Rev. Biol. Trop., 41 (3): 729-753,1993
Revision of the New World species of Ommatius Wiedemann (Diptera: Asilidae): the neotropical costatus species group
A. G. Scarbrough Department oC Biologica1 Sciences, Towson State University, Baltimore, MD 21204 USA.
(Rec.19-1-1993. Acep. 2-VI-1993)
Abstracl: Fourleen species of New World Ommalius Wiedemann are assigned to me coslalus species group. Characlers diagnostic of me group and of each species are described and illuSlrated, and keys 10 me species are presented. The new species O. achaelus. O. complanalus. O. denlalus. O. didymus. O. humalus. O. incurvalus. O. spinosus. and O. IUICatusare described. O. amula Curran and O. coslalusRondani are redescribed. Data are added 10 existing descriptions oC O. alexanderi. O. orellOquellSis. O. oreophilus. and O. piliferous. LectolypeS are designated for O. coslalus [=0. barbiellinii Curran, NEW SYNONYMYj and O. orellOquellSisBigot (=0. irifraclusScarbrough, NEW SYNONYMYj. O. spalulalus.a similar species, is compared wím me coslalus group.
Key words: Ommalius.Ommatinae, asilids,robbcr flies, Díptera, coslalusspecies group, neotropical.
Ommatius Wiedemann is one of the most here, increasing the number of neotropical widely distributed genera of the family species to 66. Asilidae, with about 200 species occurring in subtropical and tropical regions (Hull 1962). MATERIAL AND METHODS Fifty-eight neotropicaI species are presentIy known (Martín and Papavero 1970, Scarbrough General methodoIogy used in this study 1990, Scarbrough and Poinar 1992); 28modern foIlows that by Scarbrough (1990). Descriptive and two extánt species from the West Indies, 15 terminology foIlows that in McAlpine (1981). from Mexico .and Mesoameriea and 13 from Figures 1-11. 34 and 35 indicate terminology South America. Another 50 or more for structures of the terminalia, wing and mid neotropicaI species, in at least6 species groups, femur. Hind tibiae and characters of the remain to be described and reported lO science. terminalia are illustrated at the same scale. To This paper, which assigns 14 species to the conserve space, scale lengths associated with costatus species group, represents Part II of a these structures are listed only once in the series revising the New World Ommatius caption of Figures 1-11. Descriptions and Wiedemann. Part 1assigned eight spccies to the redeseriptions are composites of aH specimens pumilus species group (Scarbrough 1990). The in each species series. O. alexanderi is costatus species group ineludes six previously described in detail while only significant known species: O. oreophilus Farr and O . differences are included for the remaining alexanderi Farr (Jamaica, Farr 1965), and O . species. Characters omitted in the latter are piliferous Scarbrough (Cuba, Scarbrough essentiaIly identical to those Usted under O . 1985b); O. amula Curran (Mexico, Curran alexanderi. Only significant differences of 1928), O. coslatus Rondani (Brazil, Rondani characters areUsted for females. 1850), and O. orenoquensis Bigot (Guiana, The foIlowing ratios are used in the Bigot 1876). Eight new species are reported descriptions, and are defined here for the 730 REVISTA DE BIOLOGIA 1ROPICAL
convenience of the user. Face-to-head ratio Pesquisas da Amazonia, Manaus, Brazil; F. (FHWR) is the width of face at antenna/width Fernandez Yepes, (IZA) Instituto de Zoología of head (greatest width as measured Agrícola, University of Central Venezuela, horizontalJy from outer margin of one eye to Maracay; Donald Webb, (INHS) Illinois the other); width-to-Iength ratio of flagellum Natural History Survey, Champaign, ILL.; (FWLR) is the greatest lateral width/length (INBIO) Instituto Nacional de Biodiversidad of from base to apex; width-to-Iength of hind Costa Rica; (LJB) Larry Bezark, California femur (HFW LR) is the greatest vertical Department of Agriculture, Sacramento, CA.; width/dorsal length from the apex of the femur Charles L. Hogue, (LANHM) Los Angeles to the trochanter. Ratios are averages of 15 County Natural History Museum, CA.; J. specimens of each sex whenever possible. AH Manuel Ay ala L., (MAL) IsIe de Margarita, specimens were measured and averages Venezuela; D. J. Furth, (MCZ) Museum of calculated when the number of specimens in Comparative Zoology, Cambridge, MA.; Mike the series was less than 15. Ivie, (MIV ) Montana State University, Collection and locality data of new species Bozeman, MT.; Nelson Papavero, (MZSP) are standardized and Usted sequentially Museo de Zoologia, Universidade de Sao whenever possible, i. e. by country, sexes, Paulo, Brazil; C. A. Triplehorn, (OSU) Ohio numbers, state or province, specific location, State University, Columbus, OH.; D. Spencer elevation, date, and coIlector. Acronyms of Smith & George C. M. McGavin, (OXF) Hope museums oc private collections storing the Entomological Museum, Oxford, England; specimens are placed in parentheses following Donald Azuma, (PAS) Philadelphia Academy the locality data of each specimen or a series of of Science, Philadelphia, PA.; Hilda Atice de specimens. Curators and acronyms for O. Gasta!, (RGSZ) Museu de Ciencias Naturais collections are as follows: David Grimaldi, da Río Grande du Sol, Porto Alegre, Brazíl; (AMNH) American Museum of Natural Thomas H. Farr, (SMIJ) Science Museum, History, New York; A. G. Scarbrough, (AGS) Institute of Jamaica, Kingston, Jamaica; Todd Towson State University, Baltimore; K. V. G. Shelly, (TSC) Honolulu, HI.; R. O. Schuster, Smith & J. E. Chainey, (BMNH) Natural (UCD) University of California, Davis, CA.; History Museum, London, England; C. V. Saul Frommer, (UCR) Universíty of California, Riley, (B YU) Brigham Young University, Ríverside, CA.; L. Fernando Jiron, (UCRM) Provo, UT.; Norman Penny, (CAS) (California Universidad de Costa Rica, Ciudad Academy of Sciences, San Francisco, CA.; P. P. Universitaria, Rodrigo Facio, Costa Rica; Parrillo, (CFMNH) Chicago Field Museum of Antonio Rolondo, (UDT) Museo di Zoología, Natural History; IL.; Monty Wood & P. LaClair, DelIa Universito, Torino, Italia; G. W. Byers & (CNC) Canadian National Collection, Ottawa, R. W. Brooks, (UKN) University of Kansas, Canada: Chen W. Young, (CMNH) Carnegie Manhattan, KN.; F. Christian Thompson, Museum of Natural History, Pittsburgh, PA.; (USNM) United States National Museum of Jerry A. PoweIl, (CIS) University of California, Natural Hístory, Washington, D. C.; W. J. Berkeley, CA.; L. L. Pechuman, (COR) Cornell Hanson, (USU) Utah State Uníversity, Logan, University, lthaca, N. Y.; Eric M. Fisher, UT.; Roberto A. Zucchi, (USPP) Universidad (EMF) California Department of Agriculture, de Sao Paulo, Campus de Piracicaba, Sao Sacramento, CA.; Vanda H. P. Bueno, (ESAL) Paulo, Brazil Escola Superior de Agriculture de Lavras, Lavras, Brazil; Henry A. Hespenheide, (HAH) costatus species group University of California, Los Angeles, CA.; L. Tsacas, (NMNH) Museum National D'Historie Members of this species group are Naturelle, Paris, France; Howard L. Weems, recognized by a lancet or spear-shaped (FSCA) Florida State Collection of Arthropods, flagellum, conspicuous marginal scutellar Gainesville, FL.; Renato Contin Marinoni, bristles except O. achaetus, and one-three thin, (FUPC) Federal Universidade de Parana, proclinate postocular bristles on each side of Curitiba, Brazil; Abraham Willink, (IML) the head. In addition, the ventral setae of the Instituto Lillo Miguel, Tucuman, Argentina; J. fore femora and the posteroventral setae of the A. Rafael, (INPA) Instituto Nacional de mid femora are conspeciously long. SCARBROUGH:New World Ommalius 731
Males are recognized by the presence of a characters in combination. At least threeWest long, thin lO bristlypreapical, dorsoposterior seta Indian species (Scarbrough 1984) have similarly on the mid femora (Fig. 35; this structure is shaped mI cells but lack the thin, preapical, usuallythick and brown lOblack in other species dorsoposterior seta of the mid femur, and groups), anó the veins MI and M2 are slightly marginal scutellar bristles. Five undescribeó sigmoid, being slightly convex before the base South American species, in addition to O . of cell d and concave lOward the wing margino Iwlosenceus Schiner and O. exilis Curran, have Thus these converging veins produce a a thin mid femoral seta, and O. norma Curran constriction on the apical third of the mI cell anÓ O. neotropicus Curran have two-three long, (Fig. 34). The constriction of the mI cell is thinsetae on theepandr ium but each lacks other usually subequal lo the diameter of the cell attributes which would align them lO this species basally. The basal vein of cell mI is usually group. The costatus group may be the sister slightly oblique with its anterior end slightly grouplO the lineage giving rise lO O. spatulatus angled more lOwarÓthe wing base whereas the Curran. This relationship is evidenced by the posterior end is nearest the wing margino The general similarity of the body, the shape of cell apical margin of the wing is usually produced. mI anó the terminalia of both sexes. However, The epandrium has two lO furee long, thin setae O. spatulatus lacks the marginal scutellar or thick bristles (Figs. 2, 39, 134). The bristles, the long, proclinate, postocular bristles, gonostylus is slightly to strongly flat, and and the thin, preapical, dorsoposterior seta on usually wide in lateral profile. The rim of the the mid femur. Further stuóies are necessary lO gonocoxiteis usually a simple ridge or "elevated c1arify the relationship of the costatus species lip", although in a few species the inner margin group and O. spatulatus. Geographical range. is modified into a short to elongate digitate Cuba, Jamaica, Lesser Antilles, Mexico, process. The distiphallus is usually slightly or Mesoamerica, and South America south to strongly angled downward, and thin lo Tucuman, Argentina. Species of Ommatius are moderately thick in cross-section; the anterior thusfar not reporteófrom Chile orUruguay. marginof theaedeagal sheathis usually flaredas Remarks. Species of this group closely a low angled ridgeor a long flatprocess (Figs. 8, resemble each other. Few characters,apart from 19, 30, 43, 54, 73, 84, 95, 106, 117, 128, 139). the terminalia, can be used to construct a The female is characterized by a short dichotomous key. Dissection of the terminalia tergite 9, usually one-third or less the length of is necessary for identification of most the cercus, and the medioapical margin of specimens, especially females. In males, the sternite8 is narrowly produced with the corners epandrium, ventral lameBa, gonostylus, and angular. Spermathecae are usually elongate aedeagus are of greatimportance. Shapes of the with slightly crenulate surfaces. Two or three gonocoxal process anó hypandrium are posteroventral bristles in the posteroventralrow sorne times useful for identification of sorne of the hind femora arecontrastingly long. species. In females, relative length of tergite 9 Relationships. Sorne of the major diagnostic anó shape of the spermatheca are importantfor characters listed aboye are present in isolated identification. Shapes of the cercus and furca species of other species groups, but no known and stemite 8 usually lack significant specíes New World species possess all of these characters.
