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AN ANNOTATED CHECKLIST OF GENUS PACHYRHYNCHUS (COLEOPTERA: : )

Article in Acta Biologica Universitatis Daugavpiliensis · September 2018

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AN ANNOTATED CHECKLIST OF GENUS PACHYRHYNCHUS (COLEOPTERA: CURCULIONIDAE: PACHYRHYNCHINI)

Anita Rukmane

Rukmane A. 2018. An annotated checklist of genus Pachyrhynchus (Coleoptera: Curculionidae: Pachyrhynchini). Acta Biol. Univ. Daugavp., 18 (1): 63 – 68.

Pachyrhynchus is a genus from subfamily of which 145 species are known so far. Checklist prepared including it’s synonyms and different subspecies indicate that this genus is predominantly tropical with most of the species known from the Oriental region: Philippines (135 species or 93%), Indonesia (6 species or 4%), Japan (3 species) and Taiwan (1 species and two subspecies).

Key words: Pachyrhynchus, checklist, synonyms, distribution, Philippines.

Anita Rukmane. Institute of life sciences and technology, Daugavpils University, Coleopterological Research Center, Parādes street 1a, Daugavpils, Latvia, E-mail: [email protected].

INTRODUCTION few species distributed in Indonesia (4%) and Japan (3%). Subfamily Entiminae or broad nosed are one of the largest groups of the family Review of literature reveals lapses in the Curculionoidae and are distributed throughout descriptions justifying a substantial taxonomic the world mostly in tropics with 55 tribes, 1340 study. The present contribution enlists the genera (Nikolai et al. 2006) and more than 12,000 19 species with type locality ‘’Philippines’’ species (Alonso-Zarazaga & Lyal 1999), in with no more details. As species from genus addition, number of new taxon increases rapidly. Pachyrhynchus are highly endemic, and tend to inhabit one particular island, it is necessary During past seven years, number of species from to evaluate such cases with ample ecological genus Pachyrhynchus has increased from 104 to studies. 145 (41 new species) new species described by several local and foreign scientists (Cabras & Rukmane 2016, Rukmane & Barševskis 2016, RESULTS Barševskis 2016, Bollino et al. 2017, Chen & Lin 2017), which is almost 1/3 of number of all Genus Pachyrhynchus Germar, 1824 species in genus. Taxonomic studies are imminent = Sphaerogaster Latreile, 1825 and as a prelude to such studies an annotated = Somatodes Schoenherr, 1823 checklist had been prepared. The checklist = Sphoerogaster Latreile, 1825 reveals that it is well known in Oriental region, = Sphaerogaster Sturm, 1826 with predomination in Philippines (93%) and = Sphoenogaster Berthold, 1827

