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A New Early Cretaceous Lizard with Well-Preserved Scale Impressions from Western Liaoning , China＊

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A New Early Cretaceous Lizard with Well-Preserved Scale Impressions from Western Liaoning , China＊ PROGRESS IN NATURAL SCIENCE Vol .15 , N o .2 , F ebruary 2005 A new Early Cretaceous lizard with well-preserved scale impressions from western Liaoning , China＊ JI Shu' an ＊＊ (S chool of Earth and S pace Sciences, Peking University , Beijing 100871 , China) Received May 14 , 2004 ;revised September 29 , 2004 Abstract A new small lizard , Liaoningolacerta brevirostra gen .et sp .nov ., from the Early Cretaceous Yixian Formation of w estern Liaoning is described in detail.The new specimen w as preserved not only by the skeleton , but also by the exceptionally clear scale impressions.This lizard can be included w ithin the taxon Scleroglossa based on its 26 or more presacrals, cruciform interclavicle with a large anterior p rocess, moderately elongated pubis, and slightly notched distal end of tibia .The scales vary evidently in size and shape at different parts of body :small and rhomboid ventral scales, tiny and round limb scales, and large and longitudinally rectangular caudal scales that constitute the caudal w horls.This new finding provides us with more information on the lepidosis of the Mesozoic lizards. Keywords: new genus, Squamata, skeleton, lepidosis, Early Cretaceous, western Liaoning . Lizards are majo r groups in the Late Mesozoic Etymology:Liaoning , the province where the Jehol Biota of w estern Liaoning and the adjacent holoty pe w as collected ;lacerta (Latin), lizard . regions, no rtheastern China .Several fossil lizards Brevi- (Latin), short ;rostra (Latin), snout . have been found from the Yixian Formation , the lower unit of the Early C retaceous Jehol G roup in Holotype :An articulated skeleton w ith its rig ht w hich the feathered theropods , primitive birds , early fo relimb and mid to posterior caudals missing (GM V mamm als and angiosperms were discovered in the past 1580 ; National Geological Museum of China , decade[ 1, 2] . The reported lizards include Beijing).Although most skeletal bones them selves Yabeinosaurus tenuis[ 3—5] , Dalinghosaurus were lost , their moulds are fairly clear and very longidigitus[ 6 , 7] , and Jeholacerta formosa[ 8] , and anatomically informative (Fig .1). some other material has also been under study[ 9] . Locality and horizon:Huang banjigou , Beipiao , Recently , another new lizard specimen w as western Liaoning ;Yixian Formation . collected from the Yixian Fo rmation of a quarry near the village of Huangbanjigou , Beipiao , western Diagnosis:Skull broad with short , pointed Liaoning .This new finding further enriches our snout and large orbits .Naso-frontal suture at about know ledge on the taxonomic diversity and evolution the level of anterior bo rders of orbits .Frontals about lizards within the Jehol Biota .It should be paired.Jug al having long do rsal processes and in pointed out that the remarkable scale impressions contact w ith squamosal .Presacrals at least 26 or w ere preserved on the new specimen , representing more , w ith 19 do rsals included .Anterior caudals the second example follow ing the holoty pe of w ith one pairs of laterally directed transverse Jeholacerta formosa[ 8] . It provides important processes, first auto tomy septum on the 6th caudal. evidence for our understanding of the lepidosis Interclavicle cruciform and w ith long anterior features of the Mesozoic lizards . process.Pubis moderately elong ated .Hind limb fairly long compared with fo relimb .Distal end of tibia 1 Systematic paleontology slig htly concave .Large astragalus and relatively small calcaneum not co-ossified .M etatarsal II I equal to IV Squamata Oppel, 1811 Scleroglossa Estes, de Queiroz et Gauthier, 1988 in length .Pedal phalangeal fo rmula 2∶3∶4 ∶5 ∶4 . Liaoningolacerta gen .nov. Ventral scales small and mainly rhomboid in shape Liaoningolacerta brevirostra gen .et sp .nov. w ith the arrangement in transversal rows .Lateral ＊ Supported by National Natural Science Foundation of China (Grant No.40272008), the China Postdoctoral S cience Foundation (Grant No . 2003034012), and the Fund for Young Geologist of former Ministry of Geology and Mineral Resou rces(Grant No .Qn979827) ＊＊ Presen t address:Institute of Geology , Chinese Academy of Geological Sciences, Beijing 100037 , China.E-mail:jishu -an @sina.com Prog ress in Natural Science Vol.15 No .2 2005 w ww .tandf.co .uk/journals 163 scales very large and pavimentous.Scales covering and rectangular , forming caudal whorls . limbs tiny , round or polygonal .Caudal scales long Fig.1 . Liaoningolacerta brevirostra gen .et sp .nov., holotype and its sketch (GMV 1580).(a)—(d)indicate the regions of scales shown in Fig .4.a, astragalus;c , calcaneum ;cv1 , 1st caudal verteb ra ;dr , dorsal rib ;dv1 , 1st dorsal verteb ra ;d3 , 3rd distal tarsal; d4 , 4th distal tarsal;fe, femur ;fi, fibula;hu , humerus;mtI , metatarsal I ;mtV , metatarsal V ;ra, radius;ral, radiale ;sv1 , 1st sacral vertebra;ti, tibia;ul, ulna ;ulr , ulnare ;I —V , pedal digit I —V . 