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European Anchovy Engraulis Encrasicolus (Linnaeus, 1758) From
European anchovy Engraulis encrasicolus (Linnaeus, 1758) from the Gulf of Annaba, east Algeria: age, growth, spawning period, condition factor and mortality Nadira Benchikh, Assia Diaf, Souad Ladaimia, Fatma Z. Bouhali, Amina Dahel, Abdallah B. Djebar Laboratory of Ecobiology of Marine and Littoral Environments, Department of Marine Science, Faculty of Science, University of Badji Mokhtar, Annaba, Algeria. Corresponding author: N. Benchikh, [email protected] Abstract. Age, growth, spawning period, condition factor and mortality were determined in the European anchovy Engraulis encrasicolus populated the Gulf of Annaba, east Algeria. The age structure of the total population is composed of 59.1% females, 33.5% males and 7.4% undetermined. The size frequency distribution method shows the existence of 4 cohorts with lengths ranging from 8.87 to 16.56 cm with a predominance of age group 3 which represents 69.73% followed by groups 4, 2 and 1 with respectively 19.73, 9.66 and 0.88%. The VONBIT software package allowed us to estimate the growth parameters: asymptotic length L∞ = 17.89 cm, growth rate K = 0.6 year-1 and t0 = -0.008. The theoretical maximum age or tmax is 4.92 years. The height-weight relationship shows that growth for the total population is a major allometry. Spawning takes place in May, with a gonado-somatic index (GSI) of 4.28% and an annual mean condition factor (K) of 0.72. The total mortality (Z), natural mortality (M) and fishing mortality (F) are 2.31, 0.56 and 1.75 year-1 respectively, with exploitation rate E = F/Z is 0.76 is higher than the optimal exploitation level of 0.5. -
New and Poorly Known Perisphinctoidea (Ammonitina) from the Upper Tithonian of Le Chouet (Drôme, SE France)
Volumina Jurassica, 2014, Xii (1): 113–128 New and poorly known Perisphinctoidea (Ammonitina) from the Upper Tithonian of Le Chouet (Drôme, SE France) Luc G. BULOT1, Camille FRAU2, William A.P. WIMBLEDON3 Key words: Ammonoidea, Ataxioceratidae, Himalayitidae, Neocomitidae, Upper Tithonian, Le Chouet, South-East France. Abstract. The aim of this paper is to document the ammonite fauna of the upper part of the Late Tithonian collected at the key section of Le Chouet (Drôme, SE France). Emphasis is laid on new and poorly known Ataxioceratidae, Himalayitidae and Neocomitidae from the upper part of the Tithonian. Among the Ataxioceratidae, a new account on the taxonomy and relationship between Paraulacosphinctes Schindewolf and Moravisphinctes Tavera is presented. Regarding the Himalayitidae, the range and content of Micracanthoceras Spath is discussed and two new genera are introduced: Ardesciella gen. nov., for a group of Mediterranean ammonites that is homoeomorphic with the Andean genus Corongoceras Spath, and Pratumidiscus gen. nov. for a specimen that shows morphological similarities with the Boreal genera Riasanites Spath and Riasanella Mitta. Finally, the occurrence of Neocomitidae in the uppermost Tithonian is documented by the presence of the reputedly Berriasian genera Busnardoiceras Tavera and Pseudargentiniceras Spath. INTRODUCTION known Perisphinctoidea from the Upper Tithonian of this reference section. Additional data on the Himalayitidae in- The unique character of the ammonite fauna of Le Chouet cluding the description and discussion of Boughdiriella (near Les Près, Drôme, France) (Fig. 1) has already been chouetensis gen. nov. sp. nov. are to be published elsewhere outlined by Le Hégarat (1973), but, so far, only a handful of (Frau et al., 2014). -
Table S1.Xlsx
Bone type Bone type Taxonomy Order/series Family Valid binomial Outdated binomial Notes Reference(s) (skeletal bone) (scales) Actinopterygii Incertae sedis Incertae sedis Incertae sedis †Birgeria stensioei cellular this study †Birgeria groenlandica cellular Ørvig, 1978 †Eurynotus crenatus cellular Goodrich, 1907; Schultze, 2016 †Mimipiscis toombsi †Mimia toombsi cellular Richter & Smith, 1995 †Moythomasia sp. cellular cellular Sire et al., 2009; Schultze, 2016 †Cheirolepidiformes †Cheirolepididae †Cheirolepis canadensis cellular cellular Goodrich, 1907; Sire et al., 2009; Zylberberg et al., 2016; Meunier et al. 2018a; this study Cladistia Polypteriformes Polypteridae †Bawitius sp. cellular Meunier et al., 2016 †Dajetella sudamericana cellular cellular Gayet & Meunier, 1992 Erpetoichthys calabaricus Calamoichthys sp. cellular Moss, 1961a; this study †Pollia suarezi cellular cellular Meunier & Gayet, 1996 Polypterus bichir cellular cellular Kölliker, 1859; Stéphan, 1900; Goodrich, 1907; Ørvig, 1978 Polypterus delhezi cellular this study Polypterus ornatipinnis cellular Totland et al., 2011 Polypterus senegalus cellular Sire et al., 2009 Polypterus sp. cellular Moss, 1961a †Scanilepis sp. cellular Sire et al., 2009 †Scanilepis dubia cellular cellular Ørvig, 1978 †Saurichthyiformes †Saurichthyidae †Saurichthys sp. cellular Scheyer et al., 2014 Chondrostei †Chondrosteiformes †Chondrosteidae †Chondrosteus acipenseroides cellular this study Acipenseriformes Acipenseridae Acipenser baerii cellular Leprévost et al., 2017 Acipenser gueldenstaedtii -
