Molecular Vision 2005; 11:971-6 ©2005 Molecular Vision Received 21 June 2005 | Accepted 29 October 2005 | Published 9 November 2005

A new congenital nuclear caused by a missense in the γD- (CRYGD) in a Chinese family

Jingzhi Gu,1 Yanhua Qi,1 Li Wang,1 Jin Wang,1 Lisong Shi,2 Hui Lin,1 Xiang Li,1 Hong Su,1 Shangzhi Huang2

1Department of Ophthalmology, the Second Affiliated Hospital of Harbin Medicine University, Harbin, China; 2Department of Medical Genetics, Institute of Basic Medical Sciences, Chinese Academy of Medical Sciences and Peking Union Medical College, Peking, China

Purpose: To identify genetic defects associated with nuclear golden crystal autosomal dominant congenital cataract (ADCC) in a Chinese pedigree in the north of China. Methods: Clinical data were collected and the phenotype of the affected members in this family was recorded by slit lamp photography. Genomic DNA was isolated from peripheral blood. Linkage analyses excluded all known loci except that in 2q33-q35. Mutation analysis of CRYGs was carried by direct sequencing of the PCR products. Results: Sequencing of the coding regions of CRYGA, CRYGB, CRYGC, and CRYGD showed the presence of a heterozy- gous C>A transversion at nt109 of the coding sequence (R36S) in exon 2 of CRYGD, which co-segregated with the affected members. Conclusions: The R36S mutation in CRYGD identified in this Chinese family caused a nuclear golden crystal cataract phenotype not described before. This finding is an additional indication that there may be phenotypic heterogeneity of cataract, especially in different races.

Congenital are a significant cause of visual im- intrinsic (MIP), and one gene coding for beaded fila- pairment in childhood. They have a high incidence and are a ment structural protein 2 (BFSP2). Also, a mutation in CRYGS significant cause of vision loss world wide causing approxi- with autosomal dominant cataract in humans was recently re- mately one tenth of childhood blindness [1]. Roughly 50% of ported [26]. congenital cataracts are hereditary and family studies have re- In our study, we performed linkage analysis on a four vealed that approximately 30% of children with bilateral iso- generation Chinese family with nuclear golden crystal cata- lated congenital cataract had a genetic basis [2]. racts by using STRs markers on 12 ADCC loci and haplotype Congenital cataract is phenotypically and genetically het- analysis with makers on 2q33-q35. A missense mutation in erogeneous. To date, congenital cataracts, isolated or exon 2 of the CRYGD gene was identified, which is respon- syndromic forms, have been mapped to 27 genetic loci, and sible for the disease in this pedigree. the disease-associated have been identified in 18 , including those coding for αA-crystallin [3,4], αB-crys- METHODS tallin [5], βA1-crystallin [6,7], βB1-crystallin [8], βB2-crys- Clinical evaluation: The family was ascertained by the De- tallin [9], γC-crystallin [10], γD-crystallin [11-13], beaded fila- partment of Ophthalmology, the Second Affiliated Hospital ment structural protein 2 [14], heat shock transcription factor of Harbin Medicine University, Heilongjiang province, China. 4 [15], gap junction protein alpha-3 [16], gap junction protein Informed consent in accordance with the Declaration of alpha-8 [17], paired-like homeodomain transcription factor-3 Helsinki and the Heilongjiang Institutional Review Board ap- [18,19], ferritin [20], galactokinase 1 [21], glucosaminyl(N- proval was obtained from all participants. The pedigree was a acetyl)transferase 2 [22], major intrinsic protein of fiber four generation family with 7 members affected (Figure 1). (MIP) [23], lens intrinsic membrane protein 2 (LIM2) [24], ADCC diagnosis was established by the presence of affected and paired box gene 6 [25]. Among them, 12 distinct genes individuals in each generation and male to male transmission. have been identified to cause nonsyndromic autosomal domi- Affected status was determined by the surgical records of cata- nant cataracts, including seven genes coding for crystallin ract extraction for the patients or ophthalmologic examina- (CRYAA, CRYAB, CRYBA1/A3, CRYBB1, CRYBB2, CRYGC, tion, included slit lamp examination with dilated pupils, vi- CRYGD) and two genes coding for gap junctional channel sual acuity testing, intraocular pressure measurement, fundus protein (GJA3, GJA8), one gene coding for heat-shock tran- examination, and ultrasonic examination. Peripheral blood (5 scription factor 4 gene (HSF4), one gene coding for major ml) was collected from each of 5 available affected and 5 un- affected individuals in the family, and genomic DNA was ex- Correspondence to: Jingzhi Gu, Affiliated Second Hospital of Harbin tracted using standard protocol. Medicine, University Heilongjiang Province, 246 Baojian Road, Genotyping and linkage analysis: Twenty-one Harbin 150086, China; Phone: 0086-0451-86605851; FAX: 0086- microsatellite markers close linked to 12 known ADCC loci 0451-86605116; email: [email protected] were used to perform allele-sharing among patients in the fam- 971 Molecular Vision 2005; 11:971-6 ©2005 Molecular Vision ily, including D1S252, D1S305, D2S1782, D2S325, D3S1290, D3S1744, D11S898, D11S1986, D12S90, D12S1676, TABLE 1. PCR PRIMERS AND PRODUCT SIZES

