DOCSLIB.ORG

The Ground Beetle Supertribe Zuphiitae in the Southern Levant

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
The Ground Beetle Supertribe Zuphiitae in the Southern Levant SPIXIANA 38 2 237-262 München, Dezember 2015 ISSN 0341-8391 The ground beetle supertribe Zuphiitae in the southern Levant (Coleoptera, Carabidae) Thorsten Assmann, Kevin Austin, Estève Boutaud, Jörn Buse, Vladimir Chikatunov, Claudia Drees, Ron F. F. L. Felix, Ariel-Leib-Leonid Friedman, Fares Khoury, Tamar Marcus, Ittai Renan, Constantin Schmidt & David W. Wrase Assmann, T., Austin, K., Boutaud, E., Buse, J., Chikatunov, V., Drees, C., Felix, R. F. F. L., Friedman, A.-L.-L., Khoury, F., Marcus, T., Renan, I., Schmidt, C. & Wrase, D. W. 2015. The ground beetle supertribe Zuphiitae in the southern Levant (Coleoptera, Carabidae). Spixiana 38 (2): 237-262. Based on the study of approximately 400 specimens, we give an overview of the systematics and taxonomy, distribution, dispersal power, and habitat preferences of the carabids belonging to the tribes Anthiini, Helluonini, Dryptini, and Zuphiini in the southern Levant (Egypt: Sinai, Israel and Jordan). We provide identification keys for the members of the given taxa in this region. Eleven species of the Zuphi- itae sensu Ober & Maddison (2008) have previously been published from the southern Levant. Our study with rigorous examinations of verifiable records up- dated the known distribution ranges of six species, though the total number of species which occur in the given region remains eleven. – The following two new species are described: Zuphium orbachi spec. nov., which is similar to Zuphium nu- midicum Lucas, 1846, but differs in its elongate body shape, specific form of aedea- gus and further characters. Parazuphium salmoni spec. nov., a microphthalmic spe- cies from the superficial underground compartment or in deep soil horizons in the Upper Galilee, has robust antennae and legs, but has an aedeagus similar to that of P. chevrolatii Castelnau de Laporte, 1833. – We compiled a list of the 24 known subterranean Zuphiini species (including Parazuphium salmoni spec. nov.), all are restricted to the subtropical and tropical zones. – The following taxonomic acts are proposed: Anthia (Thermophilum) sexmaculata marginata Latreille, 1823, stat. rest.; Zuphium axaridis Iablokoff-Khnzorian, 1972, syn. nov. of Zuphium cilicium Peyron, 1858; Zuphium olens kochi Schatzmayr, 1936, syn. nov. of Zuphium olens (P. Rossi, 1790) (colour variation). – We present first records of: Macrocheilus saulcyi Chevro- lat, 1854 for Jordan; Zuphium olens (P. Rossi, 1790) for Jordan; Zuphium cilicium Peyron, 1858 for Iran; Zuphium fuscum Gory, 1931 for Yemen; Parazuphium chevrolatii (Castelnau de Laporte, 1833) for Greece; Parazuphium damascenum (Fairmaire, 1897) for Cyprus, Greece and Syria; Polistichus fasciolatus (P. Rossi, 1790) for Israel. Thorsten Assmann (corresponding author), Estève Boutaud & Tamar Marcus, Institute of Ecology, Leuphana University Lüneburg, Scharnhorststr. 1, 21314 Lüneburg, Germany; e-mail: [email protected] Kevin Austin, Cwmbychan, Llandre SY24 5BS, Ceredigion, Wales, UK Jörn Buse, Ecosystem Analysis, Institute for Environmental Sciences, University of Koblenz-Landau, Fortstr. 7, 76829 Landau, Germany Vladimir Chikatunov, Claudia Drees, Ariel-Leib-Leonid Friedman & Ittai Renan, Tel Aviv University, Department of Zoology, The National Collections of Natural History, Ramat-Aviv, Tel Aviv, 69978, Israel 237 Claudia Drees, Biocentre Grindel and Zoological Museum, University of Ham- burg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany Ron F. F. L. Felix, Hazelaarlaan 51, 5056 XP Berkel Enschot, The Netherlands Fares Khoury, Department of Biology and Biotechnology, American University of Madaba, P.O. Box 2882, Amman, 11821, Jordan Constantin Schmidt, Deichstr. 13, 21354 Bleckede, Germany David W. Wrase, Dunckerstr. 