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Tí R BOTÁNICA VOLUMEN 47 NÚMEROS 3-4 - 1 ISSN 0016-5301 1 T - tí r BOTÁNICA VOLUMEN 47 NÚMEROS 3-4 UNIVERSIDAD DE CONCEPCION-CHILE FACULTAD DE CIENCIAS BIOLÓGICAS Y DE RECURSOS NATURALES UNIVERSIDAD DE CONCEPCIÓN CHILE DIRECTOR DE LA REVISTA Andrés O. Ángulo REEMPLAZANTE DEL DIRECTOR Osear Matthei J. REPRESENTANTE LEGAL: Augusto Parra Muñoz PROPIETARIO Universidad de Concepción DOMICILIO LEGAL: Victor Lamas 1290, Concepción. Chile EDITOR EJECUTIVO SERIE BOTÁNICA Roberto Rodríguez R. COMITÉ ASESOR TÉCNICO Miren Alberdi Jurke Grau Universidad Austral de Chile Universidad de München, Alemania Sergio Avaria Mary Kalin Arroyo Universidad de Valparaíso Universidad de Chile, Santiago Danko Brncic Luis Ramorino Universidad de Chile, Santiago Universidad de Valparaíso Eduardo Bustos Bernabé Santelices Universidad de Chile, Santiago Universidad Católica, Santiago Hugo Campos Federico Schlegel Universidad Austral de Chile Universidad Austral de Chile Juan Carlos Castilla José Stuardo Universidad Católica, Santiago Universidad de Concepción Fernando Cervigon Tod Stuessy Fundación Científica Los Roques, Venezuela Ohio State University, U.S.A. MÓNICA DURRSCHMIDT Guillermo Tell Justus-Liebic Universitat Giessen. Alemania Federal Universidad de Buenos Aires, Argentina RN Wl>! / Ha roldo Toro lad de Chile, Santiago Universidad Católica, Valparaíso 51 LIBRARV JUN 2 9 igq? ISSN 0016 5301 NEW YGRfc BOTANICAl GHRV A BOTÁNICA VOLUMEN 47 NÚMEROS 3 4 1990 CONTENIDO / CONTENTS y G.A. WHEELER & M. MUÑOZ-SCHICK. Carex andina Philippi (Cyperaceae): ¡ts taxonomy, distribution and lectotypification 7 Carex andina Philippi (Cyperaceae): su taxonomía, distribución y lectotipi- ficación. /T.F. Stuessy & C. M ARTICORFNA. Orthography of some epithets honoring Ber tero in the vascular flora of the Juan Fernández Islands and continental Chile 77 Ortografía de algunos epítetos en honor a Bertero en la flora vascular de las islas de Juan Fernández y Chile continental. y M. Baeza. Rytidosperma paschalis (Pilger) Baeza, una nueva combinación para la flora agrostológica de Chile 83 Rytidosperma paschalis (Pilger) Baeza, a new combination for the agrosto- logical flora of Chile. y C. Marticorena. Contribución a la estadística de la flora vascular de Chile 85 Contribution to the statistics of the vascular flora of Chile y O. M atthei & M. Qi'ezada. Nuevos géneros para la flora advena de Chile 1 1 New genera for the adventitious flora of Chile y M.T.K. Arroyo. C. Marticorena and M. Muñoz. A checklist of the native annual flora of continental Chile 1 19 Un catálogo de la flora anual nativa de Chile continental. UNIVERSIDAD DE CONCEPCIÓN CHILE "Los infinitos seres naturales no podrán perfectamente conocerse, sino luego que los sabios del país hagan un especial estudio de ellos". CLAUDIO GAY. Hist. Chile, Zool. 1:14(1847) Portada: Galeopsis letrahil L.. planta advena de Chile ESTA REVISTA SE TERMINO DE IMPRIMIR EN IOS TALLERES DE Editora Aníbal Pinto. S.A.. MAIPU 769. CONCEPCIÓN CHILE. EN EL MES DE ABRIL DE 1991. LA QUE SOLO ACTÚA COMO IMPRESORA PARA EDICIONES UNIVERSIDAD DE ( ON< I W ION Gayana, Bol. 47 (3-4): 71 76, 1990 ISSN OOI6 530I CAREX ANDINA PHILIPPI (CYPERACEAE): ITS TAXONOMY. DISTRIBUTOR AND LECTOTYPIFICATION CAREX ANDINA PHILIPPI (CYPERACEAE): SU TAXONOMÍA, DISTRIBUCIÓN Y LECTOTIPIF1CACIÓN Gerald A. Wheeler* and Mélica Muñoz-Schick' ABSTRACT resumía Carex andina occurs in Central Chile and the northwestern Carex Andina se encuentra en Chile central y en la región part of Argentine Patagonia. where it grows on dry and noroeste de la Patagonia argentina, donde crece en laderas rock} slopes ai elevations of about 1200-3300 rh. Reports of secas \ rocosas entre 1200 3300 m. Citas de esta especie para ilus species from southern Patagonia are base/1 on misidenti- el sur de la Patagonia están basadas en especímenes mal fied specimens. Morphological differences between C. andina identificados. Se discuten las diferencias morfológicas entre and C. set¡folia are discussed. The salient features of C andi C. andina y C serifolia. Las características sobresalientes de na are: perigynia obevate. essentially beakless; achenes obo- C. andina son: periginios obovados, casi sin pico; aquenios vate; rachilla relatively broad, lanceolate, 1/3 to 3/4 the obovados; raquilla relativamente ancha, lanceolada, de 1/3 a length of the achene; and leaf sheaths with the ventral band 3/4 del largo del aquenio; vainas foliares con la banda ventral smooth at the mouth. A lectotype is chosen for C. andina. lisa en la boca. Se elige un lectotipo para C. andina. Kevwords: Carex anilina. Cyperaceae, southern South Ame rica, lectotype. INTRODUCTION Kük.) in southern South America. Examination of syntypes of C. andina reveáis that some A recent study of Carex L. (Cyperaceae) material specimens previously reported as "C andina" are from Argentina and Chile, as well as a survey of actually referable to other species; also, some South American literature, reveáis that some specimens clearly assignable to C. andina have confusión exists regarding the distribution of Ca- erroneously