Coleoptera: Cerambycidae: Cerambycinae) in the Fauna of Asia
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Russian Entomol. J. 28(3): 269–281 © RUSSIAN ENTOMOLOGICAL JOURNAL, 2019 The longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) in the fauna of Asia. 6. A new or little-known species of the genus Massicus Pascoe, 1867 Æóêè-äðîâîñåêè òðèáû Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) ôàóíû Àçèè. 6. Íîâûé è ìàëîèçâåñòíûå âèäû ðîäà Massicus Pascoe, 1867 Alexandr I. Miroshnikov1,2 À.È. Ìèðîøíèêîâ1,2 1 Russian Entomological Society, Krasnodar, Russia. E-mail: [email protected] 2 Sochi National Park, Moskovskaya str., 21, Sochi 354002, Krasnodar region, Russia 1 Русское энтомологическое общество, Краснодар, Россия 2 Сочинский национальный парк, ул. Московская, 21, Сочи 354000, Краснодарский край, Россия KEY WORDS: Coleoptera, Cerambycidae, Cerambycini, Massicus, new or little-known species, Peninsular Malaysia, Borneo. КЛЮЧЕВЫЕ СЛОВА: Coleoptera, Cerambycidae, Cerambycini, Massicus, новый и малоизвестные виды, полуостров Малакка, Борнео. ABSTRACT. A new species, Massicus regius sp.n., The present paper describes a new species that has that has been mixed with M. pascoei (J. Thomson, been mixed with the type species of the genus, M. 1857) for a long time, is described from Peninsular pascoei (J. Thomson, 1857) for a long time. Addittion- Malaysia and Borneo. Massicus fryi Gahan, 1890 and ally, it provides new records of several little-known M. ivani Miroshnikov, 2017 are recorded from Western species expanding or clarifying their distribution areas Malaysia, as generally from Indochina, for the first to some extent, as well as presents other new data. time. The data on the distribution of M. pascoei are The material treated in this work belongs to the follow- discussed. New records of M. intricatus (Pascoe, 1866) ing institutional and private collections: BMNH — and M. valentinae Miroshnikov, 2017 from Western Natural History Museum (London, United Kingdom); Malaysia, and M. punctulipennis Holzschuh, 2018 from IRSN — Institut Royal de Sciences naturelles de Bel- southern Vietnam are presented. The lectotype of M. gique (Bruxelles); MNHN — Muséum national d’Histoire intricatus is designated. naturelle (Paris, France); NHMD — Natural History Museum of Denmark, University of Copenhagen (Copen- РЕЗЮМЕ. Описан новый вид Massicus regius hagen, Denmark); cAM — collection of Alexandr Mirosh- sp.n. с Малайского полуострова и Борнео, который nikov (Krasnodar, Russia); cCH — collection of Carolus долгое время смешивался с M. pascoei (J. Thomson, Holzschuh (Villach, Austria); cDH — collection of Daniel 1857). Massicus fryi Gahan, 1890 и M. ivani Mirosh- Heffern (Houston, USA); cLD — collection of Luboš nikov, 2017 впервые отмечены в Западной Малайзии, Dembický (Brno, Czech Republic). как и в целом в Индокитае. Рассмотрены данные о Genus Massicus Pascoe, 1867 распространении M. pascoei. Приведены новые находки M. intricatus (Pascoe, 1866) и M. valentinae Massicus Pascoe, 1867: 319 [name replacement for Conotho- rax J. Thomson, 1864, non Jeckel, 1854 (Curculionidae)]. Lacor- Miroshnikov, 2017 в Западной Малайзии, а M. daire, 1868: 260; Gemminger, 1872: 2802; Gahan, 1906: 129; punctulipennis Holzschuh, 2018 — в южном Вьетнаме. Aurivillius, 1912: 55; Gressitt, Rondon, 1970: 59; Lee, 1982: 27; Обозначен лектотип M. intricatus. Kusama, Takakuwa, 1984: 252; Hüdepohl, 1990: 60; Catalogue..., 2010: 161; Heffern, 2013: 10; Nga et al., 2014: 435; Kariyanna et al., 2017: 31; Miroshnikov, 2017: 224. Until now, many groups of the tribe Cerambycini Mallambyx Bates, 1873: 152 (Pachydissus subgen.). remain poorly studied in various aspects. The genus Mallambyx: Ganglbauer, 1889: 473; Winkler, 1929: 1141; Ma- Massicus Pascoe, 1867 is no exception in this regard. tsushita, 1933: 243; Mitono, 1940: 82; Plavilstshikov, 1940: 79, 635; How to cite this article: Miroshnikov A.I. 2019. The longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) in the fauna of Asia. 6. A new or little-known species of the genus Massicus Pascoe, 1867 // Russian Entomol. J. Vol.28. No.3. P.269–281. doi: 10.15298/rusentj.28.3.06 270 A.I. Miroshnikov Gressitt, 1951: 135; Kojima, Hayashi, 1969: 47; Tsherepanov, 1981: 22. Thomson”, “Ex F.M.S. Museum B.M. 1955–354”, “NHMUK Type species: Cerambyx pascoei J. Thomson, 1857, by monotypy. 