A Distribution of Dragonflies and Damselflies in the State of Wyoming

George G. Sims1

Most recent update: August 28, 2013

This “working document” is an attempt to bring together odonate collection and distribution information from two major sources, the national odonate database at Odonata Central (Abbott 2007) and the Odonata of Wyoming (Molinar & Lavigne 1979), and present it in a clear and easily- accessible form. Every care has been taken to insure the accuracy of the information, and any errors, omissions, or oversights are completely the fault of the author. It is hoped that this effort will help to provide a starting point for odonatists who are seeking previously unrecorded species or studying the possibilities of discovering types which should be encountered in a specific county or location. No new information, other than that contained in the above references, is found here.

Methodology

In an e-mail exchange with Dr. Scott Shaw, curator of the insect museum at the University of Wyoming, Dr. Shaw made me aware of the existence of Molinar & Lavigne’s Odonata of Wyoming, a study published in 1979, which is believed to be the most recent comprehensive record of odonate collection in the state. After several weeks’ fruitless search for a copy of the book, Dr. Ken Tennessen, of Wisconsin, informed me that he possessed two copies, and kindly presented me with one for the purpose of this distribution study.

Some seventy-nine (79) odonate species have been found to occur in Wyoming—thirty-three (33) zygopteran (damselfly) and forty-six (46) anisopteran (dragonfly) varieties. Since the publication of Odonata of Wyoming, several taxonomic changes have occurred; therefore:

Aeshna californica becomes Rhioaeshna californica Aeshna multicolor Rhionaeshna multicolor Argia violacea A. fumipennis Enallagma cyathigerum E. annexum Gomphus cornutus Arigomphus cornutus Gomphus intricatus Stylurus intricatus Libellula lydia Plathemis lydia Sympetrum occidentale S. semicinctum

1 115 Dillon Drive, Lander, Wyoming 82520-2957, [email protected] 2

All records noted as originating in Molinar & Lavigne are taken from Appendix II, “List of species of Odonata in Wyoming by county.” In the case of Park County, which encompasses Yellowstone National Park, only those records specifically attributable to Park County have been included, as the National Park also includes areas outside the boundaries of the State of Wyoming. Odonata of Wyoming lists a separate classification for records within the park, which are not included.

In the last few years, the advent of Dr. John Abbott’s remarkable on-line Odonata Central (www.odonatacentral.org) has proven to be a valuable resource for listing individual collecting efforts, and documenting the presence of odonate species throughout the country (and the world).

Information from Odonata Central for the selected counties was first extracted, with the submissions from Thomas W. Donnelly’s DOT Map Project segregated from those entries representing collections by other individuals.

The author has presented the distribution data in several ways: first, by listing each of the species found in the area, showing first a distribution map, then a listing, of the counties from which the species has been reported. English names, as well as the taxonomic arrangement of the species accounts, are taken from Paulson & Dunkle, 2011. Alongside each county represented is found an alphabetic code which indicates the source of the information.

This section is immediately followed by a listing of the counties of Wyoming, with a state map showing the location of the particular county, followed by a listing of the individual species which have been reported within each county, again with each representative species followed by a source code.

The source codes for the information are as follows:

W – A specimen recorded in Odonata of Wyoming.

D – An adult specimen recorded in Thomas W. Donnelly’s DOT Map Project, as shown in the records of Odonata Central.

O – An adult specimen has been collected and/or photographed by an individual not affiliated with the DOT Map Project, with identification verified by an expert, and the record appears on Odonata Central. (Note: Odonata Central is not currently configured to maintain records of larval [nymph] specimens.)

T – Specimen collected by Dr. Lusha Tronstad, University of Wyoming.

The experts mentioned above consist of several prominent odonatologists who provide verification of all records submitted for inclusion on Odonata Central.

THE COUNTIES OF WYOMING

Albany Hot Springs Platte Big Horn Johnson Sheridan Campbell Laramie Sublette Carbon Lincoln Sweetwater Converse Natrona Teton Crook Niobrara Uinta Fremont Park Washakie Goshen Weston

Summary

Every effort has been made to ensure the accurate of this information. Any erroneous or incomplete data is the fault of the author. All suggestions, comments, or corrections are welcomed, and should be directed to:

George G. Sims 115 Dillon Drive, #313 Lander, Wyoming 82520-2957 (417) 554-1682 [email protected] Last updated: August 28, 2013

THE SPECIES DISTRIBUTIONS

Hetaerina Hagen in Selys Lestes Leach Suborder Rubyspots Pond Spreadwings ZYGOPTERA Damselflies Hetaerina americana Lestes congener American rubyspot Spotted spreadwing CALOPTERYGIDAE Broad-winged Damsels

