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rcO\Z)ILZ ?7ZQ)EIotst(tlN 266 Nx)ffia Vol. 13, No. 2 (2000) describedin George (1995). Measurementsof leavesand habit were made from dried herbarium specimensand floral characterswere measuredfrom fresh, reconstitutedand pickled (707oethanol) material. Illustrations of floral and foliage featureswere preparedby the author.

Taxonomy

Synapheabrachyc€ras R. Butcher,rp. ,1ov.

2Synapheafavosavar.lanceolataMeisn in A.L.P.P. de Candolle, Prodr. 14: 314 ( 1856).Type: Swan RiverColony [Westem Australia], J. D rummond3rd coll.258 (synrBM, K, tt.v.).

Foliaintegra velpinnatifida, ad apicem obtusavel acuta, r€ticulo tenui atque profundo vel foveato Perianthiumin demidioinferiore glabratum vel puberulum.Tepalum adaxiale 4.1-5.5 mm longum, 1.71.4 mm latum. Tepalumabaxiale 2.5-3.7 mm longum,0.7-1.9 mm latum,ad apicemvalde recuryatum.Stigma manifeste bicome, et superpagina dorsali comu parvorecurvo instructum, 1.2-1.5mm longo. 0.9-1.6 mm lato.

ryprrr. north-westof ArthurRiver [precise locality witheld], Western Australia, 29 Septemberl998, R. Butcher& J.A. WegeRB 498(holor PERTH 05495660; rso; CANB, K, MEL, NSW).

Caespitosesubshrub to 50 cm tall, 20-90 cm wide. S/emsseveral, to 15 cm long, appressed- pubescenton upper stems. PetioleJ 3.5-20 cm long, glabrous to shortlypubescent, with scatteredlong hairs;sheath appressed pubescent. l*af bladesent\re and oblanceolate orthreelobed to pinnatipartite with thelowest pair of lobesagain pinnatipartite; ultimate lobes linear to lanceolate;apices rounded to acute,shortly pungent; 6.5-17.5 cm long, 0.6-17 cm wide,sub-glabrous to shortlypubescent, with scatteredlonghairs; reticulation fine and deep topitted. Inflorescence aspike, 2.5-15 cm long, shorter thanorjust exceedingthe leaves; flowers moderately crowded, intemodes half to c' equalthe flower length;peduncle 2.5-17 cm long, simple to little-branched,appressed puberulous to pubescent;rachis appressedpubescent; bracts ovate, obtuse, 1-1.5 mm long,pubescent at base,glabrous toward apex with ciliate margin. Perianthascending, gaping, extemally glabrescent to sparselypuberulous in lowerhalf;adaxial tepal concave, margins reflexed towards apex,4.l-5.5 mm long,1.7-2.4 mm wide, sparselypuberulous intemally in thedistal half; abaxialtepal convex, 2.5-3'7 mm long,0.7-1.9 mm wide, apexacute and strongly reflexed to 0.6-1 mm of its length,pubescent intemally behind the anther;lateral tepals falcate,2.7-4 mm long, 1.5-3.1mm wide,the apexreflexed to 0.7-1.4rnn of length,puberulous intemally behind the anthers.StiSma transversely elliptic to elliptic, convex, 1.2-1.5mm long,0.9-1.6 mm wide,with two divergent,reflexed homs 0.7-1 mm long,the dorsal surfaceofthe stigmawith a short,curved projection ofc. 0.3rnm lengthl ovary cylindric to ellipsoid, with apicalbeak to 0.2mm long(after style separates), sericeous, c. 0.9mm long,0.5 mm wide'with a ring of translucentglandlike hairsat apex.Fruit elliptic,with a beakto 0.5 mm long,pubescent, c. 4.5 mn long. Seednarrowly turbinate, c. 2.2 mm long, 0.9 mm wide; testacream, smooth. (Figure1 A-J)