Key tothe adults of the costa tus species group males
1. Mid femur with a brown, moderately thick, preapical,dorsoposterior seta ...... 2 - Mid femur with a pale, long, thin, preapical, dorsoposteriorseta (Fig. 35) ...... 3
2. Hind tibia with a short, thick spurlike bristle anteriorly (Pig. 132); margin of scutellum with strong bristles; terminaliaas in Figs. 133-139; (Brazil, Colombia, Peru) ...... uncatus. N. Sp. - Hind tibia without a short, thick spurlike bristle anteriorly (Fig. 77); margin of scutellum with weak bristles; terminalia as in Figs.78-84; (Costa Rica, Guatemala, Honduras, Mexico, Nicaragua)...... incurvatus. N. Sp. 734 REVISTA DE BIOLOGIA1ROPICAL
11. Spermatheca narrowed basa1ly & apically, apex pointed (Fig. 33); sternite 8 usually with large
yellowish brown medioapical spot (Fig. 31); (Brazil)...... complanatus. N. Sp. - Spermatheca not unusually narrowed basally, each lateral gland usually narrowed apically with apex at least slightIy pointed & often directed toward median gland; median gland broadly rounded apically (Fig. 131); stemite 8 usually uniform brown (Fig. 129): (Bolivia, Brazil,
Ecuador, Pero) ...... •...... •...... spinosus. N. Sp.
Abdomen dark brown to blackish with SPECIES SYNOPSIS mostIy grayish tomentum and whitish vestiture, Ommatius alexanderi Farr all or most ter gites dorsally with brown Figures 1-11 tomentum and darkvestiture. Terminalia dark brown. Epandrium with Ommatius alexanderi Farr, 1965, 13 (2): apex acutely pointed. Gonostylus styliform. 24-25; Type locality: Sto Andrews, Ferry, Ventral lamella simple with basolateral margins Jamaica. Holotype d, and allotype Q, only slightly curved downward. Gonocoxite (USNM). Maninand Papavero 1970 (35b):59. with a prominent mediobasal process. Male. Brown to blackish. Length, body Distiphallus strongly decumbant, thin apically. 9.0-12.0 mm; wing 8.9-9.7 mm. Face yellowish Hypandrium simple. white to grayish tomentose with mostIy whitish Female. Length: body, 9.0-12.0 mm; wing, vestiture, four dark bristles present; FHWR 9.5-10.0 mm. FHWR 1/7.1; FWLR 1/1.9; 1/9.0. Palpus with whitish vestiture. Vestiture HFWLR 1/4.9. Tergite 9 about third as long as of antenna, frons, and ocellarium, blackish; cercus. Spermatheca oval, slight1y crenulate. FWLR 1/1.8. Occiput with four-five blackish Stemite 8 dark, apical third often lighter in postocular bristIes, two-three short with apical color; medioapical margin produced anterior, one-fourth toone-third of each proclinate. lateral corners subacutely rounded. Scutum mostly brownish tomentose, posthumeros, sides, grooves and prescutellum grayish; vestiture blackish, two notopleural, one supra-alar, one postalar bristle present; posteriorly, three-four dorsocentral setae present on each side, subequal to equal length and thickness of lateral bristles and greater in length and thickness than marginal scutellar bristIes.
Scutellum grayish tomentose with sparse whitish 1 - 3 pile and two marginal bristles. Pleuron mostIy grayish tomentose with whitish vestiture. Wings anteriorIy and apicalIy brownish; costal margin with weak dilatiónanteriorIy. Coxa with thin whitish to yellowish bristIes. 0\ Fore and mid femora largely brownish yellow to 4 5 -- reddish, apical one-third to one-half brown 6 - anterodorsaIly. Fore femora ventrally with several long whitish setae. Mid femora with three anterior, one dorsoposterior, and three-four anteroventral thick, blackish bristles; posteroventralIy, four-five long thin setae A presento Hind femora with apical half dark 7 8 brown, brownish yellow basaIly; aIl or most bristIes short, thick, anrl whitish; HFWLR 1/4.4. Figs.1-8. OmmatiusalexanderiFarr, maleo 1. Left hind tibia Tibiae with narrow apices and apical third of with a spurlike bristle, lateral view, 2-4. tenninalia, lateral, hind tibiae dark brown; hind tibiae with one ventral and dorsal views, S. ventral lamella, 6. left short, spurlike apical bristle. Basal tarsomere of gonostylus, 7-8. aedeagus, lateral and dorsal views. Abbr. : ce=cercus, ep=epandrium, hy=hypandrium, gs=gonostylus, fore and mid tarsi largely yellowish; fore tarsi . gcp=gonocoxal process. Sca1e: Figs. 1, S = 0.3 mm; Figs. with two-threeyellow bristles. 2-4, 7 = 0. 5 mm; Fig. 6 = 0. 1 mm; Fig. 8 = 0.2 mm. SCARBROUGH: New World Ommatius 735
Scutum with four-five dorsocentral bristles; scutellum with two-four, marginal bristles.Wing usuaUy with costal marginwealdy dilated. ¿l(\ ...... --_/ 666 large, red-black to yellow-brown, preapical or 9 11 basal spot posteriorly and ventrally; most ventral setae or bristles of fore and mid Figs. 9-11. Ommaliw alexanderi Farr, female. 9-10. femora unusually long and yellowish; míd tenninalia, ventral and dorsal views, 11. spennathecae. femur with one-two blackish, anteroventral Abbr.: ce=cercus, t9=tergite 9, s8=stemite 8. Scale: Fig. 9= bristles. Hind femur usually with basal fifth or 0.3 mm; Fig. 10 = 0.2 mm; Eg. 11 = 0.1 mm. less yellowish to reddish, all or most bristles usually brown to black, sometimes one-six Specimens examined. Holotype cJ, alIotype ventral bristles and one anterior bristle Q, 22 paratypes. whitish; one long, yellowish setae usually present basally in posterior row of bristles; Distribution: 17 April-2 October; JAMAICA. HFWLR 1/3.8. Narrow apices of fore and mid Collections with specimehs. AGS, BMNH, tibiae and apical one-third to one-half of hínd SMU, USNM. tibia blackish to reddish brown; anterior Remarks. The male of O. alexanderi is surface of each tibia usually with a recognized by a dark brown body, a clubbed brown-yellow strip, often extended entire terminalia, an epandrium which is acuminate length of tibia; hind tibia with one apical apically and abruptly angled posteriorly, and a spurlike bristle. Fore tarsus with one-four prominently developed gonocoxal pro�ess. !he whitish to yellow bristles. female is characterized by a long terglte 9, l. e. Abdomen mostly grayish lomen tose with about one-third as long as the cercus, and paJe yellow to whitish vestiture, tergites 1-5 spermathecae that are oval and slightly medially brown tomentose; tergites 6-7 usually crenulate. The female is similar to O. entirely brown with brown setae. oreophilus (Farr 1965) but is separated from Terminalia brown to black. Apical third of the latter by the uniform width of tergite 9. In each epandrium angled about 900 behind O. oreophilus. tergite 9 is much ¡onger laterally terminalia, apex flat horizontally with a shallow than medially. sinus between two differentially short lobes, and a vertical ridge which terminates in a short spine Ommatius amula Curran dorsally. Ventral lamella basolaterally C-shaped, Figures 12-22 with shallow depressíons formed by margins projecting ventrally. Gonostylus narrow and Ommati us amula Curran. 1928(327):4. subcylindrical. Distiphallus elongate, moderately Holotype Q, monotypic. Type locality: Amula, thick in cross-section, apically angled obliquely Guerrero, Mexico (S. W. Williston Coll.), downward; base of distiphallus in dorsal view (AMNH). Hull 1962, 224 (2):435; Martín and bordered laterally by two small flanges. Papavero 1970(35b):59. Gonocoxite with a large, thin, flat process Male. Length: body, 8.7-1 2.6 mm; wing, medially, usually transparent, and fused with rim 6.7-9.1 mm; body dark brown to black. Face laterad; the rim and process slightly angled yellowish gray to metallic yellow tomentose posteriorly forming a median, vertical groove in with light yellow setae and foor-eleven brown dryspecimens. bristles; FHWR 1/7.4. Palpus usually wilh Female, redescription. Length: body, mostly or entirely whitish to yellowish 1 O.3-l3.4 mm, wing, 7.9-11.0 mm; FHWR vestiture, rarely only black vestiture present 1/6.8; FWLR 1/1.8. Hind femur with one-three FWLR 1/ 1.8. Occiputwith five-sevenbrownish unusually long bristly, yellow setae basal1y; postocularbristles aboye each eye. 736 REVISTA DE BIOLOGIATROPICAL HFWLR 1/5.2. Tomentum of head and thorax usualIy grayish with only a slight tint of yellow. Specimens from the southem pan of the range have a darker yellow tomentum. Tergite 9 longin dorsalview, abouthalf as long as cercus. Stemite 8 with a large medioapical �,�� yellow spot. Spennatheca small, oval, slightly 12 13. - longer than wide, surface slightly crenulate Specimens examined. Holotype O, 96 dO',109 OO . Distribution: 6 July-12 November; MEXlCO: � Olihuahua, Guerrero, Jalisco, Michoacan, Cff!wJJ Morelas, Nayarit, Oaxaca, Sinaloa, Veracruz. BELIZE: El Cayo. GUATEMALA: Peten. EL SALVADOR: Quezaltpeque. Collections: AMNH, CNC, CMNH, EMF, FSCA, LAMNH, UCD, USNM, CFMNH, CIS. Remarks. O. amula is recognized by the A brown to dark black body and combined 18 18 -- characters of the terminalia. AIso, the male has a moderately swollen hind femur, and the Figs.12-19. Ommatius amula Curran, maleo 12. Left hind female has a large yellow spot on the tibia with a spurlike bristle, anterior view; 13-15. medioapicalmargin of stemite 8. terminalia, lateral,dorsal, and ventral views; 13a. apex of epandrium, lateral view; 16. ventral lameIla; 17. left Many specimens from the province of gonostylus; 18-19. aedeagus, lateral and dorsal views. Peten, Guatemala, and a few from the state Sc ale: Fig. 13a. = 0.1 mm. of Veracruz, Mexico, differ as follows from the "typical" black specimens: body cuticle light brown to brown, fore and mid femora yellow except for the narrow anterior and dorsal apices, brownish to slightly reddish brown; hind femur mostly brown, extreme 21fJ9 __ base yellow. The dorsoapical spine of the epandrium and the outer flange of the flat, gonocoxal process are significantly weaker or reduced. They are otherwise identical to the "typical" black specimens, and are .. dÓ included here as O. amula. The "typical" form of O. amula is similar Figs.20-22. Ommatiusam ula Curran, female. 