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ISSN 1514-4186 INSTITUTO SUPERIOR DE CORRELACIÓN GEOLÓGICA (INSUGEO) El Neógeno de Argentina Editores: F. G. Aceñolaza INSUGEO - Facultad de Ciencias Naturales e Instituto Miguel Lillo Universidad Nacional de Tucumán Miguel Lillo 205, 4000 Tucumán, Argentina y R. Herbst INSUGEO - Facultad de Ciencias Naturales e Instituto Miguel Lillo Universidad Nacional de Tucumán Miguel Lillo 205, 4000 Tucumán, Argentina Consejo Nacional de Investigaciones Científicas y Técnicas Facultad de Ciencias Naturales e Instituto Miguel Lillo Universidad Nacional de Tucumán 2000 4 CORRELACIÓN GEOLÓGICA Nº 14 Consejo Nacional de Investigaciones Científicas y Técnicas Universidad Nacional de Tucumán Instituto Superior de Correlación Geológica (INSUGEO) Director Dr. Florencio G. Aceñolaza Directores Alternos Dr. Alejandro Toselli Dr. Alfredo Tineo Consejo Editorial Dr. Florencio Aceñolaza Dr. Guillermo Aceñolaza Dr. Luis Buatois Dra. Susana Esteban Dr. Juan Carlos Gutiérrez Marco (España) Dr. Rafael Herbst Dr. Roberto Lech Dra. María Gabriela Mángano Dr. Hübert Miller (Alemania) Dra. Isabel Rábano (España) Dra. Juana Rossi de Toselli Dr. Julio Saavedra Alonso (España) Dr. Alcides N. Sial (Brasil) Dr. Alfredo Tinello Dr. Alejandro Toselli Dr. Franco Tortello Instituto Superior de Correlación Geológica Miguel Lillo 205 4000 - San Miguel de Tucumán REPÚBLICA ARGENTINA e-mail: [email protected] Tapa: Cartera Cristamine, Aldea Brasilera, Entre Ríos, afloramientos de Formación Paraná, donde se explo- tan niveles de arenas medanosas. CORRELACIÓN GEOLÓGICA Nº 14 5 Indice Prólogo ........................................................................................................................................ 7 Aceñolaza, F. G. La Formación Paraná (Mioceno Medio): estratigrafía, distribución regional y unidades equivalentes........................................................................ 9 Marengo, H. G. Rasgos micropaleontológicos de los depósitos de la transgresión Entrerriense-Paranense en la Cuenca Chaco-Paranense y Noroeste Argentino, República Argentina. ............................................................................................. 29 Sprechman, P., Fernando, L. A. y Martínez, S. Estado actual de los conocimientos sobre la Formación Camacho (Mioceno?-Superior?), Uruguay. ............ 47 Zucol, A. y Brea, M. Análisis fitolítico de la Formación Paraná en la Provincia de Entre Ríos. ........................................................................................................... 67 Del Río, C. Malacofauna de las formaciones Paraná y Puerto Madryn (Mioceno marino, Argentina): su origen, composición y .................................................... 77 significado bioestratigráfico. Barreda, V. y Palamarczuck, S. Estudio palinoestratigráfico del Oligoceno tardío-Mioceno en secciones de la costa patagónica y plataforma continental argentina. ................................................................................................................. 103 Fernández Garrasino, C. y Vrba, Ana. La Formación Paraná: aspectos estratigráficos y estructurales de la región chacoparanense. ............................................... 139 Lech, R., Aceñolaza, F. y Grizinik, M. Icnofacies Skolithos- Ophiomorpha en el Neogeno del Valle inferior del Río Chubut, provincia de Chubut, Argentina. ............................................................................................. 147 Alonso, R. El Terciario de la Puna en tiempos de la ingresión marina paranense. ....................................................................................................................... 163 Herbst, R. La Formación Ituzaingó (Plioceno). Estratigrafía y distribución. ................ 