Sigillaria, It Is Necessary to Describe the Specimens on Which These Views As to the Systematic Position of This Species Have Been Founded

Total Page:16

File Type:pdf, Size:1020Kb

Sigillaria, It Is Necessary to Describe the Specimens on Which These Views As to the Systematic Position of This Species Have Been Founded 560 MR. R. KIDSTON ON THE OCCURRENCE OF and he probably assumed the Indian specimens to be distinct. Until De Kotaris distinguished his 3. decipiens, the two species confused had passed as one ; but when they are divided, it is clear that Hedwig had already fixed the idea of P. adiantoides, by de- scribing it as dioicous ; and that the overlooked one was the one here enumerated as P. mujus, which the older authors thought to be a bog species, and which Dillenius describes, p. 264, as “Hypnunz taxiforme palustre ramosum, majus et erecturn.” The Hypnunz adiantoides of Linn. Sp. Plant., has I‘ fronde pinnata ramosa erecta medio pedunculifera.” Here the “ palustre ” is omitted. Bay had in his character “ aquaticus ;” and this mention of its habitat points rather to the species here marked as 3. majus, El adiantoides being more a rupestral moss. It is remarkable in this species that the specimens from America are all, as figured by Sullivant, very much smaller in all their parts except the floral leaves. It is probably the most generally as well as the most widely distributed species known.-The whole temperate northern hemisphere. 31. F. SUBBASILARIS, Hedw., Xull. Tcon. 26.-A small tufted species with erect fruit.--North America. 38. F. JULIANUS,Xaai (Fontinalis), Brid. ii. p. 678.-ItaIg and West France. It should be looked for in the clear streams of S.W. Britain. 39. 3’. HALLIANUS,Sull. et Lesp., Xull. Icon. p. 48, t. 28. A much shorter-stemmed and tufted species.-New Jersey and Illinois. On the Occurrence of Lycopodites (Xigillaria) Vawxemi, Gop- pert, in Britain, with remarks on its Affinities. By ROBERT EIDSTON,F.G.S. (Communicated by Dr. J. MUEIE,P.L.S.) [Read 4th June, 1885.1 (PLATEXVIII.) SINCEwriting my previous paper, ‘‘ On a new Species of Lyco- podites from the Calciferous Sandstoue Series of Scotland”’, Lesquereux, in the third volume of his Coal Flora?, has described * Ann, & Mag Nat. Hist. Bug. 1884, p. 111, t. Description of the Coal Flora of the Carboniferous Formation in Penn- aylvaniaand throughout.the United States, 701. iii. p. 778 (1884) : Harrisburg. LYCOPODITES VA’dNUXEMI IN BRITAIN. 561 four new species of Lycopodites :-Lycopodites arborescens, L. .&~ifoji~s,L. simp~ex,and L. Lacoei. Of these I think the last two must be regarded as doubtfully referable to Lycopodites, Goldenberg ; but L. arborescens and L. jkxifolius are two most interestiiy additions to Goldenberg’s (not Brongniart’s) genus. The firrt-mentioned author formed, or, more correctly, resus- citated the Brongniartian name Lycopodites in 1855* for a then new class of Palreozoic plants, “ true herbaceous Lycopods j ” but to enable Lesquereux to include his lately-discovered species in thia genus, he waa under the necemity of slightly modifying Goldenberg’s definition of Lycopodites, by adding the term “ or woody,” to enable him to place in it plants which were evidently not herbaceous, but which probably did not otherwise differ from those species previously placed in Lycopodifes by Golden- berg. The generic definition of Lycopodites, Goldenberg, so emended, must now read:- Herbaceous or woody plants zuifh dichotomous ram$cation ; branches with leaves placed spirally or in verticils, leaves of the same or of two djferent forms on the same branches; sporangia placed in the axils of the 1eaGes or forming terminal cones. The addition of the term ‘‘ woody ” was rendered necessary on account of the size of the stem of Lycopodites arborescens, Lesq., which in every other character is referable to Goldenberg’s genus : hence the slight modification of the generic terms of Golden- berg’s Lycopodites, as proposed by Lesquereux, seems quite legi- timate. It is also possible that Goldenberg’s type-specimens were only branches of a comparatively large-stemmed plant ; but whether Goldenberg’s plants were herbaceous or not, it is most improbable that Lycopodites arborescens, Lesq., or Xigillaria Vafiuxenzi, Giippert, mere herbaceous, the latter of which I now propose to include in Lycopodites, Goldenberg. Lycopodites (SiyiZZaria) Panuxemi was first figured by Vanuxern in 1842f. He gires a description of the plant without applying any name to it, but expressed the opinion that it more closely related to Zepidodendpon than to any other genus. This specimen was discovered in Allan’s Quarry, near @wego, New * 6 Flora Sarqontana Fossilis,’ Heft i. PP. 9, 10. i ‘ Gd. of New York,’ part iii. p. 184, fig. 51. 