Corrections in Recently Described Species of Ferns and Lycophytes from the Neotropics
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Insights on Reticulate Evolution in Ferns, with Special Emphasis on the Genus Ceratopteris
Utah State University DigitalCommons@USU All Graduate Theses and Dissertations Graduate Studies 8-2021 Insights on Reticulate Evolution in Ferns, with Special Emphasis on the Genus Ceratopteris Sylvia P. Kinosian Utah State University Follow this and additional works at: https://digitalcommons.usu.edu/etd Part of the Ecology and Evolutionary Biology Commons Recommended Citation Kinosian, Sylvia P., "Insights on Reticulate Evolution in Ferns, with Special Emphasis on the Genus Ceratopteris" (2021). All Graduate Theses and Dissertations. 8159. https://digitalcommons.usu.edu/etd/8159 This Dissertation is brought to you for free and open access by the Graduate Studies at DigitalCommons@USU. It has been accepted for inclusion in All Graduate Theses and Dissertations by an authorized administrator of DigitalCommons@USU. For more information, please contact [email protected]. INSIGHTS ON RETICULATE EVOLUTION IN FERNS, WITH SPECIAL EMPHASIS ON THE GENUS CERATOPTERIS by Sylvia P. Kinosian A dissertation submitted in partial fulfillment of the requirements for the degree of DOCTOR OF PHILOSOPHY in Ecology Approved: Zachariah Gompert, Ph.D. Paul G. Wolf, Ph.D. Major Professor Committee Member William D. Pearse, Ph.D. Karen Mock, Ph.D Committee Member Committee Member Karen Kaphiem, Ph.D Michael Sundue, Ph.D. Committee Member Committee Member D. Richard Cutler, Ph.D. Interim Vice Provost of Graduate Studies UTAH STATE UNIVERSITY Logan, Utah 2021 ii Copyright © Sylvia P. Kinosian 2021 All Rights Reserved iii ABSTRACT Insights on reticulate evolution in ferns, with special emphasis on the genus Ceratopteris by Sylvia P. Kinosian, Doctor of Philosophy Utah State University, 2021 Major Professor: Zachariah Gompert, Ph.D. -
Fern Classification
16 Fern classification ALAN R. SMITH, KATHLEEN M. PRYER, ERIC SCHUETTPELZ, PETRA KORALL, HARALD SCHNEIDER, AND PAUL G. WOLF 16.1 Introduction and historical summary / Over the past 70 years, many fern classifications, nearly all based on morphology, most explicitly or implicitly phylogenetic, have been proposed. The most complete and commonly used classifications, some intended primar• ily as herbarium (filing) schemes, are summarized in Table 16.1, and include: Christensen (1938), Copeland (1947), Holttum (1947, 1949), Nayar (1970), Bierhorst (1971), Crabbe et al. (1975), Pichi Sermolli (1977), Ching (1978), Tryon and Tryon (1982), Kramer (in Kubitzki, 1990), Hennipman (1996), and Stevenson and Loconte (1996). Other classifications or trees implying relationships, some with a regional focus, include Bower (1926), Ching (1940), Dickason (1946), Wagner (1969), Tagawa and Iwatsuki (1972), Holttum (1973), and Mickel (1974). Tryon (1952) and Pichi Sermolli (1973) reviewed and reproduced many of these and still earlier classifica• tions, and Pichi Sermolli (1970, 1981, 1982, 1986) also summarized information on family names of ferns. Smith (1996) provided a summary and discussion of recent classifications. With the advent of cladistic methods and molecular sequencing techniques, there has been an increased interest in classifications reflecting evolutionary relationships. Phylogenetic studies robustly support a basal dichotomy within vascular plants, separating the lycophytes (less than 1 % of extant vascular plants) from the euphyllophytes (Figure 16.l; Raubeson and Jansen, 1992, Kenrick and Crane, 1997; Pryer et al., 2001a, 2004a, 2004b; Qiu et al., 2006). Living euphyl• lophytes, in turn, comprise two major clades: spermatophytes (seed plants), which are in excess of 260 000 species (Thorne, 2002; Scotland and Wortley, Biology and Evolution of Ferns and Lycopliytes, ed. -