Otherspecimens exan ined WESTERNAUSTRALIA [preciselocalities withheld]: NW of Arthur River,28Sep. 1998,R. Butcher &J.A.WegeRB497 (PERTH);NW of ArthurRiver,29Sep 1998, R. Butcher& J.A.Wege RB 499(PERTH); NW of ArthurRiver, 30 Sep.1997 , R. Butcher,J.A. Wege & C.F. WilkinsRB 307 (PERTH)INW of Arthur River, 30 Sep.1997, R. Butcher,J.A. Wege& C.F.Wlklns RB 308 (PERTH); NW of tuthurRiver,30Sep. 1997 ,R. Butcher, J.A.Wege & C.F Wilkins 'ur. enSrc I = J?q al?rs e.req^\'g pu? Y ldatxo'tutu I =req ol?rs 66t gu ?3a,4.tvt * ra|rvlg d luo{ u^\up II 'aot gx rru?1,4{ t3 T aSaa'v'[ laqrtng u'uro4 u^\erp'I-3 ssrnSlc ,{le^pcedsar 80e su sll1ri,44 JJ ? g sem8rC aBaA Vt'raqqt1{ ypuP60€ Ad sultttl,|'fOT a7ay' Vt'ratp1t18 V wo4 ulNrrp PUeV lnu-I:spuelS 'liEurs pu€ Yr^uor 8ur^loqs lerrdtr qt!^\ ft!^o snoaiuos I :uoq lEsrop pe^nceJ poe enssp a^llseuuor r)rlnq lPrlur^ - aprs nro.U po^\er^ ?ruSrls - H :(pa^\oll€) uollreford pu? suloq lueire^P 8ul^\oqs ?u8ls Jo sreJ:lnsI€slop C :xede - paxauol qtr^\ erryns rolno pu? reqrux nln.ollun Bul^\oqs (e^oq?) ac4lns rauul lEdol lelalEl i :xadP plxauol ,{l8uo$ Surmoqsecryns:alno pue reqlup relnsolrq qll^\ (Ual) ecP.Jnslouul'pdat prxr:qt - g ixed? 1r sulSrPurpaxauel 8ul^\oqs p?rq erryns .relnopux lueu?lu raqlup ellnls ql!^\ olal) arslns leuul'Fdal lDlxlp? d :Jpq reddn eql uI rueis.rqel8 puB - qrueuod Suruado,{1apr,t aql 8ul^{oqs e.+ns FIxPqB aqt tt|o.t] p.,^\ol^ Ia^\oU J :srpq porallPts pux uorleln.llel daopJo - - erfiU lmtep qrr^l JEal ellu?dleuuld g :uoF?lntler pslrld Jo llrlap qll.a\Fol aldurs V s?lar{qrtuq oa|llouns I M @ /,fl,\l Vru &r

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(PERTH);NNEofArthur RB 309(PERTH);NNEof ArthurRiver, 30 Aug'1995'V'CrowlevDKN 140 1995'V' CrowleyDKN nin".,:b eog. l6eS,V. CrowleyDKN142 (PERTH);E ofDardadine' 22 Ocl (PERTH); BlockState i+ripenfni;NW of ArthurRiver,24 Sep.1998,M' GrahamMSG 985 Quinns Forest52, SW of Highbury,28 Sep.1998, G' Warren62 (PERTH)'

roadsideremnants in the Distribution. synapheabrachyceras is restrictedto a few smallreserves and carriedout in 1997and 1998 ertfru, niu"r-tiighUoryu,"u, ". 200k-SE of Perth(Figure 2) Fieldwork piesseville populations.Further arounclHillman, bardadine,Tarwonga and failed to locateadditional surveyin surroundingareas is urgently requiredin light of the rarity of this species'

sandover lateriteand grey Habitat. Synapheabrachycerashasbeencollected from red-brownclayey Eucalyptus openwoodland with r"-"Oygt;t"l "*. granite'Associated vegetation includes .wandoo Dryandra' Beaufortia' " f"*'rft-U undeistoreycomprising Isopogon teretifolius and speciesof Allocasuarinahumilis Bossiaea,Hibbertia, Daimpiera, Lepiospe'tmim and Hypocalymma' as well as Dryandra cuneataand ind nok* trf\unotoopen scrubover aio w heathof AdenanthosflavidiJ'lorus' attenuata' In undisturbed iortnonn*L sp' witi occasionalNuytsia floribunda and Banksia populationsit is popufu,iontS. brachycerashas a scattereddittribotion, but in highly disturbed may be a disturbance iol,-on Aong the edgesof tracksand gravelscrapes suggesting that it opportunist.

to Octoberwith fruits Phenology.Flowes of Synapheabrachycerashavebeen recorded from August recordedin Novemberand December.