562 MR. R. KIDSTON ON THE OCCURRENCE OF Pork, in rocks belonging to the Chenning Group (Upper De- vonian). In 1852 Goppert named it Siqillaria Panuxemi". Although several notes on this species occur in the literature of fossil Botany, till the discovery of the British specimens, the type figured by Vanuxem seems to have been the only individual of the species known. Before fully stating the considerations which have led me to place this plant in Lycopoditest, in preference to retaining it in Sigillaria, it is necessary to describe the specimens on which these views as to the systematic position of this species have been founded. For the specimens of Lycopodites (Sigillaria) Panuxemi, which have been submitted to me for examination, and of which are figured on PI. XVIII. some of the more characteristic and better preserved examples, I am indebted to the Council of the Kendal Museum, as also to Mr. R. Bullen Newton,F.G.S., of the British Museum, South Eensington, through whose ageucy they were communicated to me. Description of Specimens. figure 1. This specimen, of which fig. 1 shows (natural size) the upper portion, is about 13 centim. long and about 2.5 centim. wide, though its original breadth must have been greater, as its full width is not shown in the fossil. The leaf-scars are more distinct on the upper than on the lower part, where they are somewhat obscured by coaly matter. They are hexagonal, about 6 millim. long by 5 millim. broad ; the angles formed at the point of junction of the slightly raised boundary lines of the leaf-scars me acute, though the two angles at the upper part of the leaf- scar have occasionally a tendency to become rounded. No dis- tinct trace of any vascular cicatricule is shown on this specimen. The broadest part of the leaf-scar is slightly above the middle. Zigure 2. This figure, which is natural size, shows the mode of growth of the younger branches. The specimen is not very distinctly preserved, but it exhibits at various poiuts the form and size of the leaf-scars, whose outline is defined by a carbo- naceous stain. On the upper branches the leaf-scars are much longer than broad, perfectly flat, and bear a single vascular * Foss. Flora d. Uebergangsgebirges, p. 249 (1852). + I have already stated my objections to the use of the generic name Lyco- podium for these fossils in Ann. & Mag. Nat. Hist. Bug. 1884, p. 114. LYCOPODITES VANUXEMI IN BRITAIN. 5 63 cicatricule, which is indicated by a dark point situated shghtly above the centre. The leaf-scars on the lower portions of the stems are distinctly hexagonal, about as broad as long, and sliow in several cases the scar of the vascular bundle, whict is indicated by a small dark point. The branchas appear to divide dichoto- mously ; but this character is better seen on some of the other examples. This fossil is labelled ‘‘ Shap Toll-bar, J. Euthven.” -Figure 3a. This shows a small fra,ment of a branch from an- other part of the same slab as that from which fig. 2 was taken, and gives the form of the leaf-scars on the younger branches. The stem here, as in the previous specimens, is flattened by pressure. The leaf-scars are indicated by a bron 11 carbonaceous outline. Fig. 3 6 is the same enlarged. Pigure 4. This is a small portion of a young branch, natural size, given to illustrate the form of the leaf-scars, which is broadly fusiform. 9igure 5. An enlarged sketch of a portion of a small branch (marked a on fig. 2) is given at fig. 5, to show more fully the form of the leaf-scars and their relationship to each other. They are broadly fusiform; the upper and lower extremities of the leaf-scars are separated by a slight interval, though the scars laterally rest upon each other. The hexagonal form of the leaf-scars, eventually assumed on the older branches, is evidently the result of mutual pressure and the lateral tension exerted on them by the increase of the stem in girth. The leaf-scars show a single elongated vascular cicatricule ; in the older stems (fig. 2 b) the -mscular cicatricule appears to be punctiform. Pigwe 6. One of the stems on this slab exhibits very clearly the dichotomous ramifications of the species. The brmch is shown in section, as the separation of the stone has passed through the central axis of the stem. Most of the leaves with which it is clothed are likewise shown in section, but towards the upper part of the left arm of the fork their form is indicated, though not distinctly. At the part marked a (fig. 6a gives an enlarged sketch) the shape of the leaves is more clearly exhibited, as one of them extends beyond the broken extremity of a small branch. Owing to the imperfect state of its preservation, no trace of a central vein is observable. 534 MR. R. KIDSTON ON THE OCCURRENCE OF General Remarks. The leaf-scars on the young branches are Lepidodendroid in form, but do not touch each other at their upper and lower extremities, though they meet laterally.
