Illustrated Handbook of Succulent Plants
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Illustrated Handbook of Succulent Plants Illustrated Handbook of Succulent Plants: Crassulaceae Bearbeitet von Urs Eggli 1st ed. 2003. Corr. 2nd printing 2005. Buch. XIII, 506 S. Hardcover ISBN 978 3 540 41965 5 Format (B x L): 21 x 27,9 cm Gewicht: 1660 g Weitere Fachgebiete > Chemie, Biowissenschaften, Agrarwissenschaften > Biowissenschaften allgemein > Ökologie Zu Inhaltsverzeichnis schnell und portofrei erhältlich bei Die Online-Fachbuchhandlung beck-shop.de ist spezialisiert auf Fachbücher, insbesondere Recht, Steuern und Wirtschaft. Im Sortiment finden Sie alle Medien (Bücher, Zeitschriften, CDs, eBooks, etc.) aller Verlage. Ergänzt wird das Programm durch Services wie Neuerscheinungsdienst oder Zusammenstellungen von Büchern zu Sonderpreisen. Der Shop führt mehr als 8 Millionen Produkte. _______________________________________________________________________________________U. Eggli Crassulaceae Crassulaceae Subfamily Sedoideae A. Berger 1930: Fl obdiplo- stemonous (St 2× as many as Pet) (with few excep- Perennial or rarely annual or hapaxanthic herbs, tions): subshrubs to shrubs (rarely aquatics, or tree-like, or − Tribe Kalanchoeae ’t Hart 1995: L decussate or epiphytic, or scandent), usually with succulent L, alternate, flat (rarely terete), crenate or dentate (ra- sometimes with succulent stems or underground rely entire); Fl 4- or 5-merous; Pet united to form a caudices; L opposite and decussate or alternate and distinct tube; Se costate: spiral, frequently aggregated into Ros,simple, Group A: L alternate: Adromischus, Tylecodon. usually entire, or crenate to lobed, glabrous or to- Group B: L decussate: Cotyledon, Kalanchoe. mentose; stipules absent; Inf lateral or terminal − Tribe Sedeae ’t Hart 1995: L mostly alternate, many-flowered spikes or panicles, rarely few- to 1- thick, entire (if decussate Pet free and Se costate); flowered and axillary; Fl bisexual or unisexual Fl (3- to) 5- to 32-merous; Se costate, (multi-) pa- (then plants ± dioecious), actinomorphic (except pillate or reticulate: Tylecodon grandiflorus), frequently 5-merous but • Subtribe Telephinae ’t Hart 1995: Stems often varying from 3- to 32-merous; Sep free or basally basally woody, tuberous or sympodial or mono- united; Pet free or basally united to form a short to podial rhizomes; L usually alternate, flat, den- long Cl tube; St as many or 2× as many as Pet, free tate or crenate (rarely decussate, terete or en- or fused to them; Ca as many as Pet, superior, free tire); Fl 5-merous (if polymerous then L flat, or almost so, basally with a small to conspicuous opposite or whorled); Sep basally connate; Pet NSc, gradually tapering into short to long Sty, with usually free (if connate then Se multipapillate); few to many ovules; Fr usually dehiscent follicles, Se costate or multipapillate: Hylotelephium, Hy- capsular; Se smallish, to 1.5 - 3 mm, elongate, pagophytum, Orostachys, Perrierosedum, Phe- smooth, papillate to longitudinally ridged, mostly dimus, Pseudosedum, Rhodiola, Umbilicus. brownish. • Subtribe Sedinae ’t Hart 1995: Stems herbace- ous, rarely woody or rhizomatous; L often ro- Distribution: Worldwide but esp. N hemisphere sulate or subrosulate, terete or semiterete, or flat and S Africa. and / or dentate-serrate and/or decussate; Fl 4- or to 32-merous; Sep basally connate or free; Literature: Berger (1930); ’t Hart (1995); Eggli Pet usually free and stellately patent, rarely & al. (1995). united; Se reticulate, papillate or costate: Group C: Plants various; L in Ros or scattered, The family counts ± 1400 species in 33 genera glabrous or glandular-hairy; Inf terminal or here accepted. It is here treated in its entirety, ad- lateral: Pistorinia, Prometheum, Rosularia, opting the consensus classification proposed by Sedum. Eggli & al. (1995), but modified to accept the ad- Group D: Plants rosulate, from Europe, W ditional genera Afrovivella, Phedimus and Prome- Asia, the Caucasus or Macaronesia, mostly theum, and subsuming Bryophyllum under Kalan- glandular-hairy; Inf mostly terminal; Fl choe, and Jovibarba under Sempervivum. The mostly > 5-merous: Aeonium, Aichryson, family has no important crop plants, with the excep- Monanthes, Sempervivum. tion of numerous taxa of horticultural interest, Group E: N American rosulate plants, glabrous mainly as hardy perennials for rock gardens, and − but often farinose; Inf lateral; Fl 5-merous: more recently − for establishing green ‘living’ Dudleya. roofs. Hylotelephium spectabile is sometimes used Group F: American plants, mostly with well- in the cut-flower trade. Numerous species from al- defined Ros, hairy or glabrous; Inf lateral or most all genera are popular in succulent plant col- rarely terminal; Fl mostly 5-merous: Eche- lections. veria, Graptopetalum, Lenophyllum, Pachy- Due to numerous suspected parallel develop- phytum, Thompsonella, Villadia. ments, the classification of the family has been dif- ficult, and the identification of species is often pro- blematical because of hybridization (to a limited Key to the genera: extent natural, and almost without limits in cultiva- 1Flwith St as many as Pet (haplostemonous): 2 tion, esp. amongst genera in Group F). The traditi- − Fl with St 2× as many as Pet (obdiplostemon- onal circumscription of 6 subfamilies by Berger ous): 8 (1930) was known to be artificial for some time. 2Floften (2- to) 4-merous or more rarely 10- to Recently, a revised classification was proposed by 12-merous (predominantly in Africa; extra-Af- ’t Hart (1995), based on molecular and morphologi- rican taxa dwarf minute ephemeral herbs with cal data: 2- to 4-merous Fl): 3 − Fl usually 5-merous (outside Africa): 4 Subfamily Crassuloideae A. Berger 1930: Fl ha- plostemonous (St as many as Pet): Crassula. 