Bermuda InstItute of Ocean scIences

AnnuAl REpoRt 2010 17 Biological Station, St George’s, GE 01 Cover & this page, photos by tiffany Wardman ContEntS

Letter from the Director 4 Board of Trustees 6 Financial Highlights 8 Research Programs 12 Publications 15 Education Programs 17 Seminars, Lectures & Events 20 BIOS Faculty & Staff 22 Honor Roll of Donors 24

With roots dating back to 1903, the of Ocean Sciences, Inc., (BIOS) is a world class marine science research and education insti- tute, Bermuda's principal source of en- vironmental education and a global training centre for young scientists. photos by tiffany Wardman lEttER from the diRECtoR

he past year brought many challenges to BIOS, but EDucATIOn HIGHLIGHTS T also many accomplishments. BIOS remains strong, The coopers Island restoration work has been given an and continues to be an extraordinary global scientific and indirect boost through HSBc’s involvement. The bank educational institution, despite the difficult economic has sponsored the removal of derelict buildings and in- times. The business of BIOS is doing well – grant success vasive trees from the park, which will enable greater ac- is high, scientific publications are on the rise, and educa- cess for our student groups to continue their work tion programs are well subscribed. Our focus in 2010 was planting native trees. to increase funding to the endowment, and we have made significant strides on that front. We also focused on major Woodland restoration work continues as a means of pro- facility issues, however much of this will have to be tem- viding hands-on experience with ecological issues and the pered, factoring in economic challenges. local oceanic island curriculum. Several local classes are scheduled to assist at coopers Island nature Reserve, and We at BIOS have an unwavering commitment to edu- other informal education groups (Operation Raleigh, cating Bermudian youth about marine science and the BHS Duke of Edinburgh group) are assisting at Ferry local environment. The institution also continues to be Point Park. a training centre on an international scale. It was another outstanding year for BIOS education, with POGO and coral lectures and snorkel lessons continue in association the local education programs proving to be particularly with the Bermuda Sloop Foundation aboard the Spirit of successful. Bermuda. We contribute a day’s aquatic programming each week to assist the local middle school groups while The Waterstart season was fully subscribed again this year, they are aboard. This reaches all public middle school stu- and approximately 2000 local students attended the BIOS dents and seems both a valuable educational avenue as well Explorer program. In fact, due to its popularity, an addi- as good publicity for BIOS. tional day was added to the schedule to accommodate all requested visits. Efforts are being made to expand the pro- In connection with educators from the R/V Sea Dragon / gram to allow the participation of a greater number of stu- Pangaea Expedition, we have arranged field trips for local dents. A growing number of older teenaged students are students to investigate the problem of plastic trash in the looking for a “next step”, thus, the Bermuda Program is Sargasso Sea. also being expanded in 2010 in order to provide additional experiences for local students, doubling in size from four It was another impressive year for the POGO centre of scholarships to eight. The new extended program there- Excellence and we have been given the go-ahead for an- fore provides opportunities for students who have gradu- other year. The current POGOnians were selected from ated from high school but are still in the early stages of an applicant pool of 111 from thirty-eight countries, show- their undergraduate career. ing just how competitive and selective the program has be-

pg 4 Bermuda institute of ocean Sciences Annual Report 2010 lEttER from the diRECtoR come. The POGO program is an impressive range of stu- gator (PI) or co-PI for at least twenty-two active and sep- dents from around the globe and it is of great value to be arate federal agency awards and several other founda- educating young scientists about preserving the oceans tion/non-federal agency awards. This exceeds the hopes and marine life. for peer-review publication and grantsmanship outlined in the strategic plan a few years ago. Although BIOS has a 2010 also proved to be another strong year of student en- small number of faculty compared to other institutions, rollment for the fall semester courses offered at BIOS. The their per-capita contribution to the scientific and broader Marine Invertebrate Zoology and coral Reef Ecology community is impressive. classes were filled with enthusiastic students. The sum- mer internship program continues to be very popular and Finally, BIOS would like to welcome three new trustees: we receive hundreds of inquiries each year from potential neill currie, cEO of Renaissance Re, Brad Kopp the cEO interns. We now have two BIOS graduate students, one in of and Michael McGavick, cEO of XL the university of Southampton-BIOS collaborative PhD capital. It cannot go without saying that we were also sad- Program, the other in the Princeton-BIOS joint PhD Pro- dened by the untimely death of Trustee Matt Simmons gram. Both students are mentored by Dr. Mike Lomas and who was a great supporter of BIOS. co-advised by Dr. Duncan Purdie (university of Southampton, national Oceanography centre) and Drs. Our sincere gratitude for all of your support, and your Danny Sigman and Bess Ward (Princeton university). continued commitment to BIOS as we look to build our endowment and annual funds, and design a strategic plan RESEARcH HIGHLIGHTS that will make BIOS even more prominent and well-re- More than forty peer-review papers were published in spected as a global marine science institution. 2010, with several high-profile publications in Nature and Science and a very long list of co-authors at other institu- tions demonstrating the active network BIOS faculty have established.

BIOS is proud to say we received an $850,000, nSF grant for BEAcOn (BErmuda ocean Acidification and cOral reef investigation). BIOS faculty were Principal Investi-

Anthony H. Knap nicholas R. Bates Director Interim Director Bermuda Institute of Ocean Sciences Bermuda Institute of Ocean Sciences

pg 5 Bermuda institute of ocean Sciences Annual Report 2010 BoARd of tRuStEES

OFFIcers trustees Canada Chairman United States Michael A. Butt Brian Duperreault Kenneth H. Brink, PhD Buttcon Limited Woods Hole Oceanographic Institution Concord, Ontario Honorary Woods Hole, MA His Excellency the Governor Diane Francis Sir Richard Gozney, KCMG Thomas M. Church, PhD Toronto, Ontario University of Delaware Vice Chairmen Karl Wildi IdwalWyn Hughes, PhD Rita R. Colwell, PhD Toronto, Ontario Ian Strecker University of Maryland William Williams John W. Farrington, PhD Europe President and Executive Director Woods Hole Oceanographic Institution Lady (Judy) Vereker Anthony H Knap, PhD Woods Hole, MA London

Treasurer and Controller Diana H. Frazier Stephen Markman FLAG Capital Management, LLC Bermuda Stamford, CT Philip V. Bancroft Secretary ACE Limited Gillian M.S. Hollis James N. Galloway, PhD University of Virginia Philip Butterfield Charlottesville, VA HSBC Bank of Bermuda Limited

Linda K. Glover Neill Currie GloverWorks Consulting Renaissance Re Holdings Ltd Alexandria, VA Brian Duperreault Rodney H. Glover Marsh & McLennan Wiley Rein & Fielding LLP Washington, DC Richard J. Ferrett Tucker’s Town Mae C. Jemison, PhD Houston, Texas Allan W. B. Gray Orbis Investment Management Ltd Charles F. Kennel Scripps Institution of Oceanography Michael D. Hamer, PhD La Jolla, CA Smith’s

Alan McGowan Idwal Wyn Hughes, PhD New School University Hamilton Parish New York, NY Rebecca Knox-Byrne H. Rodger Melton, PhD Tucker’s Town Exxon Mobil Corporation Houston, TX Bradford Kopp Butterfield Bank William Russel, PhD Princeton University Michael S. McGavick XL Group plc Deborah K. Steinberg, PhD Virginia Institute of Marine Science Christopher Maybury Gloucester Point, VA St George’s

