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' + , 64 .L; 6294 6 ; E @ X: 5- -/: 1 9:- 2+ 6 ; <5 < --2: 6- )+ i$ ; PL5 -8 ; 1D Z $ ; + E5 ^ 1- p U9 6 ; 1 LA2U )* i-/ 65 9C "LA- > < C2$ )C A24 a $2$ - )4 1389 T-5 GN: 6-24 )L? ZU 6 ; 2`? 4 2 W 6 ; )+ 62P 1- p . )@ A 3-4 6 )* )T/ ; ,; C - UD E X 8 4 ; 1 9:- 2+ U4 )* ;#+ 6 ; 5 @- 5- w RL- U9 ,$ G -! 4 6 ; ZL A .(Potts et al., 2006) 5- q2; 4 q )L"; 2; 1 4D JA-- $ a@ W - 62$M 4 L4 a2j )C ; )+ - LA- 62 + 29! $ 5- "LA- 4 ) + ]^ --2: PL" D ,$ 4 )C ; ZL A @ )4 3- - (2L L A 80 )LA ^ 35 ) ; 2` ) ;-? 3- G OL 6 ; )+ J : L; )8- : N$ 6-24 ) 3 *- 6-24 . 6-24 ) 2.# T > -2:- -X$ H =2 – 1 Q? - ; ) . G L %75 q # $- G.@- 6LO 6 ; . pi =3- i ) )+2; A 6-V+ C B q ."$ P9 5D Southwood & ) (4) 1A a2j - \ 8 6 ; <5 5- ; ) A- 4 . $ :(Henderson, 2000 .2D z-/ C P9- X <^ n $ 5 )4 6D SQ = 2J / (a + b) (4) Bennet 2LC Michener 2LC JA$ ; ) ) #C SQ =) )X 8 4 EU4 9 E< )T * L 6 ; ) . 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L5 i$ 6- )+ 6 W ; .(SAS Institute, 2003) ) 5 _`A ) ` 5 0 )* i-/ 1 9:- 2+ 6 ;<5 Q? ) .1389 A 1- p U9 6 ; 1 LA2U )* i-/ 1 9:- 2+ 6 ; <5 < --2: 6- )+ i$ -1 8 (%)< --2: 1- p (%) < --2: U9 - ? )+ 2/97 + 10/41 + Apidae Apis mellifera 1/19 + 0 - Apidae Eucera sp.* 22/61 + 11/97 + Apidae Amegilla sp.* 0 - 0/52 + Apidae Anthophora sp.* 16/66 + 18/75 + Andrenidae Andrena sp.* 16/68 + 14/58 + Halictidae Halictus patellatus* 6/95 + 8/85 + Halictidae Halictus quadricinctus* 28/14 + 29/16 + Megachilidae Anthidium florentinum* 0/59 + 0/52 + Megachilidae Anthidium diadema** 2/97 + 1/04 + Megachilidae Megachile sp.* 0 - 2/60 + Crabronidae Bembix sp.* 0 - 0/52 + Crabronidae Philanthus sp.* 1/19 + 1/04 + Sphecidae Sceliphron sp. . 4 8 E \ 8 2 8 |2 )4 - ? 5 )4 ]#XL )+ 10 5- q2; 5- Z= 6-24 ) E4 , ^ 5 0 .2+ A 6D 6 U-24 ) 1- p U9 aRL )T/ i * 1 LA2U 5 0 i-/ 2 A 5- Z= .1389 A 1- p U9 6 ; 1 LA2U 5 0 i-/ 1 9:- 2+ 6 ; <5 < --2: -X$ -2 8 (%) --2: 1- p (%) < --2: U9 - ? )+ 2/2 + 2/5 + Apidae Xylocopa valga * 0 - 5 + Apidae Amegilla sp.* 42/2 + 30 + Andrenidae Andrena sp.* 35/6 + 25 + Halictidae Halictus patellatus* 2/2 + 5 + Halictidae Halictus quadricinctus* 0 - 2/5 + Crabronidae Cerceris sp. 2/2 + 25 + Crabronidae Philanthus sp.* 13/4 + 2/5 + Crabronidae Sphecius sp. 2/2 + 0 - Sphecidae Sceliphron sp. 0 - 2/5 + Sphecidae Podalonia sp. . 4 8 E :) ` 2o- 6-24 F 133 =185/23 P < 0/001 :NO !"# NO G4 L 2o- 6-24 F 133 =98/45 P < 0/001 )* 6- )+ 6 W 2L9 4 A24 - ) .4 1 LA2U )* i-/ K )4 1 9:- 2+ 6 ; <5 1: M 4 E; < 2P 4 .1389 A 1- p U9 6 ; 1 LA2U 5 0 )* NO f@ + 6- )+ 6 W Q? 1-/ -1 G. . 4 %5 _`A ; $ - X > "$ 2P 4 )4 9 2 W e2! NO 2; .(F 133 =46/3 P < 0/001 :) ` L?