20-21. to O. willistoni Curran, O. maculatus Banks Terminalia, ventral and dorsal views; 22. spermathecae. and O. parvulus Schaffer in that they are a11 black. This form, differs from the latter three Ommatius complanatus, N. Sp. species in the presence of scutellar bristles, Figures23-33 combined characters of the terminalia, slightly dilated costal margin of the wing, and the Male. Length: body, 10. 1-11.9 mm; wing, thin, hairlike, posterodorsal, preapical seta of 8-9. 7 mm. Body dark brown to blackish. Faee the mid femur. O. willistoni also differs in dull 10 golden yellow tomentose with six-eight having a wholIy whitish facial vestiture brown bristles; FHWR 1/8. 8. Palpus with onIy whereas O. amula has a few 10 several dark yellowish vestiture. FWLR 1/1. 8. Occiput with facial bristles. The light forms of O. amula are five-seven dark brown to blackish postocular· similar to O. incurvatus and O. humatus in bristles. color but differ in the combined characters of Scutum posteriorly with four-five long the terminalia. dorsocentral bristles in each row; five-six lateral SCARBROUGH: New World Ommatius 737 bristles present with one-two bristles subequal in thickness and length. Wing without anterior dilation, e slightly thickened medially. Legs mostly yellow with slight tint of red, browl'l to dark browl'l as follows: anterior and dorsal two-thirds to three-fourths of fore and - 2-3 mid femora with a diagonal stripe apically, narrow apices posteriorIy of fore and mid femora, apical two-thirds of hind femur, narrow apices of fore and mid tibiae, and apical half of !.�''\\ hind tibia. Mid femur with only one thin, O' í(¡ .� \ '\ ) � I, brown bristle anteroventrally; hind femur with csn \);'\,V '-'J one-two yellow bristles in each ventral row, ú¡ bristles otherwise dark brown; HFWLR 1/4.5. Hind tibia with one shon spurlike bristle apically, about one-fourth f.o one-third length of adjacent black bristle. Fore tarsus with basal segment mostly yeUow and two yellow bris!les; 211 mid tarsus with basal segment yellow-brown. Abdomen with tergítes mostly brown Figs. 23-30. OmrMlius complanalus, N. Sp" male, 23. Lef! hind tibia wÍlh a spurlike bristle, anterior view; 24-26. lomentose with brown vestiture, all stemites termínalia, lateral, ventral and dorsal víews; 27. ventral and lateral margin of tergites 1-5 yellowish lO lamella; 28. left gonostylus; 29-30. lIedeagus, lateral and slightly brownish yellow tomentose with dorsal view$. yellowish vestiture. Terminalia black. Epandrium with apex narrow, angled 900 subapically behind the terminalia, apex acuminate in dorsal view, rounded in lateral view; subapícal ventral margin of epandrium expanded as a thín, flat, subrectangular process. Ventral lamella produced ventrally in lateral view, basolateral figure-eight process elongate, antedorly with thick vertical walls. Gonostylus elongate, 31 subcylindrical. Gonocoxite basomedially with numerous bristles. Medioapical margin of hypandrium with dense yellow piJe, a sparse Figs. 31-33. Omrr.alius cOmpfafiallls, N. Sr" female. 31-32. transverse cluster of thin, yellowish sctae TerminaJia, ventral and dorsal views; 33. spermathecae. medially. Aedeagus with distiphallus moderatc!y thick in cross-section, angled ventralIy 450, sheath wings small, short and narrow. Holotype d, BRAZIL: S. P. ("",Sao Paulo), Female. Length: body, 10.9-1 3.2 mm; Est. Biol. Boraceia, Salesopolis, 850 m, wing, 8.5 mm. FHWR 1/7.9 ; FWLR 1/1.9; 1.30.68,Rabello, (MZSP); allotype Q & 1 d HFWLR 1/5.2. Scutum wíth four dorsocentral paratype, D.F .• Alto da Boa Vista Tijuca, bristles in each row, anterior setae shorter and m.1950,c, A. C, Seabra, (MZSP). Remainíng more sparse than in maleo Pore and míd paratypes: 1 d, Sao Paulo, Ferraz Vaseon, femora with díagonal brown streak 00 anterior Xn.1946,E. Rab, (l\1ZSP); 2 Q Q, Sao Paulo, and dorsal apical half or slightly less; mid Ciudade Jardim, n.20.1943,Can'era , (USNM); femur with one-two black. anterovci1tral 1 9.727,98'; 1 9, J, Paulisti, XII.5.44, bristles, Sternite 8 with medioapical surface Ramalha; 1 d . no data, (AMNH), yellowish; tergite 9 short, less lhan one Distribution: Dec.-Mar,; BRAZIL. seventh as long as cercus; spermatheca Etymology. Ladn complanatus ¡ meaning elongate, naITowed basally and apically, apex ;o flattened", refers to the ventral. subapicai pointed, surface weakly crenulate. process of the epandrium. 738 REVISTA DE BIOLOGIATROPICAL Remarks. O. complanatus is recognized by segments, vestiture otherwise yellowish to the combined characters of the terminalia. The whitish. male is further characterized by an absence of a Terminalia brown 10 dark brown. Epandrium costal dilation of the wing, entirely yellow apically slight1y angled posteriorly, with a paIpaI vestiture and only a single brownish prominent, dorsal, subapical, thom- like process; anteroventral bristle of the mid femur. apex flat, subtruncate, usually wíth a small median emargination. Gonostylus Ommatius costalus Rondani subcylindrical. Gonocoxite with a simple, flat Figures 34-46 mediobasal process. Base of distiphallus with two prominent winglike processes, apex strongly Ommatius costatus Rondani 185():188. Type curved down, thin in cross-section. Hypandrium locality: Distr. Brazil (Sao Paulo), Ilha de Sao and ventral lamella as inFigs. 38, 40. Sebastiao; Lectotype d, Museo di Zoologia, Female. Length: body, 9.5-15.0 mm; wing, DelIa Universita, Torino, Italia. 9.1-13.0 mm. FHWR 1/7.1. Scutum with Ommalius barbiellinii Curran 1934 three-four pairs of long dorsocentral bristles. (752): 18. Type locality: Sao Paulo, Brazil; Hind femur with two long, yellow, holotype d, allotype Q, 2 Q Q paratypes, posteroventral setae or bristles basally; (AMNH); New sy nonymy . Hull, 1962:435; HFWLR 1/5.4. Fore tarsus oflen entirely Martinand Papavero, 1970 (35b):59. brown. Tergite 9 short, about one-seventh or Redescription, maleo Length: body, 11.0-15.6 less as long as cercus. Spermatheca usually mm: wing, 8.8-13.0 mm. Body brown to dark spherical, rarely subspherical, surface smooth. brown. Face yellowish gray to golden yellow Material examined. Lectotype d; holotype tomentose with four-ten brown bristles; FHWR d, and 3 QQ, of O. barbiellinii Curran; 181 ln.7. Palpus with mostly or entirely yellow dd,238QQ . vestiture,often with a few to several brown Distribution: September-June; TRINIDAD bristles and setae. FWLR 1/2.3. Occiput with and SOUTH AMERICA except CHILE and five-six brown,postocular bristles. URUGUAY Scutum posteriorly with two-five pairs of Collections. AMNH, BMNH, CAS, CMNH, long dorsocentral bristles. Scutellum with CNC, CFMNH, INPA, IML, IZA, NMNH, two-four brown marginal bristles. Wing with FSCA,FUPC, MAL,MCZ, MIV, MZSP,OSU, costal margin slightly 10 moderately dilated. RGSZ,UDT, USPP, USNM. Fore coxa sometimes with one-three brown Remarks. O. costatus is recognized by the bristles. Fore and mid femora brown to subapical, spinelike process and the flat, yellowish brown, sometimes reddish,on apical subtruncate apex of the epandrium and the - two-thirds or less dorsally and anteriorly; hind subcylindrical gonostylus. The female is femur mostly brown-yellow to reddish, basal recognized by the smooth, subspherical to third or less yelIow. Mid femur with three-six spherical spermatheca and a short tergite 9, brown anteroventral bristles, posteroventral being less than one-seventh the length of the bristles either long, thin and yellowish or cercus. AH females of other species have oval moderately thick, long and brown. Hind femur or elongate spermathecae with slightly to posteroventrally with one thin