Anthony Williams Raymond E. Moore DLA Piper LLP Pembroke New York, NY

pg 6 Bermuda institute of ocean Sciences Annual Report 2010 BoARd of tRuStEES

Bermuda (continued) trustees emerItus Michael A. Butt, OBE Brian M. O’Hara AXIS Capital Holdings Limited Paget Robert E. Cawthorn Jonathan F. Reiss Warwick Ernst & Young Margot B. Perot Ralph E. Richardson Dallas,TX The ACE Foundation LIFe trustees The Honorable Gerald D.E. Simons, OBE United States The Argus Group Neil R. Andersen, PhD Foster Bam James N Stanard, PhD James N. Butler, PhD Paget Rosamond H. Butler Clayton P. Cormier Ian Strecker John A. Knauss, PhD Warwick Frederick J. Kollmorgen Cindy Lee, PhD The Honorable Sir John W. Swan, KBE, JP H. Eugene Lehman, PhD The Swan Group Frederick T. Mackenzie, PhD Frank B. Sprow, PhD Edward S. Trippe John H. Steele, PhD Harrington Sound Talbot H. Waterman, PhD W. Redwood Wright, PhD George A. Wardman Coral Beach and Tennis Club Bermuda James Burnett-Herkes, PhD William H. Williams John R.H. Lightbourn STW Fixed Income Management Ltd Terry E. Lister, JP,MP Wendell M. Smith Richard D. Spurling Brunell Spurling Ernest E. Stempel David B. Wingate, PhD,OBE

Canada Michael L. Davies

Europe F. Geoffrey Larminie Robert T. Sperring

annuaL Fund cOmmIttee We would like to thank the Annual Fund Committee for their hard work and dedication in 2010:

Committee Chair: Jonathan F. Reiss Jeffery Conyers Ann Durhager photos pg 6 by Jp Skinner, Maureen Elizabeth A.C. Durrant Conte and Rachel Evans. this pg by Gerald Simons Charlotte Best (middle) and tiffany David Skinner Wardman Amanda Swan

pg 7 Bermuda institute of ocean Sciences Annual Report 2010 December 31, 2010 anD 2009

ooking back on 2010, we realize that fundamentally BIOS is very strong. Science and edu- L cation programs continue to be our mainstay during these unpredictable economic times. Revenues derived from Program Services [scientific research and education] amounted to $9,361,000 in 2010, up $13,000 from 2009 which results are remarkable considering total revenues were down $337,000 in the same period and the challenges that BIOS faced.

This increase came in spite of the disruptions experienced on campus resulting from the forced ren- ovation of the nautilus (previously Blue Building) staff apartment complex which disruption caused significant revenue loss from educational courses and group business. Limited space yielded limitations on program enrollments. Additionally, staff were housed off-campus for a while send- ing those rents to others.

Total assets increased $500,000 from 2009 to 2010 as property and equipment increased $573,000. Most of the increase in Property and equipment resulted from upgrades to the accommodations. Investments continue to do well increasing by a net of $901,000. Receivables decreased as exist- ing $726,000 pledges were paid. Those pledge donations designated for the naess Laboratory Building, built in 2004, were used to fully repay the construction debt on that building. Plus the annual pledge payment for the ship purchase loan repaid a portion of that loan. In all $1,540,000 of long-term debt was repaid.

Development expenses were down as the new Development Officer started on September 1 filling a vacancy created at the end of the prior year. Management & General expenses increased as we embarked upon an aggressive maintenance program to restore and upgrade our facilities. new fi- nancing was in progress as of year-end to provide much of the needed funding for this effort. As has been the trend at BIOS, 88% of our expenses are for program related activities.

This report and the financial schedules herein are limited selected financial highlights and do not purport to include all information and disclosures required in accordance with accounting prin- ciples generally accepted in the u.S. A complete copy of the Financial Statements with the Inde- pendent Auditors’ Report can be obtained by following the instructions below.

Stephen M. Markman Treasurer and controller photos by Maureen Conte, Alex Hunter and tiffany Wardman Financial statements with independent auditors’ report are available on our website at www.bios.edu or by contacting Stephen M. Markman, Treasurer and Controller, telephone 441-297-1880 ext.109, fax 441-297-2222 or email [email protected].

pg 8 Bermuda institute of ocean Sciences Annual Report 2010 DECEMBER 31, 2010 AND 2009

2010 reVenues & suPPOrt Revenue and support is derived from gifts, grants and contracts received through the u.S. (60%) and Bermuda (4%) governments as well as various individual, corporate and foundation donors (22%). Additional sources of support are tuition and fees for the use of BIOS’s various scientific, marine and housing facilities and attendance at our many educational programs (14%). Investment return pertains to Endowment Funds and is not included in these percentages.

US Government and Corporate Contracts and Grants $7,174,806 (60%)

Bermuda Government $532,238 (4%)

Contributions $2,634,984 (22%)

Tuition, Guest Services and Other Income $1,654,082 (14%)

2010 eXPenses Program expenses include scientific research (69%); education activities (17%) and guest and residential services (2%). Other expenses include development, marketing and public relations (5%) and management and general (7%).

Scientific Research $8,650,314 (69%)

Education $2,097,436 (17%)

Management and General $880,384 (2%)

Development and Public Relations $619,402 (5%)

Guest and Residential Services $275,959 (7%)

pg 9 Bermuda institute of ocean Sciences Annual Report 2010 DECEMBER 31, 2010 AND 2009

SuMMARy FinAnCiAl poSition

assets 2010 2009 Cash and cash equivalents $ 1,850,158 $ 2,099,077 Grant receivables and other assets 1,098,963 1,385,413 Contributions receivable, net 1,058,730 1,498,084 Investments 10,526,597 9,625,359 Property and equipment, net 16,664,049 16,090,874 Total Assets 31,198,497 30,698,807 Liabilities and net assets Liabilities Payables, accruals, advances and deposits $ 2,383,506 $ 1,808,957 Loans payable 5,449,153 6,171,741 Total Liabilities 7,832,659 7,980,698

net assets Unrestricted 11,888,973 11,997,281 Temporarily restricted 3,857,000 3,102,213 Permanently restricted 7,619,865 7,618,615 Total Net Assets 23,365,838 22,718,109 Total Liabilities and Net Assets $ 31,198,497 $30,698,807

SuMMARy ACtivitiES StAtEMEnt

revenues & support Contributions, net $ 2,634,984 $ 2,235,635 Grants and Contracts 7,707,044 7,419,779 Tuition, guest services and other 1,654,082 1,927,967 Investment return 1,175,114 1,924,900 Total Revenue & Support 13,171,224 13,508,281

expenses Program services Scientific research 8,650,314 8,920,346 Education programs 2,097,436 2,214,989 Guest and residential services 275,959 256,956

Support Services Development and public relations 619,402 863,825 Management and general 880,384 619,074 Total expenses 12,523,495 12,875,190 Increase (decrease) in net assets $ 647,729 $ 633,091

Financial statements with independent auditors’ report are available on our website at www.bios.edu or by contacting Stephen M. Markman, Treasurer and Controller, telephone 441-297-1880 ext.109, fax 441-297-2222 or email [email protected].

pg 10 Bermuda institute of ocean Sciences Annual Report 2010 notES to SuMMARy FinAnCiAl HiGHliGHtS DECEMBER 31, 2010 AND 2009

Bermuda Institute of Ocean Sciences (BIOS), Inc., founded in 1903 as the Bermuda Biological Station and incorpo- rated in the State of new York in 1926, is an independent u.S. not-for-profit organization, and as such, is exempt from u.S. income taxes under Sections 501(c)(3) and 509(a) of the Internal Revenue code. BIOS is also a Bermuda registered charity. Our uS income tax ID is 06-0706038.