Calopteryx Leach Jewelwings

Calopteryx aequabilis River jewelwing

Crook DOW Albany DW Goshen DW Big Horn DW Niobrara DW Campbell DOW Platte DW Carbon DW Sheridan DW Converse DW Weston W Crook O Fremont DOW Goshen DW LESTIDAE Hot Springs DW Spreadwings Johnson DW Crook DW Laramie DW Platte DW Archilestes Selys Lincoln DW Natrona DW Teton DW Stream Spreadwings Niobrara DW

Calopteryx maculata Sheridan DW Archilestes grandis Sweetwater DW Ebony jewelwing Great spreadwing Teton DOW Uinta DW Weston D

Crook O Platte D Teton W

Lestes disjunctus Sweetwater DW COENAGRIONIDAE Northern spreadwing Teton DOW Pond Damsels Uinta DW Weston DW Amphiagrion Selys Lestes forcipatus Red Damsels Sweetflag spreadwing Amphiagrion abbreviatum Western red damsel

Albany DW Big Horn DW Campbell DOW Carbon D Crook DW Converse DW Fremont W Park DW Niobrara DW

Sublette DW Sheridan DW Albany DW Teton DW Big Horn DW Uinta DW Lestes unguiculatus Carbon DW

Lyre-tipped spreadwing Converse DW Lestes dryas Crook DOW Emerald spreadwing Fremont DOW Hot Springs W Johnson DW Laramie DW Lincoln DW Natrona DW Niobrara DW Platte DW Sheridan DW Sublette DW Albany DW Sweetwater DOW Big Horn D Teton DW Albany DW Campbell DW Uinta DW Campbell DOW Carbon DW Washakie DW Carbon DW Converse DW Weston DW Converse DW Crook DW Crook D Fremont DW Goshen DW Fremont DW Hot Springs DW Johnson DW Johnson DW Natrona DW Laramie DW Niobrara DW Lincoln DW Platte DW Sheridan DW Natrona DW Weston DW Niobrara DW Park D Platte DW Sheridan D Sublette DW

Argia Rambur Crook DOW Big Horn DW Carbon D Dancers Fremont DW Goshen D Crook DW

Hot Springs DW Hot Springs DW Natrona DW Argia alberta Niobrara DW Niobrara DW Paiute dancer Platte DW Platte DW Sheridan DW Sheridan DW Washakie DW Teton DW Weston DW Argia fumipennis Variable dancer Coenagrion Kirby Eurasian Bluets

Coenagrion resolutum Taiga bluet

Big Horn DW Campbell DW Niobrara DW Teton DW

Argia apicalis Albany DW Blue-fronted dancer Campbell DW Crook DOW Niobrara DW

Albany DW Argia moesta Big Horn DW Powdered dancer Carbon DW Crook DW Fremont DW Goshen D Park D Sheridan D Sublette DW Teton DW Crook O Uinta DW

Argia emma Emma’s dancer Crook O

Argia vivida

Vivid dancer

Albany DW Big Horn DW Carbon D Converse DW

Enallagma Charpentier Goshen DW Enallagma boreale Hot Springs DOW American Bluets Johnson DW Boreal bluet Laramie DW Enallagma anna Lincoln W Natrona DW River bluet Niobrara DW Park DW Platte DW Sheridan DW Sublette DW Sweetwater DW Teton DW Uinta DW Washakie DW Weston DW Albany DW Big Horn DW Enallagma antennatum Campbell DOW Carbon DW Albany DW Rainbow bluet Converse DW Big Horn DW Crook DOW Crook DOW Fremont DOW Fremont DOW Goshen DW Goshen DW Johnson DW Hot Springs DW Laramie DW Johnson DW Lincoln DW Laramie DW Natrona DW Lincoln DW Niobrara DW Natrona DW Park DW Niobrara DW Platte DW Platte DW Sheridan D Sheridan DW Sublette DW Sublette DW Campbell DW Sweetwater D Sweetwater DW Crook DOW Teton DW Teton DOW Washakie DW Uinta DW Enallagma basidens Weston DW Washakie DW Double-striped bluet Weston DW Enallagma carunculatum

Enallagma annexum Tule bluet Northern bluet

Platte O

Big Horn DW Campbell DOW Albany DW Carbon D Big Horn DW Converse DW Campbell DW Fremont DOW Carbon DW Goshen DW Converse DW Natrona DW Crook DOW Platte DW Fremont DW Sheridan DW

Washakie DW Enallagma ebrium Platte DW Weston DW Sheridan DW Marsh bluet Ischnura Charpentier Enallagma civile Forktails Familiar bluet Ischnura cervula Pacific forktail

Sheridan DW Washakie DW Weston DW

Enallagma hageni

Albany DW Hagen’s bluet

Campbell DW Big Horn DW Converse DW Campbell O Crook DOW Carbon DW Fremont DW Converse DW Goshen DW Fremont Ocervulacervula Hot Springs O Hot Springs DW Laramie DW Johnson DW Natrona DW Lincoln DW Niobrara DW Natrona DW Platte DW Park D Weston DW Sheridan DW