Diretrizes Para Auxílio Na Confecção De
Aline Possamai Della Revisão taxonômica de Jamesonia e Tryonia (Pteridaceae) ocorrentes no Brasil Taxonomic review of Jamesonia and Tryonia (Pteridaceae) occurring in Brazil São Paulo 2019 2 Aline Possamai Della Revisão taxonômica de Jamesonia e Tryonia (Pteridaceae) ocorrentes no Brasil Taxonomic review of Jamesonia and Tryonia (Pteridaceae) occurring in Brazil Dissertação apresentada ao Instituto de Biociências da Universidade de São Paulo, para a obtenção de Título de Mestre em Ciências Biológicas, na Área de Botânica. Orientador: Dr. Jefferson Prado São Paulo 2019 3 Ficha Catalográfica Della, Aline Possamai Revisão Taxonômica de Jamesonia e Tryonia (Pteridaceae) ocorrentes no Brasil / Aline Possamai Della; orientador Jefferson Prado. -- São Paulo, 2019. 121 f. Dissertação (Mestrado) - Instituto de Biociências da Universidade de São Paulo, Departamento de Botânica. 1. Eriosorus. 2. Flora. 3. Mata Atlântica. 4. Pteridoidea. 5. Samambaias. Comissão Julgadora: ________________________ Prof(a). Dr(a). ________________________ Prof(a). Dr(a). _____________________ Prof. Dr. Jefferson Prado Orientador 4 Dedicatória Dedico este trabalho aos meus pais, Rui e Albertina 5 Epígrafe “Ninguém ignora tudo. Ninguém sabe tudo. Todos nós sabemos alguma coisa. Todos nós ignoramos alguma coisa. Por isso aprendemos sempre”. Paulo Freire 6 Agradecimentos Gostaria de registrar aqui os meus sinceros agradecimentos a todas as pessoas que estiveram envolvidas direta ou indiretamente no desenvolvimento deste trabalho. Ao meu orientador, Prof. Jefferson Prado, pela confiança depositada, desde o momento inicial, sem mesmo nos conhecermos direito. Pela acolhida no Instituto de Botânica de SP durante esses dois anos, pelos ensinamentos sobre o mundo científico, pelas discussões sobre plantas e pelos conselhos tanto profissionais, quanto pessoais, que me fizeram enxergar muitas coisas de forma diferente. -
Lycopodiaceae) Weston Testo University of Vermont
University of Vermont ScholarWorks @ UVM Graduate College Dissertations and Theses Dissertations and Theses 2018 Devonian origin and Cenozoic radiation in the clubmosses (Lycopodiaceae) Weston Testo University of Vermont Follow this and additional works at: https://scholarworks.uvm.edu/graddis Part of the Systems Biology Commons Recommended Citation Testo, Weston, "Devonian origin and Cenozoic radiation in the clubmosses (Lycopodiaceae)" (2018). Graduate College Dissertations and Theses. 838. https://scholarworks.uvm.edu/graddis/838 This Dissertation is brought to you for free and open access by the Dissertations and Theses at ScholarWorks @ UVM. It has been accepted for inclusion in Graduate College Dissertations and Theses by an authorized administrator of ScholarWorks @ UVM. For more information, please contact [email protected]. DEVONIAN ORIGIN AND CENOZOIC RADIATION IN THE CLUBMOSSES (LYCOPODIACEAE) A Dissertation Presented by Weston Testo to The Faculty of the Graduate College of The University of Vermont In Partial Fulfillment of the Requirements for the Degree of Doctor of Philosophy Specializing in Plant Biology January, 2018 Defense Date: November 13, 2017 Dissertation Examination Committee: David S. Barrington, Ph.D., Advisor Ingi Agnarsson, Ph.D., Chairperson Jill Preston, Ph.D. Cathy Paris, Ph.D. Cynthia J. Forehand, Ph.D., Dean of the Graduate College ABSTRACT Together with the heterosporous lycophytes, the clubmoss family (Lycopodiaceae) is the sister lineage to all other vascular land plants. Given the family’s important position in the land-plant phylogeny, studying the evolutionary history of this group is an important step towards a better understanding of plant evolution. Despite this, little is known about the Lycopodiaceae, and a well-sampled, robust phylogeny of the group is lacking. -