been reported as other taxa (e.g., C. rex andina Philippi (sect. Junciformes (Boeckeler) setifolia Kunze ex Kunth var. neuquensis Barros). Below we attempt to: (1) clarify the distribution of C. andina in southern South America, (2) point out salient differences in Universitj of Minnesota Herbarium, Plant Biologj De- morphology between C. andina and C. setifolia U.S.A. partment, St. Paul, Minnesota, 55108. (and its varieties), species which have been Museo Nacional de Historia Natural. Casilla 787. Santiago. Chile. confused in the recent past, (3) place the ñame C. 71 1 Gavana. Bot. 47 (34), 1990 setifolia var. neuquensis in the synonymy of C. folia var. setifolia (Fig. 4A-BI the perigynia are andina, (4) discuss the status of "C. andina" sensu pyriform and distinctly beaked, the achenes Barros (1935, 1947, 1969). and (5) select a oblong, and the rachilla setiform and somewhat lectotype for C. andina. shorter than to slightly exceeding the achene. So- Full citations are given near the end of this me of the aforementioned diagnostic features of report for specimens of Carex andina collected in var. setifolia have been illustrated by Kunze Argentina and Chile; a distribution map for C. (1840-51, Tab. 26) and Kükenthal (1909, Fig. 18: andina is also provided (Fig. 1). When discussing p. 84). Furthermore, an isotype of the typical members of the C setifolia species complex, we variety of C. setifolia (Poeppig I. N° 26 [(BM!]) nave retained the nomenclature used by Kü- from Chile, clearly possesses these characters, kenthal (19091 and Barros (1935, 1947, 1969). although the perigynia and achenes of the BM specimen are somewhat immature. It is noteworthy that C. setifolia var. colchaguensis MORPHOLOGY (Philippi) Kük. and var. pungens (Boeckeler) Kük. (sensu Kükenthal 1909) also have pyriform Examination of syntypes of Carex andina (Fig. perigynia, oblong achenes, and a setiform are obovate and 2A-B) reveáis that the perigynia rachilla. A morphological comparison of 1 essentially beakless. the achenes obovate, and the characters of C. andina and C. setifolia var. 3/4 rachilla broadly lanceolate and about 1/3 to setifolia is given in Table I. the length of the achene. By contrast. in C. seti- DISCUSSION Philippi (1896. p. 485) described Carex andina from plants collected in the Andes of central Chi- le, and subsequently Kükenthal (1899, 1909) re- cognized it as a good species. More recent wor- kers have reponed C. andina from Argentina (Barros 1935. 1947. 1969) as well as from Chile (Léveillé 1915; Marticorena and Quezada 1985). Kükenthal (1910) described a variety of C andi- na, var. subabscondita Kük., from plants collec- ted in central Patagonia (Chubut Province, Ar- gentina), but these low-growing plants. which Wheeler (1986) considers to be conspecific with C. nelmesiana Barros, are more closely related (i.e.. morphologically more similar) to the other low-growing Patagonian members of sect. Junci- formes (e.g.. C. argentina Barros and C. austro- americana G. Wheeler) than to C. andina, which differs by having elongated culms (up to 35 cm tall). several to many perigynia per spike. and a relatively broad rachilla. While Philippi (1896), Kükenthal (1899, 1 909), and Léveillé (1915) reported Carex andina only from central Chile, Barros (1935, 1947, 1969) reported it also from the western portion of Argentine Patagonia. citing specimens from Neuquén {Castellaitos s.n. [BA-1892]and O'Do- FlG. 1. Map of southern South America showing the nell 2142 [LIL]). Chubut (Gerling 215 [BAF]), distribution of Carex andina. and Santa Cruz (Hogberg 42 \L?\) provinces. For 72 Carex andina, distribution: G.A. Wheeler & M. MtSo/-S< iik k Tabi i l.A selected morphological comparison oi Carex andina and C setifolia var. setifolia in South America Characters Ga\ana. Bot. 47 (3-4), 1990 Fio 3. Carex andina. Specimens from Argentina. A. Perigy- nium (ventral view); achene with attached rachilla (dorsal view); from O'Donell 2142 (LIL). B. Perigynium (ventral view); achene with attached rachilla (dorsal view); from Bur kan 9620 (Sil. C. Perigynium (ventral wiew); achene with attached rachilla (dorsal wiew); from Boelcke 10798 (BAB). In A-C: perigynium above, achene with attached rachilla below; arrow points to apex of rachilla; bar equals 1 mm. From the discussion above, it is evident that Carex andina occurs in Argentina as well as in Chile. But what is "Carex andina" sensu Barros (1935. 1947. 1969)? It is clear from Barros's Fig 4. Carex setifolia var. setifolia. Specimens from Chile. A. perigynium (ventral view); achene with rachilla illustrations of "C. andina" that more than one attached (dorsal view); from Montero 7837 (GH). B. Perigynium taxon is involved. Examination of O'Donell 2142 (ventral view); achene with attached rachilla (dorsal view); l LIL). which was the specimen cited and from E. Barros 463 (GHl. In A-B: perigynium above. achene illustrated for the species in 1947 (p. 394, Pl. with attached rachilla below: arrow points to apex of rachilla: 168), reveáis that it is indeed referable to C bar equals 1 mm.
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