ZOOTAXA 1102 Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia)—2nd edition RICHARD E. PETIT & M.G. HARASEWYCH Magnolia Press Auckland, New Zealand RICHARD E. PETIT & M.G. HARASEWYCH Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia)—2nd edition (Zootaxa 1102) 161 pp.; 30 cm. 19 Dec. 2005 ISBN 1-877407-38-0 (paperback) ISBN 1-877407-39-9 (Online edition) FIRST PUBLISHED IN 2005 BY Magnolia Press P.O. Box 41383 Auckland 1030 New Zealand e-mail: [email protected] http://www.mapress.com/zootaxa/ © 2005 Magnolia Press All rights reserved. No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any means, without prior written permission from the publisher, to whom all requests to reproduce copyright material should be directed in writing. This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose other than private research use. ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition) Zootaxa 1102: 1–161 (2005) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA 1102 Copyright © 2005 Magnolia Press ISSN 1175-5334 (online edition) Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia)—2nd edition RICHARD E. PETIT1 & M.G. HARASEWYCH2 1806 St. Charles Road, North Myrtle Beach, SC 29582-2846, USA; email: [email protected] 2 Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution,Washing- ton, DC 20013-7012, USA; email: [email protected] Table of contents Abstract . 3 Introduction . 4 Conventions used in the genus-group list . 6 Conventions used in the species-group list . 6 Notes on authors and publications . 7 World Wide Web—(Internet) . 8 Acknowledgements . 9 Genus-group Taxa . 10 Species-group taxa . 21 Literature Cited . 115 Abstract This catalogue of cancellarioidean taxa is composed of three sections. The first lists alphabetically 124 genus-group taxa originally proposed or later included within the superfamily as well as 24 misspelled or nude names. The second section lists alphabetically 1,864 species-group names proposed in or subsequently referred to genera now considered to belong in Cancellarioidea. Also listed are 392 nude names, misspellings and incorrect attributions. The third section consists of a bibliography of works referred to in this paper. Key words: Cancellarioidea, genus-list, species-list, bibliography Accepted by D. Geiger: 15 Sept. 2005; published: 19 Dec. 2005 3 ZOOTAXA Introduction 1102 The first edition of this catalogue was published in 1990. Since that time numerous new species have been named and several changes in classification have been made affecting the list of genera. Also, as with any catalogue of molluscan taxa, the first edition was not only incomplete, but contained errors, principally in determining the correct dates of some publications. The Cancellarioidea comprise a highly-specialized, poorly known, and still taxonomically enigmatic group of marine neogastropods that inhabit subtidal to bathyal sand and mud bottoms of tropical and temperate latitudes. Although most taxa at all levels within the superfamily were described on the basis of shell characters, it was a series of adaptations to suctorial feeding that resulted in a succession of adaptive radiations beginning during the late Mesozoic that gave rise to the Cancellarioidea. Diagnostic synapomorphies of the superfamily include: an anteriorly tubular, posteriorly saddle- shaped jaw that spans the dorsal surface of the buccal mass; a uniserial radula composed of extremely elongated (L/W >15), ribbon-like, tricuspid teeth with secondary dentition on one or more cusps [there are several aradulate taxa]; an extremely short anterior esophagus, with the valve of Leiblein and buccal ganglia situated at the posterior margin of the buccal mass; and the lack of a gland of Leiblein. Although the superfamily forms a monophyletic and morphologically compact natural assemblage, its relationships to other caenogastropods remains uncertain. The group had, at various times, been included in the Toxoglossa (Troschel 1865; Tryon 1882; Fischer 1883), the Volutoidea (Thiele 1929; Wenz 1943; Taylor and Sohl 1962) and its own order, Nematoglossa (Olsson 1970). Most subsequent authors (reviewed in Harasewych & Petit 1998) follow Ponder (1974) in considering the Cancellarioidea a superfamily within Neogastropoda. Taylor et al. (1980) speculated that the Cancellarioidea are an early offshoot from the Mesozoic Purpurinidae. Linnaeus (1758), in the Tenth Edition of his Systema Naturae, described a single species of cancellariid, including it in the genus Murex. In 1767 he transferred that species to Voluta and described a second species, also as Voluta. By the end of the 18th century, twelve