W S I lugens (Klug, 1815, Tenthredo ) E W S I rosae (Linnaeus, 1758, Tenthredo ) E W S colibri (Christ, 1791, Tenthredo ) spinarum (Fabricius, 1793, Tenthredo ) centifoliae (Panzer, 1798, Tenthredo ) scutellariae Cameron, 1880 E W S I galericulatae (Kontuniemi, 1951, Dentathalia )

Tribe CALIROINI CALIROA Costa, 1859 ERIOCAMPOIDES Konow, 1890 annulipes (Klug, 1816, Tenthredo ) E W S I atra (Stephens, 1835, Selandria ) cerasi (Linnaeus, 1758, Tenthredo ) E W S I limacina (Retzius, 1783, Tenthredo ) adumbrata (Klug, 1816, Tenthredo ) antipoda (W. F. Kirby, 1881, Monostegia ) 14 cinxia (Klug, 1816, Tenthredo ) E tremulae Chevin, 1974 E, added by Liston (1993) varipes (Klug, 1816): misidentification varipes (Klug, 1816, Tenthredo ) E W I

ENDELOMYIA Ashmead, 1898 aethiops (Gmelin, 1790, Tenthredo ) E W S I soror (Vollenhoven, 1869, Selandria ) testaceipes (Cameron, 1874, Eriocampa ) caninae (Cameron, 1878, Eriocampa ) nigricolle (Cameron, 1882, Poecilosoma )

Tribe EMPRIINI HARPIPHORINI AMETASTEGIA Costa, 1882 Subgenus AMETASTEGIA Costa, 1882 albipes (Thomson, 1871, Taxonus ) E W I fletcheri (Cameron, 1878, Taxonus ) Emphytus nigricans (Klug, 1818): Stephens, 1835 misidentification equiseti (Fallén, 1808, Tenthredo ) E W S I coxalis (Hartig, 1837, Tenthredo ) glabrata (Fallén, 1808, Tenthredo ) E W S I

Subgenus PROTEMPHYTUS Rohwer, 1909 carpini (Hartig, 1837, Emphytus ) E W S I glottianus (Cameron, 1874, Taxonus ) [note] pallipes (Spinola, 1808, Tenthredo ) E W S I grossulariae (Klug, 1818, Tenthredo ) perla (Klug, 1818, Tenthredo ) E S I tenera (Fallén, 1808, Tenthredo ) E W S I patellata (Klug, 1818, Tenthredo ) tener : misspelling

EMPRIA Lepeletier & Serville, 1828 POECILOSTOMA Dahlbom, 1835 POECILOSOMA Thomson, 1870: misspelling PARATAXONUS MacGillivray, 1908 LEUCEMPRIA Enslin, 1913 TRIEMPRIA Enslin, 1914 alector Benson, 1938 E W S I alpina Benson, 1938 S I gussakovskii DovnarZapolskij, 1929: Zhelochovtsev, 1988 misidentification candidata (Fallén, 1808, Tenthredo ) E W S repanda (Klug, 1816, Tenthredo ) excisa (Thomson, 1871, Poecilosoma ) E W S I fletcheri (Cameron, 1878, Poecilosoma ) S obtusa (Klug, 1817): Cameron, 1874 misidentification immersa (Klug, 1818, Tenthredo ) E W S I [note] tirolensis Enslin, 1914 liturata (Gmelin, 1790, Tenthredo ) E W S I longicornis (Thomson, 1871, Poecilosoma ) E W S I rubi Kontuniemi, 1951 pallimacula (Serville, 1823, Dolerus ) E W S I baltica Conde, 1937 parvula (Konow, 1892, Poecilosoma ) E S pumila (Konow, 1896, Poecilosoma ) E W S I [note] sexpunctata (Serville, 1823, Tenthredo ) E W S I klugii (Stephens, 1835, Selandria ) carbonaria (Konow, 1884, Poecilosoma ) klugi : misspelling 15

guttata (Fallén, 1808): Cameron, 1875 misidentification tridens (Konow, 1896, Poecilosoma ) E W S I

Unplaced species of Empria : signata (Newman, 1838, Selandria )

HARPIPHORUS Hartig, 1837 ASTICTA Newman, 1838 homonym lepidus (Klug, 1818, Tenthredo ) E I ianthe (Newman, 1837, )

MONOSTEGIA Costa, 1859 NEMATOCEROS Konow, 1896 abdominalis (Fabricius, 1798, Tenthredo ) E W S I luteola (Klug, 1816, Tenthredo )

MONSOMA MacGillivray, 1908 MONOSOMA Viereck, 1910 pulveratum (Retzius, 1783, Tenthredo ) E W S I pulverata : misspelling

Tribe ERIOCAMPINI ERIOCAMPA Hartig, 1837 BRACHYOCAMPA Zirngiebl, 1956 ovata (Linnaeus, 1761, Tenthredo ) E W S I

Subfamily Tribe BLENNOCAMPINI ARDIS Konow, 1886 pallipes (Serville, 1823, Dolerus ), E W S I brunniventris (Hartig, 1837, Tenthredo ) bipunctata (Klug, 1817, Tenthredo ) homonym sulcata (Cameron, 1882, ) E

BLENNOCAMPA Hartig, 1837 Viitasaari & Vikberg, 1985 E W S I pusilla (Klug, 1816, Tenthredo ) homonym

CLADARDIS Benson, 1952 ? elongatula (Klug, 1817, Tenthredo ) E [note] sericans (Hartig, 1837): Cameron, 1882 misidentification

CLAREMONTIA Rohwer, 1909 PSEUDOBLENNOCAMPA Malaise, 1935 alchemillae (Cameron, 1876, Blennocampa ) E W S I [note] alternipes (Klug, 1816, Tenthredo ) E W S intermedia (Kriechbaumer, 1884, Blennocampa ) tergestina (Kriechbaumer, 1888, Blennocampa ) cinereipes (Klug, 1816): Thomson, 1870 misidentification brevicornis (Brischke, 1883, Blennocampa ) E W S I confusa (Konow, 1886, Blennocampa ) geniculata (Hartig, 1837): Stephens, 1835 misidentification alternipes (Klug, 1816): misidentification puncticeps (Konow, 1886, Blennocampa ) E W ? tenuicornis (Klug, 1816, Tenthredo ) [note] ? tibialis (Stephens, 1835, Selandria ) spiraeae (Brischke, 1883, Blennocampa ) geniculata (Hartig, 1837): Enslin, 1918 misidentification uncta (Klug, 1816, Tenthredo ) E I, added by Koch (1988) [note] humeralis (Vollenhoven, 1869, Blennocampa ) waldheimii (Gimmerthal, 1847, Tenthredo ) E W S 16

subcana (Zaddach, 1859, Selandria ) subserrata (Thomson, 1870, Blennocampa )

MONARDIS Hartig, 1837 plana (Klug, 1817, Tenthredo ) W, added by Gibbs (2006) rosarum (Brischke, 1883, Blennocampa )

MONOPHADNOIDES Ashmead, 1898 PSEUDOMONOPHADNUS Malaise, 1935 rubi (Harris, 1845, Selandria ) E W S I geniculata (Hartig, 1837, Tenthredo ) homonym ruficruris (Brullé, 1832, Selandria ) E W S

PAREOPHORA Konow, 1886 pruni (Linnaeus, 1758, Tenthredo ) E I nigripes (Klug, 1816, Tenthredo ) homonym

PERICLISTA Konow, 1886 APERICLISTA Enslin, 1914 albida (Klug, 1816, Tenthredo ) E W S I melanocephala (Fabricius, 1798, Tenthredo ) homonym versicolor (Newman, 1837, Selandria ) lineolata (Klug, 1816, Tenthredo ) E S pubescens (Zaddach, 1859, Selandria ) E W

Tribe FENUSINI Westwood, 1840 PARAPHYLLOTOMA Forsius, 1930 monilicornis (Dahlbom, 1835, Phyllotoma ) S famosa Benson, 1950 minuta (Dahlbom, 1835): misidentification nigrita Westwood, 1840 E W S I [note] tormentillae (Healy, 1868, Phyllotoma ) ?agrimoniae Brischke, 1888 pygmaea (Klug, 1816): Healy, 1869 misidentification

FENUSA Leach, 1817 KALIOSYSPHINGA Tischbein, 1846 APHADNURUS Costa, 1859 PHAENUSA Cameron, 1875 CALIOSYSPHINGA Konow, 1905 KALIOFENUSA Viereck, 1910 altenhoferi (Liston, 1993, Kaliofenusa ) E S, added by Liston (1994) [note] carpinifoliae (Liston, 1993, Kaliofenusa ) dohrnii (Tischbein, 1846, Kaliosysphinga ) E W S I melanopoda (Cameron, 1876, Phaenusa ) westwoodi (Cameron, 1882, Fenella ) pumila Leach, 1817 E W S I pumila (Klug, 1818, Tenthredo ) homonym pusilla (Lepeletier, 1823): misidentification fuliginosa Healy, 1869 minima Brischke, 1883 pygmaea (Klug, 1816): Zetterstedt, 1838 misidentification ulmi Sundevall, 1847 E W S I [note] intermedia (Thomson, 1871, Fenusa ) alsia (MacGillivray, 1923, Messa )

Unplaced species of Fenusa : pusilla (Serville, 1823, Dolerus ) 17

FENUSELLA Enslin, 1914 MESSA Leach, 1817: misidentification glaucopis (Konow, 1907, Fenusa ) E W S hortulana (Klug, 1818, Tenthredo ) E I soenderupi Hering, 1935 nana (Klug, 1816, Tenthredo ) E W S I mellita (Newman, 1870, Phyllotoma ) quercus (Cameron, 1885, Fenusa ) laeta (Enslin, 1918, Scolioneura )

METALLUS Forbes, 1885 ENTODECTA Konow, 1886 POLYBATES MacGillivray, 1909 albipes (Cameron, 1875, Phaenusa ) E S tenuicornis (Hellén, 1935, Entodecta ) lanceolatus (Thomson, 1870, Blennocampa ) E W S gei (Brischke, 1883, Fenusa ) decolor (Konow, 1886, Entodecta ) bensoni Smith, 1971 pumilus (Klug, 1816, Tenthredo ) E S I pumilio (Hartig, 1837, Emphytus ) rubi (Boie, 1848, Fenusa )

PARNA Benson, 1936 apicalis (Brischke, 1888, Blennocampa ) E, added by Edmunds et al. (2007) reseri Liston, 1993 tenella (KIug, 1816, Tenthredo ) E W S tiliae (Kaltenbach, 1874, Blennocampa )

PROFENUSA MacGillivray, 1914 pygmaea (Klug, 1816, Tenthredo ) E W S I thomsoni (Konow, 1886, Fenusa ) E alumna (MacGillivray, 1923): misidentification

SCOLIONEURA Konow, 1890 [note] betuleti (Klug, 1816, Tenthredo ) E S I nigricans (Klug, 1818, Tenthredo ) homonym betulae (Zaddach, 1859, Fenusa ) vicina Konow, 1894 E S I, added by Liston & O’Connor (2005)

Tribe HETERARTHRINI HETERARTHRUS Stephens, 1835 PHYLLOTOMA Fallén, 1829 homonym DECATRIA Stephens, 1835 DRUIDA Newman, 1838 HETERARTHUS Cameron, 1882 aceris (Kaltenbach, 1856, Phyllotoma ) E W S I cuneifrons Altenhofer & Zombori, 1987, E, added by Liston & Blank (2006) microcephalus (Klug, 1818, Tenthredo ) E S I melanopyga (Klug, 1818): Healy, 1868 misidentification nemoratus (Fallén, 1808, Hylotoma ) E W S I parviceps (Newman, 1837, Fenusa ) tenellus (Zaddach, 1859, Phyllotoma ) mathesoni (MacGillivray, 1909, Phlebotrophia ) mathewsoni : misspelling ochropoda (Klug, 1818, Tenthredo ) E W S fuscipennis (Stephens, 1835, Decatria ) pinguis (Vollenhoven, 1869, Phyllotoma ) Phyllotoma maxima Strobl, 1896 vagans (Fallén, 1808, Hylotoma ) E W S I melanopygus (Klug, 1818, Tenthredo ) 18

amaurus (Klug, 1818, Tenthredo ) kamtchaticus (Malaise, 1931, Phyllotoma ) wuestneii (Konow, 1905, Phyllotoma ) E, added by Altenhofer & Zombori (1987) aceris (McLachlan, 1867, Phyllotoma ), homonym [note] healyi Altenhofer & Zombori, 1987

Unplaced species of Heterarthrus : fumipennis (Cameron, 1888, Phyllotoma )

Tribe PHYMATOCERINI Enslin, 1914 ATOMOSTETHUS Enslin, 1914 ephippium (Panzer, 1798, Tenthredo ) E W S I dubius (Gmelin, 1790, Tenthredo ) homonym inhabilis (Norton, 1861, Selandria ) gagathinus (Klug, 1816, Tenthredo ) E W S meridionalis Benson, 1954 luteiventris (Klug, 1816, Tenthredo ) E W S I fuscipennis (Serville, 1823, Tenthredo ) nigrans (Konow, 1887, ) E W S I, added by Liston & O’Connor (2005) nigrans Blank & Taeger, 1998 cinereipes (Klug, 1816): Cameron, 1882 misidentification ephippium (Panzer, 1798): misidentification punctatus (Konow, 1887, Tomostethus ) E W S brachycera (Cameron, 1893, Tomostethus ) micans (Klug, 1816): Cameron, 1877 misidentification

