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The Case of the Diminutive Trilobite Flexicalymene Retrorsa Minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati Region
EVOLUTION & DEVELOPMENT 9:5, 483–498 (2007) Evaluating paedomorphic heterochrony in trilobites: the case of the diminutive trilobite Flexicalymene retrorsa minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati region Brenda R. Hundaa,Ã and Nigel C. Hughesb aCincinnati Museum Center, 1301 Western Avenue, Cincinnati, OH 45203, USA bDepartment of Earth Sciences, University of California, Riverside, CA 92521, USA ÃAuthor for correspondence (email: [email protected]) SUMMARY Flexicalymene retrorsa minuens from the upper- rate of progress along a common ontogenetic trajectory with most 3 m of the Waynesville Formation of the Cincinnatian respect to size, coupled with growth cessation at a small size, Series (Upper Ordovician) of North America lived ‘‘sequential’’ progenesis, or non-uniform changes in the rate of approximately 445 Ma and exhibited marked reduction in progress along a shared ontogenetic trajectory with respect to maximum size relative to its stratigraphically subjacent sister size, can also be rejected. Rather, differences between these subspecies, Flexicalymene retrorsa retrorsa. Phylogenetic subspecies are more consistent with localized changes in analysis is consistent with the notion that F. retrorsa retrorsa rates of character development than with a global hetero- was the ancestor of F. retrorsa minuens. F. retrorsa minuens chronic modification of the ancestral ontogeny. The evolution has been claimed to differ from F. retrorsa retrorsa ‘‘in size of F. retrorsa minuens from F. retrorsa retrorsa was largely alone,’’ and thus presents a plausible example of global dominated by modifications of the development of characters paedomorphic evolution in trilobites. Despite strong similarity already evident in the ancestral ontogeny, not by the origin of in the overall form of the two subspecies, F. -
Available Generic Names for Trilobites
AVAILABLE GENERIC NAMES FOR TRILOBITES P.A. JELL AND J.M. ADRAIN Jell, P.A. & Adrain, J.M. 30 8 2002: Available generic names for trilobites. Memoirs of the Queensland Museum 48(2): 331-553. Brisbane. ISSN0079-8835. Aconsolidated list of available generic names introduced since the beginning of the binomial nomenclature system for trilobites is presented for the first time. Each entry is accompanied by the author and date of availability, by the name of the type species, by a lithostratigraphic or biostratigraphic and geographic reference for the type species, by a family assignment and by an age indication of the type species at the Period level (e.g. MCAM, LDEV). A second listing of these names is taxonomically arranged in families with the families listed alphabetically, higher level classification being outside the scope of this work. We also provide a list of names that have apparently been applied to trilobites but which remain nomina nuda within the ICZN definition. Peter A. Jell, Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia; Jonathan M. Adrain, Department of Geoscience, 121 Trowbridge Hall, Univ- ersity of Iowa, Iowa City, Iowa 52242, USA; 1 August 2002. p Trilobites, generic names, checklist. Trilobite fossils attracted the attention of could find. This list was copied on an early spirit humans in different parts of the world from the stencil machine to some 20 or more trilobite very beginning, probably even prehistoric times. workers around the world, principally those who In the 1700s various European natural historians would author the 1959 Treatise edition. Weller began systematic study of living and fossil also drew on this compilation for his Presidential organisms including trilobites. -
The Annals and Magazine of Natural History