Recommended publications
  • Retallack 2021 Coal Balls
    Palaeogeography, Palaeoclimatology, Palaeoecology 564 (2021) 110185 Contents lists available at ScienceDirect Palaeogeography, Palaeoclimatology, Palaeoecology journal homepage: www.elsevier.com/locate/palaeo Modern analogs reveal the origin of Carboniferous coal balls Gregory Retallack * Department of Earth Science, University of Oregon, Eugene, Oregon 97403-1272, USA ARTICLE INFO ABSTRACT Keywords: Coal balls are calcareous peats with cellular permineralization invaluable for understanding the anatomy of Coal ball Pennsylvanian and Permian fossil plants. Two distinct kinds of coal balls are here recognized in both Holocene Histosol and Pennsylvanian calcareous Histosols. Respirogenic calcite coal balls have arrays of calcite δ18O and δ13C like Carbon isotopes those of desert soil calcic horizons reflecting isotopic composition of CO2 gas from an aerobic microbiome. Permineralization Methanogenic calcite coal balls in contrast have invariant δ18O for a range of δ13C, and formed with anaerobic microbiomes in soil solutions with bicarbonate formed by methane oxidation and sugar fermentation. Respiro­ genic coal balls are described from Holocene peats in Eight Mile Creek South Australia, and noted from Carboniferous coals near Penistone, Yorkshire. Methanogenic coal balls are described from Carboniferous coals at Berryville (Illinois) and Steubenville (Ohio), Paleocene lignites of Sutton (Alaska), Eocene lignites of Axel Heiberg Island (Nunavut), Pleistocene peats of Konya (Turkey), and Holocene peats of Gramigne di Bando (Italy). Soils and paleosols with coal balls are neither common nor extinct, but were formed by two distinct soil microbiomes. 1. Introduction and Royer, 2019). Although best known from Euramerican coal mea­ sures of Pennsylvanian age (Greb et al., 1999; Raymond et al., 2012, Coal balls were best defined by Seward (1895, p.
    [Show full text]
  • The Joggins Fossil Cliffs UNESCO World Heritage Site: a Review of Recent Research
    The Joggins Fossil Cliffs UNESCO World Heritage site: a review of recent research Melissa Grey¹,²* and Zoe V. Finkel² 1. Joggins Fossil Institute, 100 Main St. Joggins, Nova Scotia B0L 1A0, Canada 2. Environmental Science Program, Mount Allison University, Sackville, New Brunswick E4L 1G7, Canada *Corresponding author: <[email protected]> Date received: 28 July 2010 ¶ Date accepted 25 May 2011 ABSTRACT The Joggins Fossil Cliffs UNESCO World Heritage Site is a Carboniferous coastal section along the shores of the Cumberland Basin, an extension of Chignecto Bay, itself an arm of the Bay of Fundy, with excellent preservation of biota preserved in their environmental context. The Cliffs provide insight into the Late Carboniferous (Pennsylvanian) world, the most important interval in Earth’s past for the formation of coal. The site has had a long history of scientific research and, while there have been well over 100 publications in over 150 years of research at the Cliffs, discoveries continue and critical questions remain. Recent research (post-1950) falls under one of three categories: general geol- ogy; paleobiology; and paleoenvironmental reconstruction, and provides a context for future work at the site. While recent research has made large strides in our understanding of the Late Carboniferous, many questions remain to be studied and resolved, and interest in addressing these issues is clearly not waning. Within the World Heritage Site, we suggest that the uppermost formations (Springhill Mines and Ragged Reef), paleosols, floral and trace fossil tax- onomy, and microevolutionary patterns are among the most promising areas for future study. RÉSUMÉ Le site du patrimoine mondial de l’UNESCO des falaises fossilifères de Joggins est situé sur une partie du littoral qui date du Carbonifère, sur les rives du bassin de Cumberland, qui est une prolongation de la baie de Chignecto, elle-même un bras de la baie de Fundy.