78, 10437 Berlin, Germany Introduction subterranean Zuphiini species from the Levant, led us to compile a brief review and comparison of the The Levant, a biogeographic region in the East Medi- world distribution ranges of subterranean Zuphiini, terranean, is characterized by a remarkable richness and to compare it with that of other subterranean of habitats as well as floral and faunal elements. ground beetles. Euro-Siberian, Central Asian, Oriental, Ethiopian, and Mediterranean species are mixed with Saharan and Arabian desert elements, giving this region a Material and methods unique biotic composition (Furth 1975, Yom-Tov & Tchernov 1988, Waitzbauer & Petutschnig 2004). This Delineation of the study area fascinating biodiversity has long attracted biologists, There are several biogeographical definitions of the including coleopterologists (e. g. Reiche & Saulcy Levant (e. g. Por 1975). As material from Lebanon and 1855, Bodenheimer 1937), however, the undertak- Syria is hardly accessible for carabidological studies, we ing of taxonomic studies in the Levant, especially focus on the southern Levant which we define as the of species-rich groups, is not an easy task due to eastern part of Egypt (Sinai Peninsula), Israel (including the biogeographic diversity. Moreover, yet many areas under Palestinian control), and Jordan. Any infor- species remain unknown to science, as can be seen mation we have about species from Cyprus, Lebanon or by the numerous species descriptions which have Syria is also given. We also add important records from been published in recent years. This is also true for outside the study area if they enlarge the known distri- one of the most species-rich families of insects, the bution range. ground beetles (Schuldt et al. 2009). The ground beetle supertribe Zuphiitae sensu Collections, distribution records Ober & Maddison (2008) is comprised of six tribes, This study is based on the examination of specimens of which four occur in the Levant: Anthiini, Zuphiini, collected during the authors’ field trips to Cyprus, Helluonini, and Dryptini (Löbl & Smetana 2003, own Israel, Jordan and Egypt (Sinai), as well as specimens observations). Although some of these tribes are spe- stored in entomological collections (including material cies rich, and species belonging to these tribes have from Europe, Africa, and other parts of Asia for com- been described as early as the 18th century, they have parisons). We studied approximately 400 specimens not undergone taxonomic revision for the Palaearctic from this group but material from Lebanon and Syria is very limited. region. Several Levantine species are known from these tribes (Bodenheimer 1932 and 1937, Baehr 2003a The material is stored in the following collections: and 2003b, Bousquet 2003, Hurka 2003, Chikatunov CAB Working collection Assmann, Bleckede (part et al. 2006, Timm et al. 2008). Over the last decade of ZSM, Germany) we analysed several hundred individuals from the CFB Working collection Felix, Berkel Enschot, The Netherlands given tribes. This material provides the basis for our ’ systematic-taxonomic analysis. Field work and a COQ Working collection Orbach, Qiryat Tiv on, (will be transferred to TAU, Israel) literature survey provide the basis for the ecological CSS Working collection Schüle, Stuttgart, Germany characterization, as already given for other ground CSW Working collection Starke, Warendorf (will be beetle groups (Assmann et al. 2008a and 2012). transferred to Westfälisches Landesmuseum The analyses of the material revealed two new Münster, Germany) taxa, one of them with strongly reduced eyes and CWB Working collection Wrase, Berlin (part of ZSM, a subterranean habitat. While, endogeic and cave- Germany) dwelling ground beetles are known from some TAU National Collections of Natural History, Tel regions of the western Palaearctic realm, there are Aviv University, Tel Aviv, Israel apparently few such species in the Middle East. AUB Natural History Museum, American Univer- Our surprising record of a previously undescribed sity of Beirut, Beirut, Lebanon 238 NHMB Natural History Museum, Entomological Col- (Lompe 1989) or in Euparal on acetate labels, and pinned lection, Budapest, Hungary beneath the specimens from which they were removed. NHMPr Natural History Museum, Entomological Col- The photographs were taken with an Olympus E-330 lection (Kunratic), Prague, Czech Republic digital camera in combination with a Leica MZ 95 ste- NHMP Muséum National d’Histoire Naturelle, Ento- reobinocular microscope. Postprocessing was done in mology Department, Paris, France Adobe Acrobat Professional 7.0. To achieve sufficient ZSM Zoological State Collection Munich (Zoologi- depth of focus, up to 40 planes were captured. These sche Staatssammlung München), München, were then copied to separate layers, and the out-of-focus Germany planes were masked using a stacking program (Com- bine ZP). We reviewed the available literature to collect data on Data on power of dispersal, phenology, habitat se- the distribution range, ecology, and biology of the given lection, and distribution ranges were