Spore Morphology of Taenitis, Syngramma and Austrogramme Species (Pteridoideae, Pteridaceae) from South-Eastern Asia
Turczaninowia 21 (3): 5–11 (2018) ISSN 1560–7259 (print edition) DOI: 10.14258/turczaninowia.21.3.1 TURCZANINOWIA http://turczaninowia.asu.ru ISSN 1560–7267 (online edition) УДК 582.394.7:581.4(59) Spore morphology of Taenitis, Syngramma and Austrogramme species (Pteridoideae, Pteridaceae) from South-Eastern Asia A. V. Vaganov1, I. I. Gureyeva2, A. I. Shmakov1, A. A. Kuznetsov2, R. S. Romanets2 1 South-Siberian Botanical Garden, Altai State University, pr. Lenina, 61, Barnaul, 656049, Russia. E-mail: [email protected] 2 Tomsk State University, pr. Lenina, 36, Tomsk, 634050, Russia. E-mail: [email protected] Keywords: Austrogramme, Pteridaceae, Pteridoideae, scanning electronic microscopy (SEM), spore morphology, Syngramma, Taenitis. Summary. A comparative study of spores of seven species: Taenitis blechnoides (Willd.) Sw., T. hookeri (C. Chr.) Holttum, T. pinnata (J. Sm.) Holttum, Syngramma alismifolia (Presl) J. Sm., S. lobbiana (Hook.) J. Sm., S. quinata (Hook.) Carruth., and Austrogramme boerlageana (Alderw.) Hennipman from South-Eastern Asia was performed us- ing the method of scanning electronic microscopy (SEM). Spores of all examined species are trilete, tetrahedral or tet- rahedral-globose. Spores of Syngramma and Austrogramme are very similar to each other in shape and ornamentation. Ornamentation of both sides of spore is similar, verrucate (microverrucate), surface covered by rodlets and granulate deposits. Spores of Taenitis species with conspicuous cingulum (Taenitis blechnoides) or without it, ornamentation of both faces of spore could be tuberculate or baculate. Spores of Taenitis hookeri and Taenitis pinnata demonstrate tendency to forming of comissural ridges. Spore size of all studied species is close: equatorial diameter of spores of Taenitis species varies within 24–43 μm, those for Syngramma species is 29.8–35.5 μm, spores of Austrogramme boerlageana are smallest, their equatorial diameter varies within 22.5–29.4 μm. -
The Community Ecology, Dynamics and Productivity of Tropical Grasslands in the Andes
The Pdramo Vegetation of Ecuador: the Community Ecology, Dynamics and Productivity of Tropical Grasslands in the Andes. by Paul Michael Ramsay A thesis submitted for the degree of Philosophiae Doctor of the University of Wales. December 1992 School of Biological Sciences, University of Wales, Bangor, Gwynedd, LL57 2UW. i Dedicated to the memory of Jack Higgins, my grandfather. "... a naturalist's life would be a happy one if he had only to observe and never to write." Charles Darwin ii Table of Contents Preface AcknoWledgements vii Summary ix Resumen Chapter 1. Introduction to the Ecuadorian P6ramos 1 Ecuador 2 The Pâramos of the Andes 2 Geology and Edaphology of the Paramos 6 Climate 8 Flora 11 Fauna 14 The Influence of Man 14 Chapter 2. The Community Ecology of the Ecuadorian P6ramos 17 Introduction 18 Methods 20 Results 36 The Zonal Vegetation of the Ecuadorian Paramos 51 Discussion 64 Chapter 3. Plant Form in the Ecuadorian Paramos 77 Section I. A Growth Form Classification for the Ecuadorian Paramos 78 Section II. The Growth Form Composition of the Ecuadorian Pâramos Introduction 94 Methods 95 Results 97 Discussion 107 Section III. Temperature Characteristics of Major Growth Forms in the Ecuadorian PSramos Introductio n 112 Methods 113 Results 118 Discussion 123 III Table of Contents iv Chapter 4. Aspects of Plant Community Dynamics in the Ecuadorian Pgramos 131 Introduction 132 Methods 133 Results 140 Discussion 158 Chapter 5. An Assessment of Net Aboveground Primary Productivity in the Andean Grasslands of Central Ecuador 165 Introduction 166 Methods 169 Results 177 Discussion 189 Chapter 6. -