    [Show full text]
  • The Joggins Cliffs of Nova Scotia: B2 the Joggins Cliffs of Nova Scotia: Lyell & Co's "Coal Age Galapagos" J.H
    GAC-MAC-CSPG-CSSS Pre-conference Field Trips A1 Contamination in the South Mountain Batholith and Port Mouton Pluton, southern Nova Scotia HALIFAX Building Bridges—across science, through time, around2005 the world D. Barrie Clarke and Saskia Erdmann A2 Salt tectonics and sedimentation in western Cape Breton Island, Nova Scotia Ian Davison and Chris Jauer A3 Glaciation and landscapes of the Halifax region, Nova Scotia Ralph Stea and John Gosse A4 Structural geology and vein arrays of lode gold deposits, Meguma terrane, Nova Scotia Rick Horne A5 Facies heterogeneity in lacustrine basins: the transtensional Moncton Basin (Mississippian) and extensional Fundy Basin (Triassic-Jurassic), New Brunswick and Nova Scotia David Keighley and David E. Brown A6 Geological setting of intrusion-related gold mineralization in southwestern New Brunswick Kathleen Thorne, Malcolm McLeod, Les Fyffe, and David Lentz A7 The Triassic-Jurassic faunal and floral transition in the Fundy Basin, Nova Scotia Paul Olsen, Jessica Whiteside, and Tim Fedak Post-conference Field Trips B1 Accretion of peri-Gondwanan terranes, northern mainland Nova Scotia Field Trip B2 and southern New Brunswick Sandra Barr, Susan Johnson, Brendan Murphy, Georgia Pe-Piper, David Piper, and Chris White The Joggins Cliffs of Nova Scotia: B2 The Joggins Cliffs of Nova Scotia: Lyell & Co's "Coal Age Galapagos" J.H. Calder, M.R. Gibling, and M.C. Rygel Lyell & Co's "Coal Age Galapagos” B3 Geology and volcanology of the Jurassic North Mountain Basalt, southern Nova Scotia Dan Kontak, Jarda Dostal,
    [Show full text]
  • JOGGINS RESEARCH SYMPOSIUM September 22, 2018
    JOGGINS RESEARCH SYMPOSIUM September 22, 2018 2 INTRODUCTION AND ACKNOWLEDGEMENTS The Joggins Fossil Cliffs is celebrating its tenth year as a UNESCO World Heritage Site! To acknowledge this special anniversary, the Joggins Fossil Institute (JFI), and its Science Advisory Committee, organized this symposium to highlight recent and current research conducted at Joggins and work relevant to the site and the Pennsylvanian in general. We organized a day with plenty of opportunity for discussion and discovery so we invite you to share, learn and enjoy your time at the Joggins Fossils Cliffs! The organizing committee appreciates the support of the Atlantic Geoscience Society for this event and in general. Sincerely, JFI Science Advisory Committee, Symposium Subcommittee: Elisabeth Kosters (Chair), Nikole Bingham-Koslowski, Suzie Currie, Lynn Dafoe, Melissa Grey, and Jason Loxton 3 CONTENTS Symposium schedule 4 Technical session schedule 5 Abstracts (arranged alphabetically by first author) 6 Basic Field Guide to the Joggins Formation______________________18 4 SYMPOSIUM SCHEDULE 8:30 – 9:00 am