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Insect Species Described from Big Bend National Park

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Insect Species Described from Big Bend National Park INSECTA MUNDI, Vol. 16, No. 1-3, March-September, 2002 125 Insect species described from Big Bend National Park Arnold F.Van Pelt 203 Howell Place Greensboro, NC 27455 Abstract: The list contains 192 insect species, of which 139 are primary types. All have been described from material collected in or near Big Bend National Park, Texas Keywords: Big Bend National Park, holotypes, paratypes, allotypes, types, primary types, insects Introduction that the investigators used, e.g., months are either in Arabic or Roman numerals. A "$" symbol follow­ Big Bend National Park is a large area of over ing an entry in References section papers cited 800,000 acres, the centerpiece of which is the Chi­ here; the other references should be of aid to inves­ sos Mountains, described as an island in the Chi­ tigators who use this list. Ifa species has become a huahuan Desert. Habitats range from the high synonym, that name will appear in brackets. Chisos Mountains, characterizedby PonderosaPine (Pinus ponderosa), Arizona Cypress (Cupressus Acknowledgements arizonica) andDouglas Fir(Pseudotsuga menziesii), through oak - pinyon - juniper and grassland asso­ Several entomologists were kind enough to aid ciations to desert and the riparian vegetation along in various ways in the preparation ofthis paper: R. the Rio Grande. From Emory Peak (7825 feet), S. Anderson; C. L. Bellamy; H. R. Burke, C. E. elevations descend steeply to foothills character­ Carlton; B. N. Danforth; A. T. and H. F. Howden; E. izedby volcanic dikes and ash (about 3500 feet), and C. Knudson, G. H. Nelson, C.W. O'Brien; W. Pula­ then a moderate slope to the Rio Grande River waski; and R. L. Westcott, and several others on (about 2000 feet). Because of the variety of habitats specific issues. Knudson has worked on the Lepi­ and microhabitats the Park provides, numerous doptera of the Park for many years, and continues plant and animal species occur, some of which are to provide valuable and extensive information on endemic. that group. I appreciate the help that these and Big Bend National Park was established in other entomologists have provided. 1944, and researchers have visited the Big Bend area and made collections of plants and animals Abbreviations since the 1800's. Over the years, species previously unknown from the Park have been collected there, Collecting sites: BBNP - in Park, area not and many of these have been new to science. designated; CA - Castolon; CB - Chisos Basin; CF About 4600 species of insects have been docu­ - Cattail Falls; CM - Chisos Mountains; CS ­ mented in the Park, along with an extensive bibli­ Croton Springs; DW - Dugout Wells; GG - Green ography (Van Pelt, 1999). From this material, 139 Gulch; GH - Grapevine Hills; GnS - Glenn Springs; specimens have been designated primary types GrG - Grama Grassland; GS -Government Springs; (three of which are syntypes) from a total type list HS - Hot Springs; JC - Juniper Canyon; KB -K of 192 designations. The orders with the largest - Bar Research Station; LBC - Lower Blue Creek; number of species are