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Available Generic Names for Trilobites
AVAILABLE GENERIC NAMES FOR TRILOBITES P.A. JELL AND J.M. ADRAIN Jell, P.A. & Adrain, J.M. 30 8 2002: Available generic names for trilobites. Memoirs of the Queensland Museum 48(2): 331-553. Brisbane. ISSN0079-8835. Aconsolidated list of available generic names introduced since the beginning of the binomial nomenclature system for trilobites is presented for the first time. Each entry is accompanied by the author and date of availability, by the name of the type species, by a lithostratigraphic or biostratigraphic and geographic reference for the type species, by a family assignment and by an age indication of the type species at the Period level (e.g. MCAM, LDEV). A second listing of these names is taxonomically arranged in families with the families listed alphabetically, higher level classification being outside the scope of this work. We also provide a list of names that have apparently been applied to trilobites but which remain nomina nuda within the ICZN definition. Peter A. Jell, Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia; Jonathan M. Adrain, Department of Geoscience, 121 Trowbridge Hall, Univ- ersity of Iowa, Iowa City, Iowa 52242, USA; 1 August 2002. p Trilobites, generic names, checklist. Trilobite fossils attracted the attention of could find. This list was copied on an early spirit humans in different parts of the world from the stencil machine to some 20 or more trilobite very beginning, probably even prehistoric times. workers around the world, principally those who In the 1700s various European natural historians would author the 1959 Treatise edition. Weller began systematic study of living and fossil also drew on this compilation for his Presidential organisms including trilobites. -
001-012 Primeras Páginas
PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. -
A Trilobite with North American Affinities from the Upper Cambrian of Sweden
A Trilobite with North American Affinities from the Upper Cambrian of Sweden By Gunnar Henningsmoen (h lo ABSTRACT.-Taenicephalus? peregrinus n.sp. is described from the Upper Cambrian Olenus beds in Sweden. l t recalls North American genera of the families Pterocephalidae and Paraboli noididae, and is regarded as one of the rare occasional invaders into the Upper Cambrian sea of Scandinavia. Introduction During an excursion to Västergötland in June 1956, Dr. V. JAANUSSON of Uppsala found a cranidium of an undescribed trilobite in the Olenus beds at Trolmen, Kinnekulle. A second, probably conspecific cranidium was collected there by me. This species, described below, is of particular interest because it apparently shows North American affinities and may thus be one of the rare occasional invaders into the Upper Cambrian sea of Scandinavia. The occur rence may be compared with that of Irvingella, and of Drepanura, described by WEsTERGÅRD (1947, 1949), also from the Upper Cambrian of Sweden. The specimens described belong to the Museum of the Palaeontological Institution of the University of Uppsala (U.M.). I wish to thank Miss B. MAURITZ for taking the photographs, Miss I. Lowzow for preparing the draw i ng, and Dr. W. C. SWEET for reading the manuscript. Description Genus Taenicephalus ULRICH & RESSER, 1924 TYPE SPECIEs.-Conocephalites shumardi HALL, 1863, by origi nal designation. Taenicephalus? peregrinus n. s p. Pl. I; Text-lig. r. DERIVATION OF NAME.-From Latin: peregrinus, foreign. MATERIAL.-Two incomplete cranidia, U.M. Nos. V g. 705 (holotype) and V g. 706. 12- .16�272 !lull. n/ (;rnl. Vol. X.\.\ VII x68 GUNNAR HENNINGSMOEN HoRIZON.-Both specimens occur in hituminous limestone (stinkstone), associated with Olenus gibbosus and Agnastus (Homagnostus) obesus, and thus belong to the Upper Cambrian zone of Olenus, subzone of Olenus gibbosus and Olenus transversus (cf. -