nomina had been proposed for cancellarioideans, allocated to the genera Murex, Voluta, Tritonium, Buccinum and Cantharus. To date, there have been over 1,800 species- group taxa proposed in the Cancellarioidea, many as ’varieties’ of Tertiary species. Lamarck (1799) was the first to recognize that the then known cancellariids comprised a distinct group and erected the genus Cancellaria. The second genus-group name to be proposed for a cancellariid was Trigona Perry, 1811 (preoccupied and replaced by the equivalent Trigonostoma Blainville, 1827). The third genus-level name to be introduced was Admete (Kröyer in Möller, 1842). A decade or so later, the flood-gates opened and many new genera and subgenera were introduced. Marks (1949) published a list of supra- specific taxa in Cancellariidae, citing 58 nominal taxa. We now list 124 nominal genus- group taxa as well as 24 misspellings and nomina nuda. Ten of these nomina are junior objective synonyms and/or junior homonyms, one has been rejected by the I.C.Z.N., one is 4 © 2005 Magnolia Press PETIT & HARASEWYCH an emendation and six were originally proposed in Cancellariidae but belong elsewhere. ZOOTAXA Of the five added since 1990, three were proposed in other families and later transferred to 1102 Cancellariidae; one Paleocene genus was described in 1989 but not seen by us until after the list was in press; and one Recent genus was described after the publication of the 1990 catalog. Some of these 124 nomina are synonyms and others are poorly known. Genus- group taxa proposed for cancellariids have not been treated uniformly in the literature as authors have differed widely in their use of available taxa. As noted by some authors, many of the nominal genera and subgenera of cancellariids seem to be unnecessary. At the same time, there are numerous species that do not readily fit any of the named taxa. Dall (1909a: 30) wrote: “I may add that what, on a cursory survey, appears to be a totally unnecessary number of names has been applied to subdivisions of the genus Cancellaria; but time fails me at present to review them in detail. Most of them are based on slight modification of form which gradually merge into one another when a full series of specimens is considered.” As an aid to the production of badly needed systematic revisions of this poorly- understood superfamily, it has been our objective to compile and maintain a list of all names proposed or currently used for cancellarioideans. We have endeavored to make this work as complete as possible, but recognize that some taxa have undoubtedly been omitted. We request that colleagues bring to our attention any errors and omissions, which will be published in a future addendum. This catalogue of superfamily Cancellarioidea is composed of three sections: an alphabetical listing of genus-group taxa, an alphabetical listing of species-group taxa, and a bibliography of the references cited herein. The genus-group list contains all taxa at the genus-group level, as defined by Article 42 of the International Code of Zoological Nomenclature (Fourth Edition 1999), that have been referred to the Cancellarioidea or any of its families or subfamilies, either originally or subsequently. Genus-group taxa originally proposed in Cancellarioidea but later referred to other superfamilies are included in our list of genera, but the species described in those genera are not included. The species-group list contains all taxa proposed at the species-group level as defined by Article 45 of the International Code of Zoological Nomenclature (Fourth Edition 1999). Names originally proposed as subspecies, varieties, forms or other infrasubspecific categories are included, with the category used by the author identified. We have listed all names originally proposed in, or now considered as being referable to, genera presently included in Cancellarioidea. Nude names (nomina nuda), excluded from some lists of taxa, are included here, as they have appeared in print and may cause confusion if their status is unclear. However, only names that appeared in print are included and names appearing only on museum labels or on websites are not. This is a non-critical list with subjective synonyms being shown only for a few taxa. We stress that it is non-critical as numerous people have inquired about the placement of various species, mistaking the original generic placement as our determination. CANCELLARIOIDEA © 2005 Magnolia Press 5 ZOOTAXA Conventions used in the genus-group list 1102 Genus-group taxa are listed alphabetically, each followed by the author(s) and an abbreviated bibliographic citation. Full citations