estimated as de- species (including an unpublished manuscript on the scribed in Assmann et al. (2012). Anthiini: Basilewsky unpublished). Where possible, the nomenclature follows the last Palaearctic Catalogue Compilation of subterranean (Löbl & Smetana 2003) or the world list of ground bee- Zuphiini species worldwide tles (Lorenz 2005). All changes of rank or nomenclature published after the Palaearctic Catalogue are ignored if We described the currently known distribution ranges they were published without appropriate scientific ar- of subterranean Zuphiini taxa based on standard data- gumentation. An example is the rank of the taxon bases (Web of Science and Zoological Record), internet Thermophilum
Load more

Recommended publications
  • New Record of Tiger Beetle Anthia Sexguttata Sexguttata (Coleoptera: Carabidea) from District Mirpurkhas Sindh
    Vol. 3, Issue 1, Pp: (14-17), March, 2019 NEW RECORD OF TIGER BEETLE ANTHIA SEXGUTTATA SEXGUTTATA (COLEOPTERA: CARABIDEA) FROM DISTRICT MIRPURKHAS SINDH Sidra-Tul Muntha, Naheed Baloch, Riffat Sultana, Javeria Shaikh Department of Zoology, University of Sindh, Jamshoro ARTICLE INFORMATION ABSTRACT Article History: Anthia sexguttata sexguttata (Febricius 1775) commonly known as tiger beetle Received: 13th Jan 2019 Accepted: : 30th Sept. 2019 or ground beetle, belong to family Carabidea. During the present study. Single Published online: 18th Oct 2019 pair was reported from Mirpurkhas. It morphometric observation is under: Author’s contribution Length of head 18mm, length of pronotum 8mm, length of antenna STM performed experiments , NB & RS organized the plan & JS complied data. 15mm,length of abdomen 2.1mm and Total body length 40mm for male, while Key words: in female it was length of head 18mm,length of pronotum 9mm,length of Anthia sexguttata sexguttata, beetle antenna 16mm,length of abdomen 23mm and total body length 41mm. Presence Carabidea, Mirpurkhas, carnivorous of six spots on 02 on thoraxic region and 04 on the elytra is key character of tiger beetle due to this character it is commonly known as six spot ground beetle. Female is larger in size then male. They are carnivorous in nature. From Sindh no work has been done on the morphological characters of tiger beetle, present study is the starting point to introduction of this beetle from Mirpurkhas. 1. INTRODUCTION Ground beetle are the member of family Carabidea. Carabids are partly diurnal, partly nocturnal (Eisner, They are usually recognized with thread-like et al., 1977; Dazzini, 1980; Detttner, 1985; Will, et antennae and have a strong mandibles and large size al., 2000, Brandmayr, et al., 2009).
    [Show full text]
  • Vol. 33, Number 3
    NO.3 May/June 1991 eDITOR o( the LEPIDOPTERISTS' SOCIETY June Preston 832 Sunset Dr Lawrence. KS 66044 U.S.A. • •• • •• • • . -. •• .-.-. ...- . • • •• ASSOCIATE eDITOR ZONE COORDINATORS 1. Ken Philip 6. Ed Knudson 10. Dave Winter 2. Jon Shepard 7. Ross Layberry 11. J.e.E. Riotte Ripples 3. Bob Langston 8. Les Ferge 12. Eduardo Welling M. Jo Brewer 4. Ray Stanford 9. Andy Beck 13. Boyce Drummond 5. Ron A. Royer • •• I • ....__... ••_ ...._._....__~._••_ ... ••_._..._._....__....__~.__•••__...__...__• BUTIERFLY DIVERSITY AND CONSERVATION IN THE RONDONIAN RAIN FOREST OF BRAZIL: A PROGRESS REPORT ON THE NEW RESERVE The state of Rondonia in west central Brazil apparently 27 lepidopterists in November 1990 and another 26 has the highest butterfly diversity per square kilometer in the lepidopterists in December 1990 has added more than 200 world, with an estimated 1,500 to 1,600 species residing additional species in these two months to the identified list, within several square kilometers in the central part of that raising the total to over 1,060 identified species inhabiting state near Ariquemes. After a major March 1989 expedition the area around the Fazenda Rancho Grande field station. We to the Rondonian rain forest around the Fazenda Rancho Grande still project that the "final" list will amount to around 1,600 involving more than two dozen members of the Lepidopterists' butterfly species! Society, it was clear that this incredible diversity - first As mentioned in Emmel (1989) and Emmel and Austin noted two years earlier in March 1987 - was about to (1990), the sampling efforts have centered on the Fazenda disappear under the hand of man, through cutting and burning Rancho Grande, a 750-hectare tract (1,875 acres) owned by of the tropical rain forest to create temporary crop land and the Harald Schmitz family.