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Records of the Western Australian Museum 20: 353-378 (2002). Lower Devonian trilobites from Cobar, New South Wales MaIte Christian Ebach Present address: School of Botany, University of Melbourne, 3010, Australia Department of Earth and Planetary Sciences, Western Australian Museum, Francis Street Perth, Western Australia 6000, Australia e-mail: [email protected] Abstract -A Lower Devonian (Lochkovian-Pragian) trilobite fauna from the Biddabirra Formation near Cobar, New South Wales, Australia includes 11 previously undescribed species (Alberticoryphe sp., Cornuproetus sp., Gerastos sandfordi sp. nov., Cyphaspis mcnamarai sp. nov., Kainops cf. ekphymus, Paciphacops sp., AcantJwpyge (Lobopyge) edgecombei sp. nov., Crotalocephalus sp. and Leonaspis sp., a styginid sp., and a harpetid sp.). Three species belonging to the genera Paciphacops, Kainops, AcantJwpyge (Lobopyge), and Leonaspis are preserved with sufficient detail to provide enough information to be coded and analysed by two cladistic analyses. The resulting cladograms provide justification for the monophyly of Paciphacops and further support for Kainops. Leonaspis sp. is placed as the most plesiomorphic species within the monophyletic Leonaspis. INTRODUCTION P. crawfordae, P. waisfeldae and P. sp.). The Leonaspis The Lochkovian-Pragian Biddabirra Formation analysis, using Ramsk6ld and Chatterton's (1991) (Glen 1987) has yielded a trilobite fauna containing characters, includes Leonaspis sp. from Cobar. The twelve species, of which eleven have previously analysis will attempt to use species with more than been undescribed. The species include the 45% of their characters coded. fragmentary material of a styginid and harpetid, All photographed and type specimens are held in internal and external moulds of Alberticoryphe sp., the Australian Museum (prefix number AMF). -
Biodiversity, Distribution and Patterns of Extinction of the Last Odontopleuroid Trilobites During the Devonian (Givetian, Frasnian)
Geol. Mag. 144 (5), 2007, pp. 777–796. c 2007 Cambridge University Press 777 doi:10.1017/S0016756807003779 First published online 13 August 2007 Printed in the United Kingdom Biodiversity, distribution and patterns of extinction of the last odontopleuroid trilobites during the Devonian (Givetian, Frasnian) ∗ RAIMUND FEIST & KENNETH J. MCNAMARA† ∗Laboratoire de Paleontologie,´ Institut des Sciences de l’Evolution, Universite´ Montpellier II, Cc 062, Place E. Bataillon, 34095 Montpellier, France †Department of Earth Sciences, University of Cambridge, Downing Street, Cambridge CB2 3EQ, UK (Received 18 September 2006; accepted 22 February 2007) Abstract – Biostratigraphical ranges and palaeogeographical distribution of mid-Givetian to end- Frasnian odontopleurids are investigated. The discovery of Leonaspis rhenohercynica sp. nov. in mid-Givetian strata extends this genus unexpectedly up to the late Middle Devonian. New material of Radiaspis radiata (Goldfuss, 1843) and the first koneprusiine in Britain, Koneprusia? sp., are described from the famous Lummaton shell-bed, Torquay, Devon. New taxa of Koneprusia, K. serrensis, K. aboussalamae, K. brevispina,andK. sp. A and K. sp. B are defined. Ceratocephala (Leonaspis) harborti Richter & Richter, 1926, is revised and reassigned to Gondwanaspis Feist, 2002. Two new species of Gondwanaspis, G. dracula and G. spinosa, plus three others left in open nomenclature, are described from the late Frasnian of Western Australia. A further species of Gondwanaspis, G. prisca, is described from the early Frasnian of Montagne Noire. Species of Gondwanaspis are shown to possess a number of paedomorphic features. A functional analysis suggests that, unlike other odontopleurids, Gondwanaspis actively fed and rested with the same cephalic orientation. The sole odontopleurid survivors of the severe terminal mid-Givetian biocrisis (‘Taghanic Event’) belong to the koneprusiine Koneprusia in the late Givetian and Frasnian, and, of cryptogenic origin, the acidaspidine Gondwanaspis in the Frasnian. -