Teton DW Enallagma clausum Campbell DW Washakie DW Crook DOW Alkali bluet Natrona DW Sheridan DW Ischnura damula Plains forktail Enallagma praevarum Arroyo bluet

Albany DW Converse DW Crook DW Fremont DW Albany DW Campbell DOW Goshen DW Converse DW Johnson DW Albany DW Crook DOW Park D Big Horn DW Goshen WW Sheridan DW Campbell DW Laramie DW Sublette DW Converse DW Natrona DW Uinta DW Crook DW Niobrara DW Weston DW Laramie DW Platte DW Niobrara DW

Sheridan DW Weston DW Aeshna eremita Weston DW Lake darner Nehalennia Selys Ischnura perparva Sprites Western forktail Nehalennia irene Sedge sprite

Albany DW Fremont DOW Goshen D Albany DW Park DW Big Horn DW Sheridan D Campbell DW Carbon D Sublette DO Carbon D Crook DW Teton D Converse DW Sheridan D Crook DOW Weston DW Aeshna interrupta Fremont DOW Goshen DW Variable darner Hot Springs DW Johnson DW Suborder Laramie DW Lincoln DW ANISOPTERA Natrona DW Dragonflies Niobrara DW Platte DW Sheridan DW AESHNIDAE Washakie DW Darners Weston DW

Ischnura verticalis Aeshna Fabricius Mosaic Darners Albany DW Eastern forktail Big Horn DW Campbell O Aeshna constricta Carbon D Lance-tipped darner Fremont DO Goshen DO Park DW Teton DW Uinta DW Weston O

Albany DW

Campbell DW

Carbon DW

Converse DW Platte D

Crook DOW

Natrona DW Niobrara DW Platte DW Sheridan DW

Aeshna juncea Aeshna sitchensis Natrona DW Washakie DW Sedge darner Zigzag darner Rhionaeschna Förster Neotropical Darners

Rhionaeschna californica California darner

Albany DW Fremont DO Carbon DW Goshen DO Fremont D Sublette DO Goshen D Teton DW Sheridan D Sublette DO Aeshna umbrosa Teton DW Shadow darner Crook DOW Natrona DW Aeshna palmata Sheridan D Paddle-tailed darner Weston DW

Rhionaeschna multicolor Blue-eyed darner

Carbon D Teton DO

Albany DW Anax Leach Campbell DOW Green Darners Carbon DW

Converse DW Crook O Anax junius Albany DW Lincoln DW Common green darner Campbell D Niobrara DW Carbon D Park DW Goshen W Platte W Hot Springs DW Sheridan DW Laramie O Sublette DW Platte DW Sweetwater DW Teton DOW Teton DW Weston DW Weston O

Albany DW Campbell O Crook DW Hot Springs DW

GOMPHIDAE Gomphus graslinellus Platte DW Sheridan D Clubtails Pronghorn clubtail Sublette DW Sweetwater DW Arigomphus Needham Teton DW Washakie DW Pond Clubtails Weston DW

Arigomphus cornutus Stylurus Needham Horned clubtail Hanging Clubtails

Stylurus intricatus Brimstone clubtail

Crook O

Ophiogomphus Selys Snaketails

Ophiogomphus morrisoni Campbell DW Great basin snaketail Crook DW

Gomphus Leach Common Clubtails Sweetwater DW

Gomphus externus CORDULEGASTRIDAE Plains clubtail Spiketailss

Cordulegaster Leach

Spiketails Sheridan W Cordulegaster dorsalis Ophiogomphus severus Pacific spiketail Pale snaketail

Niobrara DO Platte DW

Albany DW Teton O Carbon DW Converse DW Crook W Hot Springs DW Laramie DW Lincoln D Park O

CORDULIIDAE Somatochlora Selys Somatochlora minor Emeralds Striped Emeralds Ocellated emerald

Cordulia Leach Somatochlora cingulata Common Emeralds Lake emerald

Cordulia shurtleffi American emerald

Albany DW Teton DW

Albany DW Somatochlora semicircularis Somatochlora ensigera Mountain emerald Plains emerald Albany DW Sheridan D Sublette D Teton DW

Epitheca Burmeister Baskettails

Epitheca petechialis

Dot-winged baskettail Albany DW Johnson DW Carbon D Weston D Fremont D Goshen D Somatochlora hudsonica Park T Sheridan D Hudsonian emerald Sublette D Teton DW

Crook O

Albany DW Carbon D Fremont D Goshen D Park D Sublette D Teton DW

LIBELLULIDAE Leucorrhinia glacialis Park DW Sheridan DW Skimmers Crimson-ringed whiteface Washakie DW

Erythemis Hagen Leucorrhinia proxima Pondhawks Belted whiteface

Erythemis collocata Western pondhawk

Sublette D

Leucorrhinia hudsonica

Hudsonian whiteface Sheridan D Teton DW

Park O Libellula Linnaeus Teton DOW King Skimmers

Leucorrhinia Brittinger Libellula forensis Whitefaces Eight-spotted skimmer

Leucorrhinia borealis

Boreal whiteface Albany DW Big Horn DW Carbon DW Fremont DOW Goshen DO Johnson DW Teton DOW Weston DW