-. Q? 2 ):2^ u - ) 1- p 5 0 6 SLA- )4 )X@ ` 1 . 2; LC : 5- q2; 2o- )C - 1 9 f#LR 6 ; $ # 0 6 ; ZL A 8 6 ; )+ 4 E X 8 \5$ X ; )+ \5$ 24 UD G4 L 2o- U$ )4 ; .(2 G. ) 4 E?-. ĤL< 6-24 ) :) ` 2o- 6-24 F 133 =10/423 P < 0/01) 4 - NO G4 L 2o- 6-24 F 133 =25/3 P < 0/001 2 W e2! .1389 A 1- p U9 6 ; 1 LA2U 5 0 )* NO # 0 L?-. Q? 1-/ -2 G. . 4 %5 _`A ; $ - X > "$ 2P 4 )4 9 :NO 2o- 6-24) E - L5 i$ Q? - 24 5 0 2 -1 L5 i$ Q? - P < 0/001 :) ` 2o- 6-24 F 133 =329/4 P < 0/001 1/2 $1/05 4 1- p 5 0 1/60 $1/48 4 U9 24 ) ` NO G4 L 2o- • ; .( F 133 =126/9 $1/61 4 1- p )* 1/9 $1/74 4 U9 )* P < 0/01) 4 - X / ; <5 L5 i$ )C - 1 9 ):2^ u - ) .4 2 [L 1/75 6 ; <5 L5 i$ Q? - .( F 133 =24/5 6- X 2 o $ U$ )4 q2; ) ` NO i 197 ...1 9:- 2+ 6 ;<5 L5 i$ :1- . ; T nA- .(3 G. ) 4 2L C 1- p 5 0 2 A 5- 6- X `4 U9 )* 1 9:- 2+ Q? - - A24 - .4 2L9 4 ; $ U9 6 ; 1 LA2U 5 0 )* i-/ 1 9:- 2+ 6 ; <5 2 -1 L5 i$ Q? 2 -3 G. . 4 %5 _`A ; $ - X > "$ 2P 4 )4 9 2 W e2! .1389 A 1- p - #: > "$ 5- -$ )* 5 0 Anthidium florentinum 6 ; )+ A24 - T-5 ZL A )C EA- U4 . 4 T-5 ZL A i-/ Halictus patellatuds Andrena sp. 6-28- 3T 3- 6 )@ A 7 -6 q 4 )* a $2$ )4 )X@ ` 1 . 2; )* aRL 2 +24 ,C f#T 24 C 8 2 = $ # T ) ` 2; 5 0 aRL i-/ .4 a@ W ,-/:- 4 \<`@ 4 4 ; + i$ 5- 62$ \ A f ^ Philanthus Andrena sp. 6 ; )+ 1- p U9 i$ NO - 2L9 4 3- 4 3-$ ; + i$ < --2: a $2$ )4 Halictus patellatus sp. - 1 9:- 2+ 6 ; <5 UD 1 >-29! .(2 8) L - 62L9 4 a C2$ ; - 24 1/:- . 4 2$M 4 - 0 ĤL< ) A 5 $ 2" 5- 2 -1 Q? >-2 [$ 6 )- ) ` ` O J-2 e-2^- i-/ ; € 4 ; + Southwood & ) EA- 2 [L 3/5 $ 1/5 M X )L<@- )C - L5 i$ 24 6- )=!c G4 2 o $ Q? - - A24 - .(Henderson, 2000 -2 4 5- ; LC : - 2 o $ ] O$ - Up 2; 2P > G -! G -: 4 65 9C # - ) ` 2j ! - .EA- )L: 2 [$ / ZL AC- 62C C L? A 3 *- LX -. ]^ 5- 62L# C 20 $ 15 62C C L5 i$ ,-/:- )C EA- @ ! 6- )+ i$ 24 G T - )C =L- E:2+ *- 6-24 6 #C .;- 65 9C 6 ; ZL A A24 NO 5- q2; 1 9:- 2+ >-29! 6 ; A z A- ; 24 EA- @$ 6- 0 i$ 2V0 .- E ;- )4 2= .EA- )L - 62 o $ t"! 24 L< L: ; 1-T )4 - 0 65 9C 2 ?- 1D5- w#` 6-24 2U4 E‚ "! 6-24 L5 \4 a2R$ 14 65 9C 6 ; ZL AC- 6 2U4 1D -/8- 5 ƒ: +C-20 ; +K E? 4 .(Altieri, 1999) EA- )L:2+ -2 )8$ X <^ .EA- 62j |`A ) ; )+ 4 J4- + • 0 L5 i$ ).- )4 )8$ eV! ).`4 4 )X 8 q 6- 0 )5M ; $ 234 )X 8 q )4 1D ,-/:- ZL A q 5- )+ 2• :20 ) 6 . ; 5- )# A4 q 6- T 4 , 0 G4 2 W >-2o- rT 4 -$ 5- 6 ; ;- 6-24 ( .2D z-/ C P9-) 2 [$ 52- .(Pimentel et al., 1997) ZL AC- 5- L )@ - .2+ 2.9$ ; ) A 5- . LX . )4 65 9C j-- 624 C P9- )C EA- - P ) 1 0 „ L 2+ K )4 >-29! 6 ; )+ i$ ,; C GM .EA- )L:2+ 3 *- U9 A2: ]• O$ )C •*D 5- •@ wO ; 1 9:- REFERENCES 1. Altieri, M. A. (1999). The ecological role of biodiversity in agroecosystems. 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