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A New Caenagnathid Dinosaur from the Upper Cretaceous Wangshi
www.nature.com/scientificreports OPEN A new caenagnathid dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with Received: 12 October 2017 Accepted: 7 March 2018 comments on size variation among Published: xx xx xxxx oviraptorosaurs Yilun Yu1, Kebai Wang2, Shuqing Chen2, Corwin Sullivan3,4, Shuo Wang 5,6, Peiye Wang2 & Xing Xu7 The bone-beds of the Upper Cretaceous Wangshi Group in Zhucheng, Shandong, China are rich in fossil remains of the gigantic hadrosaurid Shantungosaurus. Here we report a new oviraptorosaur, Anomalipes zhaoi gen. et sp. nov., based on a recently collected specimen comprising a partial left hindlimb from the Kugou Locality in Zhucheng. This specimen’s systematic position was assessed by three numerical cladistic analyses based on recently published theropod phylogenetic datasets, with the inclusion of several new characters. Anomalipes zhaoi difers from other known caenagnathids in having a unique combination of features: femoral head anteroposteriorly narrow and with signifcant posterior orientation; accessory trochanter low and confuent with lesser trochanter; lateral ridge present on femoral lateral surface; weak fourth trochanter present; metatarsal III with triangular proximal articular surface, prominent anterior fange near proximal end, highly asymmetrical hemicondyles, and longitudinal groove on distal articular surface; and ungual of pedal digit II with lateral collateral groove deeper and more dorsally located than medial groove. The holotype of Anomalipes zhaoi is smaller than is typical for Caenagnathidae but larger than is typical for the other major oviraptorosaurian subclade, Oviraptoridae. Size comparisons among oviraptorisaurians show that the Caenagnathidae vary much more widely in size than the Oviraptoridae. Oviraptorosauria is a clade of maniraptoran theropod dinosaurs characterized by a short, high skull, long neck and short tail. -
Postcranial Anatomy of Tanius Sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Postkraniala Anatomin Hos Tanius Sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea)
Examensarbete vid Institutionen för geovetenskaper Degree Project at the Department of Earth Sciences ISSN 1650-6553 Nr 320 Postcranial Anatomy of Tanius Sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Postkraniala anatomin hos Tanius sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Niclas H. Borinder INSTITUTIONEN FÖR GEOVETENSKAPER DEPARTMENT OF EARTH SCIENCES Examensarbete vid Institutionen för geovetenskaper Degree Project at the Department of Earth Sciences ISSN 1650-6553 Nr 320 Postcranial Anatomy of Tanius Sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Postkraniala anatomin hos Tanius sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Niclas H. Borinder ISSN 1650-6553 Copyright © Niclas H. Borinder and the Department of Earth Sciences, Uppsala University Published at Department of Earth Sciences, Uppsala University (www.geo.uu.se), Uppsala, 2015 Abstract Postcranial Anatomy of Tanius Sinensis Wiman, 1929 (Dinosauria; Hadrosauroidea) Niclas H. Borinder Tanius sinensis Wiman, 1929 was one of the first hadrosauroid or “duck-billed” taxa erected from China, indeed one of the very first non-avian dinosaur taxa to be erected based on material from the country. Since the original description by Wiman in 1929, the anatomy of T. sinensis has received relatively little attention in the literature since then. This is unfortunate given the importance of T. sinensis as a possible non-hadrosaurid hadrosauroid i.e. a member of Hadrosauroidea outside the family of Hadrosauridae, living in the Late Cretaceous, at a time when most non-hadrosaurid hadro- sauroids had become replaced by the members of Hadrosauridae. To gain a better understanding of the anatomy of T. sinensis and its phylogenetic relationships, the postcranial anatomy of it is redescribed. T. sinensis is found to have a mosaic of basal traits like strongly opisthocoelous cervical vertebrae, the proximal end of scapula being dorsoventrally wider than the distal end, the ratio between the proximodistal length of the metatarsal III and the mediolateral width of this element being greater than 4.5. -
Immigrant Species, Or Native Species?