Registration 9:00 – 9:10 am Welcome by Dr. Elisabeth Kosters, JFI Science Advisory Committee Chair 9:10 – 10:30 am Talks 10:25 – 10:40 am Coffee Break and Discussion 10:45 – 12:00 pm Talks and Discussion 12:00 – 1:30 pm Lunch 1:30 – 4:30 pm Joggins Formation Field Trip 4:30 – 6:00 pm Refreshments and Wrap-up 5 TECHNICAL SESSION SCHEDULE Chair: Melissa Grey 9:10 – 9:20 am Peir Pufahl 9:20 – 9:30 am Nikole Bingham-Koslowski 9:30 – 9:40 am Michael Ryan 9:40 – 9:50 am Lynn Dafoe 9:50 – 10:00 am Matt Stimson 10:00 – 10:15 am Olivia King 10:15 – 10:25 am Hillary Maddin COFFEE BREAK Chair: Elisabeth Kosters 10:40 – 10:50 am Martin Gibling 10:50 – 11:00 am Todd Ventura 11:10 – 11:20 am Jason Loxton 11:20 – 11:30 am Nathan Rowbottom 11:30 – 11:40 am John Calder 11:40 – 12:30 Discussion led by Elisabeth Kosters LUNCH 6 ABSTRACTS Breaking down Late Carboniferous fish coprolites from the Joggins Formation NIKOLE BINGHAM-KOSLOWSKI1, MELISSA GREY2, PEIR PUFAHL3, AND JAMES M.
    [Show full text]
  • 81 Vascular Plant Diversity
    f 80 CHAPTER 4 EVOLUTION AND DIVERSITY OF VASCULAR PLANTS UNIT II EVOLUTION AND DIVERSITY OF PLANTS 81 LYCOPODIOPHYTA Gleicheniales Polypodiales LYCOPODIOPSIDA Dipteridaceae (2/Il) Aspleniaceae (1—10/700+) Lycopodiaceae (5/300) Gleicheniaceae (6/125) Blechnaceae (9/200) ISOETOPSIDA Matoniaceae (2/4) Davalliaceae (4—5/65) Isoetaceae (1/200) Schizaeales Dennstaedtiaceae (11/170) Selaginellaceae (1/700) Anemiaceae (1/100+) Dryopteridaceae (40—45/1700) EUPHYLLOPHYTA Lygodiaceae (1/25) Lindsaeaceae (8/200) MONILOPHYTA Schizaeaceae (2/30) Lomariopsidaceae (4/70) EQifiSETOPSIDA Salviniales Oleandraceae (1/40) Equisetaceae (1/15) Marsileaceae (3/75) Onocleaceae (4/5) PSILOTOPSIDA Salviniaceae (2/16) Polypodiaceae (56/1200) Ophioglossaceae (4/55—80) Cyatheales Pteridaceae (50/950) Psilotaceae (2/17) Cibotiaceae (1/11) Saccolomataceae (1/12) MARATTIOPSIDA Culcitaceae (1/2) Tectariaceae (3—15/230) Marattiaceae (6/80) Cyatheaceae (4/600+) Thelypteridaceae (5—30/950) POLYPODIOPSIDA Dicksoniaceae (3/30) Woodsiaceae (15/700) Osmundales Loxomataceae (2/2) central vascular cylinder Osmundaceae (3/20) Metaxyaceae (1/2) SPERMATOPHYTA (See Chapter 5) Hymenophyllales Plagiogyriaceae (1/15) FIGURE 4.9 Anatomy of the root, an apomorphy of the vascular plants. A. Root whole mount. B. Root longitudinal-section. C. Whole Hymenophyllaceae (9/600) Thyrsopteridaceae (1/1) root cross-section. D. Close-up of central vascular cylinder, showing tissues. TABLE 4.1 Taxonomic groups of Tracheophyta, vascular plants (minus those of Spermatophyta, seed plants). Classes, orders, and family names after Smith et al. (2006). Higher groups (traditionally treated as phyla) after Cantino et al. (2007). Families in bold are described in found today in the Selaginellaceae of the lycophytes and all the pericycle or endodermis. Lateral roots penetrate the tis detail.