This is a digital copy of a book that was preserved for generations on library shelves before it was carefully scanned by Google as part of a project to make the world's books discoverable online. It has survived long enough for the copyright to expire and the book to enter the public domain. A public domain book is one that was never subject to copyright or whose legal copyright term has expired. Whether a book is in the public domain may vary country to country. Public domain books are our gateways to the past, representing a wealth of history, culture and knowledge that's often difficult to discover. Marks, notations and other marginalia present in the original volume will appear in this file - a reminder of this book's long journey from the publisher to a library and finally to you. Usage guidelines Google is proud to partner with libraries to digitize public domain materials and make them widely accessible. Public domain books belong to the public and we are merely their custodians. Nevertheless, this work is expensive, so in order to keep providing this resource, we have taken steps to prevent abuse by commercial parties, including placing technical restrictions on automated querying. We also ask that you: + Make non-commercial use of the files We designed Google Book Search for use by individuals, and we request that you use these files for personal, non-commercial purposes. + Refrain from automated querying Do not send automated queries of any sort to Google's system: If you are conducting research on machine translation, optical character recognition or other areas where access to a large amount of text is helpful, please contact us. -
001-012 Primeras Páginas
PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. -
Cambrian Trilobite Ovatoryctocara Granulata Tchernysheva, 1962 and Its Biostratigraphic Significance
Available online at www.sciencedirect.com Progress in Natural Science 19 (2009) 213–221 www.elsevier.com/locate/pnsc Cambrian trilobite Ovatoryctocara granulata Tchernysheva, 1962 and its biostratigraphic significance Jinliang Yuan a,*, Yuanlong Zhao b, Jin Peng b, Xuejian Zhu a, Jih-pai Lin c a Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, East Beijing Road 39, Nanjing 210008, China b College of Resource and Environment Science, Guizhou University, Guiyang 550003, China c School of Earth Sciences, Ohio State University, Columbus, OH 43210, USA Received 28 March 2008; received in revised form 27 May 2008; accepted 14 August 2008 Abstract The genus Ovatoryctocara Tchernysheva, 1962, and its key species Ovatoryctocara granulata Tchernysheva, 1962, are revised. Ovatoryctocara granulata occurs near the base of the Ovatoryctocara Zone and ranges up into the lower portion of the Kounamkites Zone in the Siberian Platform. O. granulata also appears in southeastern Guizhou, South China, but O. granulata in northern Greenland may represent an indefinite species. Specimens of Ovatoryctocara from Newfoundland cannot be identified to species level. Specimens includ- ing two cranidia and three pygidia from the lower part of the Aoxi Formation at Yaxi Village, Shizhu Town, eastern Tongren, north- eastern Guizhou, were previously assigned to O. granulata, which is now reassigned as a new species O. yaxiensis sp. nov. It bears the following main features: glabella club-shaped, slightly expanded medially, with four pairs of lateral furrows, of which S1–S3 are trian- gular pits, S4 is shallow, connecting with axial furrow; shorter palpebral lobe situated a little anterior to the midway of facial suture across the fixigenae, longer posterolateral area (exsag.); semielliptical pygidium consisting of seven axial rings with a terminal piece and with eight pairs of marginal tips giving a sawtooth-like shape of the lateral margins in dorsal view. -
United States