The Journal of Paleontological Sciences: JPS.C.2017.01 TESTING THE HYPOTHESES OF THE ORIGIN OF TYRANNOSAURUS REX: IMMIGRANT SPECIES, OR NATIVE SPECIES? __________________________________________________________________________________________________________________ Chan-gyu Yun Vertebrate Paleontological Institute of Incheon, Incheon 21974, Republic of Korea & Biological Sciences, Inha University, Incheon 22212, Republic of Korea [email protected] __________________________________________________________________________________________________________________ Abstract: It is an undoubtable fact that Tyrannosaurus rex is the most iconic dinosaur species of all time. However, it is currently debatable whether this species has a North American origin or Asian origin. In this paper, I test these two hypotheses based on current fossil records and former phylogenetic analyses. Phylogenetic and fossil evidence, such as derived tyrannosaurine fossils of Asia, suggests that the hypothesis of an Asian origin of Tyrannosaurus rex is the most plausible one, but this is yet to be certain due to the scarcity of fossil records. INTRODUCTION The most famous and iconic dinosaur of all time, Tyrannosaurus rex, is only known from upper Maastrichtian geological formations in Western North America (e.g. Carr and Williamson, 2004; Larson, 2008). However, older relatives of Tyrannosaurus rex (e.g. Daspletosaurus, Tarbosaurus) are known from both Asia and North America. This leads to an evolutionary question: is the origin of Tyrannosaurus rex from Asia, or North America? About six of the currently valid tyrannosaurine taxa were described in the twenty-first century (based on parsimony analysis of Brusatte and Carr, 2016), with new species which are being described (Sebastian Dalman, Pers. Comm., 2016; Thomas Carr, Pers. Comm., 2016). It can be said that "now" is the "golden age” for studying tyrannosaurine evolution. -
New Turtle Egg Fossil from the Upper Cretaceous of the Laiyang Basin, Shandong Province, China
Anais da Academia Brasileira de Ciências (2013) 85(1): 103-111 (Annals of the Brazilian Academy of Sciences) Printed version ISSN 0001-3765 / Online version ISSN 1678-2690 www.scielo.br/aabc New turtle egg fossil from the Upper Cretaceous of the Laiyang Basin, Shandong Province, China 1 1 1 1, 2 1, 2 QIANG WANG , XIAOLIN WANG , ZIKUI ZHAO , JIALIANG ZHANG and SHUNXING JIANG 1Key Laboratory of Vertebrate Evolution and Human Origin of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, 100044, China 2University of Chinese Academy of Sciences, Beijing, 100049, China Manuscript received on June 11, 2012; accepted for publication on November 23, 2012 ABSTRACT A new type of turtle egg fossil was established: Emydoolithus laiyangensis oogen. et oosp. nov.. Based on its elliptical morphological shape, rigid eggshells, and eggshell characteristics, it is different from other types of round chelonian egg fossils. It is the second chelonian egg fossil found in Cretaceous in China. This discovery shows the Laiyang ecosystem in Late Cretaceous is more diversified than previously thought. Key words: Turtle egg fossil, Jingangkou Formation, Late Cretaceous, Laiyang Basin, Shandong Province. INTRODUCTION and Ozaki 1929), they were later recognized Turtle egg fossils are very rare. Even though they as dinosaur eggs based on their macrostructure are found from the Jurassic to the Tertiary age, turtle characteristics (Chow 1954). Recently, we have eggs are known only from a few Cretaceous deposits, restudied the macrostructure and microstructure of from localities in China (Fang et al. 2003, Jackson et these eggs found in Changtu (housed at the Dalian al. -
Caenagnathids of the Dinosaur Park Formation (Campanian) of Alberta, Canada: Anatomy, Osteo- Histology, Taxonomy, and Evolution Gregory F