    [Show full text]
  • Acari: Heterostigmata: Podapolipidae), the Ectoparasite of Carabid Beetles from Iran
    J o u r n a l o f E n t o m o l o g i c a l S o c i e t y o f I r a n 91 2011, 31(1), 91-93 Short communication New records of the genus and species Regenpolipus madrasensis (Acari: Heterostigmata: Podapolipidae), the ectoparasite of carabid beetles from Iran H. Hajiqanbar1&* and M. Khoshnevis2 1. Department of Entomology, Faculty of Agriculture, Tarbiat Modares University, 14115-336, Tehran, Iran, 2. Plant Protection Division, Institute of Jahad-e Agriculture, Chenaran, Khorasan Razawi, Iran. *Corresponding author, E-mail: [email protected] Anthia sexguttata (Fab.) (Col.; Carabidae) ƵŶǀĪģ ƱŚŤſźƸƃƝřźƏřŹŵ ƾưƬƗƭŚƳŚŝĨſƺſƽŵřŶƘţƽŹƹōƖ ưūŻřžě Podapolipidae (Acari; Heterostigmata) źºƿŻƶƧ ƽƵŵřƺƳŚųŢƿŻřŹŚěƽŚƷƶ ƴĩƾƴƬĩƽƺƋŹƱŚſřźųƱŚŤſřŹŵƖƣřƹŵŚŞƿŚţ Regenpolipus madrasensis Husband & Ramaraju, 2006 ƾƿŚſŚƴƃ ƭŚƳƶŝŵƺŝƱŚŝżǀƯƽŚƷĨ ſƺſŻřƾĪƿƽŚƷƁ ƺĜƫŚŝ ŵƺƃƾ ƯƁŹřżĭƱřźƿřŻřŹŚŝƲ ǀƫƹřƽřźŝƶƳƺĭƹžƴūƲƿřŶƃ Mites of the family Podapolipidae are highly specialized ecto- and endoparasites of different insect orders including Blattaria, Hymenoptera, Orthoptera, Hemiptera and particularly Coleoptera (Husband, 2000). Among 30 genera of the family, the following four genera exclusively parasitize ground beetles of the family Carabidae: Eutarsopolipus Berlese, 1913 (55 species); Dorsipes Regenfuss, 1968 (18 species); Regenpolipus Husband, 1986 (5 species) and Ovacarus Stannard & Vaishampayan, 1971 (3 species). Only the genus Ovacarus is the endoparasite of the ground beetles and settles in their reproductive tracts. The other three genera are ectoparasites under the elytra of carabid beetles. The mites of the genera Eutarsopolipus, Dorsipes and Ovacarus have been previously recorded from Iran (Hajiqanbar et al., 2007). During a collecting trip, the second author, discovered a large colony of podapolipid mites under the elytra of a giant carabid beetle (ca.