Albany DW Leucorrhinia intacta Campbell DOW Dot-tailed whiteface Converse DW Crook O Big Horn DW Goshen W Fremont DW Hot Springs DW Goshen D Johnson DW Park D Natrona DW Teton DW Platte DW Sweetwater O Teton DW Washakie DW Weston DW

Campbell DW Crook DW Hot Springs DW Natrona DW Niobrara W

Libellula luctuosa Carbon D Plathemis subornata Crook DOW Widow skimmer Fremont DW Desert whitetail Goshen D Hot Springs DW Johnson DW Laramie DW Natrona DW Niobrara DW Park D Platte DW Sweetwater DW Teton DW Uinta D

Washakie DW Campbell O Weston DW Platte DW

Libellula pulchella Libellula saturata Sympetrum Newman Twelve-spotted skimmer Flame skimmer Meadowhawks

Sympetrum corruptum Variegated meadowhawk

Big Horn DW Crook D Campbell DOW Park D Carbon D Teton DOW Converse DW Crook DW Plathemis Hagen Albany DW Laramie DW Whitetails Big Horn DW Natrona DW Campbell DOW

Platte DW Carbon DW Sweetwater DW Plathemis lydia Converse DW Weston DW Common whitetail Crook DOW Fremont DW Libellula quadrimaculata Goshen DW Hot Springs DW Four-spotted skimmer Johnson DW Laramie DW Lincoln D Natrona DW Niobrara DW Park D Platte DW Sheridan D Sweetwater DW Teton DW Campbell O Washakie DW Sheridan DW Weston DW

Teton DW

Albany DW

Big Horn DW

Campbell DW

Sympetrum costiferum Sympetrum internum Sympetrum obtrusum Saffron-winged Cherry-faced meadowhawk White-faced meadowhawk meadowhawk

Albany DW

Big Horn DW Campbell O Campbell DW Carbon D Big Horn DW Carbon DW Crook DOW Campbell DW Converse DW Fremont O Crook O Crook DW Goshen W Fremont O Fremont DW Niobrara O Lincoln DW Goshen DW Platte D Sheridan W Johnson D Sheridan DW Teton DOW Laramie DW Washakie DW Teton DW Lincoln DW Washakie DW Natrona DW

Sympetrum danae Niobrara DW Sympetrum pallipes Black meadowhawk Park DW Platte DW Striped meadowhawk Sublette DW Sweetwater DOW Teton DW Uinta DW

Sympetrum madidum Red-veined meadowhawk

Albany DW Big Horn DW Albany DW Carbon D Campbell DOW Fremont DW Carbon DW Goshen D Converse DW Lincoln DW Crook DOW Sublette DW Fremont O Johnson D Sweetwater DOW Laramie DOW Teton DOW Albany DW Natrona DW Campbell O Niobrara DOW Carbon DW Platte DW Crook DW Sheridan DW Niobrara DW Sublette DW Sublette DW Sweetwater DW Sweetwater DW Teton DW Teton D Uinta O

Washakie DW Weston DW

Sympetrum rubicundulum Sympetrum semicinctum Carbon DW Converse DW Ruby meadowhawk Band-winged Crook O meadowhawk Fremont DOW Goshen D Johnson DW Laramie DOW Niobrara DOW Platte DW Sweetwater O Teton D Uinta DW Weston DW

Albany W Carbon W Niobrara O Albany DW Sublette D Big Horn DW Sweetwater W Campbell DOW

ODONATE COLLECTION FREQUENCY BY COUNTIES

The map below shows the number of species recorded in each of the twenty-three (23) counties of Wyoming. In the opinion of the author, a low number of species collected in a particular county does not necessarily prove a paucity of species present, but may merely represent a low number of collectors/observers working in that county.

The information, however, may prove useful to those odonatists seeking new county records, or who desire to venture into more virgin territory, as they may wish to concentrate on the under- collected areas.

White – <20 species collected/observed Pink – 21-30 species collected/observed Orange – 31-40 species collected/observed Gray – >41 species collected/observed