Smithsonian Miscellaneous Collections
SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 53, NUMBER 6 CAMBRIAN GEOLOGY AND PALEONTOLOGY No. 6.-0LENELLUS AND OTHER GENERA OF THE MESONACID/E With Twenty-Two Plates CHARLES D. WALCOTT (Publication 1934) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION AUGUST 12, 1910 Zl^i £orb (gaitimovt (pnee BALTIMORE, MD., U. S. A. CAMBRIAN GEOLOGY AND PALEONTOLOGY No. 6.—OLENELLUS AND OTHER GENERA OF THE MESONACID^ By CHARLES D. WALCOTT (With Twenty-Two Plates) CONTENTS PAGE Introduction 233 Future work 234 Acknowledgments 234 Order Opisthoparia Beecher 235 Family Mesonacidas Walcott 236 Observations—Development 236 Cephalon 236 Eye 239 Facial sutures 242 Anterior glabellar lobe 242 Hypostoma 243 Thorax 244 Nevadia stage 244 Mesonacis stage 244 Elliptocephala stage 244 Holmia stage 244 Piedeumias stage 245 Olenellus stage 245 Peachella 245 Olenelloides ; 245 Pygidium 245 Delimitation of genera 246 Nevadia 246 Mesonacis 246 Elliptocephala 247 Callavia 247 Holmia 247 Wanneria 248 P.'edeumias 248 Olenellus 248 Peachella 248 Olenelloides 248 Development of Mesonacidas 249 Mesonacidas and Paradoxinas 250 Stratigraphic position of the genera and species 250 Abrupt appearance of the Mesonacidse 252 Geographic distribution 252 Transition from the Mesonacidse to the Paradoxinse 253 Smithsonian Miscellaneous Collections, Vol. 53, No. 6 232 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 53 Description of genera and species 256 Nevadia, new genus 256 weeksi, new species 257 Mcsonacis Walcott 261 niickwitzi (Schmidt) 262 torelli (Moberg) 264 vermontana -
Th TRILO the Back to the Past Museum Guide to TRILO BITES
With regard to human interest in fossils, trilobites may rank second only to dinosaurs. Having studied trilobites most of my life, the English version of The Back to the Past Museum Guide to TRILOBITES by Enrico Bonino and Carlo Kier is a pleasant treat. I am captivated by the abundant color images of more than 600 diverse species of trilobites, mostly from the authors’ own collections. Carlo Kier The Back to the Past Museum Guide to Specimens amply represent famous trilobite localities around the world and typify forms from most of the Enrico Bonino Enrico 250-million-year history of trilobites. Numerous specimens are masterpieces of modern professional preparation. Richard A. Robison Professor Emeritus University of Kansas TRILOBITES Enrico Bonino was born in the Province of Bergamo in 1966 and received his degree in Geology from the Depart- ment of Earth Sciences at the University of Genoa. He currently lives in Belgium where he works as a cartographer specialized in the use of satellite imaging and geographic information systems (GIS). His proficiency in the use of digital-image processing, a healthy dose of artistic talent, and a good knowledge of desktop publishing software have provided him with the skills he needed to create graphics, including dozens of posters and illustrations, for all of the displays at the Back to the Past Museum in Cancún. In addition to his passion for trilobites, Enrico is particularly inter- TRILOBITES ested in the life forms that developed during the Precambrian. Carlo Kier was born in Milan in 1961. He holds a degree in law and is currently the director of the Azul Hotel chain. -
Olenid Trilobites: the Oldest Known Chemoautotrophic Symbionts?