    [Show full text]
  • A Review of Myrmecophily in Ant Nest Beetles (Coleoptera: Carabidae: Paussinae): Linking Early Observations with Recent Findings
    Naturwissenschaften (2007) 94:871–894 DOI 10.1007/s00114-007-0271-x REVIEW A review of myrmecophily in ant nest beetles (Coleoptera: Carabidae: Paussinae): linking early observations with recent findings Stefanie F. Geiselhardt & Klaus Peschke & Peter Nagel Received: 15 November 2005 /Revised: 28 February 2007 /Accepted: 9 May 2007 / Published online: 12 June 2007 # Springer-Verlag 2007 Abstract Myrmecophily provides various examples of as “bombardier beetles” is not used in contact with host how social structures can be overcome to exploit vast and ants. We attempt to trace the evolution of myrmecophily in well-protected resources. Ant nest beetles (Paussinae) are paussines by reviewing important aspects of the association particularly well suited for ecological and evolutionary between paussine beetles and ants, i.e. morphological and considerations in the context of association with ants potential chemical adaptations, life cycle, host specificity, because life habits within the subfamily range from free- alimentation, parasitism and sound production. living and predatory in basal taxa to obligatory myrme- cophily in derived Paussini. Adult Paussini are accepted in Keywords Evolution of myrmecophily. Paussinae . the ant society, although parasitising the colony by preying Mimicry. Ant parasites . Defensive secretion . on ant brood. Host species mainly belong to the ant families Host specificity Myrmicinae and Formicinae, but at least several paussine genera are not host-specific. Morphological adaptations, such as special glands and associated tufts of hair Introduction (trichomes), characterise Paussini as typical myrmecophiles and lead to two different strategical types of body shape: A great diversity of arthropods live together with ants and while certain Paussini rely on the protective type with less profit from ant societies being well-protected habitats with exposed extremities, other genera access ant colonies using stable microclimate (Hölldobler and Wilson 1990; Wasmann glandular secretions and trichomes (symphile type).
    [Show full text]
  • The Ground Beetle Supertribe Zuphiitae in the Southern Levant 237-262 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; Download
    ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Spixiana, Zeitschrift für Zoologie Jahr/Year: 2015 Band/Volume: 038 Autor(en)/Author(s): Aßmann [Assmann] Thorsten, Austin Kevin, Boutaud Esteve, Buse Jörn, Chikatunov Vladimir, Drees Claudia, Felix Ron F. F. L., Friedman Ariel- Leib-Leonid, Khoury Fares, Marcus Tamar, Renan Ittai, Schmidt Constantin, Wrase David W. Artikel/Article: The ground beetle supertribe Zuphiitae in the southern Levant 237-262 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de SPIXIANA 38 2 237-262 München, Dezember 2015 ISSN 0341-8391 The ground beetle supertribe Zuphiitae in the southern Levant (Coleoptera, Carabidae) Thorsten Assmann, Kevin Austin, Estève Boutaud, Jörn Buse, Vladimir Chikatunov, Claudia Drees, Ron F. F. L. Felix, Ariel-Leib-Leonid Friedman, Fares Khoury, Tamar Marcus, Ittai Renan, Constantin Schmidt & David W. Wrase Assmann, T., Austin, K., Boutaud, E., Buse, J., Chikatunov, V., Drees, C., Felix, R. F. F. L., Friedman, A.-L.-L., Khoury, F., Marcus, T., Renan, I., Schmidt, C. & Wrase, D. W. 2015. The ground beetle supertribe Zuphiitae in the southern Levant (Coleoptera, Carabidae). Spixiana 38 (2): 237-262. Based on the study of approximately 400 specimens, we give an overview of the systematics and taxonomy, distribution, dispersal power, and habitat preferences of the carabids belonging to the tribes Anthiini, Helluonini, Dryptini, and Zuphiini in the southern Levant (Egypt: Sinai, Israel and Jordan). We provide identification keys for the members of the given taxa in this region. Eleven species of the Zuphi- itae sensu Ober & Maddison (2008) have previously been published from the southern Levant.