THE COUNTY DISTRIBUTIONS

Albany (39): Big Horn (25): Anax junius O Argia alberta DW Argia fumipennis DW Ariogomphus cornutus D Enallagma annexum DW Enallagma antennatum DW Enallagma boreale DOW Enallagma carunculatum DOW Enallagma civile DW Enallagma hageni DW Enallagma praevarum DW Ischnura cervula O Ischnura damula DOW Ischnura perparva DW Aeshna eremita DW Aeshna interrupta DW Ischnura verticalis DW Aeshna interrupta DW Amphiagrion abbreviatum DW Lestes congener DOW Aeshna juncea DW Argia alberta DW Lestes disjunctus DOW Aeshna palmata DW Argia emma DW Lestes dryas DOW Amphiagrion abbreviatum DW Argia vivida DW Lestes unguiculatus DW Anax junius DW Leucorrhinia intacta DW Coenagrion resolutum DW Argia emma DW Libellula forensis DOW Argia fumipennis DW Enallagma anna DW Libellula luctuosa O Coenagrion resolutum DW Enallagma annexum DW Libellula pulchella DOW Cordulia shurtleffii DW Enallagma boreale DW Libellula quadrimaculata DW Enallagma anna DW Enallagma carunculatum DW Plathemis lydia O Enallagma annexum DW Enallagma praevarum DW Rhionaeschna multicolor O Sympetrum corruptum DOW Enallagma boreale DW Ischnura cervula DW Enallagma civile DW Sympetrum costiferum DW Enallagma clausum DW Ischnura perparva DW Sympetrum internum DW Enallagma praevarum DW Lestes congener DW Sympetrum madidum O Ischnura damula DW Lestes disjunctus DW Sympetrum obtrusum O Ischnura perparva DW Lestes unguiculatus D Sympetrum pallipes DOW Ischnura verticalis DW Leucorrhinia borealis DW Sympetrum semicinctum DOW Lestes congener DW Leucorrhinia hudsonica DW Lestes disjunctus DW Libellula pulchella DW Carbon (34): Lestes dryas DW Lestes unguiculatus DW Libellula quadrimaculata DW Leucorrhinia hudsonica DW Sympetrum corruptum DW Libellula forensis DW Sympetrum costiferum DW Libellula quadrimaculata DW Sympetrum danae DW Ophiogomphus severus DW Sympetrum internum DW Rhionaeschna multicolor DW Sympetrum semicinctum DW Somatochlora cingulata DW

Somatochlora hudsonica DW Somatochlora minor DW Campbell (35): Somatochlora semicircularis DW Sympetrum corruptum DW Sympetrum danae DW Sympetrum internum DW Aeshna interrupta D Sympetrum madidum DW Aeshna juncea DW Sympetrum pallipes DW Aeshna palmata DW Sympetrum rubicundulum W Aeshna umbrosa D Sympetrum semicinctum DW Amphiagrion abbreviatum DW Argia emma D Argia vivida D Coenagrion resolutum DW Enallagma annexum DW Aeshna interrupta O Enallagma boreale DW Aeshna palmata DOW Enallagma carunculatum D

Ischnura cervula DW Crook (45): Fremont (31): Ischnura perparva D Ischnura verticalis DW Lestes congener DW Lestes disjunctus D Lestes dryas DW Lestes unguiculatus DW Leucorrhinia hudsonica DW Libellula pulchella D Libellula quadrimaculata D Nehalennia irene D Ophiogomphus severus DW Rhionaeschna multicolor D Somatochlora hudsonica D Somatochlora semicircularis D Aeshna palmata O Aeshna eremita DOW Sympetrum corruptum DW Amphiagrion abbreviatum DOW Aeshna interrupta DO Sympetrum danae D Anax junius DW Aeshna juncea D Sympetrum internum DW Argia apicalis O Aeshna sitchensis DO Sympetrum madidum DW Argia emma DOW Amphiagrion abbreviatum DOW Sympetrum obtrusum D Argia fumipennis DOW Argia emma DW Sympetrum pallipes DW Argia moesta O Coenagrion resolutum DW Sympetrum rubicundulum W Argia vivida DW Enallagma anna DOW Sympetrum semicinctum D Ariogomphus cornutus DW Enallagma annexum DW Calopteryx aequabilis DW Enallagma boreale DOW Converse (24): Calopteryx maculata O Enallagma carunculatum DOW Coenagrion resolutum DW Enallagma civile DW Enallagma anna DOW Enallagma clausum DW Enallagma annexum DOW Ischnura cervula O Enallagma antennatum DOW Ischnura perparva DOW Enallagma boreale DOW Lestes congener DOW Enallagma civile DOW Lestes disjunctus W Enallagma clausum DW Lestes dryas DW Enallagma hageni DOW Lestes unguiculatus DW Enallagma praevarum DOW Leucorrhinia borealis DW Epitheca petechialis O Leucorrhinia hudsonica DOW Gomphus graslinellus O Libellula quadrimaculata D Hetaerina americana DOW Somatochlora hudsonica D Ischnura damula DOW Somatochlora semicircularis D Aeshna palmata DW Ischnura perparva DOW Sympetrum corruptum DW Amphiagrion abbreviatum DW Ischnura verticalis DOW Sympetrum costiferum O Argia emma DW Lestes congener O Sympetrum danae DW Enallagma annexum DW Lestes disjunctus DW Sympetrum internum DW Enallagma boreale DW Lestes dryas D Sympetrum obtrusum O Enallagma carunculatum DW Lestes unguiculatus DW Sympetrum pallipes O Enallagma civile DW Leucorrhinia intacta DW Sympetrum semicinctum DOW Enallagma clausum DW Libellula forensis O Enallagma praevarum DW Libellula pulchella DW Goshen (31): Ischnura cervula DW Libellula quadrimaculata DOW Ischnura damula DW Libellula saturata D Ischnura perparva DW Nehalennia irene DW Ischnura verticalis DW Ophiogomphus severus W Lestes congener DW Rhioaeschna californica DOW Lestes dryas DW Sympetrum corruptum DOW Lestes forcipatus DW Sympetrum costiferum O Lestes unguiculatus DW Sympetrum internum DW Libellula forensis DW Sympetrum madidum DW Libellula pulchella DW Sympetrum obtrusum DOW Ophiogomphus severus DW Sympetrum pallipes DOW Sympetrum corruptum DW Sympetrum semicinctum O Sympetrum internum DW Sympetrum pallipes DW Aeshna eremita D Sympetrum semicinctum DW Aeshna interrupta DO