63 Rukmane A.

= Pachirhinus Latreile, 1828 [Philippines, Mindanao] = Pochyrhynchus Laporte, 1840 21. P. cagayanus Heller, 1929 [Philippines, = Pachyrhinchus Desmarest, 1842 Luzon] = Pachyrhincus Desmarest, 1842 22. P. chamissoi Schultze, 1922 [Philippines, = Pachyrrhynchus Gemminger & Harold, 1871 Mindanao] 23. P. chlorites Chevrolat, 1881 [Philippines] Type species – Pachyrhynchus moniliferus = rutilans Behrens, 1887 Germar, 1824 ssp. insularis Kano, 1929 [Philippines] 24. P. cingulatus Pascoe, 1873 [Indonesia, 1. P. absurdus Schultze, 1919 [Philippines, Morotai] Bucas] 25. P. circulatus Heller, 1912 [Philippines, 2. P. amabilis Schultze, 1922 [Philippines, Catanduanes] Mindanao] 26. P. conformis Yoshitake, 2017 [Philippines, 3. P. anitchtchenkoii Rukmane & Barševskis, Samar] 2016 [Philippines, Mindanao] 27. P. confusus Schultze, 1923 [Philippines] 4. P. annelifer Heller, 1912 [Philippines, Luzon] 28. P. congestus Pascoe, 1873 [Philippines, 5. P. annulatus Chevrolat, 1881 [Philippines, Luzon] Luzon] = luteoguttatus Chevrolat, 1841 = annulatus Behrens, 1887 ssp. caerulans Kraatz, 1888 [Philippines, Luzon] 6. P. antonkozlovii Rukmane & Barševskis, 2016 ssp. immarginatus Kraatz, 1888 [Philippines, [Philippines, Mindanao] Luzon] 7. P. apicatus Schultze, 1922 [Philippines, ssp. ocellatus Schultze, 1924 [Philippines, Polillo] Luzon] 8. P. apocyrtoides Schultze, 1922 [Philippines, ssp. pavonius Heller, 1921 [Philippines, Luzon] Mindanao] 29. P. consobrinus Schultze, 1922 [Philippines, 9. P. apoensis Yoshitake, 2012 [Philippines, Luzon] Mindanao] 30. P. corpulentus Schultze, 1922 [Philippines, = pseudoapoensis Rukmane & Barševskis, 2016 Mindanao] 10. P. ardentius Schultze, 1919 [Philippines, 31. P. croesus Oberthur, 1879 [Indonesia, Siargao] Sanghir] 11. P. argus Pascoe, 1873 [Philippines, Luzon] 32. P. cruciatus Schultze, 1923 [Philippines, 12. P. atrocyaneus Schultze, 1922 [Philippines, Luzon] Mindanao] 33. P. cumingi Waterhouse, 1841 [Philippines] 13. P. banglas Bollino, Sandel & Rukmane, 2017 ssp. boholensis Schultze, 1924 [Philippines, [Philippines, Mindanao] Bohol] 14. P. baluganus Schultze, 1924 [Philippines, 34. P. davaoensis Schultze, 1934 [Philippines, Luzon] Mindanao] 15. P. barsevskisii Rukmane, 2016 [Philippines, 35. P. decussatus Waterhouse, 1841 [Philippines, Luzon] Catanduanes] 16. P. basilanus Heller, 1923 [Philippines, 36. P. disgestus Heller, 1929 [Philippines, Luzon] Basilan] 37. P. dohrni Behrens, 1887 [Philippines, Luzon] 17. P. benguetanus Schultze, 1924 [Philippines, 38. P. domino Rukmane, 2016 [Philippines, Luzon] Mindoro] 18. P. bucasanus Schultze, 1922 [Philippines, 39. P. dubiosus Schultze, 1922 [Philippines, Bucas] Luzon] ssp. ornatus Schultze, 1934 [Philippines, Samar] 40. P. elegans Waterhouse, 1842 [Philippines, 19. P. cabrasae Rukmane & Barševskis, 2016 Samar] [Philippines, Mindanao] = eos Heller, 1924 20. P. caeruleovittatus Yoshitake, 2012 41. P. elenae Rukmane, 2016 [Philippines,