Olenid trilobites: The oldest known chemoautotrophic symbionts? Richard Fortey* Department of Paleontology, Natural History Museum, Cromwell Road, London SW7 5BD, United Kingdom Communicated by Lynn Margulis, University of Massachusetts, Amherst, MA, April 3, 2000 (received for review March 1, 2000) Late Cambrian to early Ordovician trilobites, the family Olenidae, Given the widespread occurrence and taxonomic spread of were tolerant of oxygen-poor, sulfur-rich sea floor conditions, and chemoautotrophic symbiosis, it is likely to have been an ancient a case is made that they were chemoautotrophic symbionts. adaptation. Because direct evidence of the fossil bacteria seldom Olenids were uniquely adapted to this habitat in the Lower preserves, evidence of such habits tends to be inferred from Paleozoic, which was widespread in the Late Cambrian over Scan- taxonomic and͞or morphological data, which is not always dinavia. This life habit explains distinctive aspects of olenid mor- reliable (11). The long-ranging solemyid and lucinid bivalves (10) phology: wide thoraces and large numbers of thoracic segments, indicate that by the Silurian (ca. 420 million years), this life mode thin cuticle and, in some species, degenerate hypostome, and the already had been adopted by some groups with living descen- occasional development of brood pouches. Geochemical and field dants. Ancient vent associations have been recognized from at evidence is consistent with this interpretation. Olenids occupied least the Devonian (12). their specialized habitat for 60 million years until their extinction I show in this paper that even by the late Cambrian period (505 at the end of the Ordovician. million years ago) certain extinct arthropods, trilobites belonging to the family Olenidae, evolved features best understood as evidence olorless sulfur bacteria, a heterogeneous category of bacte- of sulfur chemoautotrophic mode of metabolism. -
A Problematical Trilobite from the Lower Cambrian of Freuchen Land, Central North Greenland
A problematical trilobite from the Lower Cambrian of Freuchen Land, central North Greenland Philip D. Lane and Adrian W. A. Rushton A single specimen of a remarkable trilobite is described from the Early Cambrian Buen Formation of north-east Freuchen Land, central North Greenland. It is re ferred with doubt to Alacephalus Repina, 1960, as a new species A? davisi sp. nov. It possibly lacked eyes, which makes it one of, if not the, earliest non-agnostoid trilobite with this adaptation. Its thoracic segments have a unique morphology. In some respects the morphology resembles that of various trilobites adapted to low-energy benthic environments of low oxygenation; such trilobites tend to be widely distrib uted, and in agreement with this Alacephalus appears to be interprovincial. P. D. L., Department ofGeology, Vniversity ofKeele, Keele, Staffordshire ST5 5BG, V.K. A. W. A. R., British Geological Survey, Keyworth, Nottingham NG12 5GG, U.K. The trilobite described below was collected by Neil C. olenellid trilobite Buenellus higginsi Blaker, 1988 (Fam- ~ Davis from laminated, locally bioturbated, mudstones ily Nevadiidae) is associated with an abundant soft- in north-east Freuchen Land (Fig. l; Geological Survey bodied fauna (Conway Morris et al.. 1987; Conway of Greenland collection 319544) which are estimated to Morris & Peel, 1990; Peel, 1990), together with large be not more than 5 m from the top of the Buen Forma articulated sponges such as the demosponge Choia hin tion. This formation has been interpreted as a silic dei Rigby, 1986; the same kinds of sponges are associ ic1astic shelf deposit forming part of the sequence on the ated with the new trilobite in GGU collection 319544. -
Environmental Fluctuations in the Olenus Zone (Upper Cambrian), Southern Scandinavia: a Geochemical Approach
Environmental fluctuations in the Olenus Zone (Upper Cambrian), southern Scandinavia: A geochemical approach NIELS H. SCHOVSBO Schovsbo, N.H. 2000–09–18: Environmental fluctuations in the Olenus Zone (Upper Cambrian), southern Scandinavia: A geochemical approach. Bulletin of the Geological Society of Denmark, Vol. 47, pp. 53–61. Copenhagen. https://doi.org/10.37570/bgsd-2000-47-03 The Scandinavian Alum Shale Formation (Middle Cambrian to Lower Ordovician) was deposited under generally low bottom water oxygen levels. In the formation there is a covariance between faunal composition and the level of trace element enrichment caused by their mutual dependence upon changes in the bottom water oxygen levels. A detailed profile of the V/(V+Ni) ratio through the Upper Cambrian Olenus Zone in the Gislövshammar-2 core, Scania, Sweden, is described. En- vironmental information