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BULLETINS of AMERICAN PALEONTOLOGY and PALAEONTOGRAPHICA AMERICANA

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BULLETINS of AMERICAN PALEONTOLOGY and PALAEONTOGRAPHICA AMERICANA 6UL 17/t' HARVARD UNIVERSITY Library of the Museum of Comparative Zoology 3u(Qtmscf towqs) VOLUME 81, 314 NUMBER MAY 6, 1982 Latest Ordovician Solitary Rugose Corals of Eastern North America by Robert J. Elias PRI Paleontological Research Institution anniversary 1259 Trumansburg Road year Ithaca, New York, 14850 U.S.A. Library of Congress Card Number: SI-SI')!! Printed in the United States of America Allen Press, Inc. Lawrence, KS 66044 U.S.A. CONTENTS Abstract 5 Uppermost Ordovician of Oklahoma, Missouri and Illinois Introduction 5 Introduction 36 Abbreviations of Repositories 6 Arbuckle Mountains, Oklahoma 39 Acknowledgments 6 Cape Girardeau County. Missouri 39 Latest Ordovician Paleogeography. Lithofacies, and Solitary Pike County. Missouri 40 Rugose Corals of Eastern North America Will County, Illinois 40 Overview 8 Continental margin Richmond Group Introduction 40 Introduction 8 Penobscot County, Maine 41 Cincinnati Arch region Ashland, Maine 41 Stratigraphy 9 Perce, Quebec 41 Depositional environments 11 Anticosti Island. Quebec Cincinnati Arch 12 Stratigraphy 42 Solitary rugose corals Solitary rugose corals 45 Grewinf,'kici canadensis (Billings. 1862) Paleobiogeography of Latest Ordovician Solitary Rugose Corals Stratigraphic distribution 13 in Eastern North America Orientation of corals 13 Overview 47 Coral size 14 Red River-Stony Mountain Province Epizoans 17 Introduction 48 Borings 17 Maquoketa Subprovince 48 Intraspecific variation 19 Maritime Subprovince 48 Streptetasma divaricans (Nicholson, 1875b) Richmond Province Stratigraphic distribution 22 Introduction 49 Orientation of corals 23 Origin 50 Coral size 25 Subsequent history 51 Epizoans 25 Edgewood Province 51 Borings 25 Systematic Paleontology Intraspecific variation 25 Introduction 52 Summary 26 Family Streptelasmatidae Nicholson (in Nicholson and Ly- Burkesville, Kentucky 27 dekker. 1889) Goodlettsville-Gallatin area, Tennessee 27 Genus Sliepletasma Hall (1847) 52 Little Sturgeon Bay, Wisconsin 29 Genus Helicelasma B. Neuman ( 1969) 60 Little Bay de Noc, Michigan 29 Genus Deiiacoralliiim Nelson ( 1963) 63 Drummond Island, Michigan 30 Genus Grewinf;kia Dybowski (1873) 65 Manitoulin Island, Ontario 32 Genus LobocoralHiim Nelson ( 1963) 74 Meaford, Ontario 33 Genus Kenophylhim Dybowski ( 1873) 76 Streetsville. Ontario 34 Genus Bodophylluin B. Neuman ( 1969) 77 Montreal, Quebec 34 Genus Bif;lwinia Duncan (1957) 79 Lake St. John, Quebec 34 Family Paliphyllidae Soshkina (1955) Maquoketa Group Genus Paliphyllum Soshkina ( 1955) 82 Introduction 35 Appendix; Collecting Localities 83 Northeastern Iowa 35 References Cited 86 Southeastern Iowa and northwestern Illinois 36 Plates 93 Thebes, Illinois 36 Index 109 . .. LIST OF ILLUSTRATIONS 22. Upper Ordovician stratigraphic sections and distribution of solitary rugose corals in Maine and southeastern Quebec 43 23. Length of specimens of Grewingkia pidchella (Billings. Text -figure Page 1865) from the Vaureal and Ellis Bay Formations. 1. Time-stratigraphic subdivisions of the North American and Quebec 46 European Upper Ordovician including informal subdivi- 24. Paleobiogeography of latest Ordovician solitary rugose cor- sions of the Richmondian 6 als in eastern North America 47 2. Latest Ordovician paleogeography and lithofacies in east- 25. Relation between number of major septa and coral diameter ern North America 7 in Slreptelasma divaricans (Nicholson, 1875b) from the 3. Richmondian stratigraphic sections and distribution of sol- entire Richmond Group, Cincinnati Arch region 54 itary rugose corals in Ohio. Indiana, Kentucky and 26. Blastogeny in .S7repjt>/(ji7?i(j <//i(jr/((j«i (Nicholson, 1875b) 55 Tennessee foldout inside front cover 27. Relation between number of major septa and coral diameter 4. Middle Richmondian paleogeography and lithofacies in the in Slreptelasma suhregiilare (Savage. 1913) 58 greater Cincinnati Arch region 13 28. Relation between number of major septa and coral diameter 5. Directional orientation of specimens of Grewiiiiikiti ciimi- in Slreptelasma rankini n. sp 58 denxis (Billings, 1862) in the Richmond Group 14 29. Relation between number of major septa and coral diameter 6. Length of specimens o{ Greningkiu canadensis (Billings, in Slreptelasma affine (Billings. 