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EDITORS’CHOICE EDITED BY GILBERT CHIN AND JAKE YESTON B I O M E C H A N I C S No Skimming Allowed Pterosaurs were flying reptiles and the first air-borne vertebrates; they dominated the skies from the late Triassic to the end of the Cretaceous, during the epoch of their relatives, the dinosaurs. On the basis of similarities in jaw structure, it has been suggested that several pterosaurs, including Thalassodromeus and the giant Quetzalcoatlus (with a wingspan of up to 15 m), could have fed by skimming in a manner akin to that of extant ternlike shorebirds (Rynchops spp.) Skimmers fly low over calm shallow water with the tip of their lower beak dipping beneath the water surface. Humphries et al. have used full-sized mod- on August 31, 2007 els of mandibles from Thalassodromeus and the modern skimmer R. niger to demonstrate that the pterosaur bill would have generated an order of magnitude more drag in traveling through the water. Modeling indicated that the energetic cost to a shorebird of flying with its beak in the water is almost prohibitive (~20% of the total cost of flight), and the authors sug- gest this levy might explain the rarity of the skimming life-style. The substantially greater cost for a pterosaur larger than 2 kg appears to exclude outright skimming as a possible means for procuring food. Furthermore, many of the morpho- logical specializations to the head and neck seen in Rynchops are not found in pterosaurs of any size, including the ability to regenerate broken or abraded bill tips and the presence of a reinforced lower jaw. -
Pterosaurs Flight in the Age of Dinosaurs Now Open 2 News at the Museum 3
Member Magazine Spring 2014 Vol. 39 No. 2 Pterosaurs Flight in the Age of Dinosaurs now open 2 News at the Museum 3 From the After an unseasonably cold, snowy winter, will work to identify items from your collection, More than 540,000 Marine Fossils the Museum is pleased to offer a number of while also displaying intriguing specimens from President springtime opportunities to awaken the inner the Museum’s own world-renowned collections. Added to Paleontology Collection naturalist in us all. This is the time of year when Of course, fieldwork and collecting have Ellen V. Futter Museum scientists prepare for the summer been hallmarks of the Museum’s work since Collections at a Glance field season as they continue to pursue new the institution’s founding. What has changed, discoveries in their fields. It’s also when Museum however, is technology. With a nod to the many Over nearly 150 years of acquisitions and Members and visitors can learn about their ways that technology is amplifying how scientific fieldwork, the Museum has amassed preeminent own discoveries during the annual Identification investigations are done, this year, ID Day visitors collections that form an irreplaceable record Day in Theodore Roosevelt Memorial Hall. can learn how scientists use digital fabrication of life on Earth. Today, 21st-century tools— Held this year on May 10, Identification Day to aid their research and have a chance to sophisticated imaging techniques, genomic invites visitors to bring their own backyard finds have their own objects scanned and printed on analyses, programs to analyze ever-growing and curios for identification by Museum scientists. -
Titanosauriform Teeth from the Cretaceous of Japan
“main” — 2011/2/10 — 15:59 — page 247 — #1 Anais da Academia Brasileira de Ciências (2011) 83(1): 247-265 (Annals of the Brazilian Academy of Sciences) Printed version ISSN 0001-3765 / Online version ISSN 1678-2690 www.scielo.br/aabc Titanosauriform teeth from the Cretaceous of Japan HARUO SAEGUSA1 and YUKIMITSU TOMIDA2 1Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6, Sanda, 669-1546, Japan 2National Museum of Nature and Science, 3-23-1 Hyakunin-cho, Shinjuku-ku, Tokyo 169-0073, Japan Manuscript received on October 25, 2010; accepted for publication on January 7, 2011 ABSTRACT Sauropod teeth from six localities in Japan were reexamined. Basal titanosauriforms were present in Japan during the Early Cretaceous before Aptian, and there is the possibility that the Brachiosauridae may have been included. Basal titanosauriforms with peg-like teeth were present during the “mid” Cretaceous, while the Titanosauria with peg-like teeth was present during the middle of Late Cretaceous. Recent excavations of Cretaceous sauropods in Asia showed that multiple lineages of sauropods lived throughout the Cretaceous in Asia. Japanese fossil records of sauropods are conformable with this hypothesis. Key words: Sauropod, Titanosauriforms, tooth, Cretaceous, Japan. INTRODUCTION humerus from the Upper Cretaceous Miyako Group at Moshi, Iwaizumi Town, Iwate Pref. (Hasegawa et al. Although more than twenty four dinosaur fossil local- 1991), all other localities provided fossil teeth (Tomida ities have been known in Japan (Azuma and Tomida et al. 2001, Tomida and Tsumura 2006, Saegusa et al. 1998, Kobayashi et al. 2006, Saegusa et al. 2008, Ohara 2008, Azuma and Shibata 2010). -