    [Show full text]
  • Plant Evolution
    Conquering the land The rise of plants Ordovician Spores Algae (algal mats) Green freshwater algae Bacteria Fungae Bryophytes Moses? Liverworts? Little body fossil evidence Silurian Wenlock Stage 423-428mya Psilophytes Rhyniopsidsa important later in early Devonian Cooksonia Rhynia Branching stems, flattened sporangia at tips No leaves, no roots short 30 cms rhizoids Zosterophylls Early stem group of Lycopodiophytes Ancestors of Class Lycopsida (clubmosses) Prevalent in Devonian Spores at tips and on branches Lycopsids (?) Baragwanathia with microphylls in Australia Zosterophylls Silurian Cooksonia Development of Soil Fungae Bacteria Algae Organic matter Arthropods and annelids Change in erosion Change in CO2 Devonian Devonian Early Devonian simple structure Rhynie Chert (Rhyniophytes) Trimerophytes First with main shoot Give rise to Ferns and Progymnosperms Up to 3m tall Animal life (mainly arthropods) Late Devonian Forests First true wood (lignin) Forest structure develops (stories) Sphenopsids (Calamites) Lycopsids (Lepidodendron) Seed Ferns (Pteridosperm) Progymnosperm Archaeopteris Cladoxylopsid First vertebrates present Upper Devonian Lycopsida 374-360 mya Leaves and roots differentiated Most ancient with living relatives Megaphylls branching in on plane Photosynthetic webbing Shrub size vertical growth limited (weak) Lateral (secondary) growth (woody) Development of roots Homosporous Heterosporous Upper Devonian Calamites (Sphenopsid) Horestail Sphenophyta (Calamites-Annularia) Devonian Archaeopteris Ur. Devonian - Lr. Carboniferous
    [Show full text]
  • Lycophytes-1St-Semester
    Structural Botany Class Lycopsida Archana Dutta Study material for M.Sc Botany First semester Assistant Professor(Guest Faculty) Dept. of Botany, MLT College, Saharsa [email protected] Mob No. - 9065558829 5th May, 2020 _______________________________________________________________________ ___ Members of the Lycopsida have true stems, leaves and (usually) roots. Their sporangia are borne on the adaxial surface of (or in the axils of) sporophylls, and they may or may not resemble the vegetative leaves. In the Alycopods@branch gaps are present in the stele, but no leaf gaps occur in the xylem. Therefore, when the stele has parenchyma at the center, it is a medullated protostele. The leaves (with only one or two exceptions) have a single vascular strand and appear to have arisen phylogenetically from superficial emergences or enations. The extant orders of Lycopsida are: Lycopodiales, Selaginellales, and Isoetales. Fossil orders sometimes include the Protolepidodendrales, Lepidodendrales, and Pleuromeiales. However, in this course the Protolepidodendrales are included in the Lycopodiales, and both the Lepidodendrales and Pleuromeiales are included in the Isoetales. A comprehensive evaluation of lycophytes was published in the Annals of the Missouri Botanical Garden in 1992 (Vol. 79: 447-736), and the included papers still reflect current knowledge of the group. Order Lycopodiales Lycopodiales are homosporous, herbaceous, eligulate, isophyllous, and they have true roots. Traditionally, approximately 400 species of two extant genera have been recognized and assigned to the family Lycopodiaceae. More recently, however, (Lycopodium ) has been subdivided such that seven or more genera and three families are now recognized. Classifications are as follows (from Wagner and Beitel, 1992). For the purposes of (examinations in) this course we will recognize only three genera (Lycopodium, Huperzia and Diphasiastrum).
    [Show full text]
  • Stigmaria Fossils
    Lycopods Throughout geologic time the seas have undergone transgressions (high sea level) and regressions (low sea level). During times of regression land is exposed, and in the Mississippian and Pennsylvanian there was abundant plant life thriving in subtropical conditions around the world. Most of the plants that grew in swampy areas eventually fell and accumulated in large numbers to ultimately form coal, lending the name Carboniferous Period or coal measures. The dominant trees of this period were the lycopods Lepidodendron and Sigillaria. Other plants that were more common on higher ground and along floodplains are the fossil horsetails called Calamites which were tree-sized plants reaching a height of approximately 20 m. In Arkansas, the best preserved of these fossils are found in Pennsylvanian age rocks from the Arkansas River Valley. Small fragments of plant fossils can be found in Mississippian and Pennsylvanian age rocks in the Ozark Plateaus Region. Lepidodendron, an extinct coal-age tree. Illustrations by Sherrie Shepherd. Sigillaria, an extinct coal-age tree. Illustrations by Sherrie Shepherd. Calamites, an extinct horsetail. Illustrations by Sherrie Shepherd. Stigmaria fossils Plant fossils with small round indentions are usually fossils of the root system called Stigmaria, of either of the trees, Lepidodendron or Sigillaria. The bark of Sigillaria is differentiated by parallel lines between rows of round indentions. The majority of Stigmaria fossils are found in the Pennsylvanian Hartshorne Sandstone and McAlester Shale of the Arkansas River Valley. Stigmaria fossil from Bates, Arkansas Stigmaria fossil from Bates, Arkansas .