Upper Jurassicelower Cretaceous Stratigraphy in South-Eastern Tibet: a Comparison with the Western Himalayas
Author's personal copy Cretaceous Research 29 (2008) 301e315 www.elsevier.com/locate/CretRes Upper JurassiceLower Cretaceous stratigraphy in south-eastern Tibet: a comparison with the western Himalayas Xiumian Hu a,*, Luba Jansa b, Chengshan Wang c a State Key Laboratory of Mineral Deposits Research, Department of Earth Sciences, Nanjing University, Nanjing 210093, China b Department of Earth Sciences, Dalhousie University, Halifax B3H 3J5, Canada c Geological Centre for Tibetan Plateau, China University of Geosciences, Beijing 100083, P.R. China Received 24 January 2007; accepted in revised form 14 May 2007 Available online 28 November 2007 Abstract Lithostratigraphic studies of the Upper JurassiceLower Cretaceous sedimentary successions exposed in the TingrieGyangze area, south- eastern Tibet resulted in the establishment of a revised stratigraphic framework. A major crustal fault separates the southern Tibetan sedimentary successions into a Southern Zone and a Northern Zone. The Upper JurassiceLower Cretaceous strata of the Southern Zone are subdivided into the Menkadun Formation (Oxfordian, Kimmeridgian, up to lower Upper Tithonian), the Gucuo Formation (Upper Tithonian to Lower Albian) and the overlying Dongshan Formation (Upper Albian). The Gucuo Formation is further subdivided into a quartz arenite unit, which is overlain by a shale unit, in turn overlain by a volcaniclastic sandstone unit. The youngest cluster of detrital zircon absolute age data (127.7 Æ 1.8 Ma) from the lower part of the volcaniclastic unit of the Gucuo Formation place the volcanic event before the Late Barremian. In the Northern Zone, the Upper JurassiceLower Cretaceous strata have been subdivided into four formations: the Zhera Formation (Upper Jurassic), the Weimei Formation (Tithonian), the Rilang Formation (?Berriasian) and the Gyabula Formation (? post-Valanginian). -
The Scales of Mesozoic Actinopterygians
Mesozoic Fishes – Systematics and Paleoecology, G. Arratia & G. Viohl (eds.): pp. 83-93, 6 figs. © 1996 by Verlag Dr. Friedrich Pfeil, München, Germany – ISBN 3-923871–90-2 The scales of Mesozoic actinopterygians Hans-Peter SCHULTZE Abstract Cycloid scales (elasmoid scales with circuli) are a unique character of teleosts above the level of Pholidophorus and Pholidophoroides. Cycloid scales have two layers. A bony layer, usually acellular, is superimposed on a basal plate composed of partially mineralized layers of plywoodlike laminated collagen fibres. The tissue of the basal layer is refered to here as elasmodin. Basal teleosts (sensu PATTERSON 1973) possess rhombic scales with a bony base overlain by ganoin (lepidosteoid ganoid scale). Amioid scales (elasmoid scales with longitudinally to radially arranged ridges or rods on the overlapped field) are found within halecomorphs. This scale type evolved more than once within primitive actinopterygians and other osteichthyan fishes. It may have even developed twice within halecomorphs, in Caturidae and Amiidae, from rhombic scales of lepidosteoid type. Some basal genera of halecomorphs show remains of a dentine layer between ganoin and bone that is characteristic of actinopterygians below the halecostome level. The Semionotidae placed at the base of the Halecostomi, exhibit scale histology transitional between the palaeoniscoid and lepidosteoid scale type. Introduction Actinopterygians, from primitive Coccolepididae to advanced teleosts, are represented in the Solnhofen lithographic limestone. These are fishes with rhombic and round scales. Ganoid scales of the lepidosteoid type are found in the following fishes: semionotid Lepidotes and Heterostrophus, macrosemiids Histionotus, Macrosemius, Notagogus and Propterus, ophiopsid Ophiopsis, caturids Furo and Brachyichthys, aspido- rhynchid Belonostomus, pleuropholid Pleuropholis, and pholidophorid Pholidophorus. -