yellowish seta strongly crenulate surfaces. betweenbasal two bristles,length about as long This species is one of the two most common as adjacent bristles; HFWLR 1/5.0. Apical and abundant neotropical species in collections, one-third to one-half of hind tibia brown, one and one of the most variable in color and size. spurlike bristle present apically. Fore and mid The color of the tomentum ranges from a weak tarsi with basal half yellowish 10 yellow brown, yellow 10 brownish yellow or reddish; vein C is sometimes reddish, fore tarsus usually with only slightly thickened and weakly dilated in one-two yellow bristles. sorne specimens whereas in others it is quite Abdomen largely brownish lomentose thick and moderately dilated; the anterior and dorsally,sometimes reddish to grayish, sides of dorsal surfaces of the fore and mid femora tergites and all stemites usually yellowish gray range from narrowly brown apically 10 almost tomentose; vestiture of tergites largely brown entirely brown whereas the posterior and ventral dorsal1y, especially on apical three-four surfaces are usually mostly or entirely yellow. SCARBROUGH: New World Ommatius 739 The posteroventral bristles on the mid femur are moderately long and thick and usually brown in specimens from southero Brazil and Argentina whereas these bristles are mu�h thinner, slightly longer and usually yellow m specimens in other areas of South America. Similarly the scutellar bristles are usualIy short and thin or hairlike in specimens in the southern extreme of its range whereas they are significantIy thicker and slightly longer in 44 specimensfro m more northeropopulations. 46 O. costatus is separated from all others in the species group by the combined characters Figs. 44-46.0mmalius coslatus Rondani, female. 44-45 Terminalia,ventral anddorsal views; 46. spermathecae. of the terminalia. The terminalia of both sexes are almost identical to those of O. spatu/atus. A male from the Rondani collection in the O. spatulatus differs in the lack of marginal Museo di Zoologia, DelIa University, Torino, scutellar bristles, and the presence of a slender Italy, is in excellent condition, having onIy lost yellowish brown hind femora with only pale the right styIus and flagellum, left tibia and fore yellow toyellow bristles on the hind femora, a tarsus, and the two apical tarsomeres of the thick, posterodorsal, preapical bristle, and a right tarsus. The pin with the specimen bears a spatulate abdomen. The male of O. costatus hand written labeI with the name Ommatius has a long, thin posteroventralseta on the mid costatus Rondi., Lupd. S. Sebast. geni, and a remur, largely darkbrown bristles below the second label with the word Holotype, the name hind femora, and the abdomen is only slightly Ommatius tibialis Rondo, N. Papavero 72 spatulate. (misidentified). The specimen is essentially iden tical to the holotype of O. barbiellinii Curran. The latter name is thus a synonym of O. costatus, and the Rondani specimen is here ��designated Lectotype. Ommatius dentatus, N. Sp. q � LV Figures 47-57 3S 3� 37 -. - Male. Length, body 11.3 mm; wing, 8.4 mm. Face and occiput light yellowish gray tomentose with pale yellowish to whitish � vestiture; FHWR 1/8.3. Palpus yellowish ¡(I),\;\)� (� brown with brown bristles apically. FWLR � v 1/1.8. Each side of occiput with six-seven - brown postocular bristles. Y!9 Scutum laterally with four long dorsocentral bristles. Wing without an anterior dilation, C &\ only slíghtly thickenedmedialIy. AH femora mostly yelIow, apical third of � � .,- fore and mid femora with onlya faint brownish ·41- color dorsoapically; hind femur wíth most of dorsum and anterior apical half brown, Figs. 34-43. Ommatius coslalus Rondani, maleo 34. Right wing; 35. left mid femur with posterodorsal preapical thin, otherwise yellow; mid femur anteroventrally hairlike setae, dorsal view; 36. right hind tibia with a short, with onIy two long, thin, brown bristles. Hind spurlike bristle, lateral view; 37 -39. terminalia, lat�ral, femur with mostIy brown bristles, at least one ventral and dorsal views; 40. ventral lamella; 41. nght anterior and two ventralbristles yellow; HFWLR 42-43. . gonostylus; aedeagus, �ateral a�d d�rsal vie,:"s 1/5.1. Hínd tibia brown on apical third, spurlike Abbr.: C=costal dilation, MI =fust medlal vem, mI =flr8t medial cell, S= !hin hairlike seta. apical bristle absent. Basal segment of fore and 740 RBVISTA DB BIOLOOIA TROPICAL mid tarsi mostly yellow; fore tarsus with one yellow bristle. Abdomen with tergites mostly brown,a pices yellowish, sternites lighter yellow-brown; tomentum yellowish to slight1y grayish laterally and ventrally on most segments with light vestiture: dorsum of tergites 1-6 with small to large brown tomentose spot and 47 brownish setae: anterior corners of segment 6, most or a11 of tergites 7-8 sparsely tomentase. Terminalia brown. Epandrium with apex long, toothlike. Gonostylus flat and wide in lateral view, abruptly narrowed subapically. Basolateral margin of ventral1amella produced Q� ventrally, and cupshaped. Aedeagus with apex 11 curved strongly downward, apodeme wide in lateralview , moderatelythi ck distal1y; aedeagal flanges moderately strong in dorsal view. Hypandrium with dense yellow pile anteriorly, medioapical margin truncate, and a subapical transverse cluster of yellow bristly setae. 53 Female. Length: body, 12.3-16.7 mm; wing, 10.7-13.7 mm . FHWR ],,/8.2: FWLR 1/2.1; Figl. 47-S4. Ommatiua túnlatua, N. Sp., maleo 47. Leftbind HFWLR 1/5.3. Body yellow tomentose, Cace tibia with slender bristlel only, anterior view; 48-S0. 1. witlt Cour-five brown bristles and palpus with terminalia, lateral, ventral and dorsal viewI; S ventral lame11a; S2. right SonOllylul; S3-S4. aedeasus, lateral and vestiture entirely brown; brownish streak oC dorsal viewI. femora and tergites much darker brown than in maleo Mid fem ur with three-four anteroventral bristles. All bristles of hind femur black except for one anterior, yellowish red bristle; br istles absent on basal third of hind femur posteroventrally.Apical two-tltirds of hind tibia � 58 brownish yellow 10 brownish. Abdomen witlt tergites much darker brown titan in male, lateral margins of tergites 1-5 or 6 narrowly reddish to reddish yellow, sternites yellowish brown; tergites mostly brown 55 . " " .: tomentose with brown setae; sternites with ," " . sparse, yellowish tomentum. Tergite 9 short, @�. one-sixth or less as long as cercus. Spermathcca 57 . elongate, sausagelike, apically rounded, with a Fiss. SS·S7. Ommatiua túntatua, N. Sp., female. SS-S6. weak constriction admedially, surface weakly Terminalia,ventral and dorsal views; S7. spermathecae. crenulate. Holotype d, BRAZIL: Manaus, Res. 1210 RLO, Bk (X.22 85), V. P. Daniel (alcohol speci men, mounted on a pin), (INPA). Allotype O, Etymology. Latin dentatus, meaning and paratype d, BRAZIL: terr. Amapa', Serra "tootlted", refers 10 tite suong toothlike apex of do Navio, IX.28.1957, K. Lenko, (MZSP); 1 O, the epandrium. #5650, Para,Tucurui-nha Chorona, VIII. 17 .1980, Remarks. O. dentatus is recognized by tite eq. Nunes de Mello, (INPA); CAYENNE: 2 mostly yellow 10 reddish pleura and tite brow 00, V.1917,ac c. 6008, (CMNH). nish streak on the anterodorsal surface of each Distribution: May-Oct.; BRAZIL, CAYEN femora. In addition, the strong toothlike apex of NE. the epandrium and the apically truncate hy- SCARBROUGH: New World OlNPUll i"" 741 pandrium characterize the maleo TIte shape of the spennatheca separat.e it from the females of spinosusand complanatus. Two females (#5650 and 6008» are darker than the other female paratypes. TIte pleura of specimen #5650 are brown on the dorsal half � - ,, II lO and the femora are brown anteriorly and dorsally except on the basal one-fourth to one- third which is yellow to yellow-brown. The pleura of one specimen (#6008) are entirely blackand the femara are dark brown 10 blackish on the apica1 third or more. Ommatiusdidymus, N. Sp. Figures 58-65 Male. Length: body, 12.1 mm; wing, 10 mm. Face golden yellow tomentose, with six brown bristles and sparse brown setae aboye mystax; FHWR 1/8.7. Palpus brown with mostly yellow vestiture, apical bristles brown lO brownish yellow. FWLR 1/2.5. Occiput with Pi.l. 58-65. Ommati"" didymu8, N. Sp., male. 58. Left hind seven postocular bristleson each side of head. tibia withllender bri.t1ea,anterior view; 59-61. terminali.., Scutum laterally with four dorsocentral lateral, ventral .nd dorsal view.; 59 •• ventral procell of bristles. Wing without an anterior dilation, C epandrium; 62. ventral lamella; 63. left 10000ltylu.; 64-65. slightly thickened media1ly. aedeaSU1,lateral and dorsal. Scale: Pi¡. 