' Williams . Highbury NlA

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'uur8gs eql;o ecc.1rnsIES.rop eqt uo uorlcafordaq1 o1 erue:eJal ur ,.uJorl,, 3urueeursrz.rarpuu,,uoqs,,8urueeu.r[{Jrl4 {ae:g eql uo4 pa,luapsr leq]rda cU nads aq1 t3o1owt1g

'Ealg Vtd lse^\ qtnos ra^ru rnqlv !uo4 s:rJrds t\ex r sDra)t\rprq ,rqdrrf,S rrqrlng uouo,{d Vol. 13, No. 2 (2000) 2',70 Na)fta distinguishedfloms.brachycerasbytheveryfineandshallowleticulationontheleaflamina,aswell Similarly, as thJroader, lessdivided leaf lobes,the lowest of which areusually curved downwards. from the specimensof S. brachyceras having only simple leaveson the plant can be distinguished on the leaf iheatbelt speciesS. piatyphylla A.S. Georgewhich possessesvery shallow reticulation third ofits length lamina, ania stigmawhich is squareto obtrapeziformand lobed to approximatelya without a dorsal Projection.

Initsdivelgentstigmatichorns,s'braclrlcerasisaklrrtos,nexosaA.S.Georgewhichhaslonglobes dte to its at the apex oflhe stijma like membersof sect.Bicornir, but is included in sect.Synaphea from s. ne.rora deeply iivided leaf lamina (A.s. George). S. brachycerascan be easily differentiated to pitted reticulation as its ieaveshave pinnatipartitedivision, acuteapices to the lobes and fine, deep the leaf lobes and whilst thoseof the latter taxon havetripartite division, obtuseto lounded apicesto shorterthan to shortly open,shallow reticulation. Additionaliy , S. brachycerashas its inflorescences length' Although exceedingthe leaveswheteas S. l?e,roschas its flower spikesgreatly exceeding leaf s.petiola-rissubsp'trilobaA.S'GeorgeisverysimilartoS.ne.rosa,nomembersoftheS.petiolLris taxonomy of ,p""1", .ornpt"* ituve been included in this discussionfor the sakeof simplicity' as the thisgroupisfa'fromresolvedandtheirinclusiondoeslittletofurtherthepurposeofthispaper.

'tar. in synonymy with Nores. The name Synapheafavosa lanceolota Meisn' may belong and this remainsa s. brachyceras,b|lt the availablesyntype and isotypematerial examinedis sterile name of uncertain application.

"Flora Amendments to k€y in ofAustralia"

key to The recognition of S. brachyceras requiressome amendments to be made to the existing distinction of species(Geoige 1995), although the only changessuggested here are to facilitate the 3 now designated i. brorhyr"roi. Two new couplets(3 and 7), arerequired, with the original couplet andnew ,3a', Somechangesare also necessary to sizerangesin someofthecouplets; all modifications of the key in coupletsare marled with an asterisk. The following extractmay be insertedat the start George(1995).

1. Stigma divided to half-way or more into 2 narrow homs 2 Leaf lamina deeply divided into linear to lanceolate,obtuse (rarelyacute)lobes *3 Mature leavestrilobed to pinnatipartite;reticulation very fine and deep; inflorescenceshorter than or shortly exceedingleaves " """"" S' brachyc€ras [Arthuri.iver area]. *3: Mature leavesonce or twice tdlobed; reticulation open and shallow inflorescencegreatly exceedingleaves 3a Shrub to 60 cm talt; petiole to 20 cm long; peduncleto 35 cm long; S' petiolaris fruit 5-6 mm long [Perth to Busseltonand E to CapeArid] """""" 3a: Shrub to I m talll petiole often 20-30 cm tong: peduncleto 60 cm S' nexosa long; fruit 8-9 mrn long lscott River] 2: Leaf larnina entire to dentateor shortly lobed with tdangular lobes 4 Leaves obovate to oblanceolate ( eurnoqte^:OdISl) 90€-tLZ dd'9t to^,!?l?!snv Jo erct&,,:J Daqdou(S(966t) SV'o&org ( uopuo'I |oJ T e^oeu) I IoA ,! slsuoll?Isnv?lolC,, (0t81) D'uEqtuog 'f:rli""";""?r"utu",TlX*o pue.rpue uor.^rosuoc Jo ruaruu'dacE'srrsnv uelse^{ roJ )s!.r ?rors ^,,r"tra oJ:|n'o

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