NOTE 1 INVESTMENTS AND INVESTMENT RETURN Investments principally consist of funds managed by Common fund, The Investment Fund for Foundations’ (TIFF) Investments Program (TIP) and The Vanguard Group, Inc; and, certain donated equity securities, sale of which is restricted. Investments relate to Endowments. The return on these investments is appropriated for expenditure in accordance with the Board of Trustees policy of permitting endowment support spending up to 5% (in accordance with the provisions of the laws of the State of New York) of the endowment portfolio average market value at June 30 of each of the three previous fiscal years.

NOTE 2 ENDOWMENT FUNDS Donor-restricted funds whose purpose is to provide long-term support for BIOS are Endowment Funds. Following is a recapitulation of the aggregate activity in Endowment Funds for the years ended December 31:

2010 2009 Balance at January 1 $ 9,754,417 $ 8,156,125 Contributions 1,250 1,100 Investment income 244,303 264,519 Capital appreciation (depreciation) 925,226 1,656,694 Appropriated for expenditure (404,021) (324,021) Balance at December 31 $ 10,521,175 $ 9,754,417

NOTE 3 PROPERTY AND EQUIPMENT Depreciation expense amounted to $1,114,000 in 2010 and $1,049,000 in 2009. Acquisitions of fixed assets amounted to $1,686,736 and $499,000. Property and equipment, at undepreciated cost, is $26,974,000 at December 31, 2010.

NOTE 4 LOANS PAYABLE BIOS repaid and retired two bank loans in 2010 and entered into another loan for purposes of performing major maintenance and upgrades on existing properties. In addition to the major maintenance loan, the remaining loans are for the purchase and retrofitting of the R/V HSBC Atlantic Explorer, and construction of the staff apartment building. Interest rates on the bank loans float based on the three-month LIBOR rate plus up to 1%. The bank loans are generally payable on demand by the bank. Interest expense was $34,000 and $135,000 for the years ended December 31, 2010 and 2009.

NOTE 5 NET ASSETS BIOS reports its financial position and activities according to three classes of net assets: permanently restricted, temporarily restricted, and unrestricted. Permanently Restricted Net Assets comprise gifts and bequests that create or increase endowment funds. Temporarily Restricted Net Assets are those gifts which are received with stipulations that the gift be used for a particular purpose or during a specified future time. All other net assets and those for which restrictions have been met are classified as unrestricted.

pg 11 Bermuda institute of ocean Sciences Annual Report 2010 RESEARCH pRoGRAMS abundance, dynamics and diversity collaborative research: Dr. D.A. Hansell, Rosenstiel School of of Viruses in the northwestern Bacterioplankton Biology and Marine and Atmospheric Science sargasso sea Biogeochemistry at the Bermuda Funding: NSF Dr. M. Breitbart, University of South Florida atlantic time-series study station: R. Parsons, BIOS an Oceanic microbial Observatory collaborative research: development Funding: NSF Dr. C. Carlson, University of California, of the automated dissolved Inorganic Santa Barbara carbon analyzer (amIca) for seawa- air Quality Program Dr. S. Giovannoni, Oregon State University ter dIc analyses Drs. A.J. Peters and A.H. Knap, BIOS R. Parsons, BIOS Dr. N.R. Bates, BIOS Funding: The Bermuda Government Funding: NSF Prof. J.H. Sharp, University of Delaware Dr. G.E. Friederich, Monterey Bay Aquarium air-sea pcO2 Observations in the collaborative research: Best: the Research Institute north atlantic Impact of changes in sea Ice extent Funding: NSF and OTIC Dr. N.R. Bates, BIOS on Primary Production, Dr. C.L. Sabine, NOAA-Pacific Marine Phytoplankton community structure, collaborative research: dimensions in Environmental Laboratory and export in the eastern Bering sea Biodiversity: Biological controls on Funding: NOAA Dr. M.W. Lomas, BIOS the Ocean c:n:P ratios Dr. S.B. Moran, University of Rhode Island, Dr. M.W. Lomas, BIOS algal Biodiesel Graduate School of Oceanography Drs. A. Martiny and S. Allison, University of Drs. F.G. Plumley, M.W. Lomas and Funding: NSF California, Irvine A.H. Knap, BIOS Dr. S. Levin, Princeton University Funding: PartnerRe collaborative research: carbon Funding: NSF through the twilight Zone: application of the Leaf Wax-aerosol new tools to measure change collaborative research: the Great method to assess spatial and Dr. M.W. Lomas, BIOS southern coccolithophore Belt temporal Patterns of carbon Isotopic Drs. K.O. Buesseler and C.H. Lamborg, Drs. W.M. Balch and B. Twining, Bigelow Fractionation of atmospheric cO2 by Woods Hole Oceanographic Institute Laboratory for Ocean Sciences terrestrial Photosynthesis Dr. D. Siegel, University of California, Santa Dr. P. Lam, Woods Hole Oceanographic Dr. M.H. Conte, BIOS Barbara Institute Funding: NOAA, Global Carbon Cycle Dr. D. Steinberg, Virginia Institute of Marine Dr. N.R. Bates, BIOS Program Sciences Funding: NSF Funding: NSF BeacOn: Bermuda collaborative research: Plankton Ocean acidification and collaborative research: combining community composition and trophic coral reef Investigation Flow cytometry and stable Isotope Interactions as modifiers of carbon Drs. A.J. Andersson, N.R. Bates and S. de techniques: a method to measure export in the sargasso sea. Putron, BIOS Phytoplankton- and Bacteria-specific Dr. M.W. Lomas, BIOS Funding: NSF, Chemical oceanography nitrogen and carbon uptake Dr. T. Richardson, University South Dr. M.W. Lomas, BIOS Carolina the Bermuda atlantic time-series Dr. D. Bronk, Virginia Institute of Marine Dr. S. Neuer, Arizona State University study, Years 21–25 Sciences Dr. R. Condon, Dauphin Island Sea Lab Drs. A.H. Knap, N.R. Bates, M.W. Lomas and Funding: NSF Funding: NSF R.J. Johnson, BIOS Funding: NSF collaborative research: composition collaborative research: of the Plankton community and its Prochlorococcus and its contribution the Bermuda Bio-Optics Project contribution to Particle Flux in the to new Production in the sargasso (BBOP) sargasso sea sea Drs. N. Nelson and D.A. Siegel, University Dr. M.W. Lomas, BIOS Dr. M.W. Lomas, BIOS of California, Santa Barbara Dr. S. Neuer, Arizona State University Dr. A.C. Martiny, University of California, Dr. M.W. Lomas, BIOS Funding: NSF Irvine Funding: NASA Funding: NSF collaborative research: determining Bermuda reef Fish die Off study the Present and Future Ocean collaborative research: sources and R. Parsons and T. Noyes, BIOS carbon dynamics in the chukchi sea Biogeochemical cycling of Intact Dr. W. Vogelbein, The College of and Pan-arctic Ocean: a contribution membrane Lipids in the upper William and Mary, Virginia Institute of to shelf-Basin Interactions (sBI) Ocean. Marine Science Phase III Dr. M.W. Lomas, BIOS Funding: The Bermuda Government Dr. N.R. Bates, BIOS Dr. B. van Mooy, Woods Hole Dr. S.B. Moran, University of Rhode Island Oceanographic Institute Dr. J.T. Mathis, University of Alaska, Funding: NSF Fairbanks pg 12 Bermuda institute of ocean Sciences Annual Report 2010 RESEARCH pRoGRAMS