59. = 0.1 mm. Pore and mid femara mostly yellow, apical third or less brownish dorsa1ly and ant.eriorly, angled downward at 450 angle, ventrally with a n arrow apices brown post.eriorly. Mid femur flat, apically truncat.e, plat.e which bears a thick, with two dark brown anteroventral bristles. medioventral ridge t.enninating in a short, acute Hind femur with apical half brown, basal half spine; flanges of acdeagal sheath absent; yellow, all bristles dark brown or black except ejaculatory apodeme narrow, spatulate. one anterior and one ventral yellow bristle; Hypandrium with mostly yellow setae or weak post.eroventral.row wi� one long, thick, yellow bristles, subapical median margin with fine, seta present between basal two bristles; dense,yellow pile. HFWLR 1/4.3. Narrow apex of mid tibia and apical third of hind tibia, brown; spurlike apical Female unknown. bristle absent. All tarsi brown; fore tarsus with Holotype d, BRAZIL, #2876, Amazo one yellowbristle. nas-Manaus, Est. nml, km 134, VII.10.1968, Abdomen dark brown to blackish with (INPA). mostly light grayish to brownish gray Distribution: July; Brazil. lOmentum and pale yellow 10 yellow vestiture, Etymology. Latin didymus, meaning most t.ergites with brown tomentum dorsally "divided", refers lO the divided thin, ventral with increasing density of brown setae on process of the epandrium. apical four-five t.ergites. Remarks. The male of O. didymus is 'Ienninalia blackish brown10 slightly blackish recognizedby theabsence of a costal dilation and red. Epandrium with apex narrow and by thecombined charact.ers of thetenn inalia. subtruncate; ventralmargin with a flat, bifurcated process. Ventral lamella with an inverted C Ommatius humatus, N. Sp. shaped ventrolateral process. Gonostylus Figures 66-76 elongat.e and subcylindrical. Each gonocoxite with one narrow, bladelike mediobasal process. Male. Len¡th: bOdy,10.5-15.6 mm; win¡, Disdphallus moderately thick in cross-sectlon, 9.1-11.9 mm. Bady reddish brown to blackilh. 742 REVISTA DE BIOLOGIA TROPICAL Face bright to dull yellow tomentose with Medioapical margin of sternite 8 weakly two�seven long, thin, brown bristles; FHWR produced. Spermathecae oval, slightly longer l/8.0. Palpus with vestiture entirely yellow to than wide, surface smoothto weakly crenulate. mostly brown. FWLR 1/2.7. Occiput with Holotype eJ, COSTA RICA: Prov. five-sevenbrown postocular bristles. Heredia, F. La Selva, 3 km s Pto Viejo, 10026' Scutum laterally with four-five long brown N 80 01 W, IV.9.1985. H. A. Hespenheide; dorsocentral bristles. Pleuron mostly bright allotype O, sarne dataexcept V1.18.1985, (US yellow tomentose. Wing with anterior margín NM). Paratypes. BELIZE (= British Honduras): moderatelydilated. 1 O, J. M. Aldrich; 1 O, Frio Temez; 1 O, Fore coxa sometimes with one-four brown Manatos, (AMNH); COSTA RICA: 1 O, bristles. Fore and mid femora mostly yellow, Heredia Prov.• 4 km se Puerto Viejo de apical third or less brown; mid femur with Sarapiqui, Finca La Selva. IX.15.1981, Charles two-three brown anteroventral bristles. Hínd E. Griswold; 1 eJ, 1 O. Heredia. 3 km s Puerto femur with apical one-third to four-fifths dark Viejo, IV.18.72, P. A. Opler. (CIS); 1 O. Prov. brown, all or most bristles dark brown to black, Alajuela. Est. Exp. Fabio Baudrit. often one-two anterior and ventral yellow VII1.16.1967, G. Miranda. (UCRM); 1 eJ. 1 bristles present; posteroventral row of bristles O, Cart. T urrialba. CATIE. VI.26-29.1986, W. with one short, thin, yellow seta between basal Hanson, G. Bohart; 1 eJ. 2 Q O, same two bristles, length as long as adjacent bristles; location, VIII.20.l989. F. D. Parker; 25 eJeJ, HFWLR 1/5.2. Apical one-third to one-half of 24 OQ, Guan.(Guanacaste Pk), 20 km s Upala. hind tibiabrown with one short, spurlike bristle, V.6-XII.6.l990, P. D. Parker; 28 eJeJ, 24 QQ, length one-fourth to one-third nearest apical sarne location, 1.1 .-XII.5.l991; 4 eJeJ, 2 QQ, bristle. Fore tarsus with only one-two yelIow same location. X.l-1O.1992, XI.1l-20.1992; 1 bristles. eJ, 2 Q Q, Guan., 3 km se R. Naranjo. Abdomen mostly brown-yellow tomentose IV.1l-21.1992, XI.11-20.1992, F. D. Parker; 1 with lateral margins of tergite 1 and basal eJ, 2 O O, Guan., 14 km s Canas, corners of tergite 2 and all sternites yellowish II.15-24.1990. IX.1-15.1990, 1 ,1.17-24.1991, to light brownish gray tomentose, tergites 1-7 1 • I1I.16-28.1991. F. D. Parker; 1 eJ, Guan, with dark brown tomentum medially, exy sta s Canas. VIII.� .1988, F. D. Parker; 1 appearing as a narrow strip on tergite 1 and (5. 3 km se R. NaranJO, IX. 10-20. 1992 F. D. becoming wider on each succeeding tergite; Parker. (USU); 2 QQ. VIl.13,VIIIA.1957. lateral margins of tergites 6-7 shiny, with a ArnoldMenke; 1 O, I1CA grounds, Turrialba, dense brown tomentose triangle dorsalIy, base XII.8.62, C. L. Hogue, (LANHM); 2 eJeJ. 3 of triangle widest at base of tergite 6 and apex QQ. Alajuela, 20 km w San Antonio, Hwy. 3, at apical margín of tergite 7; tergite 8 600 m, VII.17.75, E. M. Fisher; 5 Q Q, narrowly tomentose apically. Vestiture of Cartago, 5 km e Turrialba, CATIE, abdomen mostly yellowish, brown tomentose VII.21 ,22.75, E. M. & J.L. Fisher; 1 eJ. latter areas with brownsetae. location and date, E. M. Fisher; 3 eJeJ, 16 Terminalia dark brown. Epandrium with an Q Q, Prov. Heredia, F. La Selva. 3 km s Pto. acuteapex. Ventrallamellabasolaterally angular, Viejo. 100 26' N 840 01' W, VII.23-31.1976, H. not unusually enlarged or projecting ventrally. A. Hespenheide/E. M. Fisher; latter location, 3 Gonostylus flat, wide medially, narrow apically. eJeJ, IV.2-12. 1983, 1 , 2 , IlI.28-30.1984, 9 DistiphalIus narrow in cross section and eJ eJ, 2 Q Q. I1I.29-VII.6.1985. H. A. cylindrical apically, strongly decurved and Hespenheide; 2 eJeJ, 5 Q Q, Puntarenas. recurved subapically; aedeagal sheath usually Prov. Parque Nacional Corcovado, Est. Sirena, with two dorsolateral flanges in dorsal view. 8° 28-31' N 830 36' W, I1I.17-25.1981, H. A. Gonocoxiteand hypandrium simple (pig.68). Hespenheide; 1 eJ, Limon Prov., Guapapiles, Female. Length: body. 10.4-14.4 mm; wing, VI.24.65, R. J. Hamton; 2 eJeJ, Puntar, F.Las 8.7-10.9 mm. FWLR 1/2.1; FHWR 1/6.6. Hind Cruces, 6 km s San Vito, 1200-1400 m, femur with basal two-three yellow VIlI.21,25.76, E. M. Fisher, (EMF); 5 eJeJ. 13 posteroventral bristles significantly longer than Q, Prov. Heredia, F. La Selva, 3 Km S PTO. remaining ventral bristles; HFWLR 1/5.5. Viejo, 100, 20' N 840 01' W, VI. 16- Tergite 9 one-third as long as cercus . VIII.13.1982. III.25,30.1984, IV.I-VII.6.1985. SCARBROUGH: New World OmmaJius 743 111.22.1987. I1I.30.1988, IV.21-IX.23.1989, Zone, Barro Colorado Island, n.22.67, Roger D. IV.28.1990, prey: 1 Psocoptera, 1 Díptera, Akre (FSCA); 3 d d , 3 O O, Canal Zone, Hemiptera (2 Fulgoridae), Hymenoptera (1 Chiva Rd. 8 air km n Ft Clayton VII.23.1978, Echaritidae & 2 Formicidae) & 6 Coleoptera (3 Eric Fisher; 1 d, PanamaPt, Cerro Jefe, 1000 m Scolytidae. 1 Chrysomelidae, 1 Platypodíae, 2 (nr C. Azul), VII.31.78, E. M. Fisher, (EMF); unid.).H. A. Hespenheide, (HAH/lNBIO); 1 d, NICARAGUA: 1 O, Chinandega, Baker, Pedregoso, D. L. Rounds; 1 O, SanJose', 10 km (USNM); ECUADOR: 2 dd, 1 O, Los Rios, n VIII.9.1972, J. Maldonado C., (USNM); EL Queredo Pichilingue, V. 1936, Fritz leg, coll SALVADOR: 1,No. 444-16B, Stelco, V.20.54, Martinez, (MAL); 1 d, Provide, Los Rios, M. S. V., (AMNH); GUATEMALA: 1 d Guat., 111.7.38, W. Clarke-McIntyre, A. E. Pritchard 31 km ne Guatemala, 915 m, VII.4.1977, E. M. CoL, (USNM); 1 d , Pich. e Sto Domingo, Fisher; 1 d, 1 O, Peten, 10/3 1 km se Flores V.6-12.1990. W. R Hanson, (USU). VIII.26,27.1974, VII.7 .1977, E. M. Fisher, Distribution: January-December; MEXICO, (EMF); 1 d, 1 O, La Providencia Obispo, MESOAMERICA,ECUADOR. IV.14. J. M. Aldrich; 1 d, Gualan, 1.12.1905, Etymology. Latin h,umatus. meaning auls.C. Deam, (AMNH); 1 d, La Providencia "hooked", refers to the shape of the distipballus. Obispo, C. M. Roiullard, A. E� Pritchard, Remarks. O. humatus differs from other (USNM); 1 O, Gualan, 1.22.05; 1 O, Santa species by the bright yellow tomentum of the Lucia, 11.2.1905, (OSU); HONDURAS: 1 San pleuron and the combined characters of the Pedro, VIII.21.05, (OSU); 1 d, Minas de Oro, male terminalia. It is most similar to O. Comay, 4000ft, V.29, J. B. Edwards (AMNH); 3 oreophilus (parr 1965) fromJamaica but differs d d, Copen, Ruines de Copan, VIII. 22,23. in its type locality, in its lighter body and 1974, E. M. Fisher, (EMF); MEXICO: 2 00, yellow tomentumand in tbe male terminalia. Xilitla S. L. P., 1450 ft., Vn.23.54, Univ. Kans. Exp., (UKN); 1 O, Chiapas, Tapachula, m.7.52, Females of O. humatusand O. orenoquen Crawford. (OSU); 1 a, Tab. (=Tabasco), sis are difficult to separate, especially in re Frontera, 6.4, Townsend; 1 O, Tabasco, Teapa, gions where tberanges of tbe two species over 11, H. H. S.; 1 O, Gro., AcaJ)lllco, VI.18.1936, lap. The light brownish color of tbe wings, tbe A. E. Pritchard, (USNM); 1 t:J, Ver. (Veracruz) FortiD de las Flores, 3000', Vm.l1.62, R. H. & E. M. Painter,(FSCA); 1 d. Veracruz, Fortin de las Flores, Vm.17.196S, A. S. Gillogly; 7 d d, 1 Q, Veracruz, 6.Sn km ne Catemaco, 300-305 m, vm.1.1977, E. M. Fisher & P. Sullivan; latter location. 