Aeshna juncea D Johnson (18): Sympetrum internum DW Aeshna sitchensis DO Sympetrum pallipes DOW Amphiagrion abbreviatum D Sympetrum semicinctum DOW Argia emma D Coenagrion resolutum D Enallagma anna DW Lincoln (14): Enallagma annexum D Enallagma boreale DW Enallagma carunculatum DW Enallagma civile DW Enallagma clausum DW Hetaerina americana W Ischnura damula W Ischnura perparva DW Lestes congener DW Amphiagrion abbreviatum DW Lestes dryas D Enallagma anna DW Lestes unguiculatus DW Enallagma annexum DW Leucorrhinia borealis D Enallagma boreale DW Leucorrhinia hudsonica DO Enallagma clausum DW Aeshna palmata DW Libellula forensis W Ischnura cervula DW Amphiagrion abbreviatum DW Libellula quadrimaculata D Ischnura perparva DW Enallagma anna DW Rhionaeshna multicolor W Lestes congener DW Enallagma annexum DW Somatochlora hudsonica D Lestes dryas DW Enallagma boreale DW Somatochlora semicircularis D Lestes unguiculatus DW Ischnura cervula DW Sympetrum corruptum DW Leucorrhinia hudsonica DW Ischnura perparva DW Sympetrum danae D Libellula forensis DW Lestes congener DW Sympetrum internum DW Libellula quadrimaculata DW Lestes dryas DW Sympetrum obtrusum W Somatochlora ensigera DW Ophiogomphus severus D Sympetrum semicinctum D Sympetrum corruptum DW Sympetrum corruptum D Sympetrum internum D Sympetrum costiferum DW Sympetrum pallipes D Sympetrum danae DW Hot Springs (17): Sympetrum semicinctum DW Sympetrum internum DW

Laramie (18): Natrona (24):

Amphiagrion abbreviatum W Anax junius DW Argia emma DW Argia vivida DW Amphiagrion abbreviatum DW Amphiagrion abbreviatum DW Enallagma anna DW Enallagma anna DW Anax junius DW Enallagma annexum DOW Enallagma annexum DW Argia vivida DW Enallagma civile O Enallagma boreale DW Enallagma anna DW Ischnura cervula DW Enallagma civile DW Enallagma annexum DW Ischnura perparva DW Enallagma praevarum DW Enallagma boreale DW Lestes congener DW Ischnura damula DW Enallagma carunculatum DW Lestes dryas DW Ischnura perparva DW Enallagma civile DW Leucorrhinia intacta DW Lestes congener DW Enallagma hageni DW Libellula forensis DW Lestes dryas DW Ischnura cervula DW Libellula quadrimaculata DW Libellula pulchella DW Ischnura damula DW Ophiogomphus severus DW Libellula quadrimaculata DW Ischnura perparva DW Rhionaeshna multicolor DW Ophiogomphus severus OW Ischnura verticalis DW Sympetrum corruptum DW Rhionaeschna multicolor O Lestes congener DW