collection of sediment cores from the effect of Ocean acidification and research experiences in Open Ocean Lac toponowini (French Guiana) to Increased temperature on the and subtropical marine environments reconstruct Holocene ecosystem and calcification of new coral recruits of Bermuda climate Variability of the Drs. A. Cohen, D. McCorkle, N. Cantin, Dr. F.G. Plumley, BIOS northeastern amazon G. Gaetani and K. Rose, Woods Hole Funding: NSF-Research Experience for Dr. M.H. Conte, BIOS Oceanographic Institute Undergraduates Funding: National Geographic Society, H. Barkley, Princeton University Committee for Research and Exploration Funding: Vita Foundation, Woods Hole sGer: exploratory analyses of Oceanographic Institute and BIOS sediment cores from Lac toponowini a comprehensive comparative (French Guiana) Genomic approach for Identifying the evaluating carbon dioxide (cO2) Dr. M.H. Conte, BIOS saxitoxin synthesis Genes sinks and sources and Impacts of Funding: NSF, Paleoclimate Dr. F.G. Plumley, BIOS Ocean acidification of Key ecosystems Dr. J. Hackett, University of Arizona Dr. N.R. Bates, BIOS sea urchins as models for aging and Dr. D. Bhattacharya, Rutgers University Funding: Exxon-Mobil cancer research Dr. D. Anderson, Woods Hole Drs. A. Bodnar and J. Loram, BIOS Oceanographic Institute Interannual Variability of the carbon Funding: Hannover Life Re, Bermuda Ltd., Funding: NSF cycle in the north atlantic Ocean Molecular Biology Endowment and Ray Drs. N.R. Bates and A.H. Knap, BIOS Moore Endowment, CABIOS

controls on and rates of air-sea cO2 Funding: Exxon Mobil Corporation Gas exchange in the Beaufort and shipboard scientific support chukchi sea: Investigations of climate marine environmental Programme equipment for the R/V and environmental change on arctic Drs. R. Jones and A.H. Knap, BIOS Atlantic Explorer Pacific shelves. Funding: The Bermuda Government Dr. N.R. Bates and Capt. R. Harelstad, BIOS Dr. N.R. Bates, (BIOS), Funding: NSF Dr. J.T. Mathis, University of Alaska, Fairbanks mg-calcite mineral dynamics in natu- Funding: NASA ral subtropical seawater systems: ship Operations, r/V atlantic explorer control of the carbonate acid system Dr. N.R. Bates, Capt R. Harelstad, BIOS copper-binding Organic Ligands in and relevance to Ocean acidification. Funding: NSF the san Francisco Bay estuary: Drs. A.J. Andersson and N.R. Bates, BIOS evaluating current and Future Funding: NOAA source tracking of sewage Likelihood of copper toxicity events contamination in Bermuda’s Inshore in a Perturbed ecosystem. Ocean carbon measurements in Waters Dr. K. Buck, BIOS support of the Joint russian-american R. Parsons, BIOS Dr. K. Barbeau, Scripps Institution of Long-term census of the arctic Funding: The Government of Bermuda Oceanography (rusaLca) research Program in the Dr. Robin Stuart, USGS Bering and chukchi seas time-series Particle Flux Funding: California Sea Grant/ CALFED Dr. N.R. Bates, BIOS measurements in the sargasso sea Bay-Delta Science Fellows Program Funding: NOAA Drs. M.H. Conte and K. Buck, BIOS Funding: NSF, Chemical Oceanography coral reproduction and Oceanographic Instrumentation for recruitment studies the r/V atlantic explorer us GeOtraces north atlantic Dr. S. de Putron, BIOS Dr. N.R. Bates and Capt. R. Harelstad, BIOS section: the chemical speciation of Dr. G. Goodbody-Gringley, Funding: NSF dissolved iron and copper. Harvard University Dr. K. Buck Prof. J. Ryland, Oceanographic technical services Funding: NSF of Wales Dr. N.R. Bates and Capt. R. Harelstad, BIOS J. Lawson, Dalhousie University Funding: NSF the us network of VOs surface

M Costa, Princeton University pcO2 sampling Funding: BIOS, Harvard Operation of a marine atmospheric Dr. N.R. Bates, BIOS sampling Facility at tudor Hill, Dr. R. Wanninkhof, NOAA-Atlantic cruise ships and air Quality at Bermuda Oceanography and Meteorology dockyard Drs. A.J. Peters adn A.H. Knap, BIOS Laboratory Dr. A.J. Peters, BIOS Funding: NSF Dr. R.A. Feely, NOAA-Pacific Marine Funding: The Bermuda Government Environmental Laboratory POGO centre of excellence in Prof. T. Takahashi, Lamont Doherty Earth Observational Oceanography Observatory Dr. F.G. Plumley, BIOS Prof. F.J. Millero, Rosenstiel School of Marine Funding: Nippon Foundation and Atmospheric Science Funding: NOAA

pg 13 Bermuda institute of ocean Sciences Annual Report 2010 RESEARCH pRoGRAMS

Graduate researcH

molecular approach to determine contributions of the Protist community to carbon Flux Jessica Mohler, PhD Candidate, Arizona State University FACULTY ADVISORS: Dr. S. Neuer, Arizona State University Dr. M.W. Lomas, BIOS Funding: NSF

Bacterial nitrogen cycling in the mesopelagic Steven Baer, PhD Candidate, College of William and Mary FACULTY ADVISORS: Dr. D. Bronk,Virginia Institute of Marine Science Dr. M.W. Lomas, BIOS

nutrient cycling in Harmful algal Blooms George Boniello, PhD Candidate, Old Dominion University FACULTY ADVISORS: Dr. M. Mulholland, Old Dominion University Dr. M.W. Lomas, BIOS

microphytoplankton of the sargasso sea: Variability and chemical composition Charlotte Best, University of Southampton FACULTY ADVISORS: Dr. D. Purdie, National Oceanography Centre, Southampton Dr. M.W. Lomas, BIOS

Population-specific nitrogen cycling in the sargasso sea Sarah Fawcett, Princeton University FACULTY ADVISORS: Drs. B. Ward and D Sigman, Princeton University Dr. M.W. Lomas, BIOS

Biogeochemistry of the Bering sea shelf Matthew Baumann, University of Rhode Island FACULTY ADVISORS: Dr. S.B. Moran, University of Rhode Island Dr. M.W. Lomas, BIOS photos by Rachel Evans (top) and tiffany Wardman