Pterosaur Distribution in Time and Space: an Atlas 61
Zitteliana An International Journal of Palaeontology and Geobiology Series B/Reihe B Abhandlungen der Bayerischen Staatssammlung für Pa lä on to lo gie und Geologie B28 DAVID W. E. HONE & ERIC BUFFETAUT (Eds) Flugsaurier: pterosaur papers in honour of Peter Wellnhofer CONTENTS/INHALT Dedication 3 PETER WELLNHOFER A short history of pterosaur research 7 KEVIN PADIAN Were pterosaur ancestors bipedal or quadrupedal?: Morphometric, functional, and phylogenetic considerations 21 DAVID W. E. HONE & MICHAEL J. BENTON Contrasting supertree and total-evidence methods: the origin of the pterosaurs 35 PAUL M. BARRETT, RICHARD J. BUTLER, NICHOLAS P. EDWARDS & ANDREW R. MILNER Pterosaur distribution in time and space: an atlas 61 LORNA STEEL The palaeohistology of pterosaur bone: an overview 109 S. CHRISTOPHER BENNETT Morphological evolution of the wing of pterosaurs: myology and function 127 MARK P. WITTON A new approach to determining pterosaur body mass and its implications for pterosaur fl ight 143 MICHAEL B. HABIB Comparative evidence for quadrupedal launch in pterosaurs 159 ROSS A. ELGIN, CARLOS A. GRAU, COLIN PALMER, DAVID W. E. HONE, DOUGLAS GREENWELL & MICHAEL J. BENTON Aerodynamic characters of the cranial crest in Pteranodon 167 DAVID M. MARTILL & MARK P. WITTON Catastrophic failure in a pterosaur skull from the Cretaceous Santana Formation of Brazil 175 MARTIN LOCKLEY, JERALD D. HARRIS & LAURA MITCHELL A global overview of pterosaur ichnology: tracksite distribution in space and time 185 DAVID M. UNWIN & D. CHARLES DEEMING Pterosaur eggshell structure and its implications for pterosaur reproductive biology 199 DAVID M. MARTILL, MARK P. WITTON & ANDREW GALE Possible azhdarchoid pterosaur remains from the Coniacian (Late Cretaceous) of England 209 TAISSA RODRIGUES & ALEXANDER W. -
First Evidence of Azhdarchid Pterosaurs from the Late Cretaceous of Hungary
First evidence of azhdarchid pterosaurs from the Late Cretaceous of Hungary ATTILA ŐSI, DAVID B. WEISHAMPEL, and CORALIA M. JIANU Ősi, A., Weishampel, D.B., and Jianu, C.M. 2005. First evidence of azhdarchid pterosaurs from the Late Cretaceous of Hungary. Acta Palaeontologica Polonica 50 (4): 777–787. New remains of an azhdarchid pterosaur were discovered from the Upper Cretaceous (Santonian) Csehbánya Formation at the Iharkút vertebrate locality in the Bakony Mountains, western Hungary. Among the isolated bones, consisting prin− cipally of 21 symphyseal jaw fragments, four cervical vertebrae, a right radius, and some fragmentary limb bones, is a complete articulated mandible that represents one of the best−preserved mandibular material of any presently known azhdarchid pterosaur. The complete edentulous jaw, referred to Bakonydraco galaczi gen. et sp. nov. posesses several fea− tures diagnostic for azhdarchids which prove that Bakonydraco belongs to this group. The cervical vertebrae exhibit azhdarchid features and consequently are referred to as Azhdarchidae indet. The discovery of these fossils helps to under− stand the construction of the azhdarchid mandible and provides new insight for studying the feeding style of the edentulous azhdarchid pterosaurs. Key words: Pterosauria, Azhdarchidae, mandible, cervical vertebrae, Cretaceous, Hungary. Attila Ősi [[email protected]], Department of Palaeontology, Eötvös Loránd University, Budapest, 1117, Hungary; Da− vid B. Weishampel [[email protected]], Center for Functional Anatomy and Evolution, Johns Hopkins Univer− sity, Baltimore, 21205, USA; Coralia M. Jianu [[email protected]], Muzeul Civilizaţiei Dacice şi Romane Deva, Deva, 2700, Romania. Introduction Arambourgiania philadelphiae (Arambourg, 1959) from the Maastrichtian of Jordan (Frey and Martill 1996; Martill et al. -
Azhdarchid Pterosaurs: Water-Trawling Pelican Mimics Or “Terrestrial Stalkers”?