5 _______________________________________________________________________________________Crassulaceae U. Eggli 3 Perennial herbs with underground tuber or tu- 15 NSc conspicuous, broader and more obvious berous rhizome; stems deciduous; L ternate; Fl than the insignificant Pet (Canary and Selvagen 10- to 12-merous; Ca 2-seeded, follicles open- Islands): Monanthes p.p. ing transversally (Ethiopia): Hypagophytum − NSc inconspicuous, much narrower than the − Not as above; Ca with > 2 Se, follicles opening showy Pet: 16 apically: Crassula 16 Fl 5- (to rarely 6-) merous; Inf corymbose to 4 Plants with persistent or monocarpic, dense to much elongated and spike-like: 17 lax Ros: 5 − Fl 6- to 32-merous; Inf corymbose to dome- − Plants annual or perennial herbs without obvi- shaped, never much elongated: 20 ous Ros: 7 17 Inf flat-topped, corymbose or cymose, few- 5 Monocarpic Ros-forming herbs (Europe and flowered: 18 Asia): 6 − Inf elongate, many-flowered: 19 − Perennial shrublets with lax Ros at the stem tips 18 Pet 2.5 - 4 mm, ascending, white (E Asia): (Mexico): Graptopetalum pentandrum Meterostachys 6Infflat-topped (corymbose): Sinocrassula − Pet > 4 mm, ascending to spreading, whitish, − Inf thyrsoid-elongate: Orostachys p.p. yellowish, red or pink (E Mediterranean, W 7 Annual to perennial herbs; Fr many-seeded fol- Asia): Prometheum licles opening along the ventral side (Europe 19 Br of the Inf helicoid (Turkey, Iraq, Turkmeni- and Asia): Sedum p.p. (e.g. S. rubens) stan): Rosularia elymaitica − Annual minute ephemeral herbs; Fr indehiscent − Br of the Inf never helicoid (C to E Asia): and nutlet-like, each follicle with a single seed Orostachys p.p. (USA: California): Sedella p.p. 20 Ros sessile, usually < 10 cm ∅; Fl 6- to 18-mer- 8 Perennial plants, stems perennating but with an- ous, often in shades of pink and purple, rarely nually deciduous L, or stems annually decidu- white or yellow: Sempervivum ous: 9 − Ros sessile or often with a conspicuous and − Annual (to biennial) plants, or perennial plants sometimes branched stem, often > 10 cm ∅; Fl with at least some perennating L: 14 (6- to) 10- to 32-merous, often in shades of 9 Stems obviously succulent, perennating; L yellow but also whitish, more rarely reddish: crowded at branch tips (S and SW Africa): Aeonium Tylecodon 21 L decussate throughout the whole length of the − Stems not succulent and regularly deciduous stems: 23 and plants perennating with underground root − L verticillate or alternate at least in the upper stocks or small caudex (mostly outside of Afri- stem parts, or in Ros: 22 ca): 10 22 L verticillate (Africa): 10 L usually distinctly peltate; Inf terminal ra- Sedum p.p. (e.g. S. epidendrum) cemes or panicles (Asia and E Africa): − L alternate at least in the upper stem parts, or in Umbilicus p.p. Ros: 30 − L flat, never peltate; Inf usually cymes or 23 Annual to biennial glabrous to glandular-hairy corymbs, ± flat- topped (outside of Africa): 11 herbs to 15 cm; Fl (4- to) 5-merous, white, pink 11 Plants with tuberous caudex and annually de- or purplish; Pet 4-5mm(Mediterranean): 24 ciduous stems; L alternate; Fl 4- to 6-merous, − Perennial herbs (sometimes monocarpic), or often unisexual (plants monoecious or shrubs or small trees, or lianas; Fl 4- to 6-mer- dioecious): Rhodiola ous; Pet > 5 mm, in various colours: 25 − Plants with or without thickened roots but never 24 Annual glabrous herbs; Inf to 5 cm tall: with a caudex; L opposite, alternate or rarely Phedimus stellatus verticillate; Fl various but plants never dioec- − Annual to biennial glandular-hairy herbs; Inf to ious: 12 60 cm tall: Sedum p.p. (e.g. S. cepaea) 12 L flat: 13 25 Fl 4-merous; herbs (sometimes monocarpic) to − L terete-subulate: Villadia p.p. shrubs or small trees, or lianas (Africa, Mada- 13 R thickened, fusiform, and stems annually deci- gascar, Asia, neophytes world-wide):Kalanchoe duous; or R fibrous and stems perennial, woody, − Fl 5- or 6-merous; shrublets or herbs: 26 forming dwarf shrublets: Hylotelephium 26 Fl 5-merous, not white (Africa, Caucasus, N − R fibrous; stems annually deciduous:Phedimus America): 28 14 Plants with perennial monocarpic Ros: 15 − Fl (5- to) 6-merous, erect, white: 27 − Plants annual (to biennial), or perennial but then 27 Shrublets to 80 cm tall (Madagascar): not with monocarpic Ros: 21 Perrierosedum − Dwarf herbs to 10 cm tall (Europe and N Afri- ca): Sedum p.p. (e.g.