DEPARTMENT OF THE INTERIOR BULLETIN OF THE UNITED STATES ISTo. 146 WASHINGTON GOVERNMENT Pit IN TING OFFICE 189C UNITED STATES GEOLOGICAL SURVEY CHAKLES D. WALCOTT, DI11ECTOK BIBLIOGRAPHY AND INDEX NORTH AMEEICAN GEOLOGY, PALEONTOLOGY, PETEOLOGT, AND MINERALOGY THE YEA.R 1895 FEED BOUGHTON WEEKS WASHINGTON Cr O V E U N M K N T P K 1 N T I N G OFFICE 1890 CONTENTS. Page. Letter of trail smittal...... ....................... .......................... 7 Introduction.............'................................................... 9 List of publications examined............................................... 11 Classified key to tlio index .......................................... ........ 15 Bibliography ............................................................... 21 Index....................................................................... 89 LETTER OF TRANSMITTAL DEPARTMENT OF THE INTEEIOE, UNITED STATES GEOLOGICAL SURVEY, DIVISION OF GEOLOGY, Washington, D. 0., June 23, 1896. SIR: I have the honor to transmit herewith the manuscript of a Bibliography and Index of North American Geology, Paleontology, Petrology, and Mineralogy for the year 1895, and to request that it be published as a bulletin of the Survey. Very respectfully, F. B. WEEKS. Hon. CHARLES D. WALCOTT, Director United States Geological Survey. 1 BIBLIOGRAPHY AND INDEX OF NORTH AMERICAN GEOLOGY, PALEONTOLOGY, PETROLOGY, AND MINER ALOGY FOR THE YEAR 1895. By FRED BOUGHTON WEEKS. INTRODUCTION. The present work comprises a record of publications on North Ameri can geology, paleontology, petrology, and mineralogy for the year 1895. It is planned on the same lines as the previous bulletins (Nos. 130 and 135), excepting that abstracts appearing in regular periodicals have been omitted in this volume. Bibliography. The bibliography consists of full titles of separate papers, classified by authors, an abbreviated reference to the publica tion in which the paper is printed, and a brief summary of the con tents, each paper being numbered for index reference. -
A Trilobite with North American Affinities from the Upper Cambrian of Sweden
A Trilobite with North American Affinities from the Upper Cambrian of Sweden By Gunnar Henningsmoen (h lo ABSTRACT.-Taenicephalus? peregrinus n.sp. is described from the Upper Cambrian Olenus beds in Sweden. l t recalls North American genera of the families Pterocephalidae and Paraboli noididae, and is regarded as one of the rare occasional invaders into the Upper Cambrian sea of Scandinavia. Introduction During an excursion to Västergötland in June 1956, Dr. V. JAANUSSON of Uppsala found a cranidium of an undescribed trilobite in the Olenus beds at Trolmen, Kinnekulle. A second, probably conspecific cranidium was collected there by me. This species, described below, is of particular interest because it apparently shows North American affinities and may thus be one of the rare occasional invaders into the Upper Cambrian sea of Scandinavia. The occur rence may be compared with that of Irvingella, and of Drepanura, described by WEsTERGÅRD (1947, 1949), also from the Upper Cambrian of Sweden. The specimens described belong to the Museum of the Palaeontological Institution of the University of Uppsala (U.M.). I wish to thank Miss B. MAURITZ for taking the photographs, Miss I. Lowzow for preparing the draw i ng, and Dr. W. C. SWEET for reading the manuscript. Description Genus Taenicephalus ULRICH & RESSER, 1924 TYPE SPECIEs.-Conocephalites shumardi HALL, 1863, by origi nal designation. Taenicephalus? peregrinus n. s p. Pl. I; Text-lig. r. DERIVATION OF NAME.-From Latin: peregrinus, foreign. MATERIAL.-Two incomplete cranidia, U.M. Nos. V g. 705 (holotype) and V g. 706. 12- .16�272 !lull. n/ (;rnl. Vol. X.\.\ VII x68 GUNNAR HENNINGSMOEN HoRIZON.-Both specimens occur in hituminous limestone (stinkstone), associated with Olenus gibbosus and Agnastus (Homagnostus) obesus, and thus belong to the Upper Cambrian zone of Olenus, subzone of Olenus gibbosus and Olenus transversus (cf. -
Smithsonian Miscellaneous Collections
SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 53, NUMBER 6 CAMBRIAN GEOLOGY AND PALEONTOLOGY No. 6.-0LENELLUS AND OTHER GENERA OF THE MESONACID/E With Twenty-Two Plates CHARLES D. WALCOTT (Publication 1934) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION AUGUST 12, 1910 Zl^i £orb (gaitimovt (pnee BALTIMORE, MD., U. S. A. CAMBRIAN GEOLOGY AND PALEONTOLOGY No. 6.