2 d d, 2 QQ,VI.27-VIII.1.1977, E. M. Fisher; latter location/date/col. except 14 km s Catemaco, Id; 1 d, Oaxaca, 42 km n Matias Romero, vn.26.1963, 1 , S. Luis Potosi, 13 km ne Xilitla, VI.24.1977, E. M. Fisher, (EMF); 1 d, Ver. (=Veracruz), Telcolopa, V.28.51, A. Berrera, (CNC); 1 d, Veracruz, Metlac, 910 m, XI.5.1975, H. Brailovsky, (MAL); PANAMA: 1 d, 1 Q, Barro Colorado l., Panama C. Z., 1.1947. N. L. H. Krauss; 1 , Taboga Is., VI.29. 1924, N. Banks, S. W. Bromley; 1 Q, Taboga Island, XI.23.1923, F 4863; 1 d, Trinidad Rio, III.6.1912, A. Busch; 1 Q, Trinidad Riv, m.30 - .1912, VIII, Busch; 1 Q, Surnmit,Panana C. Z., 72 7 S XI.1946, N. L. H. Krauss; 1 Flamenco I. X .. l946, N. L. H. Krauss, (USNM); 1 Q, 4 km Figs.66-73. Ommati'!S humatus. N. sp.• maleo 66. Left hind w Balboa, Pan Am Hwy, Canal Zone, tibia witb a spurlike bristle, anterior view; 67-69. tenninalia, lateral, ventral, and dorsal viewI; 70. ventral VI.20. 1974, M. L. Siri, (UCD); 1 d BCI, lame11a; 71. left gonostylus; 72 -73. aedeaguI, lateral and V. 17.1980, Todd Shelly, (TSC); 2 00, Canal donal views. 744 REVISTA DE BIOLOGIA TROPICAL usually slightly yellowish brown to darker brown, sometimes anterior femur largely dun OCj yellow-brown to brown anteriorly. Mid femur with anterior and anteroventral bristles long, �-- 15 two-mree bristles abaut one and one-fourth to one and one-half times width of mid femur mediany (these bristles in other species usually equal to the greatest width of the mid femur); ll �\ three-five dark brown bristles present in anteroventral row; posterodorsal, preapical 'M -�_.- l)U bristle brown, moderately thick. Hind femur --- 16 withapical half to four-fifths reddish brown to Fi,!. 74-76.0_/iU$ n_tus, N. Sp., femate. 74-75. brown and only brown bristles; HFWLR 1/4.2. Tem1i1'!1I1il!, vmln.l alld donru.Iil'ÍIeWII; 76. spermllthec!ie. Apical third of hind tibiae reddish brown; hind tibia apically without spurlike bristle. Fore darker brown color oi the femora, and the tarsus usually withone yellow bristle. narrow yellow base oí the hind femora, in Abdomen yellow to brownish yellow except combmation with other charaeters, are llsually as follows: narrow apical margin of tergite 1, reliable for separation of O. humatus and O. narrow transverse band on tergite 2, small - orenoquensis. The basal third 01' more of the medial spots on tergites 3 and 4, most 01'aIl of hind femora of O. orenoquensi.s is llsually tergites 5·8, these areas brown to black with yeUow. Spermathecae of O. huma tus are brown tomentum and usually brown to reddish consistently oval and their surfaces are only brown setae, vestiture otherwise yellow with weakly crenulated. The spermathecae of O. yellowish gray to yellow tomentum. orenoquensis are alsooval, but usually tapere yellow, and yellow vestiture; tergite 8 with (USU); GUATEMALA: 3 d d, Baja sparse brown tomentum, surface subshiny. Verapaz, 50 km s Purulha, 925 m, 20 m, Tergite9 sOOrtin dorsal view, about fifth length VII.20.1977, E. M. Fisher, P. Sullivan, of cercus.Spermailieca elongate, abouttwice as (EMF); Id, 2 OO. GuaL, 31 km ne long as wide, sunace slightly crenulate. Guatemala, 915 m, VII.l,4.1977, E. M. Fisher; 1 d, 2 O Q, Peten, Ruinas TJ.k al, 245 Holotype. d, GUATEMALA: 31 km ne m, VII.7,lO.77, E. M. Fisher, (EMF); Guatemala, 915 m, VIIA.1977, E. M. Fisher, HONDURAS: 2 d d. 3 O O, Puerto (EMF); allotype O. GUATEMALA: Peten, Castilla, VI.21 -23.26, R. H. Pa¡nter, Runias Tikal, 245 m., VII.7,1O.77, E. M. & J. (AMNH); MEXICO: 1 d, Tepotzlan, L. Fisher, (EMF). Paratypes. COSTA RICA: 1 IX.7.1951, F. M. Hull, (CNC); 1 , Agua O. Prov. La Pacifica, 4 km nw Canas, Priesta, VII.18.37, (BYU) ; 1 d, Alencingo, Guanacaste, IX.15,20.1973, P. A. Opler. (CIS); VI.2.1922, E. G. Smyth; 1 O, Mor 6 OO. Prov. Guanacaste, OTS, Palo Verde (=Morelos), Ac atlifun, X.4.49, W. G. Downe, Sta., 29 km wsw Canas, 100 21' N 850 21' W, (USNM); 1 d. Nayarit, 30 km n Puerto Vn.l-ll.1976, H. A. Hespenheide, E. M. Vallarta, VI.21.1983, L. G. Bezark, (LGB); 4 Fisher; 4 O O, Guanac., 11/32 km nw O O, Nayarit, 55/82 km sse Acaponeta, Liberia, VII.12.1975, E. M. Fisher; 1 O, VI.27.75, E. M. Fisher; 1 O, Chiapas, 6 km Cartago, 5 km e Turrialba, Catie, nw El Jaote, 610 m, VII.1.1977, E. M. Fisher; VII.21,22.75 E. M. & J. L. Fisher, (EMF); 1 2 d d, 3 O O. Tamaulipas, 7 km s O, Higuito, San Mateo, Pablo Schild, Tomaseno, 335 m, VI.22,23.77, E. M. & J. L. (USNM); 2 d d, Guanacaste Pk, 14 km s Fisher, (EMF); NICARAGUA: 4dd, 1 O, Canas, XIl,7-9 .1990, F. D. Parker; 3 d d, Leon, 36 km se Leon, VIII.22.1975, E. M. same data except m.8-31.1991; lO, same Fisher, (EMF). data as before except IV.23-30.1991; 1 cJ, lO, same data as before except V.20-29.1991 Distribution: June 2-0ct. 4; MEXICO to COSTA RICA. Etymology. Latin incurvatus, meaning "curved inward", r eferring to the emarginate apical margin of the epandrium. Remarks. O. incurvatus is recognized by its largely yellowish gray tomentum, yellow -brown to yellow body; yellow vestiture of the !�íSJ palpus; short, thin, marginal scutellar bristles; combined characters of the male terminalia; 79 and the elongate spermathecae. In addition, the male has a moderately thick, brown, O:f!>r5 posterodorsal preapical bristle. 82 _ � 83 84 Figs. 77-84. OmmatiusiflcUTvalus . N. Sp., maleo 77. Lef! hind tibia with only slender bristles, lateral view; 78-80. terrninalia, lateral, ventral, and dorsal views; 81. ventral lamella: 82. right gonostylus; 83-84. aedeagus, lateral and Figs. 85-87.0 mmatius iflCUTvalUS. N. sp., female. 85-86. dorsal views. Tenninalia. ventraland dorsal views; 87. sperrnathecae. 746 REVISTA DE BIOLOGIA TROPICAL Ommatius orenoquensis Bigot Female. Length: body, 8.5-15.0 mm; wing, Figures 88-98 7.9� 13.5 mm. FHWR !n6; FWLR 112.00 Fore femora, mid femora and hind tibiae usually Ommatius orenoquensis Bigot 1876:1xxxxv, with apica1 one-thirdto one-halfbrown to dark Type locality: Guiana (Cayenne), Distr. brown; apical two-thirds to three-fourths of Guyana. Lectotype Q. Hope Entomological hind fein ora brown-darkbrown; femora anteco Collection, (OXF) . Martin and Papavero. basalIy often reddish. Hind femora basalIy with 1970(35b):59. two long yellowish bristles, about one and Ommatius infraetus Scarbrough one-half times or more length of other longest 1985a:643-644. TyJJe locality: Sto Vincent bristIes. Tergite 9 long, at leastone-third length Island, holotype (J.(USNM) . New SY· of cercuS. Spermatheca oval, surface crenulate, NONYMY. apex somewhat pointed, often retracted. Male. Brown. Length: body, 10.6-16.5 mm; Distribution: January-December; COSTA wing, 9.7-14.9 mm. Face andfront usua1ly pale RICA and STo VINCENT ISLAND southward, brassy yellow lOmentose, with mostly pale ye SOUTH AMERICA except URUGUAY and llow vestiture; face with 4-8 brown bristles; CHILE. FHWR ln.9. Palpus usually with vestiture en Specimens examined. Lectotype '1,of O . tirely yellow lO brown, often mixed yellow and orenoquensis Bigot; type series of O. in/rae brown. FWLR 1/2.0. Occiput with four-eight tus; 220 cJcJ,226 '1'1. brown postocular bristles on eachside of head. Collections: AMNH, BMNH, CAS, CMNH, two-three moderately proclinate. CNe, CFMNH, EMF, FSCA, FUPC, INPA, Scutum laterally with three prominent dor IML, INHS, IZA, MAL, MCZ, MIV, MZSP, socentral setae, subequal in length and thick NMNH, OSU, OXF, USNM, USU. ness to lateral bristles. Pleuron mostly grayish tomentose, anepistemum and katepistemum Remarks. O. orenoquensis Bigot is recogni brownish yellow. Wing slightlyyellowish ante zed by the combined characters of the termina riorly; costal margin slightly dilated anteriorly. lia: male with epandrium curved posteriorly, Legs mostly yellow to reddish yellow; fore apex subtruncate and flattened dorsoventrally; and mid femora with apical fourth or more process of gonocoxite elongate and flat; go brownish, hind femora and hind tibiae with nostylus flat, wide most of its length, abruptly apical one-third to one-half brown. Basal half narrowed subapically; female with tergite 9 of mid femora with four brown anteroventral fourth or more the length of cercus, and sper bristIy setae or bristIes. Hind femora usually matheca oval, length