A Revised Generic Classification of Vittarioid Ferns (Pteridaceae)
Schuettpelz & al. • Revised classification of vittarioid ferns TAXON 65 (4) • August 2016: 708–722 A revised generic classification of vittarioid ferns (Pteridaceae) based on molecular, micromorphological, and geographic data Eric Schuettpelz,1 Cheng-Wei Chen,2 Michael Kessler,3 Jerald B. Pinson,4 Gabriel Johnson,1 Alex Davila,5 Alyssa T. Cochran,6 Layne Huiet7 & Kathleen M. Pryer7 1 Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington, D.C. 20013, U.S.A. 2 Division of Botanical Garden, Taiwan Forestry Research Institute, Taipei 10066, Taiwan 3 Institute of Systematic Botany, University of Zurich, 8008 Zurich, Switzerland 4 Department of Biology, University of Florida, Gainesville, Florida 32611, U.S.A. 5 Department of Biology and Marine Biology, University of North Carolina Wilmington, Wilmington, North Carolina 28403, U.S.A. 6 Department of Biology, Colorado State University, Fort Collins, Colorado 80523, U.S.A. 7 Department of Biology, Duke University, Durham, North Carolina 27708, U.S.A. Author for correspondence: Eric Schuettpelz, [email protected] ORICD ES, http://orcid.org/0000-0003-3891-9904 DOI http://dx.doi.org/10.12705/654.2 Abstract Vittarioid ferns compose a well-supported clade of 100–130 species of highly simplified epiphytes in the family Pterid- aceae. Generic circumscriptions within the vittarioid clade were among the first in ferns to be evaluated and revised based on molecular phylogenetic data. Initial analyses of rbcL sequences revealed strong geographic structure and demonstrated that the two largest vittarioid genera, as then defined, each had phylogenetically distinct American and Old World components. The results of subsequent studies that included as many as 36 individuals of 33 species, but still relied on a single gene, were generally consistent with the early findings. -
Exploring the Physiology and Evolution of Hornworts
Utah State University DigitalCommons@USU All Graduate Theses and Dissertations Graduate Studies 12-2019 Exploring the Physiology and Evolution of Hornworts Tanner A. Robison Utah State University Follow this and additional works at: https://digitalcommons.usu.edu/etd Part of the Biology Commons Recommended Citation Robison, Tanner A., "Exploring the Physiology and Evolution of Hornworts" (2019). All Graduate Theses and Dissertations. 7668. https://digitalcommons.usu.edu/etd/7668 This Thesis is brought to you for free and open access by the Graduate Studies at DigitalCommons@USU. It has been accepted for inclusion in All Graduate Theses and Dissertations by an authorized administrator of DigitalCommons@USU. For more information, please contact [email protected]. EXPLORING THE PHYSIOLOGY AND EVOLUTION OF HORNWORTS by Tanner A. Robison A thesis submitted in partial fulfillment of the requirements for the degree of MASTER OF SCIENCE in Biology Approved: ______________________ ____________________ Paul Wolf, Ph.D. Keith Mott, Ph.D. MaJor Professor Committee Member ______________________ ____________________ Bruce Bugbee, Ph.D. Richard S. Inouye, Ph.D. Committee Member Vice Provost for Graduate Studies UTAH STATE UNIVERSITY Logan, Utah 2019 ii ABSTRACT Exploring the Evolution and Physiology of Hornworts by Tanner Robison, Master of Science Utah State University, 2019 Major Professor: Dr. Paul G. Wolf Department: Biology To improve the ease, accuracy, and speed of organellar genome annotation for hornworts and ferns, we developed a bioinformatic tool which takes into account the process of RNA editing when examining annotations. This software works by checking the coding sequences of annotated genes for internal stop codons and for overlooked start codons which might be the result of RNA editing. -