Sympetrum corruptum DW Lestes dryas DW

Lestes unguiculatus DW Park (19): Gomphus externus DW Leucorrhinia intacta DW Hetaerina americana DW Libellula forensis DW Ischnura damula DW Libellula pulchella DW Ischnura perparva DW Libellula quadrimaculata DW Ischnura verticalis DW Rhioaeschna californica DW Lestes dryas DW Sympetrum corruptum DW Lestes unguiculatus DW Sympetrum internum DW Libellula forensis DW Sympetrum pallipes DW Libellula pulchella DW Libellula quadrimaculata DW Niobrara (29): Ophiogomphus severus DW Plathemis subornata DW Rhionaeschna multicolor DW Sympetrum corruptum DW Aeshna eremita DW Sympetrum internum DW Aeshna interrupta DW Sympetrum obtrusum D Aeshna palmata DW Sympetrum pallipes DW Coenagrion resolutum D Sympetrum semicinctum DW Enallagma annexum DW Enallagma boreale DW Sheridan (36): Enallagma clausum D Erythemis collocata O Ischnura cervula D Lestes disjunctus DW Aeshna palmata DW Lestes dryas D Amphiagrion abbreviatum DW Leucorrhinia borealis D Argia alberta DW Leucorrhinia intacta DW Argia emma DW Libellula quadrimaculata D Argia fumipennis DW Libellula saturata D Argia vivida DW Ophiogomphus severus O Enallagma anna DW Somatochlora hudsonica D Enallagma annexum DW Somatochlora semicircularis T Enallagma boreale DW Sympetrum corruptum D Enallagma civile DW Sympetrum internum DW Aeshna eremita D Enallagma praevarum DW Aeshna juncea D Gomphus externus DO Platte (32): Aeshna palmata DW Hetaerina americana DW Amphiagrion abbreviatum DW Ischnura damula DW Argia emma DW Ischnura perparva DW Argia vivida DW Ischnura verticalis DW Coenagrion resolutum D Lestes congener DW Cordulia shurtleffii D Lestes dryas DW Enallagma anna DW Lestes forcipatus DW Enallagma annexum DW Lestes unguiculatus DW Enallagma boreale D Leucorrhinia intacta DW Enallagma carunculatum DW Libellula quadrimaculata DW Enallagma clausum DW Sympetrum corruptum DW Enallagma ebrium DW Sympetrum internum DW Enallagma hageni DW Sympetrum madidum DW Enallagma praevarum DW Sympetrum obtrusum O Aeshna constricta D Hetaerina americana DW Sympetrum pallipes DOW Aeshna palmata W Ischnura cervula DW Sympetrum rubicundulum O Amphiagrion abbreviatum DW Ischnura damula DW Sympetrum semicinctum DOW Archilestes grandis D Ischnura perparva DW Argia emma DW Ischnura verticalis DW Argia vivida DW Lestes congener DW Calopteryx aequabilis DW Lestes dryas D Enallagma anna DW Lestes forcipatus DW Enallagma annexum DW Lestes unguiculatus DW Enallagma basidens O Leucorrhinia intacta DW Enallagma boreale DW Leucorrhinia proxima D Enallagma carunculatum DW Nehalennia irene D Enallagma civile DW Ophiogomphus morrisoni W Enallagma praevarum DW

Ophiogomphus severus D Enallagma annexum DW Somatochlora hudsonica DW Plathemis lydia DW Enallagma boreale D Somatochlora minor DW Rhioaeschna californica D Lestes congener DW Somatochlora semicircularis DW Somatochlora semicircularis D Lestes dryas DW Sympetrum corruptum DW Sympetrum costiferum DW Libellula forensis O Sympetrum costiferum DW Sympetrum obtrusum DW Libellula pulchella DW Sympetrum danae DW Sympetrum pallipes DW Libellula quadrimaculata DW Sympetrum internum DW Ophiogomphus severus DW Sympetrum madidum D Sublette (22): Stylurus intricatus DW Sympetrum obtrusum DW Sympetrum corruptum DW Sympetrum pallipes DW Sympetrum danae DOW Sympetrum semicinctum O Sympetrum internum DOW Sympetrum madidum DW Sympetrum pallipes DW Uinta (13): Sympetrum rubicundulum W Sympetrum semicinctum O

Teton (42):

Aeshna eremita DO Aeshna juncea DO Aeshna palmata DW Aeshna sitchensis DO Amphiagrion abbreviatum DW Aeshna interrupta DW Coenagrion resolutum DW Amphiagrion abbreviatum DW Cordulia shurtleffii D Coenagrion resolutum DW Enallagma anna DW Enallagma anna DW Enallagma annexum DW Enallagma annexum DW Enallagma boreale DW Aeshna eremita D Enallagma clausum DW Enallagma clausum DW Aeshna interrupta DW Lestes congener DW Lestes disjunctus DW Aeshna juncea DW Lestes disjunctus DW Lestes dryas DW Aeshna palmata DW Lestes dryas DW Leucorrhinia glacialis D Aeshna sitchensis DW Libellula quadrimaculata D Ophiogomphus severus DW Aeshna umbrosa DO Sympetrum internum DW Somatochlora hudsonica D Amphiagrion abbreviatum DW Sympetrum pallipes O Somatochlora semicircularis D Archilestes grandis W Sympetrum semicinctum DW Sympetrum danae DW Argia alberta DW Sympetrum internum DW Argia vivida DW Washakie (18): Sympetrum madidum DW Calopteryx aequabilis DW Sympetrum pallipes DW Coenagrion resolutum DW Sympetrum rubicundulum D Cordulegaster dorsalis O Cordulia shurtleffii DW Sweetwater (19): Enallagma anna DOW Enallagma annexum DW Enallagma boreale DW Erythemis collocata DOW Ischnura cervula DW Lestes congener DW Lestes disjunctus DW Lestes dryas DOW Leucorrhinia borealis DW Leucorrhinia hudsonica DOW Amphiagrion abbreviatum DW Leucorrhinia proxima DW Anax junius DW Libellula forensis DW Argia emma DW Libellula quadrimaculata DW Enallagma anna DW Libellula saturata DOW Enallagma annexum DW Aeshna palmata DW Ophiogomphus severus DW Enallagma boreale DW Amphiagrion abbreviatum DOW Plathemis lydia DW Enallagma carunculatum DW Enallagma anna DW Rhionaeschna multicolor DW Enallagma ebrium DW Ischnura cervula DW