between one and with all dark.brown bristIes ventrally; HFWLR one-fourth to one and one-half its width, surfa 1/4.7. Hind tibiae with a spur-like apica1 bris ce strongly crenulate, sometimes pointed api tIe. Fore and mid tarsi with basal segments in cally. part yellowish; fore tarsus laterally with Two females from the Bigot collection, one-two yellowbristles. in the Hope Entomological Collection, bear Abdomen dorsally with dark. brown tomen identification labels of this species. One fe tum andsparse lO entirely brown setae; sides of male has three labels with the following in tergites and stemites entirely light yellowish to formation: "714 in Coll Bigot"; "coll. Bigot yellowish brown tomentose with pale yello abt. 1845-93. preso 1914 by J. E. Collin."; wish vestiture. "Type Dip: 290 1/2, Ommatius orenoquen Terminalia brown. Epandrium strongly cur sis Bigot, Hope Dept. Oxford." The second ved posteriorly, apex flat dorsally, wider than female also has three labels with similar in constricted subapex, weak emargination pre formation: "Ommatius orenoquensis Bigot sent. Ventral lamella simple (Fig. 92 ). (hand written), 714 in Coll Bigot"; "coll Gonostylus flat, wide in lateral view. Bigot abt. 1845-93 preso 1914 by J. E. Gonocoxite basomedially with a long, flatdigi Collin".; Type Dip: 290 2/2, Ommatius ore tate process. Aedeagus thin in cross-section, noquensis Bigot, Hope Dept. Oxford." Both strongly decumbant, basally with prominent specimens are severely damaged although winglike flanges. Hypandrium simple, subtrun the wing venation and terminalia were sui cate apically. table for study. Examination of these struc- SCARBROUGH: New World OmmaJ ius 747 Holotype d and Q, alIotype (USNM), 8 d d , and 14 Q Q paratypes. Martin and Papavero, 1970 (35b): 59. Male. Length: body, 13.0-16.0 mm; wing, 10.0-11.0 mm. Face, frons and occiput grayish to slightly brownish gray tomentose with largely whitish vestiture; face with two-four brown bristles; FHWR 1/1.8. Palpus with whitish vestiture. FWLR 1/1.9. Occiput often with aH white postocular bristlesor only one or two brown. Scutum with four long, thick dorsocentral bristles, much longer and thicker than marginal scuteHar bristles. Pleuron light yellowish gray to gray tomentose. Wing with only a slight anterior dilation. es -- Fore and mid femora largely brown Figs. 88-95.0mmaJius orenoquensis Bigot, maleo 88. Left anteriorly and dorsally, brownish yellow to hind tibia with a spurlike bristle, anterior view; 89 -9 l. yellow elsewhere. Hind femora with apical terminalia, lateral, ventral and dorsal views; 92. ventral lamella; 93. right gonostylus; 94-95. aedeagus, lateral and two-thirds brown, basal third yellow; each dorsal views. ventral row with three-four brown bristles; HFWLR 1/4.5. Fore and mid tibiae brownish anteriorly; hind tibiae with apical three-fourths brown, brownish yellow basally, and a short spurlike bristle. Tarsi brownish; fore tarsi with m three-fouryellow bristles. 97 -- Abdomen largely brownish tomentose dorsallywith brown vestiture; lateral and apical margins of tergites and all sternites grayish Óóó tomentose with pale vestiture. 98 Figs. 96-98. Ommatius orenoquensis Bigot, Cemale. 96-97. Tenninalia, ventral and dorsal views; 98. spermathecae. tures revealed that the two females were different species. The wing venation and �too� � terminalia of specimen number 290 1/2 were 101 ___ identical to those of O. infractus Scarbrough, whereas those of specimen 290 2/2 were identical to O. norma Curran. The specimen numbered 290 1/2 is declared here ��� LECTOTYPE for the species and bears a label o » -- as such. The species name O. infractus thus 102 103 104 - becomesa synonym of O. orenoquensis. � IJ 105 Ommatius oreophilus Farr 105 Figures 99-109 Figs. 99-106 Ommatiusore op hilusFarr, maleo 99. Right hind tibia with a spurlike bristle, anterior view; Ommatius oreophilus Farr, 1965, 13(2): l00-102.terminalia, lateral, ventral, and dorsal views; 103. 21-22, Type locality: Hardwar Gap, Jamaica. ventral lamella; 104. leCt ¡onostylus; 105-106. aedeagus, lateral and dorsal vi ews. 748 REVISTA DE BIOLOGIA TROPICAL Tenninalia dark brown. Epandrium elongate face with six brown bristles; FHWR 1/9.1. with apex acutely pointed dorsally. Ve ntral Palpus withpale yellow vestiture. FWLR 1/1.7. lamella with basal corners rounded. Gonocoxite Occiput with one-two brown postocular and hypandrium as in Figs. 100-102. Gonostylus bristles,short and moderatelyproclinate. slightlycurved forward,broad and flat in profile. Scutum with three brown dorsocentral setae Distiphallus apically thin in cross-section, on each side. Scutellum with two brown, weak, strongly decurved with apex slightly recurved; marginal setae. Wing with costal margin'only aedegal sheath apically and distiphallus basally slightlydilated anteriorly. with a pairof short,narro w flanges. Legs mostly yellow 10 reddish-yellow, brown as follows: apical one-third 10 one-half of fore Female. Length: body, 14.0-16.0 mm; wing, and mid femora, apical one- third 10 one-half of 10.5-11.9 mm. FHWR 1/7.9; FWLR 1/2.2; hind femora, narrow apices of mid tibiae and HFWLR 1/6.1. Tergite9 laterally long, dorsally apical third of hind tibiae. Hind femora venttally short, about one-third as long as cercus. with largely black bristles; HFWLR 1/4.5. Hind Medioapical margin of sternite 8 moderately tibiae with a spurlike bristle'. Basal tarsomere of produced with large yellowish spot. fore and mid tarsi largely yellow; foretarsi with Spennathecae oval, longer than wide, surface four yellow bristles. weakly crenulate. Abdomen dorsally mostly brown tomentose with brown vestiture, ventrally and narrow lateral margins of most tergites brownish gray 10 grayish with whitishvestiture. ffi Terminalia brown. Epandrium subtruncate apicaí'ly. Ventral lamella with basolateral 108 - co�rs angular. Gonostylus flat and wide in profile with short hooked apex. Gonocoxite simple with a recessed subtruncate process. .0 Distiphallus moderately decurved and thin in 1 6 FiI•. loo-loo. Ommatilll orlophillll Farr, female. 107-108. TenninaHa,ventral and dorsal views; 109. spennathecae. Specimens examined. Holotype d, allotype Q, 22 paratypes. Distribution: June 24-September 18; 111 JAMAICA. Collection with specimens. AGS, 110 BMNH, SMIJ,USNM. Remarks. O. oreophilus, probably a sister species of O. huma tus N. Sp., differs in the darkerbody, narrow face, grayish tomentum of the pleuron, and combined characters of the terminalia. Ommatiuspilif erous Scarbrough Figures 110-120 118 Ommatius piliferous Scarbrough, 117 __ 1985b:I227-1229. � locality: Cuba, holotype (J,allotype Q, (MCZ); Fils. 110-117. Ommati'uspilifero lll Scarbrough, maleo 110. Q Left hind tibia with only slender bristles, anterior view; paratype d & "(AMNH). I1l-ll3. tenninaHa, lateral, ventral, and donal VieWI; 114. Ma1e. Length, body 12.0 mm; wing 10.7 ventral lamo1la; llS. riaht aonOlt)'luI; 116-117. udoaaU1, mm. Pace and front pale yellow tomentose, lateraland donal viewI. SCARBROUGH: New Wodd Omnllltill8 749 mostIy yellow with apical one-third to four-fifths dorsally and anteriorly brown 10 8 yellowish brown. Mid femora with two-three 111 -- long,thin, brown anteroventral setae ocbrisdes. Hind femora with apical one-half10 four-flfths dark brown. yellowish basaUy; brisdes mosdy brown, often one-two pale yellow to yellow 66 bristles present anteriorly and/or ventralIy; HFWLR 1/4.7. 