—OLENELLUS AND OTHER GENERA OF THE MESONACID^ By CHARLES D. WALCOTT (With Twenty-Two Plates) CONTENTS PAGE Introduction 233 Future work 234 Acknowledgments 234 Order Opisthoparia Beecher 235 Family Mesonacidas Walcott 236 Observations—Development 236 Cephalon 236 Eye 239 Facial sutures 242 Anterior glabellar lobe 242 Hypostoma 243 Thorax 244 Nevadia stage 244 Mesonacis stage 244 Elliptocephala stage 244 Holmia stage 244 Piedeumias stage 245 Olenellus stage 245 Peachella 245 Olenelloides ; 245 Pygidium 245 Delimitation of genera 246 Nevadia 246 Mesonacis 246 Elliptocephala 247 Callavia 247 Holmia 247 Wanneria 248 P.'edeumias 248 Olenellus 248 Peachella 248 Olenelloides 248 Development of Mesonacidas 249 Mesonacidas and Paradoxinas 250 Stratigraphic position of the genera and species 250 Abrupt appearance of the Mesonacidse 252 Geographic distribution 252 Transition from the Mesonacidse to the Paradoxinse 253 Smithsonian Miscellaneous Collections, Vol. 53, No. 6 232 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 53 Description of genera and species 256 Nevadia, new genus 256 weeksi, new species 257 Mcsonacis Walcott 261 niickwitzi (Schmidt) 262 torelli (Moberg) 264 vermontana -
Bathynotus: a Key Trilobite Taxon for Global Stratigraphic Boundary Correlation Between Cambrian Series 2 and Cambrian Series 3
Available online at www.sciencedirect.com Progress in Natural Science 19 (2009) 99–105 www.elsevier.com/locate/pnsc Bathynotus: A key trilobite taxon for global stratigraphic boundary correlation between Cambrian Series 2 and Cambrian Series 3 Jin Peng a,b,*, Yuanlong Zhao b, Jinliang Yuan c, Lu Yao b, Hong Yang b a Department of Earth Sciences, Nanjing University, Nanjing 210093, China b College of Resource and Environmental Engineering, Guizhou University (Caijiaguan Campus), Guiyang 550003, China c Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China Received 14 March 2007; received in revised form 4 March 2008; accepted 4 March 2008 Abstract The Cambrian trilobite Bathynotus is a special morphology of Redlichiid, which is characterized by a wider thorax axis and an 11th macropleural segment bearing a long, backwardly directed spine. Additionally, its 12th and 13th pleural segments are narrow and they terminate at a fulcrum where they fuse together and to the posterior edge of the long macropleural spine. The genus is widely distributed, occurring in North America, Siberia, the Altay-Sayan fold belt of Russia, South China, the Tarim Basin of China, and Australia. It occurs primarily in deposits of the upper part of the continental slope, at the transition between stable platform settings and subsiding margins of the late Early Cambrian. Here, we recognize three species of Bathynotus from China, including B. holopygus which is wide- spread around the world. Although six species of Bathynotus have been reported from Siberia and the Altay-Sayan fold belt area of Rus- sia, data presented here suggest that only three of these species are valid. -
Arthropod Pattern Theory and Cambrian Trilobites
Bijdragen tot de Dierkunde, 64 (4) 193-213 (1995) SPB Academie Publishing bv, The Hague Arthropod pattern theory and Cambrian trilobites Frederick A. Sundberg Research Associate, Invertebrate Paleontology Section, Los Angeles County Museum of Natural History, 900 Exposition Boulevard, Los Angeles, California 90007, USA Keywords: Arthropod pattern theory, Cambrian, trilobites, segment distributions 4 Abstract ou 6). La limite thorax/pygidium se trouve généralementau niveau du node 2 (duplomères 11—13) et du node 3 (duplomères les les 18—20) pour Corynexochides et respectivement pour Pty- An analysis of duplomere (= segment) distribution within the chopariides.Cette limite se trouve dans le champ 4 (duplomères cephalon,thorax, and pygidium of Cambrian trilobites was un- 21—n) dans le cas des Olenellides et des Redlichiides. L’extrémité dertaken to determine if the Arthropod Pattern Theory (APT) du corps se trouve généralementau niveau du node 3 chez les proposed by Schram & Emerson (1991) applies to Cambrian Corynexochides, et au niveau du champ 4 chez les Olenellides, trilobites. The boundary of the cephalon/thorax occurs within les Redlichiides et les Ptychopariides. D’autre part, les épines 1 4 the predicted duplomerenode (duplomeres or 6). The bound- macropleurales, qui pourraient indiquer l’emplacement des ary between the thorax and pygidium generally occurs within gonopores ou de l’anus, sont généralementsituées au niveau des node 2 (duplomeres 11—13) and node 3 (duplomeres 18—20) for duplomères pronostiqués. La limite prothorax/opisthothorax corynexochids and ptychopariids, respectively. This boundary des Olenellides est située dans le node 3 ou près de celui-ci. Ces occurs within field 4 (duplomeres21—n) for olenellids and red- résultats indiquent que nombre et distribution des duplomères lichiids. -