Vertebrate Anatomy Morphology Palaeontology 8:105–153 105 ISSN 2292-1389 Caenagnathids of the Dinosaur Park Formation (Campanian) of Alberta, Canada: anatomy, osteo- histology, taxonomy, and evolution Gregory F. Funston School of GeoSciences, University of Edinburgh, Edinburgh, Scotland, UK and Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada; [email protected] Abstract: Our understanding of caenagnathid anatomy, diversity, and ecology has improved considerably in the past twenty years, but numerous issues still remain. Among these, the diversity and taxonomy of caenag- nathids from the Dinosaur Park Formation of Alberta, Canada, have remained problematic. Whereas some authors recognize three genera, others suggest only two are present, and there is considerable disagreement about which specimens are referable to which genus. This study aims to resolve this issue by reviewing the known specimens and using osteohistology to establish a testable taxonomic framework of Dinosaur Park Formation caenagnathids. Numerous new specimens from all regions of the skeleton provide insight into morphological variation in caenagnathids, and three morphotypes are recognized based on a combination of morphological features and body size. Osteohistology shows that representatives in each body size class are at skeletal maturity, and therefore supports the delineation of three taxa: the smaller Citipes elegans gen. nov., the intermediate Chirostenotes pergracilis, and the larger Caenagnathus collinsi, new material of which shows it rivalled Anzu wyliei in size. However, these analyses also raise concerns about the referral of isolat- ed material to each taxon in the absence of skeletal overlap between specimens or osteohistological analysis. Caenagnathids are consistently recovered throughout the Dinosaur Park Formation interval, and two geo- graphic clusters of increased abundance probably reflect collection and taphonomic biases. -
Review of Historical and Current Research on the Late Cretaceous
第55卷 第2期 古 脊 椎 动 物 学 报 pp. 187-200 2017年4月 VERTEBRATA PALASIATICA figs. 1-5 Review of historical and current research on the Late Cretaceous dinosaurs and dinosaur eggs from Laiyang, Shandong ZHANG Jia-Liang1,2 WANG Qiang1 JIANG Shun-Xing1 CHENG Xin1 LI Ning1 QIU Rui1,2 ZHANG Xin-Jun1,2 WANG Xiao-Lin1,2* (1 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences Beijing 100044 * Corresponding author: [email protected]) (2 University of Chinese Academy of Sciences Beijing 100049) Abstract Here we briefly review the history of research on the Laiyang dinosaur and dinosaur egg faunas, summarize the contributions of C. C. Young and other elder paleontologists to the discoveries of the Late Cretaceous dinosaurs and dinosaur eggs from Laiyang, and introduce the new discoveries and the advances in the research on the Laiyang faunas. The new investigations in Laiyang from 2008 have found a series of valley developed in plain and more than ten new dinosaur or dinosaur egg fossil beds. In 2010, we began the massive excavations at two localities in Jingangkou and collected abundant dinosaurs and other vertebrate fossils, such as a new saurolophine, some theropod material and a new taxon of turtle egg. The bone beds in Locality 2 with the typical sedimentary and burial patterns of mudflow, and these fossil deposits are interpreted as having been carried and deposited by mudflow. The re-observation and the CT scanning data of the crest of Tsintaosaurus spinorhinus (IVPP V 725) show that the crest is fractured and solid. -
Chinaxiv:201711.01905V1
ChinaXiv合作期刊 第55卷 第2期 古 脊 椎 动 物 学 报 pp. 187-200 2017年4月 VERTEBRATA PALASIATICA figs. 1-5 Review of historical and current research on the Late Cretaceous dinosaurs and dinosaur eggs from Laiyang, Shandong ZHANG Jia-Liang1,2 WANG Qiang1 JIANG Shun-Xing1 CHENG Xin1 LI Ning1 QIU Rui1,2 ZHANG Xin-Jun1,2 WANG Xiao-Lin1,2* (1 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences Beijing 100044 * Corresponding author: [email protected]) (2 University of Chinese Academy of Sciences Beijing 100049) Abstract Here we briefly review the history of research on the Laiyang dinosaur and dinosaur egg faunas, summarize the contributions of C. C. Young and other elder paleontologists to the discoveries of the Late Cretaceous dinosaurs and dinosaur eggs from Laiyang, and introduce the new discoveries and the advances in the research on the Laiyang faunas. The new investigations in Laiyang from 2008 have found a series of valley developed in plain and more than ten new dinosaur or dinosaur egg fossil beds. In 2010, we began the massive excavations at two localities in Jingangkou and collected abundant dinosaurs and other vertebrate fossils, such as a new saurolophine, some theropod material and a new taxon of turtle egg. The bone beds in Locality 2 with the typical sedimentary and burial patterns of mudflow, and these fossil deposits are interpreted as having been carried and deposited by mudflow. The re-observation and the CT scanning data of the crest of Tsintaosaurus spinorhinus (IVPP V 725) show that the crest is fractured and solid. -