1865) 60 1 862) from the Richmond Group 15 30. Relation between number of major septa and coral diameter 7. Length of specimens of Grewingkia canadensis (Billings, in Helicelasma randi Elias ( 1981) 62 1862) from Richmond Group sections 16 3 1 Relation between number of major septa and coral diameter 8. Diameter of specimens of Giewingkia canadensis (Billings, in Helicelasma selectum (Billings, 1865) 63 1862) from Richmond Group sections 17 32. Relation between number of major septa and coral height, 9. Diameter of borings assigned to Tnpanites weisei Magde- and coral form in species of Deiracoralliiim 65 frau (1932) in Ordovician solitary rugose corals, brachio- 33. Relation between number of major septa and coral diameter pods, and bryozoans 18 in Grewingkia canadensis (Billings, 1862) from the entire 10. Relation between number of major septa and coral diameter Richmond Group, Cincinnati Arch region 68 in Grewingkia canadensis (Billings, 1862) from the Rich- 34. Relation between number of major septa and coral diameter mond Group 20 in Grewingkia deltensis n. sp 70 1 1 Relation between number of major septa and coral diameter 35. Relation between number of major septa and coral diameter in Grewingkia canadensis (Billings, 1862) from the Rich- in Grewingkia ruslica (Billings, 1858a) 71 mond Group at section I , Cincinnati Arch region 21 36. Relation between number of major septa and coral diameter 12. Percent frequency of values on the a.xial region compara- in Grewingkia penohscotensis n. sp 72 tive scale for specimens of Grewingkia canadensis (Bil- 37. Relation between number of major septa and coral diameter lings, 1862) from the Richmond Group, Cincinnati Arch in Grewingkia pidchella (Billings, 1865) 74 region 22 38. Relation between number of major septa and coral cross- 13. Percent frequency distribution showing number of coral- sectional area in Lobocorallium trilohatum trilobalum lites per corallum in Streptelasnia divaricans (Nicholson, (Whiteaves, 1895), L. trilobalam vaurealense (Twenhofel, 1875b) from the Richmond Group, Cincinnati Arch region 23 1928), and Grewingkia huysii (Meek, 1865) 76 14. Length of specimens of Slreptelasma divaricans (Nichol- 39. Relation between number of major septa and coral diameter son. 1875b) from the Richmond Group, Cincinnati Arch in Bodophylhim neiimani n. sp 78 region 23 40. Relation between number of major septa and coral diameter 15. Relation between number of major septa and coral diameter in Bodophylhim englishheadense n. sp 79 in Slreptelasma divaricans (Nicholson. 1875b) from the 41. Relation between number of major septa and coral height Richmond Group. Cincinnati Arch region 24 in Bighornia patella (A. E. Wilson, 1926) and B. cf B. 16. Percent frequency of values on the axial region compara- patella 81 tive scale for specimens of Slreptelasma divaricans (Ni- 42 Relation between number of major septa and coral diameter cholson, 1875b) from the Richmond Group, Cincinnati in Paliphylliim ellisense (Twenhofel, 1928) 83 Arch region 25 1 7. Relation between number of major septa and coral diameter LIST OF TABLES in Grewingkia canadensis (Billings. 1862) from the Rich- Table Page mond Group, Tennessee 28 18. Richmondian stratigraphic sections and distribution of sol- 1 Attachment sites of Slreptelasma divaricans (Nicholson, itary rugose corals in Illinois, Iowa, Wisconsin, Michigan. 1875b) on brachiopods from the Richmond Group, Cincin- 23 Ontario and southwestern Quebec . foldoul inside back cover nati Arch region 19. Relation between number of major septa and coral diameter Frequency of values on the axial region comparative scale in Grewingkia canadensis (Billings, 1862) from the Geor- for specimens of Grewingkia canadensis (Billings, 1862) gian Bay Formation. Michigan 31 from the Georgian Bay Formation, Michigan and Ontario 31 20. Relation between number of major septa and coral diameter Latest Ordovician solitary rugose corals in paleobiogeo- in Grewingkia canadensis (Billings. 1862) from the Geor- graphic provinces and subprovinces, eastern North gian Bay Formation, Ontario 32 America 48 21. Uppermost Ordovician stratigraphic sections and distribu- Solitary rugose coral genera in the Ashgill of Scandinavia 50 tion of solitary rugose corals in Oklahoma, Missouri and Stratigraphic position of collected intervals within the Rich- Illinois 37 mond Group, Cincinnati Arch region 83 ;;z^ 3u(Qtmsof -American yakcmtWloqy VOLUME 81, NUMBER 314 MAY 6, 1982 MU8. COMP. ZOOU i--^— I r ray JUN 1 1982 _.give:RSi T-v Latest Ordovician Solitary Rugose Corals of Eastern North America by Robert J. Elias PRI anmversarv year PALEONTOLOGICAL RESEARCH INSTITUTION Officers President John Pojeta, Jr. Vice-President Bruce M. Bell Secretary Philip C. Wakeley Acting Treasurer Robert E. Terwillegar Assistant Treasurer John L. Cisne Director Peter R. Hoover Director Emeritus Katherine V. W. Palmer Legal Counsel Armand L. Adams Trustees Bruce M. Bell (to 6/30/84) Duane O. LeRoy (to 6/30/84) Richard E. Byrd (to 6/30/83) William A. Oliver, Jr. (to 6/30/83) Kenneth E. Caster (to 6/30/82) Katherine V. W. Palmer (Life) John L. Cisne (to 6/30/82) John Pojeta, Jr. (to 6/30/82) Lee B. Gibson (to 6/30/83) Raymond Van Houtte (to 6/30/82) Rebecca S. Harris (Life) William P. S. Ventress (to 6/30/84) Porter Kier (to
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