64 An annotated checklist of genus Pachyrhynchus (Coleoptera: Curculionidae: Pachyrhynchini).

Mindanao] Panay] 42. P. esperanza Bollino, Sandel & Rukmane, 67. P. libucanus Schultze, 1923 [Philippines, 2017 [Philippines, Mindanao] Libucan] 43. P. eques Heller, 1912 [Philippines, Luzon] 68. P. loheri Schultze, 1917 [Philippines, Luzon] 44. P. equester Heller, 1929 [Philippines, Luzon] 69. P. lorquini Chevrolat, 1881 [Philippines] 45. P. erichsoni Waterhouse, 1841 [Philippines] = flavomaculatusKraatz, 1888 = chrysocompsus Heller, 1912 = flavopunctatusKraatz, 1888 ssp. eschscholtzi Waterhouse, 1841 [Philippines, 70. P. lubanganus Bollino & Sandel, 2015 Luzon] [Philippines, Lubang] 46. P. erosus Schultze, 1920 [Philippines, Luzon] 71. P. marinduquensis Rukmane & Barševskis, 47. P. felipeae Rukmane & Cabras, 2018 2016 [Philippines, Marinduque] [Philippines, Panay] 72. P. masatoshii Yoshitake et Yap, 2018 48. P. forsteni Snellen van Vollenhoven, 1864 [Philippines, Luzon] [Indonesia, Molucas] 73. P. miltoni Cabras & Rukmane, 2016 49. P. franciscoi Rukmane & Cabras, 2018 [Philippines, Mindanao] [Philippines, Panay] 74. P. mohagani Bollino & Sandel, 2015 50. P. galeraensis Schultze, 1934 [Philippines, [Philippines, Lubang] Mindoro] 75. P. mollendorffi Heller, 1898 [Philippines, 51. P. gemmatus Waterhouse, 1841 [Philippines] Panay] = atratus Heller, 1912 ssp. marinduquanus Rukmane & Cabras, 2018 ssp. purpureus Kraatz, 1888 [Philippines, Luzon] [Philippines, Marinduque] 52. P. gilvomaculatus Yoshitake, 2017 76. P. moniliferus Germar, 1824 [Philippines, [Philippines, Mindanao] Luzon] 53. P. gloriosus Faust, 1895 [Philippines, Luzon] = confinisChevrolat, 1841 ssp. abbreviatus Schultze, 1922 [Philippines, = monilifer Germar, 1871 Luzon] ssp. abranus Heller, 1934 [Philippines, Luzon] 54. P. halconensis Schultze, 1922 [Philippines, ssp. chevrolati Eydoux & Souleyet, 1839 Mindoro] [Philippines] 55. P. helleri Kuntzen, 1914 [Philippines, Luzon] ssp. stelluliufer Heller, 1912 [Philippines] 56. P. hirokii Yoshitake, 2012 [Philippines, 77. P. morio Heller, 1912 [Philippines, Luzon] Mindanao] 78. P. morotainensis Snellen van Vollenhoven, 57. P. igorota Schultze, 1917 [Philippines, Luzon] 1864 [Indonesia, Morotai] 58. P. inclytus Schultze, 1924 [Philippines, = waterhousei Faust, 1895 Luzon] 79. P. multipunctatus Waterhouse, 1841 = apicalis Kraatz, 1888 [Philippines] = modestior Behrens, 1887 = auroguttatus Chevrolat, 1881 = transversatus Heller, 1921 80. P. naokii Yoshitake, 2012 [Philippines, 59. P. infernalis Fairmaire, 1897 [Japan, Ryukyu] Mindanao] = niger Sakaguchi, 1927 81. P. negrosensis Schultze, 1924 [Philippines, 60. P. ilgas Rukmane, 2017 [Philippines, Samar] Negros] 61. P. jitanasaius Chen & Lin, 2017 [Taiwan, 82. P. neoabsurdus Rukmane, 2017 [Philippines, Green] Mindanao] 62. P. jugifer Waterhouse, 1841 [Philippines] 83. P. niisatoi Yoshitake, 2017 [Philippines, 63. P. kraslavae Rukmane & Barševskis, 2016 Luzon] [Philippines, Mindanao] 84. P. nitcisii Rukmane & Barševskis, 2016 64. P. lacunosus Heller, 1912 [Philippines] [Philippines, Mindanao] 65. P. latifasciatus Waterhouse, 1842 [Philippines, 85. P. nobilis Heller, 1912 [Philippines] Samar] ssp. yamianus Kano, 1929 [Taiwan] 66. P. layroni Rukmane & Cabras [Philippines, 86. P. notocruciatus Yoshitake, 2017 [Philippines,