MONOPHADNUS Hartig, 1837 pallescens (Gmelin, 1790, Tenthredo ) E W S I albipes (Gmelin, 1790, Tenthredo ) homonym furvus Benson, 1930

PARACHARACTUS MacGillivray, 1908 DICROSTEMA Benson, 1952 gracilicornis (Zaddach, 1859, Selandria ) E W

PHYMATOCERA Dahlbom, 1835 PECTINIA Brullé, 1846 HYPARGYRICUS MacGillivray, 1908 PHYTOMATOCERA : misspelling aterrima (Klug, 1816, Tenthredo ) E W S robinsoni (Curtis, 1850, Selandria ) fuliginosa (Schrank, 1781): misidentification

RHADINOCERAEA Konow, 1886 micans (Klug, 1816, Tenthredo ) E W

STETHOMOSTUS Benson, 1939 fuliginosus (Schrank, 1781, Tenthredo ) E W S fuscus (Serville, 1823, Tenthredo ) funereus (Klug, 1816, Tenthredo ) E

Tribe TOMOSTETHINI TOMOSTETHUS Konow, 1886 nigritus (Fabricius, 1804, Tenthredo ) E W

Tribe WALDHEIMIINI HALIDAMIA Benson, 1939 affinis (Fallén, 1807, Hylotoma ) E W S I hyalina (Klug, 1816, Tenthredo ) assimilis (Thomson, 1870, Blennocampa ) 19

Subfamily NEMATINAE Tribe CLADIINI Illiger, 1807 Subgenus CLADIUS Illiger, 1807 pectinicornis (Geoffroy, 1785, Tenthredo ) E W S I ? difformis (Panzer, 1799, Tenthredo ) crassicornis (Stephens, 1835, Nematus )

Subgenus PRIOPHORUS Dahlbom, 1835 brullei Dahlbom, 1835 E W S I immunis Stephens, 1835 melanostigma (Stephens, 1835, Nematus ) tener Zaddach, 1859 tristis Zaddach, 1859 morio : misidentification compressicornis (Fabricius, 1804, Tenthredo ) E W S I [note] albipes (Fallén, 1808, Tenthredo ) homonym pallipes Serville, 1823 padi : misidentification pilicornis Curtis, 1833 E W S I rufipes Serville, 1823 E S I ulmi (Linnaeus, 1758): misidentification ulmi (Linnaeus, 1758, Tenthredo ) E W S I eradiatus Hartig, 1837 laevifrons (Benson, 1936, Priophorus )

Subgenus TRICHIOCAMPUS Hartig, 1837 grandis (Serville, 1823, Nematus ) E S I viminalis (Fallén, 1808, Tenthredo ) homonym luteicornis Stephens, 1835

Tribe DINEURINI Marlatt, 1896 destructor Benson, 1952 E W S I duplex (Lepeletier, 1823): misidentification

DINEURA Dahlbom, 1835 stilata (Klug, 1816, Tenthredo ) E W S I testaceipes (Klug, 1816, Tenthredo ) E W S I virididorsata (Retzius, 1783, Tenthredo ) E W S I

HEMICHROA Stephens, 1835 LEPTOCERA Hartig, 1837 LEPTOCERCUS Thomson, 1871 australis (Serville, 1823, Tenthredo ) E W S I alni (Linnaeus, 1767, Tenthredo) homonym luctuosa (Hill, 1773, Tenthredo ) nomen oblitum crocea (Geoffroy, 1785, Tenthredo ) E W S I rufa (Panzer, 1799, Tenthredo ) homonym stigma Stephens, 1835

NEMATINUS Rohwer, 1911 acuminatus (Thomson, 1871, Nematus ) E S I caledonicus (Cameron, 1882, Nematus ) E S I nigrosternatus Malaise, 1931 fuscipennis (Serville, 1823, Nematus ) E W S I abdominalis (Fabricius, 1798): misidentification luteus (Panzer, 1804, Nematus ) E W S I willigkiae Stein, 1926 willigkiae ssp. pilosus Benson, 1958 20 steini Blank, 1998 E W S I alneti (Bechstein & Scharfenberg, 1805, Tenthredo ) homonym bilineatus (Klug,1819) : Cameron, 1877 misidentification ruficapillus (Gmelin, 1790): W. F. Kirby 1882, misidentification luteus (Panzer, 1804): misidentification

Unplaced species of : antennatus (Cameron, 1877, Nematus ) S [note]

PLATYCAMPUS Schiødte, 1839 LEPTOPUS Hartig, 1837 homonym CAMPONISCUS Newman, 1869 luridiventris (Fallén, 1808, Tenthredo ) E W S I niger (Stephens, 1835, Nematus ) alnivorus (Hartig, 1840, Nematus ) healaei (Newman, 1869, Camponiscus )

Tribe HOPLOCAMPINI Hartig, 1837 alpina (Zetterstedt, 1838, Tenthredo ) E S I pallida (Newman, 1837, Selandria ) homonym ariae Benson, 1933 E I brevis (Klug, 1816, Tenthredo ) E chrysorrhoea (Klug, 1816, Tenthredo ) E W S I crataegi (Klug, 1816, Tenthredo ) E W S I plagiata (Klug, 1816): Cameron,1885 misidentification flava (Linnaeus, 1761, Tenthredo ) E W S I ferruginea (Fabricius, 1804): misidentification minuta (Christ,1791): misidentification fulvicornis (Panzer, 1801, Tenthredo ) E W S I rutilicornis (Klug, 1816, Tenthredo ) pectoralis Thomson, 1871, E W S I gallicola Cameron, 1877 testudinea (Klug, 1816, Tenthredo ) E W S

Tribe MESONEURINI MESONEURA Hartig, 1837 opaca (Fabricius, 1775, Tenthredo ) E W S I verna (Klug, 1816, Tenthredo ) biloba (Stephens, 1835, Selandria ) selandriiformis (Cameron, 1875, )

Tribe Konow, 1890 abnormis (Holmgren, 1883, Nematus ) S tolli Konow, 1907 aulatus MacGillivray, 1919 amplus Konow, 1895 E S I fasciatus Konow, 1897 E W S I perkinsi Benson, 1933 variabilis Malaise, 1931: misidentification godmani Benson, 1955 S [note] hedstroemi Malaise, 1931 E W S I [note] rex Benson, 1948 tillbergi Malaise, 1920: misidentification histrio (Serville, 1823, Nematus ) E S I [note] rufescens (Hartig 1837, Nematus) glenelgensis (Cameron, 1882, Nematus ) humeralis (Serville, 1823, Nematus ) E W S terminalis Malaise, 1931 krausi Taeger & Blank, 1998 E 21

puniceus (Christ, 1791, Tenthredo ): misidentification lateralis Konow, 1896, E W S I trautmanni Enslin, 1919 cameroni Perkins, 1929 piliserra Lindqvist, 1943 imperfectus (Zaddach, 1876): Cameron, 1885 misidentification longiserra (Thomson, 1863, Nematus ) E S mcluckiei Benson, 1935, S pustulatus Lindqvist, 1962 arcticola Enslin, 1915: misidentification miltonotus (Zaddach, 1883, Nematus ) E W mundus Konow, 1895 E W S I sagmarius Konow, 1895 E S I semilacteus (Zaddach, 1883, Nematus ) S stenogaster (Förster, 1854, Nematus ) E W S I festivus Saarinen, 1950 fallax (Serville, 1823): misidentification toeniatus (Serville, 1823, Nematus ) E S taeniatus (Lepeletier, 1823, Nematus ) aemulus Konow, 1895 zetterstedti Malaise, 1920 alpicola Konow, 1895: misidentification tunicatus (Zaddach, 1883, Nematus ) E I vittatus (Serville, 1823, Nematus ) E W S I crispus Benson, 1948

BRACHYCOLUMA Strand, 1929 BRACHYCOLUS Konow, 1895 homonym DECANEMATUS Malaise, 1931 leucolenus (Brischke, 1883, Nematus ) E I Amauronematus leucolaenus : Benson, 1958 misspelling saarineni (Lindqvist, 1935): Benson, 1948 misidentification viduatus (Zetterstedt, 1838, Tenthredo ) E W S I notatus (Förster, 1854, Nematus ) longiserra (Thomson, 1863): Cameron, 1876 misidentification Amauronematus viduatus : Benson, 1958

CRAESUS Leach, 1817 CROESUS Curtis, 1824: misspelling alniastri (Scharfenberg, 1805, Tenthredo ) E W S I varus (Villaret, 1832, Nematus ) brischkei (Zaddach, 1876, Nematus ) E latipes (Villaret, 1832, Nematus ) E W S I septentrionalis (Linnaeus, 1758, Tenthredo ) E W S I stephensii Newman, 1837

EPICENEMATUS Lacourt, 1998 montanus (Zaddach, 1883, Nematus ) E W S I montanus : Benson, 1958

EUPONTANIA Zinovjev, 1985 [note] aquilonis (Benson, 1941, ) S algida (Benson,1941, Pontania ) arbusculae (Benson, 1941, Pontania ) S arcticornis (Konow, 1904, Pontania ) E S I phylicifoliae (Forsius, 1920, Pontania ) hepatimaculae (Malaise, 1920, Pontania ) brevicornis (Förster, 1854, Nematus ) E S curticornis (Cameron, 1885, Nematus ) saliciscinereae (Retzius, 1783, Tenthredo ): misidentification kriechbaumeri Konow, 1901: misidentification 22

pedunculi (Hartig, 1837): Kopelke 1991 misidentification collactanea (Förster, 1854, Nematus ) E S I vacciniellus (Cameron, 1876, Nematus ) crassipes (Thomson, 1871, Nematus ) S lapponica (Malaise, 1920, Pontania ) gallarum (Hartig, 1837, Nematus ) E S aestiva (Thomson, 1862, Nematus ) harrisoni (Benson, 1940, Pontania ) varia (Kopelke, 1991, Pontania ) herbaceae (Cameron, 1876, Nematus ) E W S I crassipes (Thomson, 1871): misidentification myrsiniticola (Kopelke, 1991, Pontania ) S, added by Liston & Blank (2006) pedunculi (Hartig, 1837, Nematus ) E S I bella (Zaddach, 1876, Nematus ) baccarum (Cameron, 1876, Nematus ) pusilla (Lindqvist, 1964, Pontania ) gallarum (Hartig, 1837): Kopelke, 1991 misidentification pustulator (Forsius, 1923, Pontania ) E S samolad (Malaise, 1920, Pontania ) S vesicator (Bremi, 1849, Nematus ) E S I helicina (Brischke, 1850, Nematus ) viminalis (Linnaeus, 1758, Cynips ) E W S I vollenhoveni (Cameron, 1874, Nematus ) ?interstitialis (Cameron, 1876, Nematus ) hungarica (Enslin, 1918, Pontania )

EUURA Newman, 1837 Subgenus Newman, 1837 CRYPTOCAMPUS Hartig, 1837 amerinae (Linnaeus, 1758, Cynips ) E S I salicispentandrae (Retzius, 1783, Tenthredo ) gallae Newman, 1837 medullarius (Hartig, 1837, Nematus ) saliceti (Fallén, 1808, Tenthredo ) mucronata (Hartig, 1837):Vollenhoven, 1871 misidentification atra (Jurine, 1807, Pteronus ) E W S I [note] auritae Kopelke, 2000 S purpureae Kopelke, 1996 I testaceipes (Brischke, 1883, Cryptocampus ) E S cynips Newman, 1837 nomen oblitum venusta (Brischke, 1883, Cryptocampus ) E W S weiffenbachii Ermolenko, 1988 S I added by Liston & O’Connor (2005)

Subgenus GEMMURA E.L.Smith, 1968 mucronata (Hartig, 1837, Nematus ) E W S I [note] ? nigritarsis Cameron, 1885 saliceti (Fallén, 1808): misidentification

LARINEMATUS Zhelochovtsev, 1988 imperfectus (Zaddach, 1876, Nematus ) E W S Pachynematus imperfectus : Benson, 1958

NEMATUS Panzer, 1801 Subgenus KONTUNIEMIANA Lacourt, 1998 leucotrochus Hartig, 1837 E W S I maculiventris Hartig, 1840 approximatus Förster, 1854 consobrinus Vollenhoven, 1871 cognatus (Lindqvist, 1957, Pteronidea ) olfaciens Benson, 1953 E S ribesii (Scopoli, 1763, Tenthredo ) E W S I 23

dimidiatus Serville, 1823 trimaculatus Serville, 1823 grossulariae Moore, 1831 ventricosus (Bouché, 1834, Tenthredo ) macrocerus Hartig, 1840 ribis Dufour, 1847

Subgenus NEMATUS Panzer, 1801 HYPOLAEPUS W. F. Kirby, 1882 HOLCOCNEME Konow, 1890 caeruleocarpus Hartig, 1837, E S I brevispinis Förster, 1854 brachyacanthus Thomson, 1863 sulcipes Hartig, 1837 coeruleocarpus : misspellling vicinus Serville, 1823 E W S I longispinus Kriechbaumer, 1885 crassus (Fallén, 1808, Tenthredo ) homonym sulcipes (Hartig, 1837): misidentification lucidus (Panzer, 1801, Tenthredo ) E W S I

Subgenus PTERONIDEA Rohwer, 1911 LINDQVISTIA Lacourt, 1998 bergmanni Dahlbom, 1835 E W S I pallicarpus Hartig, 1837 curtispina Thomson, 1871 pallens (Enslin, 1916, Lygaeonematus ) vernalis (Lindqvist, 1937, Pteronidea ) bipartitus Serville, 1823 E W S brevivalvis Thomson, 1871 E W S I kriegeri (Konow, 1903, Pteronus ) spurcus (Konow, 1904, Amauronematus ) absimilis (Lindqvist, 1939, Pteronidea ) cadderensis Cameron, 1875 E S I macroserratus (Lindqvist, 1943, Pteronidea ) dorsatus Cameron, 1875 W S fagi Zaddach, 1876 E W S fahraei Thomson, 1862 E S ferrugineus Förster, 1854 E S I glottianus Cameron, 1882 flavescens Stephens, 1835 E W S I stichi (Enslin, 1913): misidentification frenalis Thomson, 1888 S I fastosus (Konow, 1904, Pteronus ) fastuosus : misspelling fuscomaculatus Förster, 1854 E S strongylogaster Cameron, 1878 dossuarius (Konow, 1904, Pteronus ) hypoxanthus Förster, 1854 E W S I orbitalis Cameron, 1884 nigronotus (Lindqvist, 1957, Pteronidea ) incompletus Förster, 1854 E S I pulchellus Cameron, 1882 segmentarius Förster, 1854: misidentification jugicola Thomson, 1871 E S karvoneni (Lindqvist, 1969, Pteronidea ) leionotus (Benson, 1933, Pteronidea ) E S I [note] melanaspis Hartig, 1840 E W S I ? maculiger Cameron, 1882 [note] sveae (Lindqvist, 1958, Pteronidea ) melanocephalus Hartig, 1837 E W S I miliaris (Panzer, 1797, Tenthredo ) E S I capreae (Linnaeus, 1758, Tenthredo ): misidentification 24

croceus (Fallén, 1808): misidentification monticola Thomson, 1871 E S perkioemaekii (Lindqvist, 1960, Pachynematus ) ? pschornwalcheri Muche, 1972 similator Förster, 1854: misidentification myosotidis (Fabricius, 1804, Tenthredo ) E W S I ambiguus Förster, 1854 papillosus (Retzius, 1783): misidentification nigricornis Serville, 1823 E W S I nubium (Benson, 1935, Pteronidea ) S roberti (Lindqvist, 1958, Pteronidea ) oligospilus Förster, 1854 E W S I salicivorus Cameron, 1882 miliaris (Panzer, 1797): Cameron, 1885 misidentification capreae (Linnaeus, 1758): misidentification pavidus Serville, 1823, E W S I wttewaali Vollenhoven, 1862 cameronii Dalla Torre, 1894 aurantiacus Hartig, 1837: misidentification similator Förster, 1854: misidentification poecilonotus Zaddach, 1876, E W S I viridescens Cameron, 1885 subnitens (Lindqvist, 1957, Pteronidea ) palliatus Thomson, 1863: misidentification reticulatus Holmgren, 1883 S arcticus Thomson, 1871 homonym arcticola Dalla Torre, 1894 forsiusi (Enslin, 1915, Pontania ) alsius (Benson, 1935, Amauronematus ) salicis (Linnaeus, 1758, Tenthredo ) E W S spiraeae Zaddach, 1883, E W S I stichi (Enslin, 1913, Pteronidea ) E S I fuscarima (Benson, 1933, Pteronidea ) pallescens Hartig, 1837: Cameron, 1877 misidentification sylvestris Cameron, 1884, E S punctiscuta Hellén, 1948 ponojense Hellén, 1948 silvestris : misspelling tibialis Newman, 1837 E hortensis Hartig, 1837 trilineatus Norton, 1867 similaris Norton, 1880 robiniae Forbes, 1885 umbratus Thomson, 1871 E S I collinus Cameron, 1882 similis (Forsius, 1911, Pteronidea ) verrucosae (Kontuniemi, 1966, Pteronidea ) viridis Stephens, 1835 E W S I prasinus Hartig, 1837 brevivalvis Thomson, 1871: Konow, 1890 misidentification dispar Zaddach, 1876: misidentification bergmanni Dahlbom, 1835: misidentification viridissimus Möller, 1882 E W S I [note] ?glutinosae Cameron, 1882 polyspilus Förster, 1854: misidentification

PACHYNEMATUS Konow, 1890 EPITACTUS Förster, 1854 application submitted to ICZN for suppression of name calcicola Benson, 1948 E I chambersi Benson, 1948 laevigatus Zaddach, 1883: Benson, 1967 misidentification 25 clibrichellus (Cameron, 1878, Nematus ) E S thomsoni (Cameron, 1882, Nematus ) clibrichensis (Cameron, 1885, Nematus ): unjustified emendation hyperboreus (Thomson, 1871): Cameron, 1878 misidentification clitellatus (Serville, 1823, Nematus ) E W S I [note] trisignatus (Förster, 1854, Nematus ) foveolatus Konow, 1903 truncatus Benson, 1948 extensicornis (Norton, 1861): misidentification fallax (Serville, 1823, Nematus ) xanthocarpus (Hartig, 1840, Nematus ) E S I [note] virginalis Liston, 1982 kirbyi (Dahlbom, 1835, Nematus ) E W S I flaviventris (Hartig, 1840, Nematus ) melanocerus (Hartig, 1840, Nematus ) diaphanus (Eversmann, 1847, Nematus ) umbripennis (Eversmann, 1847, Nematus ) turgidus (Zaddach, 1876, Nematus ) lichtwardti Konow, 1903 E W S I apicalis (Hartig, 1837, Nematus ) homonym moerens (Förster 1854, Nematus ) E W S I pygostolus (Förster, 1854, Nematus ) pleuralis (Thomson, 1863, Nematus ) pullus Konow, 1903 torridonensis Liston, 1980 obductus (Hartig, 1837, Nematus ) E W S I conductus (Ruthe, 1859, Nematus ) graminis (Cameron, 1874, Nematus ) smithae Ross, 1945, E S angustatus Lindqvist, 1949 smithiae : misspelling sulcatus Benson, 1948, E S vagus (Fabricius, 1781, Tenthredo ) E W S I leucogaster (Hartig, 1840, Nematus ) inconspicuus (W. F. Kirby, 1882, Nematus )

PHYLLOCOLPA Benson, 1960 acutiserra (Lindqvist, 1949, Pontania ) S anglica (Cameron, 1877, Nematus ) E I [note] nigrolineata (Cameron, 1879, Nematus ) carinifrons (Benson, 1940, Pontania ) E S I [note] excavata (Marlatt, 1896): misidentification destricta (MacGillivray, 1923): misidentification apicifrons (Malaise, 1931): misidentification coriacea (Benson, 1953, Nematus ) E W S I [note] alienata (Förster, 1854, Nematus) nomen oblitum ischnocera (Thomson, 1863, Nematus ) [note] leucostigma (Cameron, 1876, Nematus ) leucapsis (Tischbein, 1846, Nematus ) E W S I leucosticta (Hartig, 1837, Nematus ) E W S I [note] sharpi (Cameron, 1876, Nematus ) piliserra (Thomson, 1863, Nematus ) E S I xanthogaster (Förster, 1854): Cameron, 1877 misidentification polita (Zaddach, 1883, Nematus ) S [note] oblita (Serville, 1823, Nematus ) E W puella (Thomson, 1871, Nematus ) E scotaspis (Förster, 1854, Nematus ) E S I westermanni (Boheman, 1852, Nematus ) nomen oblitum westermanni (Thomson, 1863, Nematus ) nomen oblitum

PIKONEMA Ross, 1937 26 scutellatum (Hartig, 1837, Nematus ) E W S I Pachynematus scutellatus : Benson, 1958

POLYNEMATUS Zhelochovtsev, 1988 albipennis (Hartig, 1837, Nematus ) E W S I sannio (Konow, 1903, Pachynematus ) Pachynematus albipennis : Benson,1958 annulatus (Gimmerthal, 1834, Nematus ) E S I Nematus flavipennis Cameron, 1876 Nematus arcticus Thomson, 1871: Cameron, 1878 misidentification rumicis (Linnaeus, 1758, Tenthredo ): Fallén, 1808 misidentification Pachynematus rumicis : Benson, 1958 glabriceps (Lindqvist, 1949, Pachynematus ) E parvilabris (Thomson, 1863): misidentification Pachynematus glabriceps : Benson, 1965

PONTANIA Costa, 1852 anomaloptera (Förster, 1854, Nematus ) E W S I [note] maidli (Zirngiebl, 1937, Amauronematus ) tuberculata (Benson, 1953, Nematus ) bridgmanii (Cameron, 1883, Nematus ) E W S I capreae (Linnaeus, 1758): misidentification dolichura (Thomson, 1871, Nematus ) E S robbinsi Benson, 1935 femoralis (Cameron, 1875, Nematus ) lapponicola Kopelke, 1994 S [note] dolichura (Thomson, 1871): misidentification nigricantis Kopelke, 1986 E S [note] dolichura (Thomson, 1871): misidentification proxima (Serville, 1823, Nematus ) E W S I gallicola (Stephens, 1835, Nematus ) flavipes (Cameron, 1885, Euura ) capreae (Linnaeus, 1758): misidentification purpureae (Cameron, 1884, Nematus ) E triandrae Benson, 1941 E S I capreae (Linnaeus, 1758): misidentification

PONTOPRISTIA Malaise, 1921 amentorum (Förster, 1854, Nematus ) E S suavis (Ruthe, 1859, Nematus ) kamtchaticus (Malaise, 1931, Pontopristia ) [note] Amauronematus amentorum : Benson, 1958

Tribe PRISTIPHORINI PRISTICAMPUS Zinovjev, 1992 arcticus (Lindqvist, 1958, Mesoneura ) S Pachynematus arcticus : Benson, 1961

PRISTIPHORA Latreille, 1810 Subgenus GYMNONYCHUS Marlatt, 1896 abbreviata (Hartig, 1837, Nematus ) E californica (Marlatt, 1896, Gymnonychus ) biscalis (Förster, 1854, Nematus ) E W I maesta (Zaddach, 1876, Nematus ) E brevicornis (Thomson, 1862, Nematus ) homonym parvicornis (W. F. Kirby, 1882, Nematus ) insularis (Malaise, 1921, Pachynematus ) moesta : misspelling retusa (Thomson, 1871, Nematus ) S 27

Subgenus LYGAEONEMATUS Konow, 1890 abietina (Christ, 1791, Tenthredo ) E W S I pini (Retzius, 1783, Tenthredo ) homonym compressa (Hartig, 1837, Nematus ) E S decipiens (Enslin, 1916, Lygaeonematus ) E, added by Liston (1981b) erichsonii (Hartig, 1837, Nematus ) E W S I leachii (Dahlbom, 1835, Nematus ): suppressed glauca Benson, 1954 E W ? laricivora (Takagi, 1931, Pachynematus ) homonym pseudodecipiens Beneš & Kristek, 1979 E S, added by Beneš & Kristek (1979) decipiens (Enslin, 1916): misidentification saxesenii (Hartig, 1837, Nematus ) E W S thalenhorsti Wong, 1975 subarctica (Forsslund, 1936, Lygaeonematus ) E pseudosaxesenii Lindqvist, 1968 wesmaeli (Tischbein, 1853, Nematus ) E W S I

Subgenus LYGAEOPHORA Hellén, 1975 LYGAEOPHORA Lindqvist, 1952 unavailable name sermola Liston, 1993 S, added by Liston (1981a) [note] variipes (Lindqvist, 1952, Lygaeonematus ) homonym lanifica (Zaddach, 1883): Liston, 1981a misidentification

Subgenus LYGAEOTUS Hellén, 1975 LYGAEOTUS Lindqvist, 1952 unavailable name albilabris (Boheman, 1852, Nematus ) E albilabris (Thomson, 1863, Nematus ) alpestris (Konow, 1903, Lygaeonematus ) E Lygaeonematus strandi Konow, 1901: Benson, 1934 misidentification borea (Konow, 1904, Lygaeonematus ) S astuta (Cameron, 1885, Nematus ) nomen oblitum [note] lapponica (Enslin, 1916, Pachynematus ) breadalbanensis (Cameron, 1882, Nematus ) E W S I tromsoensis (Kiaer, 1898, Lygaeonematus ) corpulentus (Konow, 1904, Lygaeonematus ) arcticola (Enslin, 1916, Lygaeonematus ) carinata (Hartig, 1837, Nematus ) E W S pallipes (Fallén, 1808, Tenthredo ) homonym coactula (Ruthe, 1859, Nematus ) S alpina (Thomson, 1871, Nematus ) pachyvalvis (Konow, 1904, Lygaeonematus ) groenblomi (Lindqvist, 1952, Lygaeonematus ) S [note] gronblomi : misspelling lativentris (Thomson, 1871, Nematus ) S scotica (Cameron, 1881, Nematus ) extrema (Holmgren, 1883, Nematus ) mollis (Hartig, 1837, Nematus ) E W S I whitei (Cameron, 1878, Nematus ) kontkaneni (Lindqvist, 1960, Pachynematus ) pseudocoactula (Lindqvist, 1952, Lygaeonematus ) E S pachyvalvis (Konow, 1904): Benson, 1934 misidentification

Subgenus MICRONEMATUS Konow, 1890 monogyniae (Hartig, 1840, Nematus ) E W I hibernica (Cameron, 1878, Nematus )

Subgenus OLIGONEMATUS Zhelochovtsev, 1988 friesei (Konow, 1904, Lygaeonematus ) I, added by O’Connor & Liston (1994) funerula (Costa, 1859): misidentification laricis (Hartig, 1837, Nematus ) E W S I leucocnemis (Förster, 1854, Nematus ) 28

oblonga (Cameron, 1882, Nematus ) laricivora (Brischke, 1883, Nematus ) ravida (Konow, 1903, Pachynematus )

Subgenus PRISTIPHORA Latreille, 1810 DIPHADNUS Hartig, 1837 aphantoneura (Förster, 1854, Nematus ) E [note] fulvipes (Fallén, 1808, Tenthredo ) homonym vicina Serville, 1823: misidentification appendiculata (Hartig, 1837, Nematus ) E S I pallipes Serville, 1823, homonym grossulariae Walsh, 1866 rufipes Serville, 1823: misidentification armata (Thomson, 1862, Nematus ) E W S I crassicornis (Hartig, 1837, Nematus ) homonym fletcheri (Cameron, 1884, Nematus ) nigricollis (Cameron, 1885, Nematus ) bifida (Hellén, 1948, Nematus ) S brevis (Hartig, 1837, Nematus ) E I fumipennis (Thomson, 1871, Nematus ) fuscata Benson, 1943 thalictri (Kriechbaumer, 1884): misidentification cincta Newman, 1837, E W S I [note] quercus (Hartig, 1837, Nematus ) confusa Lindqvist, 1955 E W S coniceps Lindqvist, 1955 E S conjugata (Dahlbom, 1835, Nematus ) E denudata Konow, 1902 E W S I [note] atlantica Lacourt, 1987: misidentification geniculata (Hartig, 1840, Nematus ) E I insularis Rohwer, 1910 E W S I [note] kamtchatica Malaise, 1931 mesatlantica Lacourt, 1976 luteiventris Koch, 1989 paedida (Konow, 1904): misidentification leucopus (Hellén, 1948, Nematus ) E, added by Grearson (2006) luteipes Lindqvist, 1955 E W S I, added by O’Connor et al. (1997) [note] pygmaea Lindqvist, 1964 melanocarpa (Hartig , 1840, Nematus ) E W S I wuestneii (Stein, 1885, Nematus ) pallidiventris (Fallén, 1808, Tenthredo ) E W S I ephippiger (Hartig, 1840, Nematus ) flavicoma (Tischbein, 1840, Nematus ) nigricans (Eversmann, 1847, Nematus ) caudalis (Eversmann, 1847, Nematus ) breviuscula (Eversmann, 1847, Nematus ) gemella (Förster, 1854, Nematus ) cirrhostoma (Zaddach, 1883, Nematus ) Pristiphora myosotidis (Fabricius, 1804): Stephens, 1835 misidentification Nematus obductus (Hartig, 1837): Cameron, 1874 misidentification punctifrons (Thomson, 1871, Nematus ) E W S I ruficornis (Olivier, 1811, Nematus ) E W S I fraxini (Hartig, 1837, Nematus ) rufipes Serville, 1823 E S aquilegiae (Vollenhoven, 1866, Nematus ) alnivora (Hartig, 1840): misidentification staudingeri (Ruthe, 1859, Nematus ) E W S I circularis Kincaid, 1900 ? hyperborea Malaise, 1921 [note] asperlatus Benson, 1935 subbifida (Thomson, 1871, Nematus ) E W [note] 29

carpentieri Konow, 1902 subopaca Lindqvist, 1955 E [note] testacea (Jurine, 1807, Pteronus ) E W I betulae (Retzius, 1783, Tenthredo ) homonym crassiventris (Cameron, 1878, Nematus )

SHARLIPHORA Wong, 1969 amphibola (Förster, 1854, Nematus ) E W S laeta (Cameron, 1883, Nematus ) fraterna ( Cameron, 1885, Nematus ) nigella (Förster, 1854, Nematus ) E S I ambigua (Fallén, 1808, Tenthredo ) homonym furvescens (Cameron, 1875, Nematus )

STAURONEMATUS Benson, 1953 STAURONEMA Benson, 1948 homonym platycerus (Hartig, 1840, Nematus ) E W [note] compressicornis (Fabricius, 1804): misidentification

Tribe PSEUDODINEURINI ENDOPHYTUS Hering, 1934 NEOPELMATOPUS Conde, 1934 anemones (Hering, 1924, Pelmatopus ) E tenuiserra (Lindqvist, 1949, Pelmatopus )

PSEUDODINEURA Konow, 1885 PELMATOPUS Hartig, 1837 homonym PHYLLOPAIS Hering, 1934 enslini (Hering, 1923, Pelmatopus ) E S fuscula (Klug, 1816, Tenthredo ) E S I despecta (Hartig, 1837, Dineura ) simulans (Cameron, 1877, Dineura ) mentiens (Thomson, 1871, Blennocampa ) S, added by Liston & Blank (2006)

Unplaced species of Nematinae: Nematus placidus Cameron, 1878 [note] Nematus v-flavum Cameron, 1882

Subfamily SELANDRIINAE Tribe ANEUGMENINI ANEUGMENUS Hartig, 1837 ATOPOSELANDRIA Enslin, 1913 coronatus (Klug, 1818, Tenthredo ) E S I [note] analis (Thomson, 1871, Selandria )) Selandria cereipes Vollenhoven, 1873 fuerstenbergensis (Konow, 1885, Selandria ) E S I [note] furstenbergensis : misspelling padi (Linnaeus, 1761, Tenthredo ) E W S I stramineipes (Klug, 1816, Tenthredo ) temporalis (Thomson, 1871, Selandria ) E W S I

Tribe DOLERINI DOLERUS Panzer, 1801 Subgenus ACHAETOPRION Goulet, 1986 JUNCILERUS Zhelochovtsev, 1988 ferrugatus Serville, 1823 E W S I madidus (Klug, 1818, Tenthredo ) E W S I lateritius (Klug, 1818, Tenthredo ) chappelli Cameron, 1877 schulthessi Konow, 1887: misidentification 30 pachycerus Hartig, 1837 E W S taeniatus Zaddach, 1859 tinctipennis Cameron, 1881 triplicatus (Klug, 1818, Tenthredo ) E W steini Konow, 1885

Subgenus CYPEROLERUS Zhelochovtsev, 1988 anticus (Klug, 1818, Tenthredo ) E

Subgenus DICRODOLERUS Goulet, 1986 vestigialis (Klug, 1818, Tenthredo ) E W S I genucinctus Zaddach, 1859: Benson, 1934 misidentification Loderus vestigialis : Benson, 1952

Subgenus DOLERUS Panzer, 1801 DOLERUS Jurine, 1801 suppressed DOSYTHEUS Leach, 1817 bajulus Serville, 1823 E W S I aericeps Thomson, 1871 bimaculatus (Geoffroy, 1785, Tenthredo ) E W S I tristis (Fabricius, 1804, Tenthredo ) homonym cothurnatus Serville, 1823 E W S I palustris (Klug, 1818, Tenthredo ) homonym junci (Stephens, 1835, Dosytheus ) germanicus (Fabricius, 1775, Tenthredo ) E W S I fuscipennis (Stephens,1835, Dosytheus ) fulviventris (Scopoli, 1763): Cameron, 1882 misidentification pratensis (Linnaeus, 1758): misidentification gessneri André, 1880 E W S I labiosus Konow, 1897 yukonensis Norton, 1872 E W S scoticus Cameron, 1881 [note] lateralis Konow, 1895 arcticola Kiaer, 1898 saxatilis Hartig, 1837: Benson, 1952 misidentification

Subgenus EQUIDOLERUS Taeger & Blank, 1996 DOSYTHEUS : Goulet, 1986 misidentification pratensis (Linnaeus, 1758, Tenthredo ) E W S I dubius (Klug, 1818, Tenthredo ) timidus (Klug, 1818, Tenthredo )

Subgenus LODERUS Konow, 1890 gilvipes (Klug, 1818, Tenthredo ) E S pratorum ssp. gilvipes (Klug, 1818) pratorum (Fallén, 1808, Tenthredo ) E W

Subgenus ONCODOLERUS Goulet, 1986 eversmanni W. F. Kirby, 1882 E W S I palmatus (Klug, 1818, Tenthredo ) homonym Loderus eversmanni : Benson, 1952

Subgenus POODOLERUS Zhelochovtsev, 1988 aeneus Hartig, 1837 E W S I elongatus Thomson, 1871 anthracinus (Klug, 1818, Tenthredo ) E W S asper Zaddach, 1859, E S [note] carbonarius Zaddach, 1859 oblongus Cameron, 1882 planatus Hartig, 1837: misidentification brevicornis Zaddach, 1859 E [note] 31

asper Zaddach, 1859: misidentification coracinus (Klug, 1818, Tenthredo ) E S fumosus Stephens, 1835 E W S sanguinicollis (Klug, 1818): misidentification gonager (Fabricius, 1781, Tenthredo ) E W S haematodes (Schrank, 1781, Tenthredo ) E W S I collaris (Donovan, 1808, Tenthredo ) harwoodi Benson, 1947 S liogaster Thomson, 1871 E W S I [note] niger (Linnaeus, 1767, Tenthredo ) E W S I nigratus (Müller, 1776, Tenthredo ) E W S I fissus Hartig, 1837 nitens Zaddach, 1859 E W S I coruscans Konow, 1890 wanda Ross, 1935 picipes (Klug, 1818, Tenthredo ) E W S I intermedius Cameron, 1881 possilensis Cameron, 1882 E W S I nitens Zaddach, 1859: misidentification puncticollis Thomson, 1871 E W S I schmidti Konow, 1884 E S [note] stygius Förster, 1860 E W S megapterus Cameron, 1881 varispinus Hartig, 1837 E W S I [note] rugosus Konow, 1884 rugosulus Dalla Torre, 1894 schneideri Kiaer, 1898 brevitarsis : Benson, 1947 misidentification and misspelling liogaster Thomson, 1871: misidentification

Tribe DULOPHANINI BIRKA Malaise, 1944 cinereipes (Klug, 1816, Tenthredo ) E W S I aperta (Hartig, 1837, Tenthredo )

DULOPHANES Konow, 1907 NESOSELANDRIA Rohwer, 1910 MELISANDRA Benson, 1939 morio (Fabricius, 1781, Tenthredo ) E W S I

Tribe HEPTAMELINI HEPTAMELUS Haliday, 1855 MELICERTA Stephens, 1835 homonym CAENONEURA Thomson, 1870 dahlbomi (Thomson, 1870, Caenoneura ) E S I [note] ochroleucus : misidentification ochroleucus (Stephens, 1835, Melicerta ) E W S I

Tribe SELANDRIINI BRACHYTHOPS Haliday in Curtis, 1839 CORYNA Lepeletier & Serville, 1825 homonym flavens (Klug, 1816, Tenthredo ) E W S I scapularis (Lepeletier & Serville, 1828, Tenthredo ) seminigra Haliday in Curtis, 1839 flavescens (Thomson, 1870, Selandria ) wuestneii (Konow, 1885, Selandria ) E W S flavistigma (Groenblom, 1939, Selandria )

SELANDRIA Leach, 1817 serva (Fabricius, 1793, Tenthredo ) E W S I dorsalis Stephens, 1835 32

excisa Konow, 1885 fuscitarsis Benson, 1954 melanosterna (Serville, 1823, Tenthredo ) E W S sixii Vollenhoven, 1858 grandis Zaddach, 1859 interstitialis Thomson, 1871 dorsalis Stephens, 1835: W. F. Kirby, 1882 misidentification

Tribe STRONGYLOGASTRINI PSEUDOHEMITAXONUS Conde, 1932 sharpi (Cameron, 1879, Strongylogaster ) E S exsectus Conde, 1932 mixtus (Klug, 1817): Cameron, 1874 misidentification Strongylogaster contigua (Konow, 1885): misidentification

STROMBOCEROS Konow, 1885 STROMBOCERINA Malaise, 1942 STROMBOCERUS : misspelling delicatulus (Fallén, 1808, Tenthredo ) E W S I Strongylogaster eborina (Klug, 1817, Tenthredo ) Selandria phthisica Vollenhoven, 1869

STRONGYLOGASTER Dahlbom, 1835 PSEUDOTAXONUS Costa, 1894 filicis (Klug, 1817, Tenthredo ) E extinct [note] multifasciata (Geoffroy, 1785, Tenthredo ) E W S I lineata (Christ, 1791, Tenthredo ) atricornis (Stephens, 1835, Tenthredo ) cingulata (Fabricius, 1793): misidentification macula (Klug, 1817, Tenthredo ) E W S I intermedia (Konow, 1885, Thrinax ) mixtus (Klug, 1817): Cameron, 1874 misidentification and misspelling mixta (Klug, 1817, Tenthredo ) E W S I femoralis Cameron, 1875 xanthocera (Stephens, 1835, Tenthredo ) E S geniculata Thomson, 1871 xanthoceros : misspelling

Subfamily

Tribe MACROPHYINI Dahlbom, 1835 Subgenus MACROPHYA Dahlbom, 1835 albicincta (Schrank, 1776, Tenthredo ) E W S albipuncta (Fallén, 1808, Tenthredo ) E S alboannulata Costa, 1859, E W, added by Liston (1983) annulata (Geoffroy, 1785, Tenthredo ) E W neglecta (Klug, 1817, Tenthredo ) blanda (Fabricius, 1775, Tenthredo ) E duodecimpunctata (Linnaeus, 1758, Tenthredo ) E W S I montana (Scopoli, 1763, Tenthredo ) E W S rustica (Linnaeus, 1758): misidentification ribis (Schrank, 1781, Tenthredo ) E W rufipes (Linnaeus, 1758, Tenthredo ) E W ione (Newman, 1837, Allantus ) haematopus (Panzer, 1801): Cameron, 1882 misidentification

Subgenus PSEUDOMACROPHYA Enslin, 1913 punctumalbum (Linnaeus, 1767, Tenthredo ) E W I erythropus (Schrank, 1776, Tenthredo ) 33

parvula Konow, 1884: Liston, 1987 misidentification

PACHYPROTASIS Hartig, 1837 antennata (Klug, 1817, Tenthredo ) E W S I nigronotata Kriechbaumer, 1874 W rapae (Linnaeus, 1767, Tenthredo ) E W S I simulans (Klug, 1817, Tenthredo ) E W S variegata (Fallén, 1808, Tenthredo ) E W S

Tribe SCIAPTERYGINI SCIAPTERYX Stephens, 1835 SCIOPTERYX Cameron, 1882: misspelling consobrina (Klug, 1816, Tenthredo ) E soror Konow, 1890, Sciapteryx E S costalis (Fabricius, 1775): misidentification

Tribe TENTHREDININI CYTISOGASTER Lacourt, 1996 chambersi (Benson, 1947, Rhogogaster ) E W S genistae (Benson, 1947, Rhogogaster ) E W S picta (Klug, 1817, Tenthredo ) E W

RHOGOGASTER Konow, 1884 RHOGOGASTERA Konow, 1885 dryas (Benson, 1943, Tenthredo ) E S similis Lindqvist, 1959 punctulata (Klug, 1817, Tenthredo ) E W S chlorosoma (Benson, 1943, Tenthredo ) E W S viridis (Linnaeus, 1758, Tenthredo ) E W S I scalaris (Klug, 1817, Tenthredo )

TENTHREDO Linnaeus, 1758 Subgenus CEPHALEDO Zhelochovtsev, 1988 neobesa Zombori, 1980, E ?extinct pseudorossii Taeger, 1985 rossii (Panzer, 1804): Benson, 1952 misidentification temula Scopoli, 1763: Cameron misidentification vidua Rossi, 1790: Cameron misidentification

Subgenus ELINORA Benson, 1940 CUNEALA Zirngiebl, 1956 MURCIANA Lacourt, 1988 BLANKIA Lacourt, 1998 dominiquei (Konow, 1894, Allantus ) E [note] flavipes Geoffroy, 1785: Cameron, 1882 misidentification flaveola Gmelin, 1790: misidentification

Subgenus EUROGASTER Zirngiebl, 1953 mesomela Linnaeus, 1758 E W S I mesomelas : misspelling mioceras (Enslin, 1912, Tenthredella ) E S I obsoleta Klug, 1817 E S I

Subgenus MACULEDO Zhelochovtsev, 1988 maculata Geoffroy, 1785 E W S

Subgenus OLIVACEDO Zhelochovtsev, 1988 olivacea Klug, 1817 E W S

Subgenus TEMULEDO Zhelochovtsev, 1988 temula Scopoli, 1763 E W S I 34

bicincta Linnaeus, 1767 celtica Benson, 1953

Subgenus TENTHREDELLA Rohwer, 1910 atra Linnaeus, 1758 E W S I dispar Klug, 1817 scotica Cameron, 1882 balteata Klug, 1817 E W S I colon Klug, 1817 E W S I nigricollis W. F. Kirby, 1882 fagi Panzer, 1798 E S solitaria Scopoli, 1763: Cameron, 1882 misidentification ferruginea Schrank, 1776 E W S I rufiventris Panzer, 1799 livida Linnaeus, 1758 E W S I mandibularis Fabricius, 1804 E W moniliata Klug, 1817 E W S I lachlaniana Cameron, 1878 velox Fabricius, 1798 E W S velox ab. nigrolineata Cameron, 1878 infrasubspecific name

Subgenus TENTHREDO Linnaeus, 1758 arcuata Forster, 1771 E W S I brevicornis (Konow, 1886, Allantus ) E W S I nitidior (Konow, 1888, Allantus ) aegra Enslin, 1912 acerrima Benson, 1952 sulphuripes (Kriechbaumer, 1869): Benson, 1940 misidentification notha Klug, 1817 E W S I perkinsi (Morice, 1919, Allantus ) schaefferi Klug, 1817: Benson, 1959 misidentification omissa (Förster, 1844, Allantus ) E marginella Fabricius, 1793: Thomson, 1871 misidentification schaefferi Klug, 1817 E scrophulariae Linnaeus, 1758 E W S thompsoni (Curtis, 1839, Allantus ) E W marginella Fabricius, 1793: misidentification vespa Retzius, 1783 E W S multifasciata Geoffroy, 1785: misidentification tricincta Fabricius, 1804 quadricincta Fallén, 1808 zona Klug, 1817 E quadricincta Fallén, 1808: Cameron, 1882 misidentification

Subgenus ZONULEDO Zhelochovtsev, 1988 amoena Gravenhorst, 1807 E W cingulum Klug, 1817 Allantus inversus Costa, 1894 distinguenda (Stein, R., 1885, Allantus ) E

Tribe TENTHREDOPSINI AGLAOSTIGMA W. F. Kirby, 1882 aucupariae (Klug, 1817, Tenthredo ) E W S I Tenthredo solitaria Fallén, 1808 homonym Tenthredo juvenilis Serville, 1823 Tenthredo gibbosa Fallén, 1808: Thomson, 1870 misidentification fulvipes (Scopoli, 1763, Tenthredo ) E W S I lateralis (Fabricius, 1779, Tenthredo )

PERINEURA Hartig, 1837 SYNAIREMA Hartig, 1837 35 rubi (Panzer, 1805, Allantus ) E W S I

TENTHREDOPSIS Costa, 1859 EBOLIA Costa, 1859 THOMSONIA Konow, 1884 EUTENTHREDOPSIS Enslin, 1913 coquebertii (Klug, 1817, Tenthredo ) E W S I nigricollis Cameron, 1881 opulenta Konow, 1887 ignobilis (Klug, 1817): Cameron, 1882 misidentification excisa (Thomson, 1870, Perineura ) E tesselata : Benson, 1968 misidentification and misspelling friesei (Konow, 1884, Thomsonia ) E ? palmata (Geoffroy, 1785, Tenthredo ) laticeps (Konow, 1884, Thomsonia ) korlevici Konow, 1887 arrogans Konow, 1890 pavida (Fabricius, 1775): misidentification litterata (Geoffroy, 1785, Tenthredo ) E W S carbonaria (Linnaeus, 1767, Tenthredo ) homonym 36

cordata (Geoffroy, 1785 Tenthredo ) microcephala (Serville, 1823, Tenthredo ) caliginosa (Stephens, 1835, Tenthredo ) femoralis (Stephens, 1835, Tenthredo ) nigriceps Cameron, 1881 nigronotata Cameron, 1881 pallida Konow, 1896 nassata (Linnaeus, 1767, Tenthredo ) E W S I alneti Schrank, 1781 dorsalis (Serville, 1823, Tenthredo ) fulviceps (Stephens, 1835, Tenthredo ) tristis (Stephens, 1835, Tenthredo ) albomaculata Cameron, 1881 dorsivittata Cameron, 1881 inornata Cameron, 1881 lividiventris Cameron, 1881 saundersi Cameron, 1881 elegans (Konow, 1884, Thomsonia ) gibberosa Konow, 1887 fenestrata Konow, 1890 tristior Morice, 1914 sordida (Klug, 1817): misidentification scutellaris (Fabricius, 1804, Tenthredo ) E S I spreta (Serville, 1823, Tenthredo ) picticeps Cameron, 1881 flavomaculata Cameron, 1881 austriaca Konow, 1890 dubia Konow, 1890 parvula Konow, 1890 puncticollis Konow, 1890 thornleyi Konow, 1899 nassata : misidentification campestris (Linnaeus, 1758): misidentification

Unplaced species of Tenthredinidae: Selandria ornata Newman, 1838

Superfamily XIPHYDRIOIDEA Family XIPHYDRIIDAE XIPHYDRIA Latreille, 1802 HYBONOTUS Klug, 1803 XIPHIURA Fallén, 1813 XIPHIDRIA Lamarck, 1817 HYPONOTUS Billberg, 1820 XYPHIDRIA Lepeletier, 1823 KONOWIA Brauns, 1884 PSEUDOXIPHYDRIA Enslin, 1911 camelus (Linnaeus, 1758, Ichneumon ) E W S longicollis (Geoffroy, 1785, Tenthredo ) E, added by Shaw & Liston (1985) prolongata (Geoffroy, 1785, Tenthredo ) E W dromedarius (Fabricius, 1787) 37

Superfamily XYELOIDEA Family XYELIDAE Subfamily XYELINAE Tribe Xyelini XYELA Dalman, 1819 PINICOLA Brébisson, 1818 homonym XYELATANA Benson, 1938 curva Benson, 1938 E, added by Liston & Blank (2006) julii (Brébisson, 1818, Pinicola ) E W S I longula Dalman, 1819 S piliserra Thomson, 1871 38

SPECIES REMOVED FROM BRITISH LIST

Cephalcia arvensis Panzer, 1805. Recorded by Shinohara (2002), deleted by Shaw (2003) because material probably not of British origin.

Pamphilius jucundus (Eversmann, 1847, Lyda ) ( nemorum auctt.). See Liston et al. (2006).

Pristiphora karvoneni (Lindqvist, 1952, Lygaeonematus ). Recorded by Liston (1983), but based on misidentification.

Pristiphora micronematica Malaise, 1931. Recorded by Liston (1982), but based on misidentification.

Megalodontes plagiocephalus (Fabricius, 1804). Up to three separate Megalodontes species have at times been considered to have been present in England (see also below). M. plagiocephalus , on the basis of its currently known distribution and habitat requirements, seems unlikely ever to have occurred in the British Isles.

NOTES

Orussus abietinus . Recorded from Kent and Devon by Stephens (1835), but see also comments in Benson (1951).

Megalodontes. If any member of this genus was ever present in England, then only M. cephalotes seems at all likely: according to Leach (1817) and Stephens (1835), found near Bristol and Plymouth. See Taeger (2002) for current taxonomy.

Pamphilius gyllenhali . See ICZN (2005) Opinion 2101.

Pamphilius latifrons . See ICZN (2005) Opinion 2101.

Sirex atricornis . Considered by Viitasaari & Midtgaard (1989) to be a valid species, which may occur in Scotland.

Urocerus augur , U. fantoma and U. tardigradus. Have at times been recorded as introductions to the British Isles, but only U. augur is regarded as established (O’Connor et al. 1997). All were treated as valid species by Smith (1978), although U. tardigradus had for a long time been placed as a synonym of U. fantoma . Benson (1958: ii, supplement to section a) added U. tardigradus to the British list with “fantoma Fabricius auctt. nec Fabricius” as its synonym and separated this in a key from “ U. fantoma fantoma (= augur augur Klug)”. The reasons for this treatment are not clear. No type specimens are known to exist for either U. fantoma or U. tardigradus , but Fabricius (1781) in his short original description of Sirex fantoma describes the legs as yellow, whereas U. augur has extensively black tibiae. Specimens previously identified as U. tardigradus have so far all proved to belong to U. fantoma (A. Taeger, pers. comm..). We revert here to treating U. tardigradus as a synonym of U. fantoma .

Urocerus bizonatus. This is a synonym of the Nearctic colour form also referred to as U. gigas ssp. flavicornis (Fabricius, 1781). Recorded as an introduction, but not known to be established in the British Isles.

Sirex bimaculatus , was described by Donovan (1808), according to a reference in Stephens (1829), but the original description has not been seen. Stephens placed S. bimaculatus as a synonym of Sirex augur Klug. S. bimaculatus does not seem to have been mentioned, even in catalogues, after 1829.

Arge stephensii . Differs very markedly, in the extreme development of pale body markings, from all known W. Palaearctic populations of A. pagana . Traditionally, A. stephensii has been regarded as an endemic British subspecies of A. pagana. Further studies [here a DNA analysis might offer good clues] are definitely required. The possibility can not be excluded, that A. stephensii is a separate species originally introduced from the E. Palaearctic or Oriental Regions, where Arge is both extremely species rich, and poorly investigated.

Abiinae. Benson (1951) treated the genera Abia and Zaraea as belonging to two different tribes. The differences between these certainly do not warrant this (Lorenz & Kraus 1957, Taeger 1998).

Cimbex quadrimaculatus . Presence in British Isles based on one female (NHM) said to have been collected 39 near Salisbury (Leach 1817; Stephens 1835).

Pseudoclavellaria amerinae . Reported as having been collected near Windsor (Leach 1817; Stephens 1835).

Trichiosoma. Taxonomy follows Viitasaari (1985, 1989).

Trichiosoma laterale . O’Connor et al. (1997: 17) were unable to trace any Irish specimens.

Trichiosoma pusillum . Possibly a synonym of T. tibiale (Viitasaari 1989).

Corynis crassicornis . Said to have been collected near Bristol (Leach 1817; Stephens 1835).

Corynis obscura . Said to have been collected in Lincolnshire (Leach 1817; Stephens 1835).

Allantus . According to Stephens (1835: 90), who gives a recognisable description of A. didymus (Klug, 1818), this species was “..rare: taken at Birch wood in June.” It is not included in the list,of British species, because no authenticated material has been located. A specimen without locality data in the Stephens Collection (BMNH) is A. melanarius (det. G. Knight).

Allantus basalis ssp. caledonicus . May represent a separate species (Blank & Taeger 1998) from Scottish specimens recorded as A. basalis ssp. basalis by Liston (1985).

Allantus togatus . O’Connor et al. (1997: 37) were unable to trace the single recorded Irish specimen.

Taxonus agrorum . O’Connor et al. (1997: 36) were unable to trace any Irish specimens.

Athalia ancilla . O’Connor et al. (1997: 30) were unable to trace any Irish specimens.

Athalia circularis . There are grounds for suspecting that this consists of a group of species, with more narrowly defined hostplant spectra than presently accepted.

Taxonus glottianus Cameron was justifiably synonymised with Ametastegia tenera by Konow (1899), because the original description strongly indicates the latter species, but after examination of the type Benson (1952) regarded T. glottianus as a junior synonym of A. carpini . We follow Benson’s decision on the status of Cameron’s name.

Empria immersa . The description of Emphytus immersus by Stephens (1835: 92) certainly is of Empria testaceipes (Konow, 1896) and not E. immersa as interpreted by eg.Dalla Torre (1894). Stephens referred to the taxon as “..scarce; taken in the vicinity of London in June”. E. testaceipes may therefore once have occurred in England, but until authenticated material has been found, we refrain from placing it on the list of British species.

Empria pumila . E. pumiloides Lindqvist, 1968 is distinguished from E. pumila by Heidemaa & Viitasaari (1999). Although a few Scottish and Welsh specimens checked by AL are all E. pumila , the presence of E. pumiloides in the British Isles can not be ruled out.

Cladardis elongatula . Presence in the British Isles uncertain: see Gibbs (2006).

Claremontia alchemillae . The taxonomy of this and the following two species has not yet been fully resolved (Liston et al. 2006).

Claremontia tenuicornis . Previous records of this species in the British Isles seem to refer to C. alchemillae and C. uncta .

Claremontia uncta. See above under C. alchemillae / C. tenuicornis.

Fenella nigrita . Benson (1952) had noted that biological differences possibly indicate the presence of two taxa under this name in the British Isles. This problem has still not been clarified.

Fenusa altenhoferi . Synonymy with F. carpinifoliae according to Liston (1996). Dolerus pusillus Serville can not at present be placed as conspecific with F. ulmi or F. altenhoferi , and must be treated as an unplaced species (Blank et al. 2001). 40

Fenusa ulmi . Recorded in Ireland by Knight (2004) as Kaliofenusa pusilla.

Scolioneura . Taxonomy follows Altenhofer & Taeger (1998).

Phyllotoma aceris McLachlan was described on the basis of specimens "bred by Mr Healy from larvae which make great blotches in the leaves of Acer campestre , and occasionally in Acer pseudo-platanus " (McLachlan,1867). According to present knowledge on distribution and hostplants, the syntype series probably consisted of more than one species. The description is not sufficiently detailed to determine the identity of the specimens which McLachlan examined. No type material has been located, as already stated by Altenhofer & Zombori (1987). Nevertheless, Altenhofer & Zombori (1987) chose to regard P. aceris McLachlan as being conspecific with a species which E. Altenhofer reared from A. campestre and proposed for it the new name Heterarthrus healyi . H. healyi was subsequently found to be synonymous with H. wuestneii (Konow, 1905) (Blank et al. 2001).

Cladius (Priophorus ) compressicornis . The change in nomenclature is explained in Liston (2007).

Nematinus antennatus . Placed as a synonym of N. luteus auct. (= N. steini ) by Benson (1958). At present, a syntype of N. antennatus matching the original description is not definitely identifiable, but study of this specimen and revision of the genus have not yet been completed (S. M. Blank, pers. comm.).

Amauronematus godmani. Specimens identified by Benson as this species, from Switzerland (type locality) and Scotland in RSM collection do not seem to be conspecific. They both belong to the Amauronematus variator (Ruthe) species group. Identity of Scottish specimens requires further study.

Amauronematus hedstroemi . See Lindqvist (1972) on taxonomy of this species and A. tillbergi Malaise, 1920.

Amauronematus histrio . British and Irish specimens at present identified as this species or as A. stenogaster may include other taxa not yet recorded in the British Isles. See revision of North European taxa by Schmidt (1997).

Eupontania . Subsequent to the revision of this genus by Kopelke (1991), numerous important taxonomic and nomenclatural changes, followed here, were proposed by Vikberg (2003) and Vikberg & Zinovjev (2006).

Euura atra . Regarded as a group of sibling (biological) species by Kopelke (1996, 2000). These are very difficult, perhaps impossible, to distinguish using characters of adult morphology. According to records of galls on various identified Salix species, at least the following 3 additional species belonging to the group occur in the British Isles: Euura auritae Kopelke, 2000, Euura purpureae Kopelke, 1996, Euura weiffenbachii Ermolenko, 1988.

Euura mucronata. According to Kopelke (2001), several biological species are included under this name. A number of these seem to occur in the British Isles. It should however be noted that an alternative view on some of the taxa proposed by Kopelke has been offered by Nyman (2002).

Nematus leionotus . O’Connor et al. (1997) were unable to trace any Irish specimens.

Nematus maculiger . Regarded by some authors as distinct from N. melanaspis (see for example Lindqvist 1962), but this requires investigation.

Nematus viridissimus . According to Lindqvist (1962), N. glutinosae (description probably prodates that of N. viridissimus by a few months) is not conspecific with N. viridissimus .

Pachynematus clitellatus . Taeger & Blank (1998) regard P. clitellatus and P. fallax as morphologically highly variable species, which together include most of the previously described taxa described in the West Palaearctic. Their arguments for this are well reasoned, but at present it seems prudent to continue to regard the morphological segregates (species according to Benson 1958) as separate taxa, because morphologically intermediate specimens are comparatively rare (at least in the male sex). The taxonomy of the whole group requires further intensive work, including genetic analysis. According to Taeger & Blank (1998) P. kirbyi and P. truncatus , treated here as valid, are synonyms of P. clitellatus :

Pachynematus fallax . See above, under P. clitellatus . According to Taeger & Blank (1998), the following taxa are synonyms of P. xanthocarpus (= P. fallax ): P. sulcatus , P. calcicola , P. chambersi . 41

Phyllocolpa anglica. According to V. Vikberg & A. Zinovjev (pers. comm.), three morphologically well distinguished species of Phyllocolpa feed on Salix viminalis in Europe. Benson (1958) had already distinguished these under the names P. piliserra , P. scotaspis and P. anglica . Kopelke (2007c) thought that only two European species occuir on this host, and synonymised both Nematus anglicus and N. nigrolineatus with P. scotaspis . This appears to leave the taxon formerly frequently referred to as P. anglica without a valid name, but until the taxonomy of these species is reexamined, the name is provisionally retained in use.

Phyllocolpa carinifrons. Taxonomy follows Kopelke (2007a).

Phyllocolpa coriacea . Kopelke (2007b) placed Nematus coriaceus Benson, 1953 as a synonym of P. alienata (Förster, 1854) and used the latter as the valid name. However, the conditions of ICZN Article 23.9.1. are fulfilled in this case: N. alienatus (nomen oblitum) has not been used as a valid name since 1900 and N. coriaceus (or Pontania coriacea or Phyllocolpa coriacea )(nomen protectum) has been used in at least 47 works by more than 10 authors during the last 50 years.

Phyllocolpa ischnocera . Taxonomy follows Kopelke (2007a).

Phyllocolpa leucosticta . According to Kopelke (2007a), P. erythropyga (Förster, 1854) and P. prussica (Zaddach, 1883) have previously been confused with P. leucosticta . It is not at present clear which of these species occur in the British Isles.

Phyllocolpa polita. Regarded as a valid species by Kopelke (2003, 2007a). The type localities include Scotland (Brischke, 1884).

Pontania tuberculata. Study of the types of this and several similar taxa by V. Vikberg and AL revealed that the taxonomy and nomenclature proposed by Kopelke (2007c) is faulty. Nematus anomalopterus Förster, 1854 is in fact conspecific with N. tuberculatus Benson (syn. nov.). P. joergenseni (Enslin, 1916) (unavailable name: junior primary homonym of P. joergenseni Strand, 1908) and P. cyrnea Liston, 2005, both wrongly placed in synonymy with P. anomaloptera by Kopelke, belong to a different species ( P. joergenseni Enslin in Zinovjev & Vikberg 1999) which is not known in the British Isles. A full clarification of these problems is being prepared by V. Vikberg.

Pontania lapponicola . Added on the basis of records of galls of P. dolichura from Salix lapponum in Benson (1954).

Pontania nigricantis . Added on the basis of records of galls of P. dolichura from Salix nigricans in Benson (1954).

Pontopristia amentorum . Lindqvist (1961) regards a large series collected by Benson at Inchnadamph, Sutherland, as representing A. kamtchaticus (Malaise) and not A. amentorum as stated by Benson (1959).

Pristiphora sermola . Lygaeophora variipes Lindqvist, 1952 is preoccupied in Pristiphora by P. varipes Serville, 1823. The names varipes and variipes are to be regarded as identical spellings: Article 58.14. (ICZN 1999). Pristiphora varipes Serville, 1823 is a species inquirenda, not a synonym of Cladius ( Priophorus ) pallipes Serville, 1823 or C. brullei (Dahlbom, 1835) (see Lacourt 2000). The conditions of Article 23.9., permitting continued use of the junior homonym as valid, are not fulfilled for variipes (Lindqvist ) because this has only been used as a valid name on twenty occasions in works published within the last 50 years (23.9.1.2.).

Nematus astutus . Taxonomy follows Liston et al. (2006).

Pristiphora groenblomi . Predominant usage is of this spelling, not gronblomi , which would be correct according to ICZN Article 32.5.2. Following ICZN (Article 33.3.1) the use of the predominant spelling is maintained.

Pristiphora aphantoneura . See Vikberg (2006) on taxonomy.

Pristiphora cincta . Probably two British species occur under this name: one more lowland with hostplant Betula , the other montane with host Vaccinium . Whilst P. cincta seems to be the correct name for the more brightly coloured lowland taxon, until a revision is made of the numerous supposed synonyms of P. cincta , the name for the species on Vaccinium is unclear. 42

Pristiphora denudata . Contrary to the opinion of Lacourt (1987), we do not consider this taxon to be synonymous with P. pallidiventris (Fallén, 1808) ( P. pallidiventris pallidiventris sensu Lacourt), nor with P. atlantica Malaise, 1939 (= P. atlantica Lacourt, 1987) as previously treated by Liston (1995), who replaced the name denudata with atlantica Lacourt on the grounds that the name denudata is preoccupied in Pristiphora by Nematus denudatus Hartig, 1840. Nematus denudatus was at that time considered to be a synonym of Pristiphora carinata (Hartig, 1837). According to Schmidt (in Lacourt, 1999) N. denudatus Hartig actually belongs to Amauronematus ( Brachycoluma ), thus removing the homonymy. P. denudata differs from P. pallidiventris not only in the more extensively pale abdomen (without intermediate specimens), but also in details of the structure of the saw, and is probably attached only to Rubus idaeus agg. whereas P. pallidiventris is polyphagous on various Rosaceae.

Pristiphora insularis . Synonymy follows Haris (2006).

Pristiphora luteipes . See Vikberg (2006) on taxonomy.

Pristiphora hyperborea . Recorded as British by Benson (1935), but later treated by him as a synonym of P. staudingeri . V. Vikberg considers P. hyperborea to represent a valid species (see Zhelochovtsev 1988). Probably both species occur in the British Isles.

Pristiphora subbifida . Characterisation of P. subbifida by Zhelochovtsev (1988) based on misidentification by Lindqvist (1973) of P. conjugata , or a similar species.

Pristiphora subopaca . Recorded by Lindqvist (1955) from Great Britain, but in the reprints of this paper which he distributed, “Grossbritannien” is scored out. Therefore this record is considered to be based on a misidentification. However, J. Grearson has recently taken females in England, identity confirmed by V. Vikberg after comparison with the holotype (Grearson, in preparation).

Stauronematus platycerus . Taxonomy follows Liston (2007).

Nematus placidus . Placed by Konow (1905) as a synonym of Pristiphora leucopodia (Hartig, 1837), a species which has never been found in the British Isles. Probably Konow did not examine the type, which according to Benson (1943b), is lost.

Dolerus scoticus . Haris (2000) regards this as a separate species from D. yukonensis . For the reasons stated by Goulet (1986), it seems more likely that they are conspecific.

Dolerus asper. Previous records of D. asper from Wales and Ireland require checking, because they may refer to D. brevicornis (below), recently distinguished by Heidemaa et al. (2004). English specimens of D. asper, leg. K. J. Grearson have been determined by M. Heidemaa. The only evidence for the presence of D. asper in Scotland is the determination of a syntype specimen of Dolerus oblongus in the BM(NH) London examined by Heidemaa et al. (2004) as D. asper . Cameron (1882) described D. oblongus from “Braemar, Rannoch, Clydesdale”.

Dolerus brevicornis . Only recently distinguished from D. asper (see above). A specimen of D. brevicornis collected by K. J. Grearson in England has been determined as D. brevicornis by M. Heidemaa.

Dolerus liogaster . Heidemaa & Saarma (2005) established that D. schmidti (below) has hitherto been confused with D. liogaster . D. liogaster is confirmed as present in Britain by M. Heidemaa (pers. comm.) based on material collected by A. Barker.

Dolerus schmidti . See under D. liogaster (above). D. schmidti is present in Britain according to determinations made by M. Heidemaa: England, leg. K. J. Grearson and Scotland, leg. A. Barker.

Dolerus varispinus . Taxonomy follows Heidemaa & Saarma (2005).

Heptamelus dahlbomi . Species revocata (Liston & Vikberg, in prep.).

Aneugmenus coronatus . Not a synonym of A. padi . See Chevin (1980). There is a major divergence in opinion here regarding association of the sexes of these species (compare, for example, Blank (1998), Chevin (1980) and Benson (1952, 1968).

Aneugmenus fuerstenbergensis . O’Connor et al. (1997: 21) were unable to trace the Irish specimens. 43

Strongylogaster filicis . Recorded only on the basis of a single specimen by Bold (1873). The present whereabouts of the specimen are unknown.

Tenthredo (Elinora) dominiquei. Placed for a time by Benson as a subspecies of the more continental T. flaveola Gmelin, 1790. This decision is not supported by current data on distributon and morphology. Lacourt (1999) treats T. dominiquei as a northern subspecies of T. baetica Spinola, 1843 (confined to Spain and Portugal), but until these taxa are studied in more detail, it seems best to regard T. dominiquei as a distinct species.

References

Altenhofer, E. & Taeger, A. 1998 Zur Kenntnis der Gattung Scolioneura Konow, 1890 (Hymenoptera: Tenthredinidae). In Pflanzenwespen Deutschlands (Hymenoptera, Symphyta). Kommentierte Bestandsaufnahme (eds A. Taeger & S. M. Blank), Goecke & Evers, Keltern: pp. 225226.

Altenhofer, E. & Zombori, L. 1987 The species of Heterarthrus Stephens, 1835 feeding on maple (Hymenoptera, Tenthredinidae). Annales historico-naturales Musei Nationalis Hungarici 79 , 185197.

Beneš, K. & Krístek, J. 1976 A new species of the genus Pristiphora (Hymenoptera, Tenthredinidae) feeding on Norway spruce. Acta entomologica bohemoslovaca 73 , 404415.

Benson, R. B. 1934: Additions to the list of British Dolerinae (Hymenoptera Symphyta). Entomologist's Monthly Magazine 70 , 1113.

Benson, R. B. 1935: The high mountain sawflies of Britain (Hymenoptera Symphyta). Transactions of the Royal Entomological Society of London 83 , 2339.

Benson, R. B. 1943a Some reputed British sawflies not found since Stephens's days (Hym., Symphyta). Entomologist's Monthly Magazine 79 , 57.

Benson, R. B. 1943b More about the sawflies of Hertfordshire and Buckinghamshire and the list of British species (Hym., Symphyta). Entomologist's Monthly Magazine 79 , 712.

Benson, R. B. 1948. Some further British species of Amauronematus Konow (Hym., Tenthredinidae). Entomologist's Monthly Magazine 84 , 2832.

Benson, R. B. 1951 Hymenoptera, Symphyta. Handbooks for the Identification of British 6(2a) , 149.

Benson, R. B. 1952 Hymenoptera, Symphyta. Handbooks for the Identification of British Insects 6(2b) , 51137.

Benson, R. B. 1953 Some changes and additions to the list of British sawflies with the descriptions of two new species (Hym., Tenthredinidae). Entomologist's Monthly Magazine 89 , 150154.

Benson, R. B. 1954 British sawfly galls of the genus Nematus [ Pontania ] on Salix . (Hymenoptera, Tenthredinidae). Journal of the Society for British Entomology 4, 206211.

Benson, R. B. 1958 Hymenoptera, Symphyta. Handbooks for the Identification of British Insects 6(2c) , 139 258.

Benson, R. B. 1959 Sawflies (Hym., Symphyta) of Sutherland and Wester Ross. Entomologist's Monthly Magazine 95 , 101104

Benson, R. B. 1968 Hymenoptera from Turkey, Symphyta. Bulletin of the British Museum (Natural History), Entomology 22 , 111207.

Blank, S. M. 1998 Die mittel und nordeuropäischen Selandriinae (Hymenoptera: Tenthredinidae). In Pflanzenwespen Deutschlands (Hymenoptera, Symphyta). Kommentierte Bestandsaufnahme (eds A. Taeger & S. M. Blank), Goecke & Evers, Keltern: pp. 207224. 44

Blank, S. M., Deters, S., Drees, M., Jänicke, M., Jansen, E., Kraus, M., Liston, A. D., Ritzau, C. & Taeger, A. 2001 Symphyta. In Verzeichnis der Hautflügler Deutschlands (Entomofauna Germanica 4) (eds H. H. Dathe, A. Taeger & S. M. Blank), Entomologische Nachrichten und Berichte . Beiheft 7 : pp. 827.

Blank, S. M. & Taeger, A. 1998 Comments on the taxonomy of Symphyta (Hymenoptera) (Preliminary studies for a catalogue of Symphyta, part 4). In Pflanzenwespen Deutschlands (Hymenoptera, Symphyta). Kommentierte Bestandsaufnahme (eds A. Taeger & S. M. Blank), Goecke & Evers, Keltern: pp. 141174.

Bold, T. J. 1873 Capture in Northumberland of a Sawfly new to Britain. Entomologist's Monthly Magazine 10 , 69.

Brischke, C. G. A. 1884 Beobachtungen über die Arten der Blatt und Holzwespen von C. G. A. Brischke, Hauptlehrer a. D. in Langfuhr und Dr. Gustav Zaddach, Professor in Königsberg, mitgetheilt von Brischke aus Zaddachs Manuskripten [Schluss, Nematus Teil 3]. Schriften der physikalisch-ökonomischen Gesellschaft zu Königsberg 24 [1883], 121173, plate I (8).

Cameron, P. 1882 A Monograph of the British Phytophagous Hymenoptera. (Tenthredo, Sirex and Cynips, Linné.) London 1: 1340; plates 121.

Chevin, H. 1980 Notes sur les Hymenopteres Tenthredoides. Bulletin mensuel de la Société Linnéenne de Lyon 49 , 453456.

Dalla Torre, C. G. de 1894 Tenthredinidae incl. Uroceridae (Phyllophaga & Xylophaga). - Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus, Lipsiae 1: 1459.

Edmunds, R., Hind, S., Liston, A. & Palmer, K. 2007 Parna apicalis (Brischke, 1888) (Symphyta:Tenthredinidae), a leafmining sawfly new to Great Britain. Entomologist’s Record and Journal of Variation 119 , 223226.

Enslin, E. 19121918 Die Tenthredinoidea Mitteleuropas. Deutsche entomologische Zeitschrift Beihefte 1- 7[19121917], 1790.

Fabricius, J. C. 1781 Species Insectorum exhibentes eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosin adiectis observationibus, descriptionibus . Hamburgi et Kilonii 1: IVIII, 1552.

Fitton, M. G., Graham, M. W. R. de V., Bouček, Z. R. J., Fergusson, N. D. M., Huddleston, T., Quinlan, J. & Richards, O. W. 1978 Kloet and Hincks. A check list of British insects. Part 4: Hymenoptera. Handbooks for the Identification of British Insects 11(4) , ix + 159 pp.

Gibbs, D. 2006 Rose Bud Sawfly, plana (Klug) (Hymenoptera: Tenthridinidae), new to Britain, discovered in South Wales. British Journal of Entomology and Natural History 19 , 105108.

Goulet, H. 1986 The genera and species of the Nearctic Dolerini (Symphyta: Tenthredinidae: Selandriinae): classification and phylogeny. Memoirs of the Entomological Society of Canada 135 , 1208.

Grearson, K. J. 2006 A sawfly, Pristiphora leucopus (Hellén) (Hymenoptera: Tenthredinidae), new to Britain. British Journal of Entomology and Natural History 19 , 15, 1 plate.

Halstead, A. J. 2004 Sawfly, Schrank (Hymenoptera: Argidae), a pest new to Britain. British Journal of Entomology and Natural History 17 , 131135.

Haris, A. 2000 Study on the Palaearctic Dolerus Panzer, 1801 species (Hymenoptera: Tenthredinidae). Folia Entomologica Hungarica 61 , 95148.

Haris, A. 2006 Study on the Palaearctic Pristiphora species (Hymenoptera: Tenthredinidae). Natura Somogyiensis 9, 201277.

Heidemaa, M., Nuorteva, M., Hantula, J. & Saarma, U. 2004 Dolerus asper Zaddach, 1859 and Dolerus brevicornis Zaddach, 1859 (Hymenoptera: Tenthredinidae), with notes on their phylogeny. European Journal of Entomology 101 , 637650. 45

Heidemaa, M. & Saarma, U. 2005 Phylogenetic relationships in Dolerus gibbosus species group and Dolerus varispinus complex (Hymenoptera: Tenthredinidae) as inferred from molecular and morphological data, with revisory notes on D. varispinus complex (unpublished PhD thesis).

Heidemaa, M. & Viitasaari, M. 1999 Taxonomy of Empria hungarica speciesgroup (Hymenoptera, Tenthredinidae) in Northern Europe. Entomologica Fennica 10 , 95101.

Hellén, W. 1975 Die Nematinen Finnlands IV (Hymenoptera, Tenthredinidae). Gattung Pristiphora Latreille. Notulae Entomologicae 55 , 97128.

ICZN 2005, Opinion 2101 (Case 3243). Lyda latifrons Fallén, 1808 and L. gyllenhali Dahlbom,1835 (currently Pamphilius latifrons and P. gyllenhali ; Insecta, Hymenoptera): usage of the specific names conserved by designation of a neotype for Lyda latifrons . Bulletin of Zoological Nomenclature 62 , 4950.

Knight, G. T. 2004 Kaliofenusa pusilla (Serville 1823) (Hymenoptera: Tenthredinidae). A species of leafmining sawfly new to Ireland. Bulletin of the Irish Biogeographical Society 28 , 6162.

Knight, G. T. 2006 First British records of the sawfly Emphytus laticinctus (Serville) (Hymenoptera: Tenthredinidae). British Journal of Entomology and Natural History 19 , 193198.

Koch, F. 1988 Die Gattung Claremontia Rohwer (Hymenoptera, Tenthredinidae). Deutsche entomologische Zeitschrift 35 , 311330.

Konow, F. W. 1899 Neuer Beitrag zur Synonymie der Chalastogastra. Entomologische Nachrichten (F. Karsch), Berlin 25 , 8696.

Konow, F. W. 1905 Hymenoptera. Fam. Tenthredinidae. Genera Insectorum 29 , 1 176.

Kopelke, J.P. 1991 Die Arten der viminalis Gruppe, Gattung Pontania O. Costa 1859, Mittel und Nordeuropas (Insecta: Hymenoptera: Tenthredinidae). Senckenbergiana biologica 71 , 65128.

Kopelke, J.P. 1996 Die Euura atra und amerinae Gruppe in Nord und Mitteleuropa (Insecta: Hymenoptera: Tenthredinidae: Nematinae). Senckenbergiana biologica 76 , 93113.

Kopelke, J.P. 2000 Euura auritae sp. n. ein neuer Gallenerzeuger der atra Gruppe in Europa (Insecta, Hymenoptera, Tenthredinidae, Nematinae). Senckenbergiana biologica 80 , 159163.

Kopelke, J.P. 2001 Die Artengruppen von Euura mucronata und E. laeta in Europa (Insecta, Hymenoptera, Tenthredinidae, Nematinae). Senckenbergiana biologica 81 , 191225.

Kopelke, J.P. 2003 Gallforming Nematinae, their hosts ( Salix spec.) and biological strategies (Insecta, Hymenoptera, Symphyta, Tenthredinidae, Nematinae: Euura, Phyllocolpa, Pontania ). Senckenbergiana biologica 82 , 163189.

Kopelke, J.P. 2007a The European species of the genus Phyllocolpa , part I: the leucosticta group (Insecta, Hymenoptera, Tenthredinidae, Nematinae). Senckenbergiana biologica 87 , 75109.

Kopelke, J.P. 2007b The European species of the genus Phyllocolpa , part II: the leucapsis group (Insecta, Hymenoptera, Tenthredinidae, Nematinae). Senckenbergiana biologica 87 , 149161.

Kopelke, J.P. 2007c The European species of the genus Phyllocolpa , part III: the species groups of crassispina, scotaspis, and piliserra (Insecta, Hymenoptera, Tenthredinidae, Nematinae). Senckenbergiana biologica 87 , 163183.

Lacourt, J. 1987 Note sur Pristiphora pallidiventris (Fallén) (= Pristiphora denudata Konow) avec description de deux nouvelles sousespèces (Hymenoptera, Tenthredinidae). Nouvelle Revue d’Entomologie 4, 259264.

Lacourt, J. 1999 Répertoire des Tenthredinidae ouestpaléarctiques (Hymenoptera, Symphyta). Mémoires de la Société Entomologique de France 3, 1432. 46

Lacourt, J. 2000 Liste des espèces de la famille des Tenthredinidae décrites par J. G. AudinetServille, en Mai 1823 et par A. L. M. Le Peletier Comte de SaintFargeau, en Août 1823, avec désignation de lectotypes [Hymenoptera, Symphyta]. Revue francaise d’Entomologie (N. S.), 22, 77108.

Leach, W. E. 1817 The Zoological Miscellany. Being Descriptions of New or Interesting . R. and A. Taylor, London 3, 1151.

Lindqvist, E. 1952 Über alte und neue Lygaeonematus Arten (Hym., Tenthredinidae). Notulae Entomologicae 32 , 80119.

Lindqvist, E. 1955 Über einige Pristiphora Arten aus Fennoskandien (Hym., Tenthredinidae). Notulae Entomologicae 35 , 3550.

Lindqvist, E. 1961 Die Pontopristia Arten Finnlands (Hym., Tenthr.). Notulae Entomologicae 41 , 6979.

Lindqvist, E. 1962 Bemerkungen über paläarktische Blattwespen (Hym. Symph.). Notulae Entomologicae 42 , 105127.

Lindqvist, E. 1972 Zur Nomenklatur und Taxonomie einiger Blattwespen (Hymenoptera, Symphyta). Notulae Entomologicae 52 , 6577.

Lindqvist, E. 1973 Taxonomische Bemerkungen über einige Blattwespen I (Hymenoptera, Symphyta). Notulae Entomologicae 53 , 3339.

Liston, A. D. 1981a Pristiphora (Lygaeophora) lanifica (Zaddach, 1882) new to Britain (Hym., Tenthredinidae). Entomologist's Gazette 32 , 181184.

Liston, A. D. 1981b Notes on little known British Pristiphora Lat. (Hym., Tenthredinidae) of the abietina group. Entomologist's Monthly Magazine 117 , 7375.

Liston, A. D. 1982 Some sawflies from Whitlaw Moss Nature Reserve, Southern Scotland, with a species new to Britain (Hym.: Symphyta). Entomologist’s Record and Journal of Variation 94 , 175179.

Liston, A. D. 1983 Distribution and Ecology of the Sawflies Macrophya alboannulata Costa and M. albicincta (Schrank) in the West Palaearctic. Faunistische Abhandlungen 10 , 151153.

Liston, A. D. 1983 Pristiphora karvoneni (Lindqvist) (Hym., Tenthredinidae) new to Britain. Entomologist's Monthly Magazine 119 , 135136.

Liston, A. D. 1985 On some sawflies (Hymenoptera, Symphyta) from Upper Deeside. Entomologist’s Record and Journal of Variation 97 , 9495.

Liston, A. D. 1993 Caliroa tremulae Chevin in the British Isles and Central Europe (Hymenoptera: Tenthredinidae). Entomologist 112 , 6768.

Liston, A. D. 1993 Nomenclature and taxonomy of a Nematine sawfly occurring in Britain (Hymenoptera: Tenthredinidae). British Journal of Entomology and Natural History 6, 103106.

Liston, A. D. 1994 Kaliofenusa carpinifoliae Liston (Hymenoptera: Tenthredinidae), a newly recognized leaf miner on Field Elms in Britain. British Journal of Entomology and Natural History 7, 1518.

Liston, A. D. 1995 Compendium of European Sawflies. List of species, modern nomenclature, distribution, foodplants, identification literature . Gottfrieding, Chalastos Forestry 1190.

Liston, A. D. 1996 Compendium of European Sawflies. Supplement. Gottfrieding, Chalastos Press: 116.

Liston, A. D. 2007 Revision of Stauronematus Benson, 1953 and additions to the sawfly fauna of Corsica and Sardinia (Hymenoptera, Tenthredinidae). Beiträge zur Entomologie 57 , 135150.

Liston, A. D. & Blank, S. M. 2006 New and littleknown British Xyelidae and Tenthredinidae (Hymenoptera, 47

Symphyta). Entomologist's Monthly Magazine 142 , 219227.

Liston, A. D. & O'Connor, J. P. 2005 Records of Irish sawflies (Hymenoptera: Symphyta) including eight species new to Ireland. Irish Naturalist’s Journal 28 , 159161.

Liston, A. D., Taeger, A. & Blank, S. M. 2006 Comments on European Sawflies (Hymenoptera: Symphyta). In Recent Sawfly Research: Synthesis and Prospects (eds S. M. Blank; S. Schmidt & A. Taeger), Goecke & Evers, Keltern: pp. 245263.

Lorenz, H. & Kraus, M. 1957 Die Larvalsystematik der Blattwespen (Tenthredinoidea und Megalodontoidea). Abhandlungen zur Larvalsystematik der Insekten 1, 1389.

McLachlan, R. 1867: Additional notices respecting the maplemining sawfly ( Phyllotoma aceris ). Entomologist's Monthly Magazine 4: 123.

Nyman, T. 2002 The willow bud galler Euura mucronata Hartig (Hymenoptera: Tenthredinidae): one polyphage or many monophages? Heredity 88 , 288295.

Nyman, T.; Zinovjev, A. G.; Vikberg, V. & Farrel, B. D. (2006): Molecular phylogeny of the sawfly subfamily Nematinae (Hymenoptera: Tenthredinidae). Systematic Entomology 31 , 569583.

O'Connor, J. P. & Liston, A. D. 1994 Pristiphora friesei (Konow) (Hymenoptera: Symphyta), a sawfly new to the British Isles. Entomologist's Gazette 45 : 6972.

O'Connor, J. P., Liston, A. D. & Speight, M. C. D. 1997 A Review of the Irish sawflies (Hymenoptera: Symphyta) including a checklist of species. Bulletin of the Irish Biogeographical Society 20 , 299.

Pesarini, F. 2002 Contributo alla conoscenza dei sinfiti della regione balcanicoegea (Hymenoptera, Symphyta). Bollettino del Museo regionale di Scienze Naturale Torino 19 , 121183.

Schmidt, S. 1997 Amauronematus Konow, 1890 Ökologie und Taxonomie der nordeuropäischen Arten des fallaxKomplexes (Hymenoptera, Tenthredinidae). Beiträge zur Entomologie 47 , 227326.

Shaw, M. R. 2003 Adverse comment on the supposed British status of the webspinning sawfly Cephalcia arvensis Panzer (Hymenoptera: Pamphiliidae). British Journal of Entomology and Natural History 16 , 2.

Shaw, M. R. & Liston, A. D. 1985 Xiphydria longicollis (Geoffroy) (Hymenoptera: Xiphydriidae) new to Britain. Entomologist's Gazette 36 , 233235.

Sheppard, D. A. 1990. A review of sawflies (Hymenoptera: Symphyta) from Jersey. Societe Jersiaise Annual Bulletin 25, 367372.

Shinohara, A. 1998 Pamphilius albopictus (Hymenoptera, Pamphiliidae) and its Close Relatives. Bulletin of the National Science Museum, Series A, Zoology 24 , 225252.

Shinohara, A. 2002 A spruce webspinning sawfly, Cephalcia arvensis (Hymenoptera: Pamphiliidae) from Britain. British Journal of Entomology and Natural History 15 , 134135.

Smith, D. R. 1978 Suborder Symphyta. (Xyelidae, Pararchexyelidae, Parapamphiliidae, Xyelydidae, Karatavitidae, Gigasiricidae, Sepulcidae, Pseudosiricidae, Anaxyelidae, Siricidae, Xiphydriidae, Paroryssidae, Xyelotomidae, Blasticotomidae, Pergidae). Hymenopterorum Catalogus (nova editio) 14, 1193.

Stephens, J. F. 1835 Illustrations of British Entomology; or, a Synopsis of Indigenous Insects: containing their generic and specific distinctions;with an account of their metamorphosis, times of appearance, localities, food, and economy, as far as practicable. Mandibulata . Baldwin & Cradock, London 7, 1312, plates XXXVXLVII.

Taeger, A. 1998 Bestimmungsschlüssel der Keulhornblattwespen Deutschlands (Hymenoptera: Cimbicidae). In Taeger, A. & Blank, S. M. 1998 (Hrsg.): Pflanzenwespen Deutschlands (Hymenoptera, Symphyta). Kommentierte Bestandsaufnahme (eds A. Taeger & S. M. Blank), Goecke & Evers, Keltern: pp. 193205.

Taeger, A. 2002 The Megalodontesidae of Europe (Hymenoptera, Symphyta). In Sawflies (Hymenoptera, 48

Symphyta) I. A review of the suborder, the Western Palaearctic taxa of Xyeloidea and Pamphilioidea (ed. M. Viitasaari), Tremex , Helsinki: pp. 461480.

Taeger, A. & Blank, S. M. 1996 Kommentare zur Taxonomie der Symphyta (Hymenoptera) (Vorarbeiten zu einem Katalog der Pflanzenwespen, Teil 1). Beiträge zur Entomologie 46 , 251275.

Taeger, A. & Blank, S. M. 1998 Beitrag zur Kenntnis einiger Nematinae (Hymenoptera: Tenthredinidae). In: Taeger, A. & Blank, S. M. 1998 (eds): Pflanzenwespen Deutschlands (Hymenoptera, Symphyta). Kommentierte Bestandsaufnahme . Goecke & Evers, Keltern : 247277; Taf. 3, 7

Taeger, A. & Blank, S. M. 2006: Bibliographic Notes. Pp. 319330 In: Blank, S. M.; Schmidt, S. & Taeger, A. (eds): Recent Sawfly Research: Synthesis and Prospects. 704 pp., 16 pl. Goecke & Evers, Keltern.

Taeger, A. & Blank, S. M. 2008 : ECatSym Electronic World Catalog of Symphyta (Insecta, Hymenoptera). Program version 3.9, data version 34 (05.09.2008) . Digital Entomological Information, Müncheberg, http://www.zalf.de/home_zalf/institute/dei/php/ecatsym/ecatsym.php.

Viitasaari, M. 1985 Taxonomic notes on the genus Trichiosoma Leach (Hymenoptera, Cimbicidae) I. Annales Entomologici Fennici 51 , 2526.

Viitasaari, M. 1989 Taxonomic notes on the genus Trichiosoma Leach (Hymenoptera, Cimbicidae) II. Annales Entomologici Fennici 55 , 111129.

Viitasaari, M. & Midtgaard, F. 1989 A contribution to the taxonomy of horntails with notes on the genus Sirex Linnaeus (Hymenoptera, Siricidae). Annales Entomologici Fennici 55, 103110

Viitasaari, M. & Vikberg, V. 1985 A checklist of the sawflies (Hymenoptera, Symphyta) of Finland. Notulae Entomologicae 65 , 117.

Vikberg, V. 2003 Taxonomy of the species of Eupontania crassipes and aquilonis groups (Hymenoptera: Tenthredinidae: Nematinae). Entomologica Fennica 14 , 129172.

Vikberg, V. 2004 Seasonal head dimorphism and taxonomy of some European species of Aprosthema (Hymenoptera: Symphyta: Argidae). Beiträge zur Entomologie 54 , 107125.

Vikberg, V. 2006 Pristiphora Latreille, 1810 (Hymenoptera: Tenthredinidae: Nematinae) Two Closely Related Species Feeding on Lathyrus and Salix and Notes on Species Feeding on Leguminous . In Recent Sawfly Research: Synthesis and Prospects (eds S. M. Blank; S. Schmidt & A. Taeger), Goecke & Evers, Keltern: pp. 105128.

Vikberg, V. & Zinovjev, A. 2006 On the taxonomy and the host plants of North European species of Eupontania (Hymenoptera: Tenthredinidae: Nematinae). Beiträge zur Entomologie 56 , 239268.

Zhelochovtsev, A. N. 1988 Pereponchatokrylye. Shestaya chast. In: Medvedev, G. C. (ed.): Opredelitel' nasekomykh evropeyskoy chasti SSSR . Nauka, Leningrad 3(6) , 3237.

Zinovjev, A. G. & Vikberg, V. 1999 The sawflies of the Pontania crassispina group with a key for the genera of the subtribe Euurina (Hymenoptera: Tenthredinidae, Nematinae). Entomologica scandinavica 30 , 281298.