D13S1236, D13S175, D16S3034, D16S421, D17S1294, Fragment D17S1288, D21S212, D22S1174, D22S315, TOP1P2, Gene Exon Strand Sequence (5'-3') size (bp) ------CRYBB2 and three additional markers, D2S2318, D2S1384, CRYGA 1 Forward GTCAGCTGGAAGGAACATCC 291 and D2S1385, linked to the CRYGs. The oligonucleotide Reverse TGAGGACAACTCGAAAATGC 2 Forward GGGTCAGGCCTTGCTATTCT 450 primer sequences were selected from The GDB Human Ge- Reverse CCATGTCTATTGGGGGTCTG nome Database. The order and genetic distances of the mark- 3 Forward GGATTGATTGAACCTGGGAG 659 Reverse GGTGAAAGTTGCAGTGAGCA ers were derived from the Marshfield database. Microsatellites CRYGB 1,2 Forward GGTGGTGCATGCCTGTAA 679 were amplified by polymerase chain reaction (PCR) follow- Reverse GCCCTTTTGTGTGATTTCCT 3 Forward AAACTTGGCCTGGGAGAACT 553 ing standard methods. The genotypes were obtained by silver Reverse GCTTCCCATCATGAAAACAT stain and manual inspection. The pedigree and haplotype was CRYGC 1 Forward CATTTCCAGTGAATGCAGGA 442 Reverse CGCAGCAAGTATTGTTGACC constructed by Cyrillic version 2.1 (MathStat Software 2 Forward GGAAGGTGAGCAGAACACAA 476 ;Victoria, Australia). Reverse TGGCTTATTCAGGTCTCTGATG 3 Forward AATGACAATTCCATGCCACA 534 Mutation analysis: DNA samples from all available af- Reverse CCCACCCCATTCACTTCTTA fected and unaffected family members of the family were CRYGD 1,2 Forward CTTATGTGGGGAGCAAACT 620 Reverse CAGCAGCCCTCCTGCTAT screened for mutations in CRYGA, CRYGB, CRYGC, and 3 Forward GAAACAGCTATGACCATGCACA CRYGD by direct cycle sequencing of the PCR products. The CTTGCTTTTCTTCTCTTT 435 Reverse ATACGACTCACTATAGGGCAAG genomic sequence of CRYGs was obtained from the Ensemble ACACAAGCAAATCAGTGCC genome data resources. Gene specific PCR primers were used to amplify the three exons and flanking introns sequences of Gene specific PCR primer sequences for all exons and flanking in- CRYGA, CRYGB (sequences available upon request), CRYGC, trons sequences of CRYGA, CRYGB, CRYGC, and CRYGD and the size of each PCR product.

Figure 1. Pedigree and haplotype of the autosomal dominant congenital cataract. Black and white symbols denote affected and unaffected individuals, respectively. The affected haplotype is indicated by a black vertical bar. The sequence of mark- ers is from centromere to telomere. The pedigree and haplotype was constructed by Cyrillic version 2.1.

972 Molecular Vision 2005; 11:971-6 ©2005 Molecular Vision and CRYGD (Table 1). PCR products were sequenced in both otide 109 in exon 2 of CRYGD in all affected members, but directions, and the data were collected and analyzed using ABI was not observed in any of the unaffected family members. 3730XL sequencer (Perkin-Elmer, Applied Biosystems, Fos- Fifty unrelated control individuals were also sequenced and ter City, CA). the possibility of a rare polymorphism was excluded.