Vertebrate Remains Are Relatively Well Known in Late Jurassic Deposits of Western Cuba. the Fossil Specimens That Have Been Coll
Paleontología Mexicana, 3 (65): 24-39 (versión impresa), 4: 24-39 (versión electrónica) Catalogue of late jurassiC VerteBrate (pisCes, reptilian) speCiMens froM western CuBa Manuel Iturralde-Vinent ¹, *, Yasmani Ceballos Izquierdo ² A BSTRACT Vertebrate remains are relatively well known in Late Jurassic deposits of western Cuba. The fossil specimens that have been collected so far are dispersed in museum collections around the world and some have been lost throughout the years. A reas- sessment of the fossil material stored in some of these museums’ collections has generated new data about the fossil-bearing lo- calities and greatly increased the number of formally identified specimens. The identified bone elements and taxa suggest a high vertebrate diversity dominated by actinopterygians and reptiles, including: long-necked plesiosaurs, pliosaurs, metriorhynchid crocodilians, pleurodiran turtles, ichthyosaurs, pterosaurs, and sauropod dinosaurs. This assemblage is commonly associated with unidentified remains of terrestrial plants and rare microor- ganisms, as well as numerous marine invertebrates such as am- monites, belemnites, pelecypods, brachiopods, and ostracods. This fossil assemblage is particularly valuable because it includes the most complete marine reptile record of a chronostratigraphic interval, which is poor in vertebrate remains elsewhere. In this contribution, the current status of the available vertebrate fossil specimens from the Late Jurassic of western Cuba is provided, along with a brief description of the fossil materials. Key words: Late Jurassic, Oxfordian, dinosaur, marine reptiles, fish, western Cuba. I NTRODUCTION Since the early 20th century, different groups of collectors have discovered 1 Retired curator, Museo a relatively rich and diverse vertebrate assemblage in the Late Jurassic stra- Nacional de Historia Natural, ta of western Cuba, which has been only partially investigated (Brown and Havana, Cuba. -
Jurassic-Cretaceous Tectonic and Depositional Evolution of the Forearc
International Geology Review ISSN: 0020-6814 (Print) 1938-2839 (Online) Journal homepage: https://tandfonline.com/loi/tigr20 The Byers Basin: Jurassic-Cretaceous tectonic and depositional evolution of the forearc deposits of the South Shetland Islands and its implications for the northern Antarctic Peninsula Joaquin Bastias, Mauricio Calderón, Lea Israel, Francisco Hervé, Richard Spikings, Robert Pankhurst, Paula Castillo, Mark Fanning & Raúl Ugalde To cite this article: Joaquin Bastias, Mauricio Calderón, Lea Israel, Francisco Hervé, Richard Spikings, Robert Pankhurst, Paula Castillo, Mark Fanning & Raúl Ugalde (2019): The Byers Basin: Jurassic-Cretaceous tectonic and depositional evolution of the forearc deposits of the South Shetland Islands and its implications for the northern Antarctic Peninsula, International Geology Review, DOI: 10.1080/00206814.2019.1655669 To link to this article: https://doi.org/10.1080/00206814.2019.1655669 View supplementary material Published online: 21 Aug 2019. Submit your article to this journal View related articles View Crossmark data Full Terms & Conditions of access and use can be found at https://tandfonline.com/action/journalInformation?journalCode=tigr20 INTERNATIONAL GEOLOGY REVIEW https://doi.org/10.1080/00206814.2019.1655669 ARTICLE The Byers Basin: Jurassic-Cretaceous tectonic and depositional evolution of the forearc deposits of the South Shetland Islands and its implications for the northern Antarctic Peninsula Joaquin Bastias a,b, Mauricio Calderónc, Lea Israela, Francisco Hervéa,c, Richard -
Remarks on the Tithonian–Berriasian Ammonite Biostratigraphy of West Central Argentina