Fern Gazette
THE FERN GAZETTE VOLUME ELEVEN PARTS TWO AND THREE 1975 THE JOURNAL OF THE BRITISHPTERIDOLOGICAL SOCIETY FERN GAZETTE VOLUME l1 PAR1S2 & 3 1975 CONTENTS Page A re-defintion of the Gymnogrammoid genus Austrogramme Fournier - E. Hennipman 61 The biogeography of endemism in the Cyatheaceae- R. Tryon and G Gastony 73 wnathyrium in the Azores- W.A. Sledge 81 The gametophyte of Chingia pseudoferox- Lennette R. Atkinson 87 Ta xonomic notes on some African species of Elaphoglossum - R.E. G Pichi Sermolli 95 Observations on the spread of the American fern Pityrogramma calomelanos -E.A. CL. E. ::Che/pe 101 A phytogeographic analysis of Choc6 Pteridophytes- D. B. Le/linger 105 Studies in the systematics of filmy ferns: I. A note on the identity of Microtrichomanes - K. Jwatsuki 115 A hybrid polypody from the New World tropics - W.H. Wagner and Florence Wagner 125 Aspidistes thomasii- a jurassic member of the Thelypteridaceae- }.£?. Lovis 137 A new arrangement for the pteridophyte herbarium - j.A. Crabbe, A.C jermy and }.M. Mickel 141 A note on the distribution of lsoetes in the Cadiz Province, Spain - Betty Moles worth A lien 163 Lecanopteris spinosa; a new ant-fern from Indonesia - A.C jermy and T. G Walker 165 Dryopteris tyrrhena nom. nov. - a misunderstood western Mediterranean species - C.R. Fraser jenkins and T. Reichstein 177 THE BRITISH FERN GAZETTE Volume 11 Part 1 was published 6 February 1975 Published by THE BRITISH PTERIDOLOGICAL SOCIETY, c /o Department of Botany, Museum {Natural History} . London SW7 5BD. \ )�eA·r""-�\t". 197�· This issue is dedicated to RICHARD ERIC HOLTTUM Honorary Member and Past- President of the British Pteridological Society, Director of the Singapore Botanic Gardens 1925- 1949 and Professor of Botany, Universit y of Malaya, Singapore 1949- 1954 on the occasion of his Eightieth Birthday 20th July 1975 FERN GAZ. -
A Classification for Extant Ferns
55 (3) • August 2006: 705–731 Smith & al. • Fern classification TAXONOMY A classification for extant ferns Alan R. Smith1, Kathleen M. Pryer2, Eric Schuettpelz2, Petra Korall2,3, Harald Schneider4 & Paul G. Wolf5 1 University Herbarium, 1001 Valley Life Sciences Building #2465, University of California, Berkeley, California 94720-2465, U.S.A. [email protected] (author for correspondence). 2 Department of Biology, Duke University, Durham, North Carolina 27708-0338, U.S.A. 3 Department of Phanerogamic Botany, Swedish Museum of Natural History, Box 50007, SE-104 05 Stock- holm, Sweden. 4 Albrecht-von-Haller-Institut für Pflanzenwissenschaften, Abteilung Systematische Botanik, Georg-August- Universität, Untere Karspüle 2, 37073 Göttingen, Germany. 5 Department of Biology, Utah State University, Logan, Utah 84322-5305, U.S.A. We present a revised classification for extant ferns, with emphasis on ordinal and familial ranks, and a synop- sis of included genera. Our classification reflects recently published phylogenetic hypotheses based on both morphological and molecular data. Within our new classification, we recognize four monophyletic classes, 11 monophyletic orders, and 37 families, 32 of which are strongly supported as monophyletic. One new family, Cibotiaceae Korall, is described. The phylogenetic affinities of a few genera in the order Polypodiales are unclear and their familial placements are therefore tentative. Alphabetical lists of accepted genera (including common synonyms), families, orders, and taxa of higher rank are provided. KEYWORDS: classification, Cibotiaceae, ferns, monilophytes, monophyletic. INTRODUCTION Euphyllophytes Recent phylogenetic studies have revealed a basal dichotomy within vascular plants, separating the lyco- Lycophytes Spermatophytes Monilophytes phytes (less than 1% of extant vascular plants) from the euphyllophytes (Fig. -