    [Show full text]
  • Arborescent Lycopsid Productivity and Lifespan: Constraining the Possibilities
    Review of Palaeobotany and Palynology 227 (2016) 97–110 Contents lists available at ScienceDirect Review of Palaeobotany and Palynology journal homepage: www.elsevier.com/locate/revpalbo Arborescent lycopsid productivity and lifespan: Constraining the possibilities C. Kevin Boyce a,⁎, William A. DiMichele b a Department of Geological Sciences, Stanford University, Stanford, CA, USA b Department of Paleobiology, NMNH Smithsonian Institution, Washington, DC, USA article info abstract Article history: One of the most enigmatic components of early terrestrial vegetation was the arborescent lycopsids. Because of Received 25 March 2015 the sheer abundance of their biomass in many wetland environments of the Late Paleozoic, they may have been Received in revised form 22 September 2015 an important variable in the global carbon cycle and climate. However, their unusual structure has invited ex- Accepted 9 October 2015 traordinary interpretations regarding their biology. One idea that has persisted in the literature for over forty Available online 9 November 2015 years is that these trees had extremely short lifespans, on the order of ten years. Such an accelerated lifecycle Keywords: would require growth rates twenty times higher than modern angiosperm trees (and at least 60 times higher — Carboniferous than modern lycopsids). Here, we evaluate the morphology and anatomy of lycopsid trees including aerenchy- Lepidodendron ma, phloem, leaf base distributions, leaf structure, rootlet anatomy, and the demography of the preserved Lepidophloios fossils—with comparison to modern plants with some similarity of overall form, most notably the palms. The en- Oldest palm vironmental context of lycopsid trees also is considered in the light of the vegetation of modern water-saturated Sigillaria substrates.
    [Show full text]
  • The Gametophyte-Sporophyte Junction in Isoetes Boliviensis WEBER (Isoetales, Lycopodiophyta) By
    ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Phyton, Annales Rei Botanicae, Horn Jahr/Year: 2002 Band/Volume: 42_1 Autor(en)/Author(s): Hilger Hartmut H., Weigend Maximilian, Frey Wolfgang Artikel/Article: The Gametophyte-Sporophyte Junction in Isoëtes boliviensis WEBER (Isoëtales, Lycopodiophyta). 149-157 ©Verlag Ferdinand Berger & Söhne Ges.m.b.H., Horn, Austria, download unter www.biologiezentrum.at Phyton (Horn, Austria) Vol. 42 Fasc. 1 149-157 29. 7. 2002 The Gametophyte-Sporophyte Junction in Isoetes boliviensis WEBER (Isoetales, Lycopodiophyta) By Hartmut H. HILGER, Maximilian WEIGEND and Wolfgang FREY*) With 10 Figures Received January 25, 2002 Key words: Isoetes, Isoetales, Lycopodiophyta. - Anatomy, gametophyte- sporophyte junction, placental space. - Electron microscopy. Summary HILGER H. H., WEIGEND M. & FREY W. 2002. The gametophyte-sporophyte junction in Isoetes boliviensis WEBER (Isoetales, Lycopodiophyta). - Phyton (Horn, Austria) 42(1): 149-157, 10 figures. - English with German summary. The gametophyte-sporophyte junction in Isoetes boliviensis WEBER consists of a sporophytic conical foot embedded in the maternal gametophytic tissue. Both gene- rations are separated by a narrow placental space filled with thin-walled collapsed cells of gametophytic origin. Gametophytic and sporophytic placental cells lack wall ingrowths, or transfer cells. Interdigitation or intermingling of placental cells is not observed. Cell wall
    [Show full text]
  • Lepidocarpon), (1956