Volumina Jurassica, 2015, Xiii (2): 23–52 DOI: 10.5604/17313708 .1185692 Remarks on the Tithonian–Berriasian ammonite biostratigraphy of west central Argentina Alberto C. RICCARDI 1 Key words: Tithonian–Berriasian, ammonites, west central Argentina, calpionellids, nannofossils, radiolarians, geochronology. Abstract. Status and correlation of Andean ammonite biozones are reviewed. Available calpionellid, nannofossil, and radiolarian data, as well as radioisotopic ages, are also considered, especially when directly related to ammonite zones. There is no attempt to deal with the definition of the Jurassic–Cretaceous limit. Correlation of the V. mendozanum Zone with the Semiforme Zone is ratified, but it is open to question if its lower part should be correlated with the upper part of the Darwini Zone. The Pseudolissoceras zitteli Zone is characterized by an assemblage also recorded from Mexico, Cuba and the Betic Ranges of Spain, indicative of the Semiforme–Fallauxi standard zones. The Aulacosphinctes proximus Zone, which is correlated with the Ponti Standard Zone, appears to be closely related to the overlying Wind hauseniceras internispinosum Zone, although its biostratigraphic status needs to be reconsidered. On the basis of ammonites, radiolarians and calpionellids the Windhauseniceras internispinosum Assemblage Zone is approximately equivalent to the Suarites bituberculatum Zone of Mexico, the Paralytohoplites caribbeanus Zone of Cuba and the Simplisphinctes/Microcanthum Zone of the Standard Zonation. The C. alternans Zone could be correlated with the uppermost Microcanthum and “Durangites” zones, although in west central Argentina it could be mostly restricted to levels equivalent to the “Durangites Zone”. The Substeueroceras koeneni Zone ranges into the Occitanica Zone, Subalpina and Privasensis subzones, the A. -
Re-Evaluation of Pachycormid Fishes from the Late Jurassic of Southwestern Germany
Editors' choice Re-evaluation of pachycormid fishes from the Late Jurassic of Southwestern Germany ERIN E. MAXWELL, PAUL H. LAMBERS, ADRIANA LÓPEZ-ARBARELLO, and GÜNTER SCHWEIGERT Maxwell, E.E., Lambers, P.H., López-Arbarello, A., and Schweigert G. 2020. Re-evaluation of pachycormid fishes from the Late Jurassic of Southwestern Germany. Acta Palaeontologica Polonica 65 (3): 429–453. Pachycormidae is an extinct group of Mesozoic fishes that exhibits extensive body size and shape disparity. The Late Jurassic record of the group is dominated by fossils from the lithographic limestone of Bavaria, Germany that, although complete and articulated, are not well characterized anatomically. In addition, stratigraphic and geographical provenance are often only approximately known, making these taxa difficult to place in a global biogeographical context. In contrast, the late Kimmeridgian Nusplingen Plattenkalk of Baden-Württemberg is a well-constrained locality yielding hundreds of exceptionally preserved and prepared vertebrate fossils. Pachycormid fishes are rare, but these finds have the potential to broaden our understanding of anatomical variation within this group, as well as provide new information regarding the trophic complexity of the Nusplingen lagoonal ecosystem. Here, we review the fossil record of Pachycormidae from Nusplingen, including one fragmentary and two relatively complete skulls, a largely complete fish, and a fragment of a caudal fin. These finds can be referred to three taxa: Orthocormus sp., Hypsocormus posterodorsalis sp. nov., and Simocormus macrolepidotus gen. et sp. nov. The latter taxon was erected to replace “Hypsocormus” macrodon, here considered to be a nomen dubium. Hypsocormus posterodorsalis is known only from Nusplingen, and is characterized by teeth lacking apicobasal ridging at the bases, a dorsal fin positioned opposite the anterior edge of the anal fin, and a hypural plate consisting of a fused parhypural and hypurals. -
Copyrighted Material
06_250317 part1-3.qxd 12/13/05 7:32 PM Page 15 Phylum Chordata Chordates are placed in the superphylum Deuterostomia. The possible rela- tionships of the chordates and deuterostomes to other metazoans are dis- cussed in Halanych (2004). He restricts the taxon of deuterostomes to the chordates and their proposed immediate sister group, a taxon comprising the hemichordates, echinoderms, and the wormlike Xenoturbella. The phylum Chordata has been used by most recent workers to encompass members of the subphyla Urochordata (tunicates or sea-squirts), Cephalochordata (lancelets), and Craniata (fishes, amphibians, reptiles, birds, and mammals). The Cephalochordata and Craniata form a mono- phyletic group (e.g., Cameron et al., 2000; Halanych, 2004). Much disagree- ment exists concerning the interrelationships and classification of the Chordata, and the inclusion of the urochordates as sister to the cephalochor- dates and craniates is not as broadly held as the sister-group relationship of cephalochordates and craniates (Halanych, 2004). Many excitingCOPYRIGHTED fossil finds in recent years MATERIAL reveal what the first fishes may have looked like, and these finds push the fossil record of fishes back into the early Cambrian, far further back than previously known. There is still much difference of opinion on the phylogenetic position of these new Cambrian species, and many new discoveries and changes in early fish systematics may be expected over the next decade. As noted by Halanych (2004), D.-G. (D.) Shu and collaborators have discovered fossil ascidians (e.g., Cheungkongella), cephalochordate-like yunnanozoans (Haikouella and Yunnanozoon), and jaw- less craniates (Myllokunmingia, and its junior synonym Haikouichthys) over the 15 06_250317 part1-3.qxd 12/13/05 7:32 PM Page 16 16 Fishes of the World last few years that push the origins of these three major taxa at least into the Lower Cambrian (approximately 530–540 million years ago). -