pg 14 Bermuda institute of ocean Sciences Annual Report 2010 puBliCAtionS Bates, n.r., Amat, A., and andersson, Jones, R., Parsons, r., Watkinson, E. and Mathis, J.T., Cross, J., Bates, n.r., Moran, a.J., 2010. The interaction of carbonate Kendall, D., 2010. Sewage contamination S.B., Lomas, m.W., and Stabeno, P.J., chemistry and coral reef calcification: of a densely populated coral ‘atoll’ 2010. Seasonal distribution of dissolved the carbonate chemistry coral reef (Bermuda) Environ Monit Assess. Online inorganic carbon and net community ecosystem feedback (CREF) hypothesis. 1-16 production on the Bering Sea shelf. Bio- Biogeosciences, 7 (5), 2509-2530. geosciences, 7(5), 1769-1787, doi: doi:10.5194/bg-7-2509-2010. Corrigen- Jutterström, S., Anderson, L.G., Bates, 10.5194/bg-7-1769-2010. dum: Biogeosciences, 7 (10), 3005-3006, n.r., Bellerby, R., Johannessen, T., Jones, doi: 10.5194/bg-7-3005-2010. E.P., Key, R.M., Lin, X., Olsen, A., and Oh, H.-M., Kang, I., Ferriera, S., Giovan- Omar, A.M., 2010. Arctic Ocean data in noni, s.J., and Cho, J.-C., 2010. Com- Beszteri, B., Temperton, B., Frickenhaus, CARINA. Earth System Science Data, 2, plete genome sequence of Croceibacter S., and Giovannoni, s. J., 2010. Average 71-78, Feb. 10, 2010. atlanticus HTCC2559T. J. Bacteriol. genome size: a potential source of bias 192(18): 4796-4797. in comparative metagenomics. ISME J. Kang, I., Oh, H.-M., Lim, S.-I., Ferriera, S., 4:1075-7.doi:10.1038/ismej.2010.29 Giovannoni, s. J., and Cho, J.-C., 2010. Orchard, E.D., Ammerman, J.W., Lomas, Genome sequence of Fulvimarina pelagi m.W., and Dyhrman, S.T., 2010. Dis- Bradley, P. B., Lomas, m.W., and Bronk, HTCC2506T, a Mn(II)-oxidizing al- solved inorganic and organic phosphorus D.A., 2010. Phytoplankton and bacterial phaproteobacterium possessing an aero- uptake in Trichodesmium and the micro- nitrogen use in Chesapeake Bay meas- bic anoxygenic photosynthetic gene bial community: The importance of phos- ured using a flow cytometric sorting ap- cluster and xanthorhodopsin. J. Bacteriol. phorus ester in the Sargasso Sea. proach. Estuaries and Coasts, DOI 192(18): 4798-4799. Limnology and Oceanography, 55:1390- 10.1007/s12237-009-9252-y. 1399. Krause, J.W., Nelson, D.M., and Lomas, Buck, K.n., Selph, K., and Barbeau, K., m.W., 2010. Production, dissolution and Orchard, E.D., Benitez-Nelson, C.R., Pel- 2010. Iron-binding ligand production and potential export of biogenic silica in a lechia, P.J., Lomas, m.W., and Dyhrman, copper speciation in an incubation ex- Sargasso Sea mode-water eddy. Limnol- S.T., 2010. Polyphosphate in Tri- periment of Antarctic Peninsula shelf wa- ogy and Oceanography, 55: 569-579. chodesmium from the low phosphorus ters from the Bransfield Strait, Southern Sargasso Sea. Limnology & Oceanography, Ocean. Marine Chemistry. Lomas, m.W., Burke, A., Lomas, d.a., 55(5), 2161-2169. doi:10.1016/j.physletb.2003.10.071 Bell, D.W., Shen, C., Ammerman, J.W., and Dyhrman, S.T., 2010. Sargasso Sea phos- Pause-Tucker, K., Parsons, r., Symonds, condon, r.H., Steinberg, D.K., and phorus biogeochemistry: An important E.M. and Breitbart, M., 2010. Diversity Bronk, D.A., 2010. Production of dis- role for dissolved organic phosphorus and distribution of single-stranded DNA solved organic matter and inorganic nu- (DOP). Biogeosciences 7: 695-710. phages in the North Atlantic Ocean. The trients by ISME Journal. 2010, 1-9 gelatinous zooplankton in the York River Lomas, m.W., Steinberg, D.K., Dickey, estuary, USA. Journal of Plankton Research. T.D., carlson, c.a., Nelson, N.B., con- don, r.H., and Bates, n.r., 2010. In- Dupont, C.L., Buck, K.n., Palenik, B. and creased ocean carbon export in the Barbeau, K., 2010. Contrasting scenarios Sargasso Sea is countered by its en- of Ni uptake and limitation in microbial hanced mesopelagic attenuation. Biogeo- assemblages from different biogeochemi- sciences, 7 (1), 57-70. cal regimes. Deep Sea Research. I. 57: 553- 556. Longnecker, K., Lomas, m.W., and Van Mooy, B.A.S., 2010. Characterizing the Helmke, P., Lomas, m.W., conte, m., abundance and diversity of het- Köster, J., and Neuer, S., 2010. What de- erotrophic bacterial cells assimilating termines the variability of organic car- phosphate in the sub-tropical North At- bon export and flux attenuation in the lantic Ocean. Environmental Microbiology, subtropical North Atlantic gyre? Deep doi:10.1111/j.1462-2920.2010.02247.x. Sea Research I, 57: 213-227.

pg 15 Bermuda institute of ocean Sciences Annual Report 2010 puBliCAtionS

Saba, V.S., Friedrichs, M.A.M., Carr, M.-E., Thrash, J. C., Stingl, U., Cho, J.-C., Ferri- Antoine, D., Armstrong, R.A., Asanuma, era, S., Johnson, J., Vergin, K. L., and I., Aumont, O., Bates, n.r., Behrenfeld, Giovannoni, s. J., 2010. Genome se- M.J., Bennington, V., Bopp, L., Bruggeman, quence of the novel marine Gammapro- J., Buitenhuis, E.T., Church, M.J., Ciotti, teobacterium strain HTCC5015. J. Bac. A.M., Doney, S.C., Dowell, M., Dunne, J., 192(14): 3838-3839. Dutkiewicz, S., Gregg, W., Hoepffner, N., Hyde, K.J.W., Ishizaka, J., Kameda, T., Thrash, J. C., Cho, J.-C., Vergin, K. L., and Karl, D.M., Lima, I., Lomas, m.W., Giovannoni, s. J., 2010. The genome Marra, J., McKinley, G.A., Mélin, F., sequences of Oceanicola granulosus Moore, J.K., Morel, A., O’Reilly, J., Sali- HTCC2516T and O. batsensis hoglu, B., Scardi, M.J., Smyth, T.J., Tang, S., HTCC2597T. J. Bacteriol. 192 (13): Tjiputra, J., Uitz, J., Vichi, M., Waters, K., 3549-50. doi:10.1128/JB.00412-10. Westberry, T.K., and Yool, A., 2010. The challenges of modeling depth-integrated ussher, s.J., Achterberg, E.P., Sarthou, marine primary productivity over multi- G., Laan, P., de Baar, H.W.J., and Wors- ple decades: A case study at BATS and fold, P.J., 2010. Distribution of size frac- HOT. Global Biogeochemical Cycles, 24, tionated dissolved iron in the Canary GB3020, doi:10.1029/2009GB003655, Basin. Marine Environmental Research, Sep 15, 2010. (2010) 70(1), 46-55 doi:10.1016/j.marenvres.2010.03.001 Shoeib, M., Vlahos, P., Harner, T., Peters, a., Graustein, M., and Narayan, J., 2010 ussher, s.J., Manning, A.J., Tappin, A.D., Survey of polyfluorinated chemicals and Fitzsimons, M.F., 2010. Hydrobiologia (PFCs) in the atmosphere over the (2010). Observed dissolved and particu- northeast Atlantic Ocean - Bermuda, late nitrogen concentrations in a mini Sable Island and coastal USA. Atmos- flume. doi:10.1007/s10750-011-0759-4 pheric Environment 44:2887-2893 ussher, s.J., Petrov, I., Quétel, C., and Sigler, M.F., Harvey, H.R., Ashjian, C.J., Worsfold, P., 2010. Validation of a Lomas, m.W., Napp, J.M., Stabeno, P.J., portable flow injection-chemilumines- and Van Pelt, T.I., 2010. How does cli- cence (FI-CL) method for the determi- mate change affect the Bering Sea nation of dissolved iron in Atlantic open ecosystem. EOS 91:457-458. ocean and shelf waters by comparison with isotope dilution - inductively cou- Stewart, G., Moran, S.B., and Lomas, pled plasma mass spectrometry (ID- m.W., 2010. Seasonal POC fluxes at ICPMS) (2010) Environmental BATS estimated from 210Po deficits. Chemistry 7(2), 139 doi Deep Sea Research I, 57:113-124. 10.1071/EN09092