Azhdarchid pterosaurs: water-trawling pelican mimics or “terrestrial stalkers”? MARK P. WITTON and DARREN NAISH Witton, M.P. and Naish, D. 2015. Azhdarchid pterosaurs: water-trawling pelican mimics or “terrestrial stalkers”? Acta Palaeontologica Polonica 60 (3): 651–660. The lifestyles of all pterosaurs are contentious, but those of the pterodactyloid clade Azhdarchidae are particularly debat- ed. A 2008 review of the functional morphology of azhdarchid pterosaurs concluded that they were probably terrestrial foragers, as evidenced by their long limbs, generalised skull construction, the arthrological limitations of their cervical series, trackway data indicating terrestrial proficiency, a strong continental skew in the depositional context of their fossils, and several additional lines of corroborating evidence. This hypothesis was recently challenged on three counts: (i) azhdarchid fossils routinely occur in aquatic deposits; (ii) terrestrially-foraging pterosaurs were highly vulnerable to predation and (iii), aerial “water trawling”, where the mandible is pulled though water to catch food in a distended throat pouch, is a more likely foraging strategy. Pelican-like jaw mechanics were suggested for azhdarchids because of the asymmetrical jaw joints in these pterosaurs, which permit lateral deflection of the mandibular rami during jaw extension. We evaluate these three claims and conclude that all are flawed. The frequent occurrence of azhdarchid fossils in aquatic sedimentary systems is not significant with regard to ecology or behaviour, since these provide the overwhelming mech- anism for the preservation of all fossil terrestrial animals. Likely pterosaur takeoff abilities and the ubiquitous nature of modern, terrestrially-foraging birds indicate that predation risks on ground-foraging pterosaurs are probably overstated. -
New Information on the Tapejaridae (Pterosauria, Pterodactyloidea) and Discussion of the Relationships of This Clade
AMEGHINIANA (Rev. Asoc. Paleontol. Argent.) - 41 (4): 521-534. Buenos Aires, 30-12-2004 ISSN 0002-7014 New information on the Tapejaridae (Pterosauria, Pterodactyloidea) and discussion of the relationships of this clade Alexander Wilhelm Armin KELLNER1 Abstract. A phylogenetic analysis indicates that the Tapejaridae is a monophyletic group of pterodactyloid pterosaurs, diagnosed by the following synapomorphies: premaxillary sagittal crest that starts at the anterior tip of the premaxilla and extends posteriorly after the occipital region, large nasoantorbital fenestra that reaches over 45% of the length between premaxilla and squamosal, lacrimal process of the jugal thin, distinct small pear- shaped orbit with lower portion narrow, and broad tubercle at the ventroposterior margin of the coracoid. Several cranial and postcranial characters indicate that the Tapejaridae are well nested within the Tapejaroidea, in sister group relationship with the Azhdarchidae. A preliminary study of the ingroup relationships within the Tapejaridae shows that Tupuxuara is more closely related to Thalassodromeus relative to Tapejara. At present tape- jarid remains have been found in the following deposits: Crato and Romualdo members of the Santana Formation (Aptian-Albian), Araripe Basin, Brazil; Jiufotang Formation (Aptian), Jehol Group of western Liaoning, China; and in the redbeds (Cenomanian) of the Kem Kem region, Morocco. An incomplete skull found in the Javelina Formation (Maastrichtian), Texas also shows several tapejarid features and might be a member of this clade. Although information is still limited, the present distribution of the Tapejaridae indicates that this clade of pterosaurs was not exclusive of Gondwana, and was more widespread than previously known. Resumen. NUEVA INFORMACIÓN SOBRE LOS TAPEJARIDAE (PTEROSAURIA, PTERODACTYLOIDEA) Y DISCUSIÓN SOBRE LAS RELACIONES DE ESTE CLADO. -
On the Osteology of Tapejara Wellnhoferi KELLNER 1989 and the first Occurrence of a Multiple Specimen Assemblage from the Santana Formation, Araripe Basin, NE-Brazil