2 Description 2 .1 Skull and low er jaw The skull is broad w ith sho rt and somew hat pointed snout , large o rbits , and open supratemporal fenestrae (Fig .2). The skull length measures roug hly 9 .6 mm , 29 % the snout-vent leng th (33 .4 mm).The skull w idth is the g reatest (6 .7 mm)at the level just behind orbits, making the skull width / leng th ratio about 70 %.The o rbit is round and large w ith its diameter 2 .7 mm (28 % skull length).The right supratemporal fenestra appears oval and relatively large . The shape of premaxilla is unknow n , w hile the maxilla is also difficult to be outlined exactly .There Fig .2 . Liaoningolacerta brevirostra gen .et sp .nov ., skull and are eig ht monocuspid teeth preserved on the left mandible.den , dentary ;fr , frontal;j, jugal;mx , maxilla ;n , premaxilla and maxilla with the anterior teeth nasal;orb , orbit;pal, palatine;po, postorbital;sq, squamosal; sm aller . stf, supratemporal fenestra . In the skull roof , the nasal is long and broad at its posterio r portio n .The straight naso-frontal suture 164 w ww .tandf.co .uk/journals Prog ress in Natural Science Vol .15 No .2 2005 slightly slopes medio-caudally , and lies about at the the dorsal ribs are nearly com pletely preserved .They level of anterior borders of orbits .The paired frontals are long and very slender .The relationships of the are distinctive in being relatively broad at the naso- anterio r several pairs with the sternum or frontal suture , but slightly constricted between the xiphisternum are uncertain because the latter tw o o rbits and becoming wider posteriorly . The were not identified .The middle pairs are longer , morphology of parietal remains unknow n because of follow ed by six pairs of shorter ribs on the posterior its damaged preservation . dorsals . The sutures between the prefrontal and its adjacent 2 .3 Pectoral girdle and fo relimb bones are not identified .The same situation is also seen for the lacrimal, so the presence or absence of a lacrimal is The pectoral girdle is partially preserved , but the uncertain.The postorbital seems to be a slender bone interclavicle is quite complete .Only a small posterior tapering posteriorly .The very long and curved bone portion of each coracoid is clearly distinguishable , which constituting the lateral border of orbit is undoubtedly the shows an obviously arched ventral margin and a hook-like jugal.Its dorsal process is narrow and bears a pointed end process at the junction between the ventral and posterior margins.The left clavicle is a long and curved bone that is in contact with the postorbital and squamosal.The without other notable features to be identified . right squamosal is exposed with a long and tapering anterior process.Posteriorly , this bone extends medially The interclavicle is ty pically crucifo rm (3 .9 m m into a small parietal process and posterolaterally into a in total leng th ) w ith slender anterior and lateral small process for the quadrate .Fragments of the palatine processes.The anterior process is long (1 m m), and are able to be distinguished at the lateral border of left occupies roug hly 37 % of the length of the posterior orbit . process (2 .7 m m), the latter ends in a slig htly The left low jaw is quite slender w ith less pointed caudal end .The lateral process (1 .5 mm)is preserved morphological info rmation .The dentary is one and a half times the leng th of the anterio r process shorter than half the w hole jaw .Only three posterio r (Fig .3(a)). dentary teeth are preserved , showing visibly conical Only the left forelimb w as preserved with some shape . manual elements absent .The humerus is a sturdy 2 .2 Axial skeleton bone (75 %fumer)with distinctly expanded proximal and distal ends :the proximal width mo re than twice The vertebral column appears to consist of at the mid-shaft w idth .The straig ht ulna and radius are least 26 presacral vertebrae (including 19 dorsals), g racile and sho rter (68 % humerus) w ith slig htly tw o sacrals and an unknow n number of caudals .All broad ends proximally .These tw o bones are almost the vertebrae are distinctly procoelous .The last five the same in length and w idth .Only radiale and ulnare cervicals are visible .Although no mo re detailed are clear in carpus, the former is elongated and the features can be identified on them , it is suggested latter nearly round .The preserved metacarpals are that the first preserved cervical is not atlas or axis .So more robust than the preserved manual phalanges. the number of cervicals would be estim ated at least The phalanges are gracile , and the claw s quite small . seven or more .Dorsal vertebrae are generally longer than cervicals , and the last five or six do rsals more massive .Sacrals possess broad and long transverse processes that may be fused at their tips .The first sacral is much larger than the second .Only eight vertebrae of the caudal series w ere preserved .The anterior caudals are non-autotomous and bear developed transverse processes pointing laterally , decreasing in length quite rapidly from the first to the Fig .3 .
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