New Ootype Prismatoolithids from the Late Cretaceous, Laiyang Basin
第56卷 第3期 古 脊 椎 动 物 学 报 pp. 264–272 2018年7月 VERTEBRATA PALASIATICA figs. 1–3 DOI: 10.19615/j.cnki.1000-3118.180708 New ootype prismatoolithids from the Late Cretaceous, Laiyang Basin and its significance WANG Qiang1,2 LI Yu-Guang3 ZHU Xu-Feng1,2,4 FANG Kai-Yong1,5 WANG Xiao-Lin1,2,4 (1 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences Beijing 100044) (2 CAS Center for Excellence in Life and Paleoenvironment Beijing 100044 [email protected]) (3 Capital Normal University Beijing 100048) (4 University of Chinese Academy of Sciences Beijing 100049) (5 School of Earth Sciences and Resources, China University of Geosciences, Beijing Beijing 100083) Abstract Here we discribe a new ootypes of prismatoolithids found from the Late Cretaceous Jiangjunding Formation in the Laiyang Basin, Shandong Province. On the basis of characters such as elongate ovoids, smooth eggshell surface, relatively thinner eggshell thickness, prismatic eggshell units, slender pores in radial section of eggshell and little pores with round or irregular shapes in tangeital section of eggshell, we erect one new oogenus and one new oospecies: Laiyangoolithus lixiangensis oogen. et oosp. nov. The discovery of L. lixiangensis not only enriches the diversity and composition of the Laiyang Dinosaur Egg Fauna, but the paleogeographic distribution of primatoolithids as well. In addition, it provides more paleontological materials for the study of the diversity and paleogeographic distribution of troodontids in China. Key words Laiyang, Shandong; Late Cretaceous; Jiangjunding Formation; dinosaur eggs, Prismatoolithidae Citation Wang Q, Li Y G, Zhu X F et al., 2018. -
(J50E021024), Nancun Town (J51E021001), Gaomi City (J50E022024) and Lancun Town (J51E022001) Map-Sheets, Shandong Province[J]
Vol. 47 Supp.(1) GEOLOGY IN CHINA Jun., 2020 doi: 10.12029/gc2020Z115 Received: 05-04-2020 Article Citation: Xu Weixiang, Zhang Deming, Gao Meixia, Tang Wenlong, Hu Bing, Zhang Kai. 2020. Spatial Accepted: 03-05-2020 Database of 1∶50 000 Digital Geological Maps of Wanjia Town (J50E021024), Nancun Town (J51E021001), Gaomi City (J50E022024) and Lancun Town (J51E022001) Map-sheets, Shandong Province[J]. Geology in China, Fund Project: 47(S1):226−242. a project titled 1∶50 000- Scale Regional Geological Dataset Citation: Xu Weixiang; Gao Meixia; Zhang Deming; Hu Bing; Zhang Kai. Spatial Database of 1∶50 000 Survey of Wanjia Town, Digital Geological Maps of Wanjia Town (J50E021024), Nancun Town (J51E021001), Gaomi City (J50E022024) and Nancun Town, Gaomi City Lancun Town (J51E022001) Map-sheets, Shandong Province(V1). Shandong Provincial No.4 Institute of Geological and Lancun Town Map and Mineral Survey; Key Laboratory of Coastal Zone Geological Environment Protection, Shandong Provincial Sheets, Shandong Province Bureau of Geology & Mineral Resources; Tianjin Center, China Geological Survey; Weifang Foundation Engineering initiated by the China Geolo- Corporation, Shandong[producer], 2018. National Geological Archives of China[distributor], 2020-06-30. 10.35080/ gical Survey (No.: DD2016 data.A.2020.P15; http://dcc.cgs.gov.cn/en//geologicalData/details/doi/10.35080/data.A.2020.P15. 0044-4) Spatial Database of 1∶50 000 Digital Geological Maps of Wanjia Town (J50E021024), Nancun Town (J51E021001), Gaomi City (J50E022024) and Lancun Town (J51E022001) Map-sheets, Shandong Province XU Weixiang 1,2, ZHANG Deming 1,2, GAO Meixia 1,2, TANG Wenlong 3, HU Bing 1,2, ZHANG Kai 4 (1. -
This Article Appeared in a Journal Published by Elsevier. the Attached
This article appeared in a journal published by Elsevier. The attached copy is furnished to the author for internal non-commercial research and education use, including for instruction at the authors institution and sharing with colleagues. Other uses, including reproduction and distribution, or selling or licensing copies, or posting to personal, institutional or third party websites are prohibited. In most cases authors are permitted to post their version of the article (e.g. in Word or Tex form) to their personal website or institutional repository. Authors requiring further information regarding Elsevier’s archiving and manuscript policies are encouraged to visit: http://www.elsevier.com/authorsrights Author's personal copy Disponible