181 Cione, A. L., Azpelicueta, M. M., Bond, M., Carlini, A. A., Casciotta, J. R., Cozzuol, M. A., de la Fuente, M., Gasparini, Z., Goin, F. J., Noriega, J., Scillato-Yané, G. J., Zoibelzon, L., Tonni, E. P., Verzi, D. y Vucetich, M. G. Miocene vertebrates from Entre Ríos province, Argentine. .............................................. 191 Argañaraz, B. y Piña, C. Quelonios de la Formación Ituzaingó (Mioceno superior-Plioceno), en Villa Urquiza, Entre Ríos, Argentina. .......................... 239 6 CORRELACIÓN GEOLÓGICA Nº 14 Brea, M. y Zucol, A. Lignoflora del Cenozoico superior del Noroeste Argentino. ........................................................................................................... 245 Piña, C. y Argañaraz, B. Presencia del género Caiman (Crocodylia: Alligatoridae) en la Formación Ituzaingó (Mioceno Superior Plioceno), Entre Ríos, Argentina .......................................................... 255 Herbst, R., Anzótegui, L. M., Estéban, G. Mautino, L. R., Morton, S. y Nasif, N. Síntesis paleontológica del Mioceno de los valles Calchaquíes, noroeste argentino. .............................................................................................. 263 Instrucciones a los autores ........................................................................................................ 289 CORRELACIÓN GEOLÓGICA Nº 14 7 Prólogo Los depósitos del Neógeno tienen una muy amplia distitribución en el territorio argentino, pudiendo decirse que el debate de sus componentes desde una visión integradora regional re- cién está comenzando. Ocurre que quienes han abordado el estudio de esta parte de la columna geológica normalmente lo han hecho desde la óptica de cada especialidad motivo por el cual muchas veces se han brindado conceptos o interpretaciones distorsionadas. Con el objeto de iniciar un debate sobre este tema, en noviembre de 1999 llevamos adelante una reunión de expertos en el Centro de Investigaciones del CONICET radicado en la ciudad de Diamante (Entre Rios). Para ello se hizo adelante un análisis comparativo y de correlación geológica tomando en consideración los sedimentos marinos y continentales que afloran en la región mesopotámica y del Uruguay. A partir de allí ,con los trabajos expuestos más otros que fueron invitados, iniciamos la tarea de compilar una serie de artículos destinados a compatibilizar las ideas sobre esta temática. El resultado queda expuesto en la presente obra y, naturalmente, los lectores encontrarán muchos puntos de concordancia pero también de discrepancia entre los autores, situación que no nos alarma ya que interpretamos que ello servirá como punto de partida para nuevas discusiones y debates. Desde luego también, que los artículos aquí presentados de ninguna manera agotan ni los temas ni las opiniones. La obra comprende trabajos referidos a la geología regional, paleontología y paleoambientes sobre las formaciones Paraná e Ituzaingó, ambas objeto del debate mencionado. Como interesante ejemplo no queremos dejar pasar por alto la circunstancia que la International Stratigraphic Commission (ISC) y la International Union of Geological Sciences (IUGS), luego de varios años de debate, ambiciosamente definieron la Global Stratigraphic Section and Point (GSSP) en el metro 35 de la sección de Lemme-Carosio, en la provincia de Alexandría, cuenca del Pimonte en el norte de Italia. Allí no solo se fijaron las pautas bioestratigráficas sino que también las resultantes de la magnetoestratigrafía y geología isotópica (Steininger et al, 1997). Asimismo, entre los criterios están, por un lado, los que surgen de la información del nannoplacton y foraminíferos, sino que se los vincula con los que proporcio- nan los vertebrados terrestres, y estudios isotópicos. La precisión lograda en tan arduos debates permitieron lograr, para el continente europeo un