Tentative Correlation of Cambrian Series 2 Between South China and Other Continents
Tentative correlation of Cambrian Series 2 between South China and other continents JINLIANG YUAN, XUEJIAN ZHU, JIHPAI LIN & MAOYAN ZHU The apparent absence, in Cambrian Series 2, of widespread and rapidly evolving organisms comparable to the later agnostids, graptolites, conodonts, ammonites, or planktonic foraminifers, has prevented a consistent intercontinental biostratigraphy. Occasional genera, and (rarely) species, of trilobites, archaeocyathans and other groups may be found in more than one region. Nevertheless, based on the complete trilobite and archaeocyathan successions in the shallow water Yangtze Platform and in deeper water Jiangnan Slope environment, correlation of Cambrian Series 2 between South China and other continents is discussed in detail. The oldest trilobite Parabadiella and Tsunyidiscus in South China can be correlated with the oldest trilobite Abadiella in Australia, Profallotapis in Siberia and Eofallotaspis in Morocco. • Key words: Cambrian Series 2, correlation, South China, other continents. YUAN, J.L., ZHU, X.J., LIN,J.P.&ZHU, M.Y. 2011. Tentative correlation of Cambrian Series 2 between South China and other continents. Bulletin of Geosciences 86(3), 397–404 (2 tables). Czech Geological Survey. Prague, ISSN 1214-1119. Manuscript received December 17, 2010; accepted in revised form July 1, 2011; published online Septem- ber 22, 2011; issued September 30, 2011. Jinliang Yuan, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China; [email protected] • Xuejian Zhu, Jihpai Lin & Maoyan Zhu, State Key Laboratory of Palaeontology and Stratigra- phy, Nanjing Institute of Geology and Palaeontology,Chinese Academy of Sciences, Nanjing, 210008, China International correlation of Cambrian Series 2 has been a nomenclature lead to paucity of biostratigraphically useful major problem since the concept of four series within the data in key regions (Palmer 1998, Geyer 2001). -
Th TRILO the Back to the Past Museum Guide to TRILO BITES
With regard to human interest in fossils, trilobites may rank second only to dinosaurs. Having studied trilobites most of my life, the English version of The Back to the Past Museum Guide to TRILOBITES by Enrico Bonino and Carlo Kier is a pleasant treat. I am captivated by the abundant color images of more than 600 diverse species of trilobites, mostly from the authors’ own collections. Carlo Kier The Back to the Past Museum Guide to Specimens amply represent famous trilobite localities around the world and typify forms from most of the Enrico Bonino Enrico 250-million-year history of trilobites. Numerous specimens are masterpieces of modern professional preparation. Richard A. Robison Professor Emeritus University of Kansas TRILOBITES Enrico Bonino was born in the Province of Bergamo in 1966 and received his degree in Geology from the Depart- ment of Earth Sciences at the University of Genoa. He currently lives in Belgium where he works as a cartographer specialized in the use of satellite imaging and geographic information systems (GIS). His proficiency in the use of digital-image processing, a healthy dose of artistic talent, and a good knowledge of desktop publishing software have provided him with the skills he needed to create graphics, including dozens of posters and illustrations, for all of the displays at the Back to the Past Museum in Cancún. In addition to his passion for trilobites, Enrico is particularly inter- TRILOBITES ested in the life forms that developed during the Precambrian. Carlo Kier was born in Milan in 1961. He holds a degree in law and is currently the director of the Azul Hotel chain.