from this zone is particularly relevant since the Olenus species in the zone apparently undergo iterative changes in morphology that may be linked to environmental changes. Moreover, the occurrences of Olenus species and Homagnostus obesus are almost mutually exclusive. The results indicate that the trilobitic intervals coincide with increases in oxygen levels as monitored by the V/(V+Ni) ratio. Peak oxygen levels do not appear at the base of each interval, suggesting that colonisation of the sea-floor took place when the oxygenation level reached a cer- tain threshold but before maximum oxygen levels were reached. The distribution patterns of Olenus and Homagnostus are interpreted as reflecting differences in substrate preference since samples containing Homagnostus have statistically significantly higher TOC levels. The V/(V+Ni) varia- tions in the Olenus bearing intervals suggest that oxygen levels fluctuated more than would be expected from morphological changes apparent in the Olenus species. -
Late Silurian Trilobite Palaeobiology And
LATE SILURIAN TRILOBITE PALAEOBIOLOGY AND BIODIVERSITY by ANDREW JAMES STOREY A thesis submitted to the University of Birmingham for the degree of DOCTOR OF PHILOSOPHY School of Geography, Earth and Environmental Sciences University of Birmingham February 2012 University of Birmingham Research Archive e-theses repository This unpublished thesis/dissertation is copyright of the author and/or third parties. The intellectual property rights of the author or third parties in respect of this work are as defined by The Copyright Designs and Patents Act 1988 or as modified by any successor legislation. Any use made of information contained in this thesis/dissertation must be in accordance with that legislation and must be properly acknowledged. Further distribution or reproduction in any format is prohibited without the permission of the copyright holder. ABSTRACT Trilobites from the Ludlow and Přídolí of England and Wales are described. A total of 15 families; 36 genera and 53 species are documented herein, including a new genus and seventeen new species; fourteen of which remain under open nomenclature. Most of the trilobites in the British late Silurian are restricted to the shelf, and predominantly occur in the Elton, Bringewood, Leintwardine, and Whitcliffe groups of Wales and the Welsh Borderland. The Elton to Whitcliffe groups represent a shallowing upwards sequence overall; each is characterised by a distinct lithofacies and fauna. The trilobites and brachiopods of the Coldwell Formation of the Lake District Basin are documented, and are comparable with faunas in the Swedish Colonus Shale and the Mottled Mudstones of North Wales. Ludlow trilobite associations, containing commonly co-occurring trilobite taxa, are defined for each palaeoenvironment. -
Ge of Silurian Trilobites
185 MORPHO~tETRIC ASPECTS OF GROWTH 4t\ND V4l\RlATION IN Al'1 ASSEMBL.-\GE OF SILURIAN TRILOBITES . HlJGHES*. Nigel C ... Geier Collections and Research Center, Cincinnati Museum of Natural History~ 1720 Gilbert Avenue, Cincinnati, OH 45202; CHAPMAJ.'I, Ralph. E. Applied Morphometries Laboratory, ADP NHB MRC 136" Smithsonian Institution, Washington. D.C. 20560. The tuffaceous Homerian (Upper Wenlock) "Aulacopleura Shale" from Na Cemidlech Hill" Lod~nice, Czech Republic contains 11 common trilobite species belonging to a wide variety of clades. The majority of post-Cambrian trilobite morphotypes are represented by well-preserved, complete specimens. This situation provides a unique opportunity to compare patterns of growth across a broad range of trilobites .. normalized for preservational style and taphomomic condition. Analyses of morphologic landmarks and multivariate relationships among metric distances of the proetide Aulacopleura konincki show that morphologic variabilty of overall shape is approximately constant from early meraspis onwards .. increasing slightly in the holaspid growth phase. Addition of segments to the thorax progressed steadily throughout meraspis" but the rate slackened at a point in ontogeny marked by a sharp change in the growth trajectory of the pygidium. This transition is analagolls to the meraspis-holaspis transition in other trilobites. A. konincki, unlike any other described trilobite, continued to add segments to the thorax throughout holaspid growth, and holaspid specimens may contain a range of 5 segments" with between 18 - 22 thoracic tergites. Variability of segment number in holaspis is significantly greater than in meraspis. Morphological variabilty in shape and segment numbers of tpe odontopleuride Odontopleura ovata is approximately constant throughout growth, and the meraspid phases of both O.