Coleoptera and Lepidoptera. MS - Mariola Scrub; NM - Nugent Mountain; OC The present list is organized phylogenetically by - Oak Creek; OS - Oak Springs; PC - Pine Canyon; orders, and alphabetically by families and genera. PJ - Panther Junction; RGV - Rio Grande Village; The entries in the list contain the type locality, and RM - Rosillos Mountains; RMS - Rosillos Moun­ references to the appropriate papers are appended. tains Summit; SEC - Santa Elena Canyon; TC ­ Full citations are listed at the end of this paper Tornillo Creek; TeC - Terlingua Creek; TF under References. Dates are included in the format 126 Volume 16, No. 1-3, March-September, 2002, INSECTA MUNDI Tornillo Flat; UTC - Upper Tornillo Creek; WT­ al Museum of Natural History (United States Na­ Window Trail. tional Museum), Washington, DC; UMMZ - Mu­ seum of Zoology, University of Michigan, Ann Collectors: AB - Andre Blanchard; AVP - A. F. Arbor; WBWC - W.E. Warner Collection, Phoenix, Van Pelt; CAT - Charles A. Triplehorn; ECK - Ed AZ C. Knudson; GHN - G. H. Nelson; HFH - Henry F. Howden; HRB - Horace R Burke; JA - J. Apper­ son; JEH - J. E. Hafernik; JNK - Josef N. Knull; Table II. Insect species RHB - Rollin H. Baker; RLW -R L. Westcott; RSA - Robert S. Anderson; WFB - W. F. Barr Heteroptera Table I. Depositories Phytocoris cinereus Stonedahl - Miridae [PARATYPE] (TAMU) (Stonedahl, 1988); GG, AMNH - American Museum of Natural History, 5700', 08 - 14 - 68, JEH (Holotype from Colo­ NYC; ANSP - Academy of Natural Sciences of rado) Philadelphia; BARC - Beltsville Agricultural Re­ Menecles portacrus Rolston - Pentatomidae [HO­ search Center; BBNP - Big Bend National Park, LOTYPE, male] (AMNH) Lost Mine Trail, 5800 TX; BMNH - British Museum of Natural History, feet [PARATYPE] GG, 5700', 07 - 24 - 68, JEH London; CASC - California Academy of Science (Rolston, 1973) Collection, San Francisco; CDAS - California De­ partment of Agriculture, Sacramento; CMNC ­ Homoptera Canadian Museum of Nature Collection, Ottawa; CNCI - Canadian National Collection of Insects, Athysanella spatulata Ball & Beamer - Cicadel­ Ottawa; CSWC - C. S Wolfe Collection, Fort Worth, lidae [PARATYPES] (SEMC) Marathon, 07 - 09 TX; DCCC - D.C. Carlson Collection, Orangevale, - 38, RH Beamer (Ball & Beamer, 1940); Burro CA; DGMC - D. G. Marqua Collection, Fort Davis, Mesa, on Heteropogon contortus, R Whitcomb, TX; DWSC - D.W. Sundberg Collection, San Anto­ det: Hicks (Whitcomb, BBNP file letter, 1985) nio, TX; DEFW - Department of Entomology, Flexamia bandarita Whitcomb & Hicks - Cicadel­ Fisheries and Wildlife, University of Minnesota, lidae [HOLOTYPE, male] CM, on Bouteloua St. Paul; EGRC - E.G. Riley Collection, College curtipendula, 7 Aug 1987, 5200 ft, IPL 003262, Station, TX; ELSC - E. L. Sleeper Collection, Long Whitcomb, R, (NMNH, BARC); [PARATYPES]: Beach, CA; FSCA - Florida State Collection of 2m, If, same collection data, Texas; Marathon Arthropods, Gainesville; FMNH - Field Museum of Basin, 8 Aug 1984 (NMNH, BARC, KMNH, Natural History, Chicago; HAHC - H. and A. KSUC); CM, 17 July 1946, D.J. & J.N. Knull Howden Collection, Ottawa ON (Types deposited in (OSUC) (Whitcomb and Hicks, 1988) CMNC); JEWC - J.E. Wappes Collection, Bulver­ Flexamia zacata Whitcomb & Hicks - Cicadellidae de, TX; IPL - accession #'s of computerized collec­ [HOLOTYPE] PJ, on Muhlenbergia porteri, 26 tion, BARC; KSBS - Biological Survey of Kansas Aug 1985, 3600 ft, Whitcomb & Hicks, IPO Invertebrate Collection, University of Kansas, 002093 (NMNH, BARC); [PARATYPES]: 33m, Lawrence; KSUC - Department of Entomology 10f, same locality and date, IPO 002057,25 Aug Collection, Kansas State University, Manhattan; 1985; 13m, 4f, same locality and date (NMNH, LACM - Los Angeles County Museum of Natural BARC, CNCI, KMNH, KSUC, OSUC) (Whit­ History; MSUC - Michigan State University, East comb and Hicks, 1988) Lansing; MCZC - Museum of Comparative Zoolo­ gy, Cambridge, MA; NCSU - North Carolina State Neuroptera Museum, Raleigh; OSUC - Ohio State University, Columbus; OSUO - Oregon State University, Cor­ Brachynemurus marshi Stange - Myrmeleontidae vallis; SEMC - Snow Entomological Museum, [HOLOTYPE] (UCDC) [PARATYPE] (UCDC) University of Kansas, Lawrence; TAMU - Texas A (FSCA) both 22 mi. S. Marathon (Stange, 1970) & M University, College Station; UCDC - Univer­ Brachynemurus nigrescens Stange - Myrmeleon­ sityofCalifornia, Davis; NMNH (USNH) - Nation- tidae [PARATYPES] (UCDC) 22 mi. S. Mara- INSECTA MUNDI, Vol. 16, No. 1-3, March-September, 2002 127 thon, 09 - 03 - 60, L. Stange; Tornillo Flat, 08 ­ OSUC) CM, 07 - 23, IN Knull; Castolon, 04 - 21 12 - 48, W. Nutting & F. Werner; Terlingua, 07 - 73, S. Wellso; near Boquillas, 04 - 16 - 83, dead - 03 - 48, C & P Vaurie (Stange, 1970) mesquite, S. Wellso (Knull, 1952); 06 - 27 - 63, on Acacia constricta, GHN, det. GHN '82 (ex Coleoptera TAMU) (Knull, 1952) Agrilus obscurilineatus Vogt - Buprestidae Elonuschisosensis Werner-Aderidae [HOLOTYPE, [=A.chisosanus Knull] [HOLOTYPE] (FMNH) male] (MCZC) CM, vii - 9 - 1948, F. Werner; CM, 07 - 08 - 55, DJ&JN Knull (Knull, 1956) [PARATYPES] (BMNH, CNCI & Univ. Arizo­ (Vogt, 1949) na, Tucson) one same data as holotype + two Agrilus putillus paraputillus Knull - Buprestidae from CB at light, May 1, 1959 andMay 29, 1959, [HOLOTYPE] (FMNH) PC, 05 - 10 - 59; Pulliam Howden/Becker (Werner, 1992) Canyon, 500', 05 - 17 - 59, H Howden, det: Eugnamptus pseudonigriuentris Hamilton - Atte­ Knull; CM, 05 - 08 - 74, G.Manley, det: Manley; labidae [HOLOTYPE] BBNP, 05 - 05 - 69, FJ [PARATYPE] (CNCI & OSUC) on maple (Acer Moore & RL Berry + [ALLOTYPE] CM, 05 - 14 grandidentatum) PC, 5 - 10 - 59, Becker & HFH - 78, AE&DS Lewis + [PARATYPE] PC, 06 - 05 (paratypes coll. 5 - 4&7 - 59, HFH & Becker) - 70, 5100', UV trap, CW O'Brien+ [PARATYPE] Boot Spring, 05 - 18 - 59, Pulliam Canyon, 5000', CB, Apr '63, JB Karren (all NMNH) (all Hamil­ 05 - 17 - 59, in dead maple twigs, Howden/ ton 1990) Becker, det: JNK (Knun, 1960a) Haplorhynchites pseudomexicanus Hamilton - At­ Brachys querci Knull - Buprestidae [PARATYPE] telabidae [PARATYPE] (NMNH) Marathon, 08 (FMNH) CM, 06 - 09 - 31, JNKI; CM, 07 - 17 - 46, - 05 - 64, HRB & JA (Hamilton, 1974) JNK [PARATYPES] (FMNH) (Knull, 1952) Pselaphorhynchites dilatarostris Hamilton - Atte­ Buprestis parmaculiuentris Knull - Buprestidae labidae [HOLOTYPE AND ALLOTYPE] [HOLOTYPE, male] (FMNH) CM, 06 - 30 - 57, (NMNH) CM, R&L Hamilton, July 16,18,19 on pine, DJ&JN Knun (Knull, 1958); CB, 06 - 2 (Hamilton, 1971) -
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