    Acta Botanica Neerlandica 7 (1958) 217 —222 On a Megaspore-bearing Lycopod Strobilus S.J. Dijkstra [Geol. Bur., Heerlen) (:received Februiry 11th, 1958) Introduction the last 30 of Carboniferous During years many species megaspores have been described, but until now not much is known with regard to the nature of the fructifications by which they were produced. This be due the fact that all the were may partly to nearly megaspores obtained by macerating lumps of coal, whereas the fructifications are generally derived from the roof of a coal seam. Moreover it is note- that the fructifications contain had worthy rarely megaspores; they shed their before themselves became detached. apparently spores they Some investigators of the past century sometimes found megaspores the in the strobili they were studying, but not realizing importance their small of findings, they unfortunately figured them on a very that it is scale. Their descriptions too are insufficient, so now very of these difficult or impossible to establish the identity spores. number of have However, during recent years a investigators paid attention to the fructifications well to the that are included as as spores in them. The most important communications on this subject are: Arnold (1930), Lepidostrobus (?), (1933, ’35), Lepidostrobus or Sigil- lariostrobus, (1938), Lepidostrobus (Lepidocarponi?), (1949), Lepidostrobus; Bochenski (1936) Lepidostrobus (Lepidocarpon!?), (1939), Sigillariostrobus; Ghaloner (1952), Lepidocarpon, (1953a) Lepidostrobus (?), (1953b), Lepidostrobus, (1953c), Sigillariostrobus, (1954), Selaginellites, Lepidostrobus (Lepidocarpon), (1956), Sporangiostrobus; Felix (1954), Lepidostrobus, Lepidocarpon; Hoskins and Cross (1940, ’52), Lepidostrobus; Leqlerq, Mathew (1938), Sigillariostrobus (Lepidostrobusr?) ; (1940), Lepidostrobus', Nemejg (1931), Sigillariostrobus (?); Schopf (1938), Lepidocarpon; Wicker Porostrobus. A detailed of the American (1934), very survey literature can be found in Felix (1954).
    [Show full text]
  • Alethopteris and Neuralethopteris from the Lower Westphalian
    Document generated on 10/01/2021 8:14 p.m. Atlantic Geology Journal of the Atlantic Geoscience Society Revue de la Société Géoscientifique de l'Atlantique Alethopteris and Neuralethopteris from the lower Westphalian (Middle Pennsylvanian) of Nova Scotia and New Brunswick, Maritime Provinces, Canada Carmen Álvarez-Vázquez Volume 56, 2020 Article abstract A systematic revision of Alethopteris and Neuralethopteris from upper URI: https://id.erudit.org/iderudit/1069888ar Namurian and lower Westphalian (Middle Pennsylvanian) strata of Nova DOI: https://doi.org/10.4138/atlgeol.2020.005 Scotia and New Brunswick, eastern Canada, has demonstrated the presence of eight species: Alethopteris bertrandii, Alethopteris decurrens, Alethopteris cf. See table of contents havlenae, Alethopteris urophylla, Alethopteris cf. valida, Neuralethopteris pocahontas, Neuralethopteris schlehanii and Neuralethopteris smithsii. Restudy of the Canadian material has led to new illustrations, observations and Publisher(s) refined descriptions of these species. Detailed synonymies focus on records from Canada and the United States. As with other groups reviewed in earlier Atlantic Geoscience Society articles in this series, it is clear that insufficient attention has been paid to material reposited in Canadian institutions in the European literature. The ISSN present study emphasizes the similarity of the North American flora with that of western Europe, especially through the synonymies. 0843-5561 (print) 1718-7885 (digital) Explore this journal Cite this article Álvarez-Vázquez, C. (2020). Alethopteris and Neuralethopteris from the lower Westphalian (Middle Pennsylvanian) of Nova Scotia and New Brunswick, Maritime Provinces, Canada. Atlantic Geology, 56, 111–145. https://doi.org/10.4138/atlgeol.2020.005 All Rights Reserved ©, 2020 Atlantic Geology This document is protected by copyright law.
    [Show full text]