Evodevo: Past and Future of Continuum and Process Plant Morphology
philosophies Review EvoDevo: Past and Future of Continuum and Process Plant Morphology Rolf Rutishauser Department of Systematic and Evolutionary Botany, University of Zurich, CH-8008 Zurich, Switzerland; [email protected] Received: 7 October 2020; Accepted: 26 November 2020; Published: 1 December 2020 Abstract: Plants and animals are both important for studies in evolutionary developmental biology (EvoDevo). Plant morphology as a valuable discipline of EvoDevo is set for a paradigm shift. Process thinking and the continuum approach in plant morphology allow us to perceive and interpret growing plants as combinations of developmental processes rather than as assemblages of structural units (“organs”) such as roots, stems, leaves, and flowers. These dynamic philosophical perspectives were already favored by botanists and philosophers such as Agnes Arber (1879–1960) and Rolf Sattler (*1936). The acceptance of growing plants as dynamic continua inspires EvoDevo scientists such as developmental geneticists and evolutionary biologists to move towards a more holistic understanding of plants in time and space. This review will appeal to many young scientists in the plant development research fields. It covers a wide range of relevant publications from the past to present. Keywords: process philosophy; scientific perspectivism; developmental genetics; plant structure ontology; homology; land plant phylogeny; morphological misfits; flower; phyllotaxis; Utricularia 1. Introduction Plant morphology is the study of the physical form and structure of plants. Literally, the term “morphology” is derived from the Greek roots: morphe, which means form and/or structure, and logos, meaning discourse or investigation [1]. Plant morphology usually includes plant anatomy, which is the study of the internal structure of plants, as seen at the microscopic level. -
PTERIDOLOGIST the Fern Magazine 2014 Edition [Volume 6, Part 1]
PTERIDOLOGIST The Fern Magazine 2014 Edition [Volume 6, Part 1] EDITED BY A.E. GREENING ISSN 0266-1640 Editorial Preview It would appear that if you want to grow ferns from spores successfully, then you need to drink plenty of milk! Not only that, the milk should be bought in plastic 4 pint bottles and from the same retail outlet each time so that they are all the same size and shape. You don’t believe me? Check out page 10 and see what Fiona Layton manages to do with hers. There are some amazing articles in this issue. They range from ferns in far flung places to those nearer home. The Carribean islands 39 of Antigua and Barbuda have had an incredible resurgance amongst the fern world, thanks to Kevel Lindsay. He has managed to find 109 species since the last major survey in 1993 which only listed 45. That’s an increase of nearly 250%! His work continues and there is a book about to be published later this year. Please read his article starting on page 14. We had hoped to review his book in this issue but it has been slightly delayed and will hopefully be published very soon. A little bit nearer home, the island of Madeira has prompted 2 articles from Neil Timm that tell of the local tradition of using ferns in Christmas decorations (Page 20) as well as those that survive underground. (Page 33) We also have a very interesting article about a fern raised from detritus from an Antarctic glacier. Have a look at Page 58 and see what Ron Lewis-Smith has done.