Swiss J Palaeontol (2011) 130:277–296 DOI 10.1007/s13358-011-0024-5 On the osteology of Tapejara wellnhoferi KELLNER 1989 and the first occurrence of a multiple specimen assemblage from the Santana Formation, Araripe Basin, NE-Brazil Kristina Eck • Ross A. Elgin • Eberhard Frey Received: 28 May 2011 / Accepted: 9 August 2011 / Published online: 26 August 2011 Ó Akademie der Naturwissenschaften Schweiz (SCNAT) 2011 Abstract The postcranial elements of two similar sized ocular lobes indicate that Tapejara possessed both excel- and juvenile individuals, along with a partial skull, are lent balancing and visual systems as a consequence of its attributed to the Early Cretaceous pterosaur Tapejara aerial lifestyle. wellnhoferi. The remains, recovered from a single con- cretion of the Romualdo Member, Santana Formation, Keywords Brazil Á Lower Cretaceous Á Santana NE-Brazil, represent the first account of multiple specimens Formation Á Pterosauria Á Tapejaridae Á Osteology having settled together and allow for a complete review of postcranial osteology in tapejarid pterosaurs. A comparison Abbreviations of long bone morphometrics indicates that all specimens BSP Bayerische Staatammlung fu¨r Pala¨ontologie und attributed to the Tapejaridae for which these elements are historische Geologie, Munich, Germany known (i.e. Huaxiapterus, Sinopterus, Tapejara) display D Dalian Natural History Museum, Dalian, China similar bivariate ratios, suggesting that Chinese and Bra- IMNH Iwaki City Museum of Coal and Fossils, Iwaki, zilian taxa must have exhibited similar growth patterns. An Japan unusual pneumatic configuration, whereby the humerus is IVPP Institute for Vertebrate Palaeontology and pierced by both dorsally and ventrally located foramina, is Palaeoanthropology Beijing, P. -
New Azhdarchoid Pterosaur (Pterosauria
Anais da Academia Brasileira de Ciências (2017) 89(3 Suppl.): 2003-2012 (Annals of the Brazilian Academy of Sciences) Printed version ISSN 0001-3765 / Online version ISSN 1678-2690 http://dx.doi.org/10.1590/0001-3765201720170478 www.scielo.br/aabc | www.fb.com/aabcjournal New azhdarchoid pterosaur (Pterosauria, Pterodactyloidea) with an unusual lower jaw from the Portezuelo Formation (Upper Cretaceous), Neuquén Group, Patagonia, Argentina ALEXANDER W.A. KELLNER1 and JORGE O. CALVO2 1Laboratório de Sistemática e Tafonomia de Vertebrados Fósseis, Departamento de Geologia e Paleontologia, Museu Nacional/ Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20940-040 Rio de Janeiro, RJ, Brazil 2Grupo de Transferencia Proyecto Dino, Universidad Nacional del Comahue, Parque Natural Geo- Paleontológico Proyecto Dino, Ruta Provincial 51, Km 65, Neuquén, Argentina Manuscript received on June 22, 2017; accepted for publication on September 4, 2017 ABSTRACT A new azhdarchoid pterosaur from the Upper Cretaceous of Patagonia is described. The material consists of an incomplete edentulous lower jaw that was collected from the upper portion of the Portezuelo Formation (Turonian-Early Coniacian) at the Futalognko site, northwest of Neuquén city, Argentina. The overall morphology of Argentinadraco barrealensis gen. et sp. nov. indicates that it belongs to the Azhdarchoidea and probable represents an azhdarchid species. The occlusal surface of the anterior portion is laterally compressed and shows blunt lateral margins with a medial sulcus that are followed by two well- developed mandibular ridges, which in turn are bordered laterally by a sulcus. The posterior end of the symphysis is deeper than in any other azhdarchoid. -
Determining the Largest Known Land Animal: a Critical Comparison of Differing Methods for Restoring the Volume and Mass of Extinct Animals
ANNALS OF CARNEGIE MUSEUM VOL. 85, NUMBER 4, PP. 335–358 31 DECEMBER 2019 DETERMINING THE LARGEST KNOWN LAND ANIMAL: A CRITICAL COMPARISON OF DIFFERING METHODS FOR RESTORING THE VOLUME AND MASS OF EXTINCT ANIMALS GREGORY PAUL 3100 St. Paul Street 604, Baltimore, Maryland, 21218 [email protected] ABSTRACT Recent claims regarding what is and is not the largest known sauropod dinosaur are tested via dimensional comparisons of the most critical metrics of relative size—especially, when possible, the functional lengths of the dorsal vertebral centra and the articulated length of the combined trunk vertebrae—and analog volumetric models based on technical skeletal restorations. The Cretaceous Argentinosaurus massed 65–75 tonnes, and its dorsal vertebrae and dorsal–sacral series are much larger than those of any other described titanosaur. Specimens of Patagotitan indicate a 50–55 tonne titanosaur, and the less complete Notocolossus, Puertasaurus, and ‘Antarctosaurus’ giganteus appear to have occupied a similar size range. Paralititan weighed between 30 and 55 tonnes. The juvenile Dreadnoughtus, as well