A Synoptic Review of the Vertebrate Fauna from the “Green Series
A synoptic review of the vertebrate fauna from the “Green Series” (Toarcian) of northeastern Germany with descriptions of new taxa: A contribution to the knowledge of Early Jurassic vertebrate palaeobiodiversity patterns I n a u g u r a l d i s s e r t a t i o n zur Erlangung des akademischen Grades eines Doktors der Naturwissenschaften (Dr. rer. nat.) der Mathematisch-Naturwissenschaftlichen Fakultät der Ernst-Moritz-Arndt-Universität Greifswald vorgelegt von Sebastian Stumpf geboren am 9. Oktober 1986 in Berlin-Hellersdorf Greifswald, Februar 2017 Dekan: Prof. Dr. Werner Weitschies 1. Gutachter: Prof. Dr. Ingelore Hinz-Schallreuter 2. Gutachter: Prof. Dr. Paul Martin Sander Tag des Promotionskolloquiums: 22. Juni 2017 2 Content 1. Introduction .................................................................................................................................. 4 2. Geological and Stratigraphic Framework .................................................................................... 5 3. Material and Methods ................................................................................................................... 8 4. Results and Conclusions ............................................................................................................... 9 4.1 Dinosaurs .................................................................................................................................. 10 4.2 Marine Reptiles ....................................................................................................................... -
Body-Shape Diversity in Triassic–Early Cretaceous Neopterygian fishes: Sustained Holostean Disparity and Predominantly Gradual Increases in Teleost Phenotypic Variety
Body-shape diversity in Triassic–Early Cretaceous neopterygian fishes: sustained holostean disparity and predominantly gradual increases in teleost phenotypic variety John T. Clarke and Matt Friedman Comprising Holostei and Teleostei, the ~32,000 species of neopterygian fishes are anatomically disparate and represent the dominant group of aquatic vertebrates today. However, the pattern by which teleosts rose to represent almost all of this diversity, while their holostean sister-group dwindled to eight extant species and two broad morphologies, is poorly constrained. A geometric morphometric approach was taken to generate a morphospace from more than 400 fossil taxa, representing almost all articulated neopterygian taxa known from the first 150 million years— roughly 60%—of their history (Triassic‒Early Cretaceous). Patterns of morphospace occupancy and disparity are examined to: (1) assess evidence for a phenotypically “dominant” holostean phase; (2) evaluate whether expansions in teleost phenotypic variety are predominantly abrupt or gradual, including assessment of whether early apomorphy-defined teleosts are as morphologically conservative as typically assumed; and (3) compare diversification in crown and stem teleosts. The systematic affinities of dapediiforms and pycnodontiforms, two extinct neopterygian clades of uncertain phylogenetic placement, significantly impact patterns of morphological diversification. For instance, alternative placements dictate whether or not holosteans possessed statistically higher disparity than teleosts in the Late Triassic and Jurassic. Despite this ambiguity, all scenarios agree that holosteans do not exhibit a decline in disparity during the Early Triassic‒Early Cretaceous interval, but instead maintain their Toarcian‒Callovian variety until the end of the Early Cretaceous without substantial further expansions. After a conservative Induan‒Carnian phase, teleosts colonize (and persistently occupy) novel regions of morphospace in a predominantly gradual manner until the Hauterivian, after which expansions are rare.