65 Rukmane A.

Mindanao] = bakeri Heller, 1921 87. P. ohbayashii Yoshitake, 2017 [Indonesia, = modestioroides Schultze, 1922 Biak] 102. P. rebus Rukmane, 2016 [Philippines, 88. P. occidentalis Rukmane, 2017 [Philippines, Luzon] Mindanao] 103. P. regius Schultze, 1922 [Philippines, Leyte] 89. P. ochroplagiatus Heller, 1912 [Philippines, ssp. boronganus Schultze, 1934 [Philippines, Luzon] Samar] ssp. multiplagiatus Schultze, 1924 [Philippines, 104. P. reicherti Schultze, 1929 [Philippines, Luzon] Mindanao] 90. P. orbifer Waterhouse, 1841 [Philippines, 105. P. reticulatus Waterhouse, 1841 [Philippines, Luzon] Luzon] = alboguttatus Chevrolat, 1841 106. P. rizali Schultze, 1934 [Philippines, Luzon] = chlorilineatus Waterhouse, 1841 107. P. roseomaculatus Waterhouse, 1841 = fahrei Schoenherr, 1845 [Philippines] = fimbriatus Chevrolat, 1841 108. P. rufopunctatus Waterhouse, 1842 = globulipennis Chevrolat, 1841 [Philippines] = pretiosus Chevrolat, 1841 109. P. rugicollis Waterhouse, 1841 [Philippines] = scintilans Chevrolat, 1841 ssp. aurinius Heller, 1921 [Philippines, Luzon] ssp. ardens Chevrolat, 1841 [Philippines, Luzon] 110. P. rukmaneae Barševskis, 2016 [Philippines, ssp. azureus Schultze, 1922 [Philippines, Luzon] Marinduque] ssp. circulifer Chevrolat, 1841 [Philippines] = takakuwai Yoshitake, 2016 ssp. gemmans Chevrolat, 1841 [Philippines, ssp. paucisignatus Yoshitake, 2018 [Philippines, Luzon] Marinduque] ssp. inornatus Waterhouse, 1841 [Philippines] 111. P. sakaii Yoshitake, 2017 [Philippines, ssp. murinus Heller, 1934 [Philippines, Luzon] Samar] ssp. striatomaculatus Yoshitake, 2017 112. P. samarensis Schultze, 1923 [Philippines, [Philippines, Luzon] Samar] 91. P. orientalis Rukmane, 2017 [Philippines, 113. P. sanchezi Heller, 1912 [Philippines, Mindanao] Luzon] 92. P. perpulcher Waterhouse, 1841 [Philippines] 114. P. sarcitis Behrens, 1887 [Philippines] 93. P. phaleratus Waterhouse, 1841 [Philippines] ssp. kotoensis Kano, 1930 [Japan, Lanyu] 94. P. pinorum Pascoe, 1873 [Philippines, Luzon] 115. P. schoenherri Waterhouse, 1841 = dimidiatus Heller, 1912 [Philippines] ssp. transversalis Heller, 1912 [Philippines] 116. P. schultzei Schultze, 1917 [Philippines, 95. P. postpubescens Schultze, 1922 [Philippines, Luzon] Mindanao] 117. P. semiignitus Schultze, 1922 [Philippines, ssp. confluens Janczyk, 1959 [Philippines, Luzon] Mindanao] 118. P. semperi Heller, 1912 [Philippines] 96. P. pseudamabilis Yoshitake, 2012 [Philippines, 119. P. septentrionalis Yoshitake, 2017 Mindanao] [Philippines, Luzon] 97. P. pseudhalconensis Rukmane, 2016 120. P. shavrinii Rukmane & Barševskis, 2016 [Philippines, Mindoro] [Philippines, Samar] 98. P. pseudoproteus Schultze, 1922 [Philippines, 121. P. signaticollis Schultze, 1922 [Philippines, Luzon] Mindanao] 99. P. psittacinus Heller, 1912 [Philippines, = transversarius Heller, 1923 Luzon] 122. P. signatus Schultze, 1919 [Philippines, 100. P. psitaculus Heller, 1921 [Philippines, Siargao] Luzon] 123. P. smaragdinus Behrens, 1887 [Philippines] 101. P. pulchellus Behrens, 1887 [Philippines] = carnosus Kraatz, 1888

66 An annotated checklist of genus Pachyrhynchus (Coleoptera: Curculionidae: Pachyrhynchini).

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