Thrash, J. C., Cho, J.-C., Ferriera, S., Zielke, S. and Bodnar, a., 2010. Telom- Johnson, J., Vergin, K. L., and Giovan- eres and telomerase activity in sclerac- noni, s. J., 2010. Genome sequences tinian corals and Symbiodinium spp. of Pelagibaca bermudensis HTCC2601T Biological Bulletin 218, 113-121. and Maritimibacter alkaliphilus HTCC2654T, the type strains of two marine Roseobacter genera. J. Bac. 192(20): 5552-5553.

photos by Charlotte Best, Rachel Evans and tiffany Wardman

pg 16 Bermuda institute of ocean Sciences Annual Report 2010 EduCAtion pRoGRAMS summer cOurse PrOGram C. Best, Southampton University Year 3 POGO Scholars Coral Reef Ecology S. Fawcett, Princeton University Lazare Kouame Akpetou, Center for Dr. S. de Putron, BIOS Dr. A. Peters, BIOS Oceanographic Research of Abidjan, Cote d'Ivoire Marine Biology Courses ADVISOR: Dr. K. Buck Dr. S. de Putron, BIOS General Oceanography, Core Skills Gayatri Dudeja, Indian National Centre for Dr. J. Gould, Princeton University Drs. F.G. Plumley & Faculty, BIOS Ocean Information Services, Hyderabad, India Microbial Oceanography R/V Atlantic Explorer Cruise to ADVISOR: Dr. T. Platt, Dalhousie University Dr. C. Carlson, University of California, Hydrostation S & BATS Maria Fernanda Coló Giannini, Universidade Santa Barbara Federal do Rio Grande, Brazil R. Parsons, BIOS Chemical Oceanography ADVISOR: Dr. M.W. Lomas Dr. K. Buck, BIOS Fabricio Luis Guaman-Guevara, University of Concepcion, Ecuador FaLL semester PrOGram Use of Satellites, Ocean Color, Pigments, and ADVISOR: Dr. A. Andersson Coral Reef Ecology Models of Phytoplankton Physiological Evgeniya Klimchuk, Moscow State University Dr. S. de Putron, BIOS Ecology in Long-term Ocean Monitoring of Lomonosov, Russian Federation K. Baird, BIOS Drs. T. Platt and S Sathyendranath, ADVISOR: Dr. R. Johnson Dalhousie University and Plymouth Marine Yosra Khammeri, National Institute Marine Biology and Oceanographic Research Laboratory Agronomy, Tunisia Dr. S. de Putron, BIOS Dr. H. Bouman, Oxford University ADVISOR: Dr. S. Ussher C Newcomb-Hodgetts and K. Baird, Drs. G. White & L. Zhai, Bedford Institute of Maziar Khosravi, I.R. of Iran Meteorological BIOS Oceanography Org, Iran Dr. A. Mead, University of Cape Town ADVISOR: Dr. R. Johnson Physical Oceanography Rafael Rasse Boada, Instituto Venezolano de Marine Invertebrate Zoology Drs. M. Andres and E Shroyer, Investigaciones Cientificas, Venezuela Dr. A. Mead, University of Cape Town Woods Hole Oceanographic Institute ADVISOR: Dr. M.W. Lomas C. Newcomb-Hodgetts, BIOS Muhammed Fatih Sert, Middle East Technical Biological Oceanography University/Institute of Marine Sciences, Research Diving Methods Dr. M.W. Lomas, BIOS Turkey A. Chequer, BIOS C. Best, University of Southampton ADVISOR: Dr. M.W. Lomas S. Fawcett, Princeton University Pham Thi Phuong Thao, Institute of POGO centre OF Oceanography, Vietnam eXceLLence In OBserVatIOnaL Moored Observatories ADVISOR: Dr. N.R. Bates OceanOGraPHY Dr. M. Conte, BIOS eLderHOsteL/rOad scHOLar Faculty Ocean Observatories: Ecological, Physical, Real Bermuda: History and Heritage Dr. A Andersson, BIOS and Climate Modeling: (Powersim) Snorkeling the Coral Island Dr. T. Anderson, University of Dr. T. Anderson, Southampton University Science in Southampton Birding Bermuda Dr. M. Brown, Intergovernmental Organic Pollutants in the Marine Oceanographic Commission Environment and Marine Policy Graduate Interns Dr. K. Buck, BIOS Dr. A. Peters, BIOS Lilian Krug, BIOS-POGO Scholar Dr. M. Conte, BIOS ADVISOR: Dr. A. Andersson Dr. F.G. Plumley, BIOS Ocean Cycle and Ocean Acidification Olga Shatova, BIOS-POGO Scholar C. Newcomb-Hodgetts, BIOS Observatories ADVISOR: Dr. R Johnson Dr. S. Sathyendranath, Dalhousie Drs. A. Andersson & N.R. Bates, BIOS Amanda Vandermark, University of Delaware University ADVISOR: Drs.T. Church, K. Buck, S. Ussher Dr. T. Platt, Dalhousie University Ocean Data Management Susan Allender Dr. H. Bauman, Oxford University Dr. M. Brown, Intergovernmental ADVISOR: Dr. M.W. Lomas Dr. G. White, Bedford Institute of Oceanographic Commission, Paris, France Dean Lea Oceanography, Plymouth Marine Laboratory ADVISOR: J.P. Skinner L. Zhai, Bedford Institute of Oceanography, Charlotte Best, Southampton University Dartmouth, Nova Scotia ADVISOR: Dr. M.W. Lomas Dr. M. Andres, Woods Hole Oceanographic Institution Dr. E. Shroyer, Woods Hole Oceanographic Institution