Ischnura perparva DW Weston (30): Ischnura perparva DW Leucorrhinia intacta DW Ischnura verticalis DW Libellula forensis DW Lestes congener D Libellula quadrimaculata DW Lestes dryas DW Ophiogomphus severus DW Lestes unguiculatus DW Sympetrum corruptum DW Leucorrhinia hudsonica DW Sympetrum costiferum DW Libellula forensis DW Sympetrum obtrusum DW Libellula pulchella DW Sympetrum pallipes DW Libellula quadrimaculata DW Nehalennia irene DW Ophiogomphus severus DW Rhioaeschna californica DW Rhionaeschna multicolor DW

Somatochlora ensigera D Aeshna interrupta O Sympetrum corruptum DW Aeshna palmata O Sympetrum pallipes DW Amphiagrion abbreviatum DW Sympetrum semicinctum DW Argia vivida DW Enallagma anna DW Enallagma annexum DW Enallagma boreale DW Enallagma carunculatum DW Enallagma civile DW Enallagma clausum DW Enallagma ebrium DW Hetaerina americana W Ischnura damula DW

Acknowledgements

I am deeply indebted to many people who contributed information toward this project. I am most particularly grateful to Ken Tennessen for the gift of the hard-to-find Odonata of Wyoming, which provides the base data for the study. Thomas W. “Nick” Donnelly’s groundbreaking DOT Map Project formed the foundation for much of the distribution information contained on Odonata Central, the brainchild of Dr. John Abbott at the University of Texas. This effort would have been impossible without the convenient databank found therein, encompassing both the DOT Map data plus personal collection data from odonatists throughout the country. Ron Lyons, of the Oregon Entomology Society, reviewed the final document and made many valuable suggestions and corrections. His kind assistance is gratefully welcomed. Thanks also to Drs. Lusha Tronstad and Scott Shaw, of the University of Wyoming for their kind help and guidance. Bob DuBois’s excellent guide to the Dragonflies and Ddamselflies of the Rocky Mountains has proved to be an indispensible resource, as is Dennis Paulson’s masterful Dragonflies and Damselflies of the West. Jim Johnson, of Vancouver, Washington, has exhibited kindness, patience, and congenial assistance in helping me to become more proficient in the identification of northwestern species.

Most of the data contained herein comes from the DOT Map Project, with only a small portion being contributed by individual collectors on Odonata Central. Of the 790 confirmed Odonata Central records, as of the most recent revision of this document (August 21, 2013), fully 577 (74.7%) are the result of the DOT Map Project, leaving only 213 submissions by individual collectors: Nicholas Block, Randall Cox, Randy Emmitt, James Fox, Terry Hibbits, Troy Hibbits, Harold Howell, Jim Johnson , Scott King, Greg Lasley, Kelby Ouchley, Dennis Paulson, George Sims, Susan Louise Sims, University of Texas Collection.

I am especially thankful for these folks who have been out and swinging nets in the State of Wyoming, and hope that their efforts will encourage others to add to the body of odonate knowledge from this beautiful state.

References

Abbott, J.C. 2005. Dragonflies and Damselflies of Texas and the South-Central United States. Princeton University Press.

Abbott, J.C. 2007. Odonata Central: An online resource for the distribution and identification of Odonata. Texas Natural Science Center, The University of Texas at Austin. Available at http://www.odonatacentral.org. (Last accessed: August 28, 2013).

Abbott, John C. 2011. Damselflies of Texas: A Field Guide. University of Texas Press.

DuBois, Robert 2010. Dragonflies & Damselflies of the Rocky Mountains. Kollath+Stensaas Publishing.

Lam, Ed 2004. Damselflies of the Northeast: A Guide to the Species of Eastern Canada and the Northeastern United States. Biodiversity Books.

Molinar, Donald R. and Robert J. Lavigne. 1979. Odonata of Wyoming. Wyoming Agricultural Experiment Station, Science Monograph 37.

Paulson, Dennis 2009. Dragonflies and Damselflies of the West. Princeton University Press.

Paulson, Dennis R. and Sidney W. Dunkle. 2011. A Checklist of North American Odonata: Including English Name, Etymology, Type Locality, and Distribution.