    [Show full text]
  • A Study of the Coleopteran Fauna of Two Agricultural
    Animal Research International (2008) 5(3): 913 – 916 913 THE EFFECT OF LAND USE ON THE COMMUNITY STRUCTURE DISTRIBUTION AND ABUNDANCE OF GROUND BEETLES (INSECTA: COLEOPTERA) IN A GUINEA SAVANNA IN NIGERIA EWUIM, Sylvanus Chima Department of Zoology, Nnamdi Azikiwe University, Awka, Nigeria. Email: [email protected] ABSTRACT A study was carried out to investigate the ground beetles of four habitats - cultivated farmland, Ifallow plot, a marshy plot and a secondary regrowth forest at the permanent site of Nnamdi Azikiwe University, Awka, Nigeria for a twelve-month period using the pitfall technique. Species of Coleoptera obtained from the four plots include Macrocheilus labrosus, Hyparpalus sp., Carpophilus fumatus (Boh.), Podagrica uniforma (Jac.), Tetragonothorax sp., Chlaenius sp., Pheropsophus parallus (Dej.), Silidus apicalis (Waterh), Tenebroides mauritanicus (L), Heteroderes sp., Heterorynchus licas (Klug.) and Mylabris sp., Analysis of Variance (ANOVA) indicate that the abundance of beetles wee significantly different in the cultivated farmland (F = 6.062). The result of Fisher’s Least Significance Difference (F-LSD) test shows that the pitfall catches of beetles from the four sampling sites were also significantly different. The heterogeneity of the coleopteran species at the cultivated plot was traced to cultivation. The role of certain coleopteran families as faunal indicators was highlighted. Other factors, which influenced the Coleopteran species at the four plots, were also discussed. Keywords: Insecta, Coleoptera, Agroecosystems,
    [Show full text]
  • Ecology and Behavior of Ground Beetles (Coleoptera: Carabidae)
    Annual Reviews www.annualreviews.org/aronline Annu. Rev. Entomol. 1996. 41:231-56 Copyright © 1996 by Annual Reviews Inc. All rights reserved ECOLOGY AND BEHAVIOR OF GROUND BEETLES (COLEOPTERA: CARABIDAE) Gabor L. Lovei Horticulture and Food Research Institute of New Zealand, Batchelar Science Centre, Private Bag 11030, Palmerston North 5301, New Zealand Keith D. Sunderland Horticulture Research International, Wellesboume, Warwick CV35 9EF, United Kingdom KEY WORDS: Coleoptera, Carabidae, bionomics, populations, assemblages ABSTRACT The ground beetles form the speciose beetle family Carabidae and, since their emergence in the Tertiary, have populated all habitats except deserts. Our knowl­ edge about carabids is biased toward species living in north-temperate regions. Most carabids are predatory, consume a wide range of food types, and experience food shortages in the field. Feeding on both plant and animal material and scavenging are probably more significant than currently acknowledged. The most important mortality sources are abiotic factors and predators; pathogens and parasites can be important for some developmental stages. Although competition among larvae and adults does occur, the importance of competition as a com­ munity organization is not proven. Carabids are abundant in agricultural fields all over the world and may be important natural enemies of agricultural pests. ----·---- Annu. Rev. Entomol. 1996.41:231-256. Downloaded from www.annualreviews.org Access provided by University of Connecticut on 03/18/19. For personal use only. INTRODUCTION The family Carabidae, the ground beetles, contains more than 40,000 described species classified into some 86 tribes (66). It is the largest adephagan family and one of the most speciose of beetle families.
    [Show full text]
  • Diversity of Ground Beetles and Saproxylic Beetles (Coleoptera: Carabidae + Div
    Diversity of Ground Beetles and Saproxylic Beetles (Coleoptera: Carabidae + div. Saproxylic) in East Mediterranean Ecosystems (Israel) - Dispersal, Habitat, Activity and Reproduction – Academic Dissertation Doktor der Naturwissenschaften (Dr. rer.nat.) Presented by Anika Timm with the permission of the Faculty of Environmental Science of the Leuphana University of Lüneburg Diversity of Ground Beetles and Saproxylic Beetles (Coleoptera: Carabidae + div. Saproxylic) in East Mediterranean Ecosystems (Israel) - Dispersal, Habitat, Activity and Reproduction – Kumulative Dissertationsschrift zur Erlangung des akademischen Grades Doktor der Naturwissenschaften (Dr. rer. nat.) Vorgelegt von Anika Timm Institut für Ökologie und Umweltchemie an derFakultät Umwelt und Technik der Leuphana Universität Lüneburg Gutachter: Prof. Dr. Thorsten Aßmann, Leuphana Universität Lüneburg Prof. Dr. Tamar Dayan, Tel Aviv Universität Eingereicht am: 28.04.2010 Artikel Die folgende Tabelle stellt die Beiträge der jeweiligen Autoren für die dieser Arbeit zugrunde liegenden wissenschaftlichen Zeitschriftenartikel dar. Die verwendeten Kürzel folgen den Anfangsbuchstaben der Autoren; die römischen Zahlen geben den jeweiligen Artikel an. Die Autoren wurden gemäß ihres Arbeitsanteils an den Entstehungsschritten der Artikel eingeordnet. I II III IV V VI Initiative AT, TA, TD AT, TA, TD AT, TA, TD JB, TA, TD JB, TA, TD TA, AF, TD Untersuchungsdesign AT, TA, TD AT, TA, TD, AT, TA, TD JB, TA, TD JB, TA, TD TA, AF WH Datenerhebung AT, TA, AT, TL, JB AT, TL, JB JB, TL JB, TL TA, AT, DW JB, AF, TL, DW Analyse AT AT AT, WP JB, BF, TD, JB, BF, TD, TA, CD, TL, AT, TA TL, AT, TA AM Verfassen des AT AT AT JB JB TA Manuskriptes Autoren: Thorsten Assmann (TA)1, Jörn Buse (JB)1,2, Tamar Dayan (TD)3, Claudia Drees (CD)1, Benedikt Feldmann (BF)6, Ariel-Leib-Leonid Friedmann (AF)3 , Werner Härdtle (WH)1, Tal Levanony (TL)3, Andrea Matern (AM)1, Wilfried Paarmann (WP)4, Anika Timm (AT)1, David W.