as Futalognkosaurus and Alamosaurus, were in the area of 30 tonnes, with the possibility that the last was substantially larger. Entirely analog, skillfully produced, high-anatomical-fidelity skeletal restorations and volumetric models representing a prime-lean condition are approximately as scientifically objective and accurate, as well as more realistic than, analog-digital, crudely-formed convex hull volumetric models, which are based on subjectively and often inconsistently or erroneously mounted skeletons and digitized skeletal reconstructions. The need to ensure that skeletal restorations are as anatomically correct and consistent as the data allow is stressed, which requires that researchers and illustrators be sufficiently skilled in animal and especially dinosaur anatomy, and the proce- dures and standards for achieving the best possible results are detailed. -
Ptero-Draw Activity Sheets 1.2MB .Pdf File
Trace the outline of the fossilised remains and use Pterosaurs (winged lizards) were flying reptiles that ruled your imagination to recreate your very own pterosaur. the skies in the time of the dinosaurs, 251 to 65 million years ago. Pterosaurs were not dinosaurs but were closely Congratulations, you have discovered a new related to them. They occupied every habitat, and ranged species of pterosaur! from tiny insectivores to the largest-ever flying creature. SPECIMEN DETAILS SCIENTIFIC NAME MEANING OF NAME DISCOVERED BY DIET TIME PERIOD (TICK BOX) HABITAT Triassic (251–200mya) Jurassic (200–145 mya) 0 Cretaceous (145–65 mya) SCALE Pteranodon sternbergi Quetzalcoatlus northropi Anhanguera blittersdorffi – an expert flyer (ter-ran-oh-don) (ket-sal-co-atil-us) (ann-yang-wear-ah) Meaning of name: Winged and toothless Meaning of name: Named after Aztec feathered Meaning of name: Old devil serpent god, Quetzalcoatl These large pterosaurs had a short tail, a long neck and skull, Anhanguera was a fish eating pterosaur. and a small body that palaeontologists think was partly covered This enormous pterosaur, or flying reptile, may have been with hair. Palaeontologists think that they mainly ate fish. the largest flying animal ever. Its body was quite small, but Anhanguera had long sturdy jaws, with a crest on top Pteranodon fossils have been found in areas that would once provided an anchor for its huge leathery wings, which stretched of its snout. This feature may have provided stability have been wetlands and coastal lagoons, several with fossilised from a very long fourth finger to the top of its leg. -
Evolution of Morphological Disparity in Pterosaurs Katherine C
Journal of Systematic Palaeontology, Vol. 9, Issue 3, September 2011, 337–353 Evolution of morphological disparity in pterosaurs Katherine C. Prentice, Marcello Ruta∗ and Michael J. Benton School of Earth Sciences, University of Bristol, Wills Memorial Building, Queens Road, Bristol, BS8 1RJ, UK (Received 9 November 2009; accepted 22 October 2010; printed 15 September 2011) Pterosaurs were important flying vertebrates for most of the Mesozoic, from the Late Triassic to the end of the Cretaceous (225–65 Ma). They varied enormously through time in overall size (with wing spans from about 250 mm to about 12 m), and in features of their cranial and postcranial skeletons. Comparisons of disparity based on discrete cladistic characters show that the basal paraphyletic rhamphorhynchoids (Triassic–Early Cretaceous) occupied a distinct, and relatively small, region of morphospace compared to the derived pterodactyloids (Late Jurassic–Late Cretaceous). This separation is unexpected, especially in view of common constraints on anatomy caused by the requirements of flight. Pterodactyloid disparity shifted through time, with different, small portions of morphospace occupied in the Late Jurassic and Late Cretaceous, and a much larger portion in the Early Cretaceous. This explosion in disparity after 100 Ma of evolution is matched by the highest diversity of the clade: evidently, pterosaurs express a rather ‘top heavy’ clade shape, and this is reflected in delayed morphological evolution, again an unexpected finding. The expansion of disparity among pterodactyloids was comparable across subclades: pairwise comparisons among the four pterodactyloid superfamilies show that, for the most part, these clades display significant morphological separation, except in the case of Dsungaripteroidea and Azhdarchoidea.