pg 17 Bermuda institute of ocean Sciences Annual Report 2010 EduCAtion pRoGRAMS

nsF-Funded researcH Ann Lovely, Connecticut College Graduate, VIsItInG academIc GrOuPs eXPerIences FOr Staff at Tabor Academy Amity High School underGraduates ADVISOR: Dr. K. Buck and J.P. Skinner Chamberlain School Hillevi Elena Morris Jaegerman, Tufts Mae Lortie, University of Southampton Clark University University ADVISOR: R. Parsons Columbia University ADVISOR: Dr. M.W. Lomas Simon Laugier East Lyme High Eric David Dexter, Portland State University ADVISOR: T. Noyes Fitch Senior High ADVISOR: Dr. A. Andersson Global Journeys Kristen Marie Sims, Texas Woman's University Bermuda PrOGramme Interns Hamden High ADVISOR: C. Newcomb-Hodgetts Kascia White Har-Bur Middle School Samantha Loomis, Eckerd College ADVISOR: S. de Putron JP Stevens High School ADVISOR: Dr. A. Peters Jae-Tremae Smith-Young Montclair Kimberly Vathani Logendran, Oregon State University ADVISOR: Dr. M.W. Lomas ADVISOR: N. MacDonald Sawyer Boyle Oxford University Christine Van Duyn, Virginia Polytechnic ADVISOR: J.P. Skinner Princeton University Institute and State University Queens University ADVISOR: C. Newcomb-Hodgetts canadIan assOcIates OF BIOs Sacred Heart Melissah Lynn Perkins, University of Central Interns St. Anne's Belfield School Missouri Mark Dowar, York University Sound School ADVISOR: Dr. F.G. Plumley ADVISOR: Dr. S. de Putron Southampton University Janelle Marie Steffen, Indiana University- Mark McInnes, Trent University Tolland Middle School Bloomington ADVISOR: Dr. A. Peters U32 High School ADVISOR: Drs. K. Buck and S. Ussher University of North Carolina Pembroke ecKerd/GaLBraItH scHOLar University of North Carolina Wilmington VOLunteer Interns Nathan Van Bibber Vassar College Mark Dowar, York University ADVISOR: R. Parsons Vernon Middle School ADVISOR: Dr. S. de Putron Wall High Mark McInnes, Trent University Waynflete School ADVISOR: Dr. A. Peters Ocean academY VOLunteers Aaron Cevallos, Princeton University Allister Stewart, Bermuda Sloop snorkel trips VIsItInG Bermuda scHOOLs ADVISOR: Drs. A Andersson and Allister Stewart, Bermuda School field trips Jeffrey Chen, Princeton University Alexander Hunter, Woodland Restoration ADVISOR: Drs. A. Bodnar and J. Loram field trips Bermuda Institute Hannah Barkley, Princeton University ADVISOR: Dr. S. de Putron BIOs eXPLOrer VOLunteers Clearwater Middle School Graham Peigh, Princeton University Samantha de Putron, Matt Bradfield, Tiffany ADVISOR: J.P. Skinner Wardman and Vanessa Shorto, BIOS Dellwood Middle School Matt Costa, Princeton University Sara Scovronick, Columbia University ADVISOR: Dr. S. de Putron Sandys Middle School Matthew Humphreys, Cambridge University Somersfield Academy ADVISOR: Dr. A. Andersson TN Tatem Middle School Dean Lea ADVISOR: J.P. Skinner Whitney Institute Nathan Van Bibber, Eckerd College Primary Schools: St Georges Prep, East End ADVISOR: R. Parsons Primary, St.Davids Primary, Francis Patton, Susan Allender Harrington Sound, Paget, Gilbert, Purvis, ADVISOR: Dr. M.W. Lomas Northlands,West Pembroke, Prospect, Lilian Krug, POGO Scholar Heron Bay, Southampton Glebe, West End ADVISOR: Dr. A. Andersson Primary, Somerset Primary Olga Shatova, POGO Scholar ADVISOR: Dr. R. Johnson Amy Gobel, Princeton University ADVISOR: Dr. A. Peters Samuel Dresner, Princeton University ADVISOR: Dr. M.W. Lomas Amanda Vandermark, University of Delaware ADVISOR: Drs. T. Church, K. Buck and S. Ussher

pg 18 Bermuda institute of ocean Sciences Annual Report 2010 EduCAtion pRoGRAMS

marIne scIence daY Robbie Smith VOLunteers & GIFts Nancy Stevens BAC St. John’s Ambulance Johnny Barnes St. George’s Club Griffins Restaurant John Barritt and Son Swizzle Inn BELCO Matt Tiahlo Bermuda Natural History Museum The Book Cellar Bermuda Regiment The Reefs Bermuda Trader Triangle Diving Bermuda Zoological Society Vince Tweed BUEI Undercover Tent Rentals Melanie Burrows Alex Venn Tia Burrows Wahoos Zak Burrows Jonathan Watts Burrows Lightbourn Kathleen Watts Butterfield & Vallis Helga Wingate Capital G C Travel Sean Correia BioS is sincerely grateful to all our Department of Youth, Sport & Recreation volunteers and we would like to Travis De Sousa thank all the companies and indi- Fairmont Hamilton Princess Hotel viduals who donated goods, serv- Brian Fox ices and time. We offer our Dellesha Fox apologies to anyone who may have Alex Gerenda been inadvertently left off the list. Graphix Signs Grotto Bay Hotel Henry VIIIth Restaurant Kaelah Hines Alice Jolley Wolde Jones Keep Bermuda Beautiful (KBB) Mai Knap Jim McLaren Ministry of Works & Engineering Kathleen Munson Liz Nystrom MarketPlace Anthony Mocklow Capt John Moore Alexis Moreno KJ Moreno Noah’s Ark Sandra Outerbridge Paradise Gift Shop Parks Department Helle Patterson Lily Peters Pier Vu Rayclan Donald Richardson Riddells’ Bay Golf Club Rickai Robinson Jorge Sanchez Crystal Schultz Hamish Shorto Joshua Simoes photos by tiffany Wardman Sinclairs

pg 19 Bermuda institute of ocean Sciences Annual Report 2010 SEMinARS & EvEntS January 5 Biologically driven vertical water transport at the July 9 Coral strategies for survival when it gets too hot Sargasso Sea. Bermuda Program Interns Seminars Dr. Lisa Rodrigues Mimoy Silvano Sea Urchins as a Model for Research March 22 Biological contributions to the carbon inventory in Jecar Chapman Coupling flow cytometry with natural abundance the North Atlantic subtropical mode waters dur- The Biological Carbon Pump nitrogen isotope analysis in the Sargasso Sea: ing its formation and dissipation. Claire Grenfell Sources of N for algal growth Lica Krug Sarah Fawcett, Princeton-BIOS PhD Bermuda Program Summer of 2010 Air-sea CO2 exchange and net calcification on Sarah Gosling April 22 the Bermuda coral reef platform. The Nitrogen Dilemma: Feed the World or Protect Gustavo Fuentes Bermuda Reef fish guilds and abundance at the Environment? different reef types. Dr. James N. Galloway, Sidman P. Poole Pro- Copper and Zinc distribution in Castle Harbor Jorge Sanchez fessor of Environmental Sciences and Associ- and Bermuda. ate Dean for the Sciences, College and Bennet Foli How corals survive: The effects of temperature, Graduate School of Arts and Sciences, at the feeding and acidification on coral polyps. University of Virginia June 18 Kascia White Shimmering Sea Soirée Benefit April 23 Coral Beach & Tennis Club Aug 13 Board of Trustee Seminar Bermuda Program Intern Seminars Ocean Academy July 1 JP Skinner Waterstart Marine Science Internship (MSI) Dive Safety and Boat Preparation Final Presentations and Graduation Forrest Williams Jr. NF-POGO Scholar Independant Research Presentations July 13 Waterstart Survival Guide Detecting stress in corals and the physiological Sawyer Boyle May 20 history that is retained within coral skeletons. NF-POGO Scholars Final Seminars Dr Neal Cantin, Woods Hole Oceanographic Particulate Organic Phosphorus. Institute Claire Grenfell Primary production in spring bloom in the Sargasso Sea: An integration of time-series in-situ July 22 Marine Environmental Program. data and ocean color remote sensing Intern Seminars Sarah Gosling observations. Hoang Cong Tin Wet and dry nitrogen deposition in the North At- Analysis of coprostanol and other sterols to lantic with air mass trajectory analysis. determine fecal contamination in sediment Variability of phytoplankton with relation to C Amy Gobel, Princeton samples around Bermuda. flux at BATS site. Kiersten Vance Nilanthy Priyadarshani Growth rates and Feeding in Corals Kascia White, Bermuda Program Source analysis and quantification of wet and dry Seasonal and inter-annual Pattern of Coccol- nitrogen deposition in the Sargasso Sea. ithophore pigment and ocean acidification. Sea Urchin's Resistance to UV Radiation and Amy Gobel, Princeton University Hui Zhao H2O2 (Results and Findings). Jecar Chapman, Bermuda Program August 11 Oligotrophic jelly-plankton blooms: implications POGO Scholar Introduction presentations for carbon cycling in the Sargasso Sea. Analysis of coprostanol and other sterols to deter- Kentaro Suzuki mine fecal contamination in sediment samples August 28 around Bermuda. Intern Seminars Assessment of zooplankton contribution to parti- Kiersten Vance, Bermuda Program cle fluxes using image analysis. Hannah Barkley, Nathan VanBiber, Olga Shatova Microplankton analysis in the Bering Sea. Liz Drenkard Jorge Sanchez, Bermuda Program Ocean-atmosphere CO2 flux in North Atlantic September 9 Ocean. Waterstart Outreach POGO Scholar Final Chemical Oceanography Lica Krug Sawyer Boyle, Bermuda Program Presentations