    [Show full text]
  • Species of Carabidae (Insecta: Coleoptera) in the University of Arkansas Insed Collection
    Species of Carabidae (Insecta: Coleoptera) in the University of Arkansas Insed Collection By Robert T. Allen Calo8oma externum AGRICULTURAL EXPERIMENT STATION Division of Agriculture University of Arkansas, Fayetteville John W. White, vice president for agriculture and acting director June, 1973 Special Report 19 Species of Carabidae (Insecta: Coleoptera) in University of Arlta.nsas Insect Collection By Robert T. Allen Entomology Department Since the early 1960's the University of Arkansas Insect Collection (UAIC) has had a formal curator and has enjoyed support from the Entomology Department of the Arkansas Agricultural Experiment Station. In recent years, 1967 to date, in addition to E. p. Rouse, the curator, two pro· fessional taxonomists, Rodney Kirkton (deceased) and Robert T. Allen, have served on the staff. The collection has grmm rapidly and is now an important part of the research and teaching program in the Depart­ ment of Entomology. In 1969 a very valuable addition was made to the collection by the purchase of part of the private collection of Carabidae belonging to Dr. S. L. Straneo of Milano, Italy. This added to the UAIC approximately 4,000 species representing most of the major tribes of Carabidae from throughout the world. The purchase was made possible by a personal gift from Dr. David Rockefeller and matching funcls from the Arlta.nsas Agricultural Experiment Station. In addition to the Straneo collection the author has donated his private collection. During the past five years the author has also been able to collect extensively in the Arkansas, Louisiana, and Mississippi area as well as in the Latin American countries of Bolivia and Panama.
    [Show full text]
  • SYNTHESIS and PHYLOGENETIC COMPARATIVE ANALYSES of the CAUSES and CONSEQUENCES of KARYOTYPE EVOLUTION in ARTHROPODS by HEATH B
    SYNTHESIS AND PHYLOGENETIC COMPARATIVE ANALYSES OF THE CAUSES AND CONSEQUENCES OF KARYOTYPE EVOLUTION IN ARTHROPODS by HEATH BLACKMON Presented to the Faculty of the Graduate School of The University of Texas at Arlington in Partial Fulfillment of the Requirements for the Degree of DOCTOR OF PHILOSOPHY THE UNIVERSITY OF TEXAS AT ARLINGTON May 2015 Copyright © by Heath Blackmon 2015 All Rights Reserved ii Acknowledgements I owe a great debt of gratitude to my advisor professor Jeffery Demuth. The example that he has set has shaped the type of scientist that I strive to be. Jeff has given me tremendous intelectual freedom to develop my own research interests and has been a source of sage advice both scientific and personal. I also appreciate the guidance, insight, and encouragement of professors Esther Betrán, Paul Chippindale, John Fondon, and Matthew Fujita. I have been fortunate to have an extended group of collaborators including professors Doris Bachtrog, Nate Hardy, Mark Kirkpatrick, Laura Ross, and members of the Tree of Sex Consortium who have provided opportunities and encouragement over the last five years. Three chapters of this dissertation were the result of collaborative work. My collaborators on Chapter 1 were Laura Ross and Doris Bachtrog; both were involved in data collection and writing. My collaborators for Chapters 4 and 5 were Laura Ross (data collection, analysis, and writing) and Nate Hardy (tree inference and writing). I am also grateful for the group of graduate students that have helped me in this phase of my education. I was fortunate to share an office for four years with Eric Watson.