120 -- Fore and mid tibiae yellow except for narrow brown apices and a slender brownish Fi¡I. 118-120. Omntlltill8 pilif,roll8 Scarbrough, female. streak on the mid tibia; brisdes mostly yellow, 118-119. Terminalia, ventral and dorsal views; 120. those at apices, brown; hind tibia brown on spermathecae. apical half or slightly more anteriorly. only brown brisdes presento Tarsal segments usually cross-section. and basolaterally with a pair of brownish withbrown brisdes, basal segment of small, thin winglike process oc tlange; aedeagaJ fore tarsus usually yellow. sometimes only sheath anterolaterally with a pair of larger weakly yellowish basally ; fore tarsus wingHke processes. Hypandrium medioapically sometimes with one yeUow brisde. with a dense pad of pale piJe. and medially a Abdomen mosdy light brown to yellowish sparseband of shortbrown setae. brown tomentose. small to large brown Female. Length. body 10.0-11.0 mm; wing lOmentose sPOtson tergites 1-6 medially; apical 9 .2-10.3 mm. Palpus with two-three brown two-tbree tergites shiny. sparsely tomentose. setae.FHWR 118.0; FWLR 1/1.7. Hind femora lateral margin of tergite 1. stemite 1 and basal with 4/5 brown; black ventral bristles present third of lateral margin of tergite 2 and stemite 2 exceptbasalIy two long yellow setae; HFWLR densely gray tomentose. Ve stiture brown in 1/4.8. Hind tibiae with apical one-half to dark tomentoseareas. with yeUowish 10 whitish two-thirds brown. Tergite 8 about half as long vestitureel sewhere.' as cercus. Sternite 8 as in Fig. 118. Terminalia brown lO blackish. Epandrium Spermathecaewith oval crenu1ate surfaces. with one-two long. thick, brown or black Distribution: June 26-August27; Cuba. brisdes and one longer, thinner yellowish seta; Specimens examined. Type series. apex of epandrium short, pointed. Ventral CoUections with specimens. AMNH.MCZ. lamella with baso lateral margin flat and Remarks. O. piliferous differs from other projecting ventrally, vertical width abaut half species in its type location and the combined its length. Gonostylus flat and moderately wide charactersof the terminalia. medialIy, strongly narrowed subapicalIy. Gonocoxite simple. Aedeagus strongly curved downward, moderately thick apically; Ommatiussp inosus. N. Sp. anterolateral flanges of aedeagal sheath only Figures 121-131 slightly developed. Hypandrium with � short medioapical spinelike process, postenorly a Male. Length: body. 10.0-14.5 mm; wing. preapical thin. often contrasting lighter in color, 9.3-12.5 mm. Face with four-five brownish flexible cuticle and a transverse cluster of bristles; FHWR In.7. Palpus with vestiture yellow to brownishbrisdy setae. . entirely or mostly yellowish, few to several Female. Length: body, 9.1-13.3 mm; wmg, yeUowish brown tobrown brisdes ofteo present. 9.5-11.7 mm. Face with 4-9 brownish bristles FWLR 1/2.2. Occiput with six-twelve dark present; FHWR 1/6.8; FWLR 1/2.3. Legs �k brown postocularbristles on eachside of head. brown anteriorly and dorsally. Basal one-thll'd Scutum with one extra but weaker postalar of hind femur with two long, thin, setae; and three-five dorsocentral brisdes present on HFWLR 1/4.6. Sternite 8 often with a large, each side. Pleuron mostly yellowish gray thin, anteromedial spot. Tergite 9 unusually na tomenlOse. Wingwith costal margin weakly lO rrow dorsally, about one-eight length of cercus; moderately dilated. Fore and ,mid fe mora spermatheca elongate, outer two spermathecae 752 REVISTADE BIOLOGIA1ROPICAL Shannon, (USNM); COLOMBIA: 1 d, Rio Tacana, Amazonas,XI.I- 13.1946, L. Richter & Frank lo1mson donors, (AMNH); BRAZIL: 1 Q, Faseuda Taperinha, Santarem, PA , X-XI.l970, Exp. Perm. Amaz.,(MZSP) Distribution: Oct.-luly; BRAZIL, COWMBIA, PERU Etymology. Latin uncatus, meaning "hooked", refers lO the shape of the apex of the epandrium. R�marks. O. uncatus is recognized by the reddish yellow colored cuticle of the pleuron and the combined characters of the terminalia. In addition, the male has a bristly, posterodorsal, preapical seta onthemid femur. Ommatius achaetus, N. Sp. Figures 143-150 150 1.1 Male. Length, body 12.1 mm; wing 8.8 mm. Body brown. Face and front yellowish gray tomenlOse; face, palpus and scape ventrally Fi.¡s. 143-151. OmmatillS acluietllS,N. Sp., maleo 143. Left . 144-146. W1th onIy pale yellow vestiture; FHWR In .6. hind tibia with a spudike bristle, anterior view' tenninalia, lateral, ventral and dorsal views; 147. apex of FWLR 1/1.7. Ve stiture of antenna largely, front epandrium with a median procesa; 148. ventral lamella; and ocellarium entirely, and three-six 149. left gonostylus; 150-151. aedeagus, lateral and dorsal �stocular bristles, brown. Occiput with apical views. Salle: Fig. 147 = 0. 1 mm. thtrd of one-two lateral postocular bristles stronglyproclinate. 147. Gonostylus slender in lateral view, weakly Scutum with three-four dorsocentral setae curved Corward, apex narrow, truncate. � latter thick and subequaI in length 10 laterai Gonocoxite with nat, apically rounded bnstles Scutellum and pleuron yellowish gray : basomedial process and a flat lateral flange. 10 graylSh tomentose; scutellum with only short Aedeagus decurved, moderately thick in pite, bristles absent. Wing hyaline with anterior cross-section; two elongate lateral flanges on margin weakly dilated. each side oC distipha1lus. Hypandrium apica1ly Fore coxae with several long, slightly thick subtruncate, a wide band oC scattered brownish bristle�. Fore and mid femora largely brown, bristles present medially. �stenorly and ventralIy largely yellowish. Female unknown. Hmd Cemora largely brown with narrow bases yellow with only brown bristles; HFWLR Holotype d, MEXICO: Oaxaca, 7 mi n 1/4.6. Tibiae largely yellow with narrow luchatengo, 3700', 10 VIII. 1970, E. Fisher, P. ap�ces brow�; hind tibiae apically with one . . Sullivan. splDebk� bnstle. Tarsi entirely brown; fore Distribution. August; MEXICO. tarsus Wlth two yellow bristles. Etymology. Latin achaetus, "a" meaning A�domen with basal Cour segments thin absent and "chaetus" bristles, reCers to the graylsh tomentum and pale vestiture, apical absence oC marginal scutellar bristles. Cour segments with brownish to brown Remarks. Ommatius achaetus is similar 10 tornentum and mostly brown vestiture. O. costatus, O. orenoquensis and O. humatus Termin�lia brown. Epandrium apically but can be distinguished Crom them by the narrow wlth a shallow emargination and absence oC marginal scutellar bristles and the pointed corners, a prominent spine present combined characters oC the terminalia. This which originates Crom apical inner surCace speciesis sympatric with the blackform of O. of emargination. Ventral lamella as in Fig. SCARBROUGH: New World Ommilli", 753 humatus.and canbe further separated fram that Curran. C. H. 1928. New species of Om"u"i"" from speciesby its lightercolor. America, with key. (Asilidae, Diptera). Amer. Mus. Novitates 327:1-6. ACKNOWLEDGEMENTS Curran, C. H. 1934. New American Asilidae (Diptera). m. Amer. Mus. Novitates 752:1-18. Thanks are extended to tite individuals and Farr, Tbomas H. 1965. Tbe robber-flie. of Jamaica institutions in the Materials and Methods sec (Diptera, Asilidae). Part 2. The subfamilies tion for arranging loans of specimens used in Dasypogoninae, Laphriinae and Asilinae. Bull. Inst. this study. 1 extend my appreciation to Robert Jamaica, Sci..Ser. 13: 17-26. Peterson and Christian Thompson of the Hull, F. M. 1962. Robber t1ies of the world. The genera of Systematic Entomology Laboratory, ARS, US thefamily Asilidae. Bull. Smithsonian Institution, U. S. DA, Wa shington. D. e .. Stephen Bullington, National Museum 224(2):433-435. Department of Entomology, Pennsylvania State University, and two anonymous reviewers for Marun, Charles H. &; Nelson Papavero. 1970. A cata their helpful comments and suggestions. logue of the Diptera ol ·the Americas south ol the United States, Family Asilidae. Bull. Mus. ZooI., Univ. Sao Paulo 35b: 59-60. McAlpine, J. F. 1981. RESUMEN Morphology and terminology - adults, p. 9-63. 111 J. F. Catorce especies de Ommatius Wiedemann del Nuevo Mundo son asignadas al grupocos McA1pine, B. V. Petenon, G. E. Shewell, H. J. Te skey, J. R. Vockeroth D. M. Wood (eds.). Manual ol Neartic tatus. Este articulo describe e ilustra los carac Diptera. Biosys. Res. InSL, Agr. Can., Ottawa. Monog. teres diagnósticos del grupo y de cada especie. '27. Ademásse presentauna clave para lasespecies. Se describen en detalle las siguientes especies: Rondani, A. C. 1850. Observazioni sopraalqante speci.e di esapodi ditteri del Museo Torinese. Nuovi Ann. Sci. O. achaetus. O. complanatus. O. dentatus. O. NaL 801ogna,ser. 3, 2:188. didymus. O. humatus. O. incurvatus. O. spino sus. y O. uncatus. Las especies O; amula Scarbrough, A. G. 1984. Synopsis ol Ommatius Curran yO. costatus Rondani fueron redescri Wiedemann lrom Hispaniola. J. N. Y. Entornol. Soc. taso Tam bién se aftaden datos a las descripcio 92: 131-149. nes existentes de O. alexanderi. O. orenquen Scarbrough, A. G. 1985a. Ommati",Wledemann (Diptera: siso O. oreophilus y O. piliferous. Se designan Asilidae) in the Lesser Antilles. Proc. EntornoI. Soc. nuevos lectotipos para O. costatus (=0. bar Wa sh. 87:641-655. biellinü Curran - sinónimo nuevo) y O. ore n quensis Bigot (=O. infractus Scarbrough - sinó Scarbrough, A. G. 1985b. New Ommati""Wiedemann J nimo nuevo). A la especie parecida, O. spatu (Diptera: Asilidae) lrom Cuba and the Babamas. . N. Y. EntomoI. Soco 93:1226-1239. Scarbrough, A. G. latusse la compara con el grupo O. costatus. 1990. Revision ol the New World Ommatill.s Wiedemann (Diptera: Asilidae). L Thepumil"" species group. Trans. Amer. EntomoL Soco 116:65-102. REFERENCES Scarbrough, A. G. &; George O. Poinar, Jr. 1992. Upper Bi¡ot. J. M. F. 1876. [Note: Descriptions of four new spe Eocene robber flies of the genus Ommati"" (Diptera: cie. of Asilidae). Bull. Soco Ent. France. ler. 5. Asilidae) in Dominican Amber. Insecta Mundi 6: lxxxv-lxxxvi. 6:13-18.