en ligne sur www.sciencedirect.com Annales de Paléontologie 99 (2013) 243–259 Original article Lindholmemydid turtles (Cryptodira: Testudinoidea) from the Late Cretaceous of Shandong Province, China Tortues Lindholmemydidés (Cryptodira: Testudinoidea) du Crétacé supérieur de la province de Shandong, Chine a a,b,∗ c c a Lu Li , Haiyan Tong , Kebai Wang , Shuqing Chen , Xing Xu a Key Laboratory of Vertebrate Evolution and Human Origin of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China b Palaeontological Research and Education Centre, Mahasarakham University, Kantarawichai, Mahasarakham 44150, Thailand c Zhucheng Dinosaur Museum, Bureau of Tourism, Zhucheng, Shandong, China Available online 27 August 2013 Abstract A new genus and new species of lindholmemydid turtle (Cryptodira: Testudinoidea), Shandongemys dongwuica n. g. and n. sp. are described on the basis of a partial skeleton with incomplete shell and skull, complete lower jaws and disarticulated limb bones from the Upper Cretaceous Wangshi Group of Zhucheng, Shandong Province, China. -
(Jiaodong Peninsula): New Insights on Late Mesozoic Continental Extension in Eastern Asia
Palaeomagnetic constraints from granodioritic plutons (Jiaodong Peninsula): New insights on Late Mesozoic continental extension in eastern Asia. Nicolas Charles, Yan Chen, Romain Augier, Charles Gumiaux, Wei Lin, Michel Faure, Patrick Monié, Flavien Choulet, Fuyuan Wu, Rixiang Zhu, et al. To cite this version: Nicolas Charles, Yan Chen, Romain Augier, Charles Gumiaux, Wei Lin, et al.. Palaeomagnetic con- straints from granodioritic plutons (Jiaodong Peninsula): New insights on Late Mesozoic continental extension in eastern Asia.. Physics of the Earth and Planetary Interiors, Elsevier, 2011, 187 (3-4), pp.276-291. 10.1016/j.pepi.2011.05.006. insu-00599546 HAL Id: insu-00599546 https://hal-insu.archives-ouvertes.fr/insu-00599546 Submitted on 10 Jun 2011 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. 1 Palaeomagnetic constraints from granodioritic plutons (Jiaodong Peninsula): New 2 insights on Late Mesozoic continental extension in eastern Asia. 3 4 Nicolas CHARLES 1,* , Yan CHEN 1, Romain AUGIER 1, Charles GUMIAUX 1, Wei LIN 2, 5 Michel FAURE 1, Patrick MONIÉ 3, Flavien CHOULET 1, Fuyuan WU 2, Rixiang ZHU 2, 6 Qingchen WANG 2. 7 8 1. Institut des Sciences de la Terre d’Orléans (ISTO) 9 Université d’Orléans-CNRS/INSU (UMR6113) 10 1A, rue de la Férollerie 11 45071 Orléans cedex 2 12 France 13 2. -
Internal Morphology of Nasal Spine of Tsintaosaurus Spinorhinus (Ornithischia: Lambeosaurinae) from the Upper Cretaceous of Shandong, China
Historical Biology An International Journal of Paleobiology ISSN: 0891-2963 (Print) 1029-2381 (Online) Journal homepage: https://www.tandfonline.com/loi/ghbi20 Internal morphology of nasal spine of Tsintaosaurus spinorhinus (Ornithischia: Lambeosaurinae) from the upper cretaceous of Shandong, China Jialiang Zhang, Xiaolin Wang, Shunxing Jiang & Guobiao Li To cite this article: Jialiang Zhang, Xiaolin Wang, Shunxing Jiang & Guobiao Li (2020): Internal morphology of nasal spine of Tsintaosaurusspinorhinus (Ornithischia: Lambeosaurinae) from the upper cretaceous of Shandong, China, Historical Biology, DOI: 10.1080/08912963.2020.1731804 To link to this article: https://doi.org/10.1080/08912963.2020.1731804 Published online: 19 Mar 2020. Submit your article to this journal View related articles View Crossmark data Full Terms & Conditions of access and use can be found at https://www.tandfonline.com/action/journalInformation?journalCode=ghbi20 HISTORICAL BIOLOGY https://doi.org/10.1080/08912963.2020.1731804 ARTICLE Internal morphology of nasal spine of Tsintaosaurus spinorhinus (Ornithischia: Lambeosaurinae) from the upper cretaceous of Shandong, China Jialiang Zhanga,b, Xiaolin Wanga,c,d, Shunxing Jianga,c and Guobiao Lib aKey Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China; bSchool of the Earth Sciences and Resources, China University of Geosciences (Beijing), Beijing, China; cCAS Center for Excellence in Life and Paleoenvironment, Beijing, China; dCollege of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing, China ABSTRACT ARTICLE HISTORY Tsintaosaurus spinorhinus is famous for the unique rod-like nasal spine, but there has been a long debate about Received 13 November 2019 whether its nasal spine is hollow, since it is difficult to examine the nasal inside through direct observation.