    [Show full text]
  • Coleoptera: Carabidae: Helluonini) of the Indian Subcontinent: Taxonomy with Notes on Habits and Distributional Patterns
    University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Center for Systematic Entomology, Gainesville, Insecta Mundi Florida 2-29-2012 The apterous endemic genus Omphra Dejean (Coleoptera: Carabidae: Helluonini) of the Indian subcontinent: taxonomy with notes on habits and distributional patterns Shiju T. Raj The New College, Royapettah, India, [email protected] Thomas K. Sabu St. Joseph’s College, Devagiri, India, [email protected] Zhao Danyang South China Agricultural University, Wushan, China Follow this and additional works at: https://digitalcommons.unl.edu/insectamundi Part of the Entomology Commons Raj, Shiju T.; Sabu, Thomas K.; and Danyang, Zhao, "The apterous endemic genus Omphra Dejean (Coleoptera: Carabidae: Helluonini) of the Indian subcontinent: taxonomy with notes on habits and distributional patterns" (2012). Insecta Mundi. 724. https://digitalcommons.unl.edu/insectamundi/724 This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA A Journal of World Insect Systematics MUNDI 0206 The apterous endemic genus Omphra Dejean (Coleoptera: Carabidae: Helluonini) of the Indian subcontinent: taxonomy with notes on habits and distributional patterns Shiju T. Raj Post Graduate and Research Department of Zoology The New College, Royapettah Chennai-670014, Tamil Nadu, INDIA [email protected] Thomas K. Sabu Litter Entomology Research Unit Post Graduate and Research Department of Zoology St. Joseph’s College, Devagiri Calicut-673008, Kerala, INDIA [email protected] Zhao Danyang Department of Entomology College of Natural Resources and Environment South China Agricultural University, Wushan Guangzhou-510640, CHINA Date of Issue: February 29, 2012 CENTER FOR SYSTEMATIC ENTOMOLOGY, INC., Gainesville, FL Shiju T.
    [Show full text]
  • Coleoptera: Carabidae: Helluonini) of China, with Descriptions of Seven New Species
    POLISH JOURNAL OF ENTOMOLOGY POLSKIE P I S M O ENTOMOLOGICZNE VOL. 79 : 3-23 Bydgoszcz 30 March 2010 A review of the genus Macrocheilus HOPE (Coleoptera: Carabidae: Helluonini) of China, with descriptions of seven new species DANYANG ZHAO 1, 2 , MINGYI TIAN 1, * 1) Department of Entomology, College of Natural Resources & Environment, South China 2) Agricultural University, Wushan, Guangzhou, 510640, China, * e-mail: [email protected] ABSTRACT. In the present paper a review of the genus Macrocheilus HOPE , 1838 (Coleoptera: Carabidae: Helluonini) of China is provided, with descriptions of seven new species, viz. M. parvimacu- latus sp. nov. , M. gigas sp. nov. , M. fuscipennis sp. nov. , M. solidipalpis sp. nov. , M. cheni sp. nov., M. quadratus sp. nov. , and M. sinuatilabris sp. nov. A key to all species found in China is also given. In addition, M. asteriscus White is reported from Laos, Indonesia and India for the first time. KEY WORDS: Coleoptera, Carabidae, Helluonini, Macrocheilus , review, new species, China. INTRODUCTION Macrocheilus HOPE , 1838 is the largest genus within the tribe Helluonini. Forty-eight spe- cies of the genus have been reported so far (LORENZ 2005), with thirty-four from the Afro- tropical Region, twelve from the Oriental Region and two from the Palaearctic Region. The Oriental fauna of Macrocheilus have been treated by ANDREWES (1920) and JEDLI ČKA (1963). Up to now only four species of Macrocheilus are known from China (ANDREWES , 1930, CSIKI 1933, JEDLI ČKA 1963, HŮRKA 2003). The first record of Macrocheilus from China was M . asteriscus from Hong Kong (WHITE 1844). BATES (1892) reported the occu- rrence of M.
    [Show full text]