alto grado de certidumbre que luego expan- dieron hacia Asia y Norteaérica. El ejemplo del Neógeno holártico, debería ser un desafío para comenzar una tarea similar en el hemisferio sur, que permita calibrar en un futuro no muy lejano nuestras secuencias sedimentarias junto a las de Antártida, Australia y Africa, con aquellas. Es nuestra esperanza que lo expuesto en esta obra sea un comienzo y sirva para potenciar el interés de los colegas para lograrlo. F. G. Aceñolaza y R. Herbst Editores En: Aceñolaza F. G. y Herbst R. (Eds.). INSUGEO, Serie Correlación Geológica, 14: 9-27 El Neógeno de Argentina Tucumán, 2000 - ISSN 1514-4186 La Formación Paraná (Mioceno medio) : estratigrafia, distribucion regional y unidades equivalentes. Florencio G. ACEÑOLAZA1 Abstract.- THE PARANÁ FORMATION (MIDDLE MIOCENE), STRATIGRAPHY AND REGIONAL DISTRIBUTION: A regional- stratigraphical analysis for the Paraná Formation (Middle Miocene) is presented. This is based on historical data in the type localitiy and some other new sections done on several localities of Argentina. The Paraná Formation was deposited during an important transgression over the pampean and patagonian regions, close to the end of the Lower to Middle Miocene. The transgression covered most of central to eastern Argentina, reaching as far as the Andean and Patagonian border. Lime- stones, sandy to argillaceous facies with abundant fossils (molluscs, echinoids, trace fossils, foraminifera, conchostraca, fish and others marine vertebrates etc.) were developed, along the Uruguayan border to the westwards Entre Rios province, in Argentina. In several localities (El Cerro and Paraná city, Molino Doll, etc) the limestone levels do have reef barrier type structure. Some others do present sandy barriers (Cristamine quarry). The fauna and sedimentological characters have been interpreted to be representative of temperate to warm waters and show correlation with outocrops in Patagonia . Key Words: Miocene.
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    Genus/Species Skull Ht Lt Wt Stage Range Abalosia U.Pliocene S America Abelmoschomys U.Miocene E USA A. simpsoni U.Miocene Florida(US) Abra see Ochotona Abrana see Ochotona Abrocoma U.Miocene-Recent Peru A. oblativa 60 cm? U.Holocene Peru Abromys see Perognathus Abrosomys L.Eocene Asia Abrothrix U.Pleistocene-Recent Argentina A. illuteus living Mouse Lujanian-Recent Tucuman(ARG) Abudhabia U.Miocene Asia Acanthion see Hystrix A. brachyura see Hystrix brachyura Acanthomys see Acomys or Tokudaia or Rattus Acarechimys L-M.Miocene Argentina A. minutissimus Miocene Argentina Acaremys U.Oligocene-L.Miocene Argentina A. cf. Murinus Colhuehuapian Chubut(ARG) A. karaikensis Miocene? Argentina A. messor Miocene? Argentina A. minutissimus see Acarechimys minutissimus Argentina A. minutus Miocene? Argentina A. murinus Miocene? Argentina A. sp. L.Miocene Argentina A. tricarinatus Miocene? Argentina Acodon see Akodon A. angustidens see Akodon angustidens Pleistocene Brazil A. clivigenis see Akodon clivigenis Pleistocene Brazil A. internus see Akodon internus Pleistocene Argentina Acomys L.Pliocene-Recent Africa,Europe,W Asia,Crete A. cahirinus living Spiny Mouse U.Pleistocene-Recent Israel A. gaudryi U.Miocene? Greece Aconaemys see Pithanotomys A. fuscus Pliocene-Recent Argentina A. f. fossilis see Aconaemys fuscus Pliocene Argentina Acondemys see Pithanotomys Acritoparamys U.Paleocene-M.Eocene W USA,Asia A. atavus see Paramys atavus A. atwateri Wasatchian W USA A. cf. Francesi Clarkforkian Wyoming(US) A. francesi(francesci) Wasatchian-Bridgerian Wyoming(US) A. wyomingensis Bridgerian Wyoming(US) Acrorhizomys see Clethrionomys Actenomys L.Pliocene-L.Pleistocene Argentina A. maximus Pliocene Argentina Adelomyarion U.Oligocene France A. vireti U.Oligocene France Adelomys U.Eocene France A.