013386253”; 1# (BMNH), “Malay Penin: Perak F.M.S., ex coll. Perak Mus., C. Warang, 19[?]”, “Ex F.M.S. Museum B.M. 1955– Massicus pascoei (J. Thomson, 1857) 354”, “NHMUK 013386280”; 1# (IRSN), “Coll. Nonfried. Borneo”. Figs 1–2, 4–6, 11, 13–14, 17, 20, DIAGNOSIS. This new species is very similar to M. 23, 27, 31, 33, 37, 41, 45, 48. pascoei, but differs clearly by the less convex eyes, more Cerambyx pascoei Thomson, 1857: 122, pl. 14, fig. 6. Type weakly developed at least on the ventral side of the head, as in locality: “Inde” (according to the original description) or “India or.” Figs 15–16, 18–19 (cf. Figs 14, 17); the length ratio of (according to the label of the holotype) (see also Remarks below). isthmus between the lower lobes of the eyes and the trans- Conothorax pascoei: Thomson, 1864: 230 (“Malasia”). verse diameter of the lower eye lobe, as in Figs 15–16, 18–19 Massicus [pascoei]: Pascoe, 1867: 319. (cf. Figs 14, 17); the obliterated or at least distinctly less Massicus pascoei: Lacordaire, 1868: 260 (“Malasie”); Gem- coarse sculpture of the submentum, but clearer dense punctu- minger, 1872: 2802 (“Archip. ind.”); Gahan, 1906: 129 (“Malay ration, as in Figs 15–16, 18–19 (cf. Figs 14, 17); the shorter Peninsula”); Aurivillius, 1912: 55 (“Malacca”); Heffern, 2013: 10 median groove on the vertex, as in Figs 7–9, 32 (cf. Figs 1–2, (“E. India; W Malaysia; SE Asia [probably within the limits of Indochina]”); Kariyanna et al., 2017: 31 (“India & Malaysia”). 5, 31); the ratio width of the pronotum at base and at apex in MATERIAL. Holotype by monotypy, $ (MNHN) (Figs 5–6), the male, as in Figs 7–8 (cf. Fig. 1); the presence of rough, “India or.”, “Ex Musæo James Thomson”, “444”, “Conothorax distinct, sparse punctures (besides a small dense punctura- Thoms. s.c.230. (Cône, Thorax.), Massicus Pasc. An. M. n. h. XIX, tion) on several basal antennomeres of the male, starting from 319.”, “Pascoei Thoms. Type, Malas. Ind.?”, “Th. / Type” (Fig. 4); antennomere 3; on the average a larger body; the structure of 2# (cAM) (Fig. 1), Laos, Houa Phan Prov., Phou Pane Mt., 20°11'N the genitalia, mainly in the male. / 104°00'E, 2060 m, 06.2017 (local collector); 2$ (cAM) (Fig. 2), Additionally, in Massicus regius sp.n., compared to M. same locality, but taken on 05.2017; 1$ (BMNH) (Fig. 3), “Malac- pascoei, the pronotum is usually less convex, with two more ca”, “Ex Mus Parry”, “Fry Coll. 1905.100.”, “43 … [illegible further or less well-developed spots of dense, light setae (besides on]”, “NHMUK 013386288” (see Remarks below). numerous sparse setae) on its sides, as in Fig. 10 (cf. Fig. 11); REMARKS. Apparently this species was described from the prosternal process at the very apex and dorsally is usually a single female originating from India [Thomson, 1857: 122: narrower and, accordingly, the apical tubercles are more “Patrie: Inde”]. However, in the subsequent Thomson’s work closely spaced, as in Figs 24–26, 28–30 (cf. Figs 6, 23, 27); [1864: 230] it was given as from “Malasia” without any the mesosternal process in the vast majority of cases is comments. On one labels of the holotype is indicated “India narrower, as in Figs 24–25, 28–30 (cf. Figs 6, 23, 27); the or.”, while on the other — “… Malas. Ind.?”. antennal tubercles are usually stronger, often transverse, at I have studied two males and two females from northeas- least so in the male, as in Fig. 12 (cf. Fig. 13); the sculpture of tern Laos that are very similar to the holotype and that I male antennomere 1 is usually coarser, as in Fig. 32 (cf. Fig. identify here as M. pascoei. Some specimens from China (Yunnan) and northern Myanmar, known to me only from 31); the constriction behind the temples in the male is less their photographs belong to this species too. sharp, as a rule, as in Figs 15–16 (cf. Fig. 14); the apical At the same time, almost all specimens I have revised external angle of the elytra is often more obtuse or even from Western Malaysia, externally similar to M. pascoei, are nearly rounded; the recumbent light setation of the submen- actually a new species described below. tum is usually more strongly developed, as in Figs 15–16, Only one female (Fig. 3) from the old collections at the 18–19 (cf. Figs 14, 17); and the antennae of the female are BMNH with the “Malacca” label almost certainly belongs to usually slightly shorter. M. pascoei. However, its geographical label is possibly errone- The new species can also be compared to M. ivani and M. ous, and therefore it cannot be considered indisputable evi- valentinae, but is distinguished very distinctly from both, dence of the distribution of M. pascoei in Western Malaysia. partly like from M. pascoei, in the completely different shape MORPHOLOGICAL NOTES. Body length 52.2–58.8 of the prosternal process at the apex and dorsally, as in Figs mm, humeral width 14.5–15.7 mm, thereby the holotype is 24–26, 28–30 (cf. Figs 52–54); the wider submentum, its 57.3 mm and 15.7 mm, respectively. obliterated or at least clearly less coarse sculpture; the less Judging from some photographs, the body length of this convex pronotum, its somewhat peculiar sculpture and usual- species can reach about 61 mm. ly also shape; the usually stronger antennal tubercles, often DISTRIBUTION. Eastern India, China (Yunnan), My- transverse, at least so in the male; the more strongly protrud- anmar, Laos; almost without any doubt, Vietnam and Thai- ing apical external angle of antennomere 1, the coarser sculp- land; ?Western Malaysia.