RESULTS DISCUSSION Clinical findings: All affected members showed nuclear cata- In this report, after excluding all known loci corresponding ract, presented at birth or developed during infancy, progress- to ADCC in a Chinese family except on 2q33-35, we have ing slowly with age. One of them has had cataract extraction. identified a locus on 2q33.3-q34 associated with the nuclear In the other four patients without cataract extraction, the cata- golden crystal cataract and found that a C>A transversion in ract was bilateral and consisted of a central pulverulent opac- exon 2 of CRYGD in all affected members of the family. This ity affecting the embryonal, fetal, and infantile nucleus of the mutation co-segregated with the cataract in the family and no lens, characterized by golden crystal punctuate, with metal- mutation was identified in the 100 independent control allele. like refractivity in the opaque nucleus (Figure 2, Figure 3). It is known that only CRYGC and CRYGD encode abun- From the hospital records, the best corrected visual acuity of dant lens γ- in humans [27,28] and almost 90% of the affected eyes varied from 20/200 to 40/200 before cata- the γ-crystallins synthesized in human lens are the products of ract extraction. There was no family history of any other ocu- these two genes [29]. CRYGD is one of only two γ-crystallins lar or systemic abnormalities. Autosomal dominant inherit- to be expressed at high concentrations in the fiber cells of the ance of the phenotype was supported by the presence of af- human embryonic lens and these cells subsequently form the fected individuals in each of the four generations and with a lens nucleus fibers. Many identified mutations in CRYGD have male to male transmission. been and have proven to be cataractogenic. We think that the Mutation detection: Allele-sharing analysis among the heterozygous mutation was responsible for the congenital cata- affected members in the family excluded linkage with the 11 ract in this family. This is because first, Kmoch et al. [30] has ADCC loci other than those on 2q33-35. Haplotype analysis identified this mutation in a Czech 5-year-old boy who suf- on 2q33-35 showed that a block of 5 markers fered from photophobia and decreased visual acuity due to (black bar in Figure 1) was co-segregated with the disease in symmetrical crystal deposition and grayish opacities in both this family. Direct sequencing was performed to cover exons lenses, and then proved that cataract was caused by deposi- and flanking intron-exon boundary sequences. A heterozygous tion of defined crystallized protein, γD crystallin. Second, we C>A transversion (Figure 4, Figure 5) was identified at nucle- sequenced 100 alleles of 50 unrelated control individuals and exclude the possibility of a rare polymorphism. Third, the single transversion in the heterozygous state (cDNA 109C>A)

Figure 2. Frontal view photograph of the eye of the proband. Photo- graph of the anterior segment of the eye of the proband using a TOPCON Retinal camera TRC-NW6S. Opacities were located in Figure 3. Slit lamp photographs of the eye of the proband. There the embryonal, fetal, and infantile nucleus of the lens while the cor- were many crystal dots in the grey-white opaque nucleus, with golden tex was transparent. metal-like refractivity. 973 Molecular Vision 2005; 11:971-6 ©2005 Molecular Vision predicted a R37S substitution at the protein level. The argin- Zenteno [31]. In contrast to the Czech boy’s crystals [30], char- ine residue is highly conserved, scoring 35 among 36 CRYGD acterized by deposition of numerous birefringent, pleiochroic, homologs cloned from various species using the MAXHOM and macroscopically prismatic crystals in the nucleus and cor- alignment [30]. This is the first report that the R36S caused tex, our cases had many crystal dots in the lens with golden autosomal dominant congenital cataract in China. metal-like refractivity in the grey-white opaque nucleus. The How does R36S mutation cause crystals in the lens nucleus punctuate crystal and pulverulent opacity affected the embryo- and affect lens transparency? Kmoch et al. [30] analyzed this nal, fetal, and infantile nucleus of the lens, but the cortex was problem by protein crystallography and found that the crystal transparent. This, perhaps, is a phenotypic polymorphism of structure at 2.25 Å suggested that the R36S mutation in γ- congenital cataract. It may indicate that or crystallin has an unaltered protein fold and it was thought that the formation of the protein structure for cataract might be the absence of the Arg36 charge led to the redistribution of the influenced by many other factors, which include unidentified surface charges in the mutants which could decrease steric modifier genes and other sequence variations. Further studies hindrances, promoting the permanent mutual contacts and will help us to better understand the mechanism of cataract aggregations of the protein molecules and decrease the solu- formation and processing of gene expression. bility of the R36S mutated protein, leading to crystal forma- tion. However, congenital cataract is phenotypically and ge- ACKNOWLEDGEMENTS netically heterogeneous. The same mutation in cataract genes We are grateful to the members of the family for their partici- may result in different phenotypes under different ethic back- pation in this study. We also acknowledge the financial sup- ground and environment, even the same mutation can cause port of the Natural Scientific Fund of Heilongjiang Provincial different cataractous phenotypes in a family, such as the Mexi- Scientific and Technical Bureau (Grant D0218). can pedigree with congenital hereditary cataract reported by

Figure 4. Forward sequence analysis of CRYGD at exon 2. A: Sequence of unaf- fected individual (individual III:1 in Fig- ure 1). B: Sequence of affected (indi- vidual III:4 in Figure 1). A single trans- version is observed at position 109 (C>A).

Figure 5. Reverse sequence analysis of CRYGD at exon 2. A: Sequence of unaf- fected individual (individual III:1 in Fig- ure 1). B: Sequence of affected (indi- vidual III:4 in Figure 1). A single trans- version is observed at position 109 (G>T).

974 Molecular Vision 2005; 11:971-6 ©2005 Molecular Vision

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The print version of this article was created on 9 Nov 2005. This reflects all typographical corrections and errata to the article through that date. Details of any changes may be found in the online version of the article. α 976