  • Short Review of Dental Microstructure and Dental Microwear in Xenarthran Teeth

    Short Review of Dental Microstructure and Dental Microwear in Xenarthran Teeth

    231 Short review of dental microstructure and dental microwear in xenarthran teeth Daniela C. Kalthoff Introduction Dentin is one of three tooth tissues occurring in mammals, Also using SEM technique, the most comprehensive and the other two being the hypermineralized tooth enamel detailed histological study to date is from Kalthoff (2011), and tooth cementum. In respect to volume, dentin usu- comprising teeth from about 50 xenarthran taxa dating ally makes up the largest part of the tooth. Interestingly, from late Eocene to Holocene. a number of mammal taxa have reduced the resistant Dental microwear analysis is the study of microscopic layer of tooth enamel during their evolutionary history, scars on the chewing surface of cheek teeth. Last consumed thus focusing on dentin as the main tooth building tissue. food items (and potentially also exogenous grit; Hoffman Enamelless teeth can be found in various extant and ex- et al. 2015) leave these scars, mainly different kinds of tinct taxa of odontocete whales (Odontoceti), aardvarks pits and scratches, during mastication and different food (Tubulidentata), armadillos, glyptodonts and pampatheres categories (leaves, grass, seeds, fruits, meat, insects, (Cingulata), sloths (Folivora), walruses (Odobenidae), sea etc.) produce different kinds of scars. Extant species with cows (Dugongidae), and elephants (Proboscidea). known diets are used to calibrate microwear scars. From This review will give an overview of recent results on the beginning of dental microwear analysis, the highly histology as well as dietary adaptations using the distinctly mineralized enamel had been the target tissue, and this built dentin teeth in armadillos and sloths and their fossil technique has been employed to various mammal clades relatives (in the following referred to with the informal term (e.
  • Fossils from India (Late Pliocene)

    Fossils from India (Late Pliocene)

    RESEARCH ARTICLE The first darter (Aves: Anhingidae) fossils from India (late Pliocene) Thomas Stidham1*, Rajeev Patnaik2, Kewal Krishan3, Bahadur Singh3, Abhik Ghosh3, Ankita Singla2, Simran S. Kotla2 1 Key Laboratory of Vertebrate Evolution and Human Origins of the Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China, 2 Department of Geology, Panjab University, Chandigarh, India, 3 Department of Anthropology, Panjab University, Chandigarh, India a1111111111 * [email protected] a1111111111 a1111111111 a1111111111 a1111111111 Abstract New fossils from the latest Pliocene portion of the Tatrot Formation exposed in the Siwalik Hills of northern India represent the first fossil record of a darter (Anhingidae) from India. The darter fossils possibly represent a new species, but the limited information on the fossil OPEN ACCESS record of this group restricts their taxonomic allocation. The Pliocene darter has a deep pit Citation: Stidham T, Patnaik R, Krishan K, Singh B, on the distal face of metatarsal trochlea IV not reported in other anhingids, it has an open Ghosh A, Singla A, et al. (2017) The first darter groove for the m. flexor perforatus et perforans digiti II tendon on the hypotarsus unlike New (Aves: Anhingidae) fossils from India (late World anhingid taxa, and these darter specimens are the youngest of the handful of Neo- Pliocene). PLoS ONE 12(5): e0177129. https://doi. org/10.1371/journal.pone.0177129 gene records of the group from Asia. These fossil specimens begin to fill in a significant geo- graphic and temporal gap in the fossil record of this group that is largely known from other Editor: Thierry Smith, Royal Belgian Institute of Natural Sciences, BELGIUM continents and other time periods.