Ocean-atmosphere CO2 flux in North Atlantic Cu (copper) Ocean. Maziar Khosravi Kirill Kivva

pg 20 Bermuda institute of ocean Sciences Annual Report 2010 SEMinARS & EvEntS

Al (aluminum) November 19 Dissolved Manganese in the Arctic and Antarctic Fatih Sert Final REU Presentations Oceans. Hydrothermal and fluvial input and bio- logical uptake. Cd (cadmium) November 20 Rob Middag Evgeniya Klimchuk Marine Science Day Open House June 11 Zn (zinc) November 30 High throughput studies of Fe speciation in sea- Fernanda Coló Giannini Sargasso Sea Alliance Presentation water: methodology and applications. Oceanography of the Sargasso Sea – Dr. Kristen Buck, BIOS Hg (mercury) Drs. Nick Bates, Simon Ussher, Andreas An- Rafael Rasse Boada dersson and Samantha de Putron July 2 Tropospheric Flux of Nonylphenol via Aqueous Mn (manganese) December 3 Aerosol Production. Thao Pham POGO Scholars Ocean Observatories Mark McInnes, Trent University and The Presentations Centre for Environmental Modelling and Ni (nickel) Chemistry (CEMC) Gayatri Dudeja Sr (strontium) scIence FrIdaY at tHe WInd July 2 Fabricio Guaman-Guevara Atmospheric Delivery of Trace Metals to February 12 Bermuda. Co (cobalt) Comparing indigenous and western scientific Amanda Vandermark, University of Lazare Akpetou knowledge of the ocean. Delaware. Anna Cummins and Marcus Eriksen, 5 Gyres September 17 July 9 Intern Seminars and POGO Scholar March 17 Seagrass Herbivores: Biting Off More Than They Presentations The Tsunami Project- Interdisciplinary Can Chew. Approaches to Tsunami Risk Reduction. Dr. Kimberly K. Holzer, University of Virginia Intern Presentations Dr. Brian McAdoo Mark Dowar, Matthew Humphreys, Susan July 15 Allender Going Broke: Transcriptome-Level Responses of Plastic Marine Debris in the North Atlantic: Re- Corals to Stress. sults from an ongoing 22-year study at Sea Edu- REU Presentations Dr. Jodi Schwarz cation Association, and fresh data from a recent Eric Dexter, Samantha Loomis, Hillevi cruise on the SSV Corwith Cramer. Jaegerman, Vathani Logendran, Kristen Sims, March19 Giora Proskurowski, Chief Scientist of SSV Christine Van Duyn, Melissah Perkins, Janelle Me, Myself and Iron Corwith Center and Faculty Oceanographer Steffen Dr. Simon Ussher, BIOS at Sea Education Association

POGO Scholar Presentations March 25 August 13 Gayatri Dudeja, Caju Giannini, Thao Pham The Composition of Rainwater Organic Matter: Determining Factors in Mangrove Species Evgeniya Klimchuk, Maziar Khosravi, Rafael Caffeine, Cigarettes and Drugs Zonation. Rasse Boada, Fatih Sert, Fabricio Guaman- Katye Altieri, Princeton Matt Costa, Princeton University Guevara, Lazare Akpetou April 9 Quantification of Hypoxia-Inducible Factor 1α in October 15 Iron, copper, and the ligands that bind them. Sea Urchin Tissues. Effects of Coral Diseases from Individual Host to Dr. Kristen Buck, BIOS Jeffrey Chen, Princeton Coral Reef Ecosystem. Dr. Croquer's April 26 August 20 GEOTRACES cruise, Reserch team on the The Bermuda Weather Service - Introductions, November 2 Dutch research vessel, Pelagia Collaborations and Research. Board of Trustees Presentations Dr. Mark Guishard, Director Bermuda NF-POGO Introduction Presentations GEOTRACES: The West Atlantic Ocean Leg 1. Weather Service, Adjunct Faculty, BIOS Loes Gerringa November 18 August 27 Red or Blue? Darwin’s Bermuda Dilemma and Natural radionuclides in the GEOTRACES Anthropogenic Ocean Acidification: cautionary his Enduring Subsidence Theory of Coral Reefs. program. tales from the past? Dr Brian Rosen, Museum of Natural History, Michiel Rutgers van der Loeff Dr. Caroline Dawber, Cambridge University London

pg 21 Bermuda institute of ocean Sciences Annual Report 2010 FACulty & StAFF scIentIsts Philippe Rouja, PhD, Bermuda manaGement senior scientists Government President and Director Nicholas R. Bates, PhD David Siegel, PhD, University of California, Anthony H. Knap, PhD, Anthony H. Knap, PhD Santa Barbara Fredric Lipschultz, PhD Thomas Sleeter, PhD, Bermuda Assistant to the President and Director, Michael W. Lomas, PhD Government and Secretary to the Corporation Richard J. Murnane, PhD Deborah Steinberg, PhD, Virginia Institute Gillian M.S. Hollis F. Gerald Plumley, PhD of Marine Science Wolfgang Sterrer, PhD, Bermuda Deputy Director associate scientists Aquarium, Natural History Museum & F. Gerald Plumley, PhD Andrea G. Bodnar, PhD Zoo Maureen H. Conte, PhD Peter G. Wells, PhD, Environment Canada Chief Operating Officer Samantha J. de Putron, PhD William Graham-Welton Andrew J. Peters, PhD scientific technical staff Steven J. Bell assistant scientists Margaret HP Best marIne OPeratIOns Andreas J. Andersson, PhD Rachel J. Biggs Science Coordinator Jerome P. Aucan, PhD Charles F. King Nicholas R. Bates, PhD Kristen N. Buck, PhD Natasha McDonald Robert H. Condon, PhD Tim Noyes Marine Superintendent Rodney Johnson, PhD Matthew Tiahlo Ronald H. Harelstad Jeannette E. Loram, PhD research technicians Administrative Assistant Postdoctoral scientists Douglas W. Bell Kathryn A. McNaughton Robert H. Condon, PhD Matthew S. Bradfield John R. Casey Marine Operations Coordinators adjunct Faculty Andrew U. Collins Susan T. Brittner Penelope A. Barnes, PhD, Malaspina Elizabeth D. Condon University-College Helen Drost Technical Services Manager Craig Carlson, PhD, University of Dafydd Gwynn Evans James Jr. D. Caison California, Santa Barbara Rachel Evans Cyr Couturier, PhD, Memorial University Rebecca Garley Marine Technicians of Newfoundland Stacey R. Goldberg Stuart C. Johnson Michael Depledge, PhD, United Kingdom Lilia M. Jackman Matthew J. Wilkinson Environment Agency David A. Koweek Charles Derby, PhD, Georgia State Kevin Lew University Debra A. Lomas r/V atLantIc eXPLOrer Éric Dewailly, PhD, Laval University Alastair J. Marks Captains Richard Dodge, PhD, NOVA University Keven Neely Quinton S. Lewis Angela Douglas, PhD, James W. Sadler Ralph H. van Hoek Steven