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“Subfossil” Koala Lemur Megaladapis Edwardsi
Evolutionary and phylogenetic insights from a nuclear genome sequence of the extinct, giant, “subfossil” koala lemur Megaladapis edwardsi Stephanie Marciniaka, Mehreen R. Mughalb, Laurie R. Godfreyc, Richard J. Bankoffa, Heritiana Randrianatoandroa,d, Brooke E. Crowleye,f, Christina M. Bergeya,g,h, Kathleen M. Muldooni, Jeannot Randrianasyd, Brigitte M. Raharivololonad, Stephan C. Schusterj, Ripan S. Malhik,l, Anne D. Yoderm,n, Edward E. Louis Jro,1, Logan Kistlerp,1, and George H. Perrya,b,g,q,1 aDepartment of Anthropology, Pennsylvania State University, University Park, PA 16802; bBioinformatics and Genomics Intercollege Graduate Program, Pennsylvania State University, University Park, PA 16082; cDepartment of Anthropology, University of Massachusetts, Amherst, MA 01003; dMention Anthropobiologie et Développement Durable, Faculté des Sciences, Université d’Antananarivo, Antananarivo 101, Madagascar; eDepartment of Geology, University of Cincinnati, Cincinnati, OH 45220; fDepartment of Anthropology, University of Cincinnati, Cincinnati, OH 45220; gDepartment of Biology, Pennsylvania State University, University Park, PA 16802; hDepartment of Genetics, Rutgers University, New Brunswick, NJ 08854; iDepartment of Anatomy, Midwestern University, Glendale, AZ 85308; jSingapore Centre for Environmental Life Sciences Engineering, Nanyang Technological University, Singapore 639798; kDepartment of Anthropology, University of Illinois Urbana–Champaign, Urbana, IL 61801; lDepartment of Ecology, Evolution and Behavior, Carl R. Woese Institute for -
Diagnosis and Differentiation of the Order Primates
YEARBOOK OF PHYSICAL ANTHROPOLOGY 30:75-105 (1987) Diagnosis and Differentiation of the Order Primates FREDERICK S. SZALAY, ALFRED L. ROSENBERGER, AND MARIAN DAGOSTO Department of Anthropolog* Hunter College, City University of New York, New York, New York 10021 (F.S.S.); University of Illinois, Urbanq Illinois 61801 (A.L. R.1; School of Medicine, Johns Hopkins University/ Baltimore, h4D 21218 (M.B.) KEY WORDS Semiorders Paromomyiformes and Euprimates, Suborders Strepsirhini and Haplorhini, Semisuborder Anthropoidea, Cranioskeletal morphology, Adapidae, Omomyidae, Grades vs. monophyletic (paraphyletic or holophyletic) taxa ABSTRACT We contrast our approach to a phylogenetic diagnosis of the order Primates, and its various supraspecific taxa, with definitional proce- dures. The order, which we divide into the semiorders Paromomyiformes and Euprimates, is clearly diagnosable on the basis of well-corroborated informa- tion from the fossil record. Lists of derived features which we hypothesize to have been fixed in the first representative species of the Primates, Eupri- mates, Strepsirhini, Haplorhini, and Anthropoidea, are presented. Our clas- sification of the order includes both holophyletic and paraphyletic groups, depending on the nature of the available evidence. We discuss in detail the problematic evidence of the basicranium in Paleo- gene primates and present new evidence for the resolution of previously controversial interpretations. We renew and expand our emphasis on postcra- nial analysis of fossil and living primates to show the importance of under- standing their evolutionary morphology and subsequent to this their use for understanding taxon phylogeny. We reject the much advocated %ladograms first, phylogeny next, and scenario third” approach which maintains that biologically founded character analysis, i.e., functional-adaptive analysis and paleontology, is irrelevant to genealogy hypotheses. -
LCFPD Track Identification and Walking Gaits
Track Identification Cat Rabbit ➢ 4 toes on each foot ➢ 4 toes on each foot ➢ round tracks ➢ oval tracks ➢ no claw marks ➢ front of skid has claw marks ➢ hind paws narrower ➢ almost impossible to see pads Dog, Fox, Coyote Raccoon ( Dog family) ➢ 5 toes on each foot ➢ 4 toes on each foot ➢ toes close together ➢ tracks maple leaf shaped or egg shaped ➢ toes long and finger shaped ➢ claw marks appear ➢ claw marks appear ➢ front feet larger Weasel, Skunk, Mink Opossum (Weasel Family) ➢ 5 toes on each foot ➢ 5 toes on each foot ➢ toes spread widely ➢ toes close together ➢ toes long and finger shaped ➢ toes short and round ➢ no claw mark on hind thumbs ➢ some partially webbed Mice, Squirrels, etc. (Rodent Family) Deer ➢ 4 toes on front feet, 5 toes on hind ➢ 2 toes on each foot feet ➢ tracks are paired when bounding ➢ front feet show dew claws ➢ hind feet step in fore feet ➢ size of track is important in separating species tracks ➢ may leave tail mark Walking Gaits Pace Diagonal Walk (Raccoon) (Dog Family, Cat, Deer, Opossum) Move both left feet together, Left front, right hind, right then both right feet front, left hind Bound Gallop (Weasel Family) (Rodent Family, Rabbit) Move both front feet together, Move both feet together, then then both hind feet move both hind feet Place hind feet behind front feet Place hind feet ahead of front feet . -
Slow Loris Forest Protector Teacher's Pack
2 www.nocturama.org https://www.facebook.com/pages/Little-Fireface-Project/ littlefi[email protected] INTRODUCTION Welcome! Welcome to the Slow Loris—Forest Protector’s Teacher’s Pack and Learning Exercise Book. The purpose of this short booklet is to explain how to use the book and to help to reinforce its message through a series of fun and easy-to- We at the Little Fireface Project (LFP) our so glad you are using this teacher’s use exercises. pack. First we want to tell you about ourselves and our passion about one of the world’s most unique little primates. LFP is a UK Charity based out of Ox- ford Brookes University set up to help save the slow loris through studying In this pack, you will find the following materials to help you and your students them in the wild and through education projects. explore the story of two night-active (nocturnal) primates, slow lorises: a moth- er (Tereh—speedy) and her young son (Bunga—flower). Tereh lovingly teaches Why the slow loris and why a charity dedicated to one group of animals? Well, her son the life skills he needs to be a grown-up slow loris. At the same time, the eight species of slow lorises, found only in Asia, are facing a tough time. Bunga learns that by doing his job in the forest, he helps the forest to grow, They are threatened for many reasons beyond the habitat loss causing many of while helping protect the crops grown by people. Asia’s species to go extinct. -
Lemur Bounce!
Assets – Reections Icon Style CoverAssets Style – Reections 1 Icon Style Y E F O N Cover StyleR O M M R A A Y E F D O N C A S R O GA M M R A D A AGASC LemurVISUAL Bounce! BRAND LearningGuidelines and Sponsorship Pack VISUAL BRAND Guidelines 2 Lemur Bounce nni My name is Lennie Le e Bounce with me B .. ou and raise money to n c protect my Rainforest e L ! home! L MfM L 3 What is a Lemur Bounce? A Lemur Bounce is a sponsored event for kids to raise money by playing bouncing games. In this pack: * Learning fun for kids including facts and quizzes Indoor crafts and outside bouncing games for * the Lemur Bounce Day * Links to teaching resources for schools * Lesson planning ideas for teachers * Everything you need for a packed day of learning and fun! Contents Fun Bounce activities Page No. Let’s BounceBounce – YourLemur valuable support 5 n n Lemur Bouncee i e L – Basics 7 B u o Planningn for a Lemur Bounce Day 8 c L e Lemur! Bounce Day Assembly 8 Make L a Lemur Mask 9 Make a Lemur Tail & Costume 10 Games 11-12 Sponsorship Forms 13-14 Assets – Reections M f Certificates 15-20 Icon Style M Cover Style L Y E F O N R Fun Indoor activities O M M R A Planning your lessons for a Lemur Bounce Day 22 D A A SC GA Fun Facts about Madagascar 23 Fun Facts about Lemurs 24 Colouring Template 25 VISUAL BRAND Guidelines Word Search 26 Know your Lemurs 27 Lemur Quiz 28 Madagascar Quiz 29-31 Resources 32 Song and Dance 33 Contact us 34 LEMUR BOUNCE BOUNCE fM FOR MfM LEMUR BASICS Y NE FO O R WHAT ‘LEM OUNC“ ‘ DO YOU KNOW YOUR LEMUR M Have you ever seen a lemur bounce? Maaasar is ome to over 0 seies o endemi lemurs inluding some M very ouncy ones lie the Siaa lemur ut tese oreous rimates are highl endangere e need to at R Let’s Bounce - Your valuable support A no to sae teir aitat and rotect tem rom etin tion arity Mone or Maaasar is alling out to A D C Wherechildren does everywhere the money to organizego? a fun charity ‘lemur bounce’. -
Ecology and Behaviour of Tarsius Syrichta in the Wild
O',F Tarsius syrichta ECOLOGY AND BEHAVIOUR - IN BOHOL, PHILIPPINES: IMPLICATIONS FOR CONSERVATION By Irene Neri-Arboleda D.V.M. A thesis submitted in fulfillment of the requirements for the degree of Master of Applied Science Department of Applied and Molecular Ecology University of Adelaide, South Australia 2001 TABLE OF CONTENTS DAge Title Page I Table of Contents............ 2 List of Tables..... 6 List of Figures.... 8 Acknowledgements... 10 Dedication 11 I)eclaration............ t2 Abstract.. 13 Chapter I GENERAL INTRODUCTION... l5 1.1 Philippine Biodiversity ........... t6 1.2 Thesis Format.... l9 1.3 Project Aims....... 20 Chapter 2 REVIEIV OF TARSIER BIOLOGY...... 2t 2.1 History and Distribution..... 22 2.t.1 History of Discovery... .. 22 2.1.2 Distribution...... 24 2.1.3 Subspecies of T. syrichta...... 24 2.2 Behaviour and Ecology.......... 27 2.2.1 Home Ranges. 27 2.2.2 Social Structure... 30 2.2.3 Reproductive Behaviour... 3l 2.2.4 Diet and Feeding Behaviour 32 2.2.5 Locomotion and Activity Patterns. 34 2.2.6 Population Density. 36 2.2.7 Habitat Preferences... ... 37 2.3 Summary of Review. 40 Chapter 3 FßLD SITE AI\D GEIYERAL METHODS.-..-....... 42 3.1 Field Site........ 43 3. 1.1 Geological History of the Philippines 43 3.1.2 Research Area: Corella, Bohol. 44 3.1.3 Physical Setting. 47 3.t.4 Climate. 47 3.1.5 Flora.. 50 3.1.6 Fauna. 53 3.1.7 Human Population 54 t page 3.1.8 Tourism 55 3.2 Methods.. 55 3.2.1 Mapping. -
GLOSSARY of MEDICAL and ANATOMICAL TERMS
GLOSSARY of MEDICAL and ANATOMICAL TERMS Abbreviations: • A. Arabic • abb. = abbreviation • c. circa = about • F. French • adj. adjective • G. Greek • Ge. German • cf. compare • L. Latin • dim. = diminutive • OF. Old French • ( ) plural form in brackets A-band abb. of anisotropic band G. anisos = unequal + tropos = turning; meaning having not equal properties in every direction; transverse bands in living skeletal muscle which rotate the plane of polarised light, cf. I-band. Abbé, Ernst. 1840-1905. German physicist; mathematical analysis of optics as a basis for constructing better microscopes; devised oil immersion lens; Abbé condenser. absorption L. absorbere = to suck up. acervulus L. = sand, gritty; brain sand (cf. psammoma body). acetylcholine an ester of choline found in many tissue, synapses & neuromuscular junctions, where it is a neural transmitter. acetylcholinesterase enzyme at motor end-plate responsible for rapid destruction of acetylcholine, a neurotransmitter. acidophilic adj. L. acidus = sour + G. philein = to love; affinity for an acidic dye, such as eosin staining cytoplasmic proteins. acinus (-i) L. = a juicy berry, a grape; applied to small, rounded terminal secretory units of compound exocrine glands that have a small lumen (adj. acinar). acrosome G. akron = extremity + soma = body; head of spermatozoon. actin polymer protein filament found in the intracellular cytoskeleton, particularly in the thin (I-) bands of striated muscle. adenohypophysis G. ade = an acorn + hypophyses = an undergrowth; anterior lobe of hypophysis (cf. pituitary). adenoid G. " + -oeides = in form of; in the form of a gland, glandular; the pharyngeal tonsil. adipocyte L. adeps = fat (of an animal) + G. kytos = a container; cells responsible for storage and metabolism of lipids, found in white fat and brown fat. -
Do Saki Monkeys Possess a Grooming Claw?
Short communication Primate Biol., 7, 19–23, 2020 https://doi.org/10.5194/pb-7-19-2020 © Author(s) 2020. This work is distributed under the Creative Commons Attribution 4.0 License. Do saki monkeys possess a grooming claw? Constanze Ohlendorf1,2,a and Eckhard W. Heymann2 1Soziobiologie/Anthropologie, Georg-August-Universität Göttingen, 37077 Göttingen, Germany 2Verhaltensökologie & Soziobiologie, Deutsches Primatenzentrum – Leibniz-Institut für Primatenforschung, 37077 Göttingen, Germany acurrent address: Planungsgemeinschaft LaReG, Helmstedter Straße 55A, 38126 Braunschweig, Germany Correspondence: Eckhard W. Heymann ([email protected]) Received: 19 February 2020 – Revised: 27 July 2020 – Accepted: 10 August 2020 – Published: 15 September 2020 Abstract. The presence of a grooming claw on the second toe is a characteristic of Strepsirrhini and tarsiers. There is also some evidence for the presence of a grooming claw in Platyrrhini. Here we report qualitative findings from different species of saki monkeys, genus Pithecia, on the presence of modified nails on the second toe. These observations suggest that a grooming claw or a grooming claw-like nail occurs in different Pithecia species, but that it does not consistently occur in all individuals. 1 Introduction Platyrrhini retain several plesiomorphic primate traits, e.g., the presence of three premolars and the fusion of the ec- totympanic ring with the side of the auditory bulla (Flea- Grooming claws are a well-known morphological charac- gle, 2013). Bluntschli (1929) was the first to suggest that teristic of Strepsirrhini and tarsiers. Strepsirrhini possess Platyrrhini may also retain a grooming claw. He reported grooming claws on the second toe, tarsiers also on the third modified nails on the second toe of wild individuals of the toe. -
Scramble Puzzle | Suborder Prosimians
Scramble Puzzle | Suborder Prosimians In taxonomy, primates are divided into two major groups: the Prosimians and the Anthropoids. Prosimians (lemurs, lorises, and tarsiers) are a diverse group of primates that appear to retain more primitive features compared to Anthropoids (monkeys, apes and humans). Several unique characteristics separate prosimians from anthropoids including the presence of a post orbital bar (i.e. lack of postorbital closure), a rhinarium (i.e. wet nose), relatively longer snout, and relatively smaller brain compared to body size. Other specialized features include a 2.1.3.3. dental formula, the presence of a tooth comb and a grooming claw on the second digit of the foot. Tarsiers show a combination of prosimian and anthropoid features, and are sometimes categorized into a different phyletic classification that uses the suborder Haplorhini. Most prosimians have large eyes and are nocturnal. Although fossils prosimians have been found in North America and Europe, living prosimians are typically found in the tropical environments of African and Asia, though five of the eight living taxonomic families live exclusively in Madagascar. Instructions for Printing Puzzle: 1. Print this page on white 8 ½” x 11” heavy cardstock, although regular printer paper will also work. Using a higher printing resolution will improve the quality of the images. 2. Cut each puzzle piece along the black lines provided. 3. Mix up the pieces, the try to reassemble by matching up the correct part of the skulls. Remember that you can rotate the puzzle pieces. 4. Have fun and learn! This puzzle contains the species (not to scale): Lesser Bushbaby Ruffed Lemur Slow Loris Tarsier (Varecia variegata) (Galago senegalensis) (Nycticebus coucang (Tarsius bancanus coucang) borneanus) ©eSkeletons 2009 www.eSkeletons.org . -
Occurrence of Asian Small-Clawed Otter Aonyx Cinereus (Illiger, 1815) in Eastern India
SCIENTIFIC CORRESPONDENCE 5. McLennan, S. M. and Taylor, S. R., J. Geol., 1991, 99, 1–21. 6. Sensarma, S. Hoernes, S. and Muk- hopadhyay, D., Proc. Indian Acad. Sci. (Earth Planet. Sci.), 2004, 113(4), 619– 648. 7. McKelvey, V. E., Everhart, D. L. and Gar- rels, R. M., Econ. Geol., 1955, 15th anni- versary volume, p. 491. ACKNOWLEDGEMENTS. We thank the Director, Atomic Minerals Directorate for Exploration and Research (AMD) for encour- agement and granting permission to publish this paper. We thank our colleagues from the Physics, Chemistry, XRF and Petrology labo- ratories, AMD, Nagpur for providing labora- tory support. Received 2 February 2014; revised accepted 16 June 2014 Figure 5. a, Sericite masked with limonite/iron oxide and very fine size quartz in radioactive 1, shale. b, Alpha track matching with limonite and iron oxide on radioactive shale. c, Adsorbed U. P. SHARMA * 1 uranium on radioactive phyllite corresponding with alpha track. d, Alpha track on radioactive S. SHUKLA 1 phyllite. P. K. SINHA 1 R. K. PUROHIT 1 basement and overlying sediments. These within rocks of Chilpi Group and under- A. MAJUMDAR 2 faults probably acted as conduits for lying basement rocks of Nandgaon A. K. RAI transfer of uranium-bearing solution Group. from basement rocks. Kanhari and 1 adjoining areas can be looked for struc- Atomic Minerals Directorate for turally controlled and fracture-bound 1. Basu, A. K., Geol. Surv. India, Spec. Publ., Exploration and Research, unconformity type of uranium minerali- 2001, 55, 181–204. AMD Complex, Civil Lines, 2. Thorat, P. K., Natrajan, A., Guha, K. -
Verreaux's Sifaka (Propithecus Verreauxi) and Ring- Tailed Lemur
MADAGASCAR CONSERVATION & DEVELOPMENT VOLUME 8 | ISSUE 1 — JULY 2013 PAGE 21 ARTICLE http://dx.doi.org/10.4314/mcd.v8i1.4 Verreaux’s sifaka (Propithecus verreauxi) and ring- tailed lemur (Lemur catta) endoparasitism at the Bezà Mahafaly Special Reserve James E. LoudonI,II, Michelle L. SautherII Correspondence: James E. Loudon Department of Anthropology, University of Colorado-Boulder Boulder, CO 80309-0233, U.S.A. E - mail: [email protected] ABSTRACT facteurs comportementaux lors de l’acquisition et l’évitement As hosts, primate behavior is responsible for parasite avoid- des parasites transmis par voie orale en comparant le compor- ance and elimination as well as parasite acquisition and trans- tement des Propithèques de Verreaux (Propithecus verreauxi) et mission among conspecifics. Thus, host behavior is largely des Makis (Lemur catta) se trouvant dans la Réserve Spéciale du responsible for the distribution of parasites in free - ranging Bezà Mahafaly (RSBM) à Madagascar. Deux groupes de chacune populations. We examined the importance of host behavior in de ces espèces étaient distribués dans une parcelle protégée et acquiring and avoiding parasites that use oral routes by compar- deux autres dans des forêts dégradées par l’activité humaine. ing the behavior of sympatric Verreaux’s sifaka (Propithecus L’analyse de 585 échantillons fécaux a révélé que les Makis verreauxi) and ring - tailed lemurs (Lemur catta) inhabiting the de la RSBM étaient infestés par six espèces de nématodes et Bezà Mahafaly Special Reserve (BMSR) in Madagascar. For trois espèces de parasites protistes tandis que les Propithèques each species, two groups lived in a protected parcel and two de Verreaux ne l’étaient que par deux espèces de nématodes. -
GAIT PATTERNS Pacer Diagonal Walker Bounder Galloper RABBITS
GAIT PATTERNS RODENTS Shows - 4 toes front, 5 toes rear, claws General Shape Normal Pace Gait: Galloper Pacer Diagonal Walker Indirect Register Gallopers: Squirrels, Ground Squirrels, Mice Rats, Bounder Chipmunks, Ground Hog, Marmot. Cross Pattern Tree dwellers show both pairs of feet parallel. Ground dwellers show dominant foot landing first. Squirrel Pacers: Porcupine, Muskrat, Beaver, Mountain Beaver Galloper Porcupine, Muskrat, Beaver - in deep mud show 5 toes in front (a hidden thumb). Mountain Beaver - always shoes 5 toes in front. RABBITS & HARES Shows - 4 toes front, 4 toes rear CAT FAMILY Shows - 4 toes front, 4 toes rear, claws (rarely) General Shape Normal Pace Gait: Galloper General Shape Normal Pace Gait: Diagonal Walker Rear Indirect Register Direct Register Elbow on the rear foot may or may not show. Front feet 1/2 larger than rear No claws (95% of time) - sometimes out during a hunt. Rabbit - rear feet 2 times larger than front feet Round Zero straddle Hare - rear feet 4-5 times larger than front. Zero pitch Front Rear The small heel pad helps to distinguish between a showshoe hare with no elbow showing and a Feral Cat - 4 toes equal size with elbow dog galloping Rabbit Mountain Lion - 4 toes equal size Cat Bobcat - inner toes larger, cleft in heel pad Lynx - outer toes larger DOG FAMILY Shows - 4 toes front, 4 toes rear, claws WEASEL FAMILY Shows - 5 toes front, 5 toes rear, claws General Shape Normal Pace Gait: Diagonal Walker General Shape Normal Pace Gait: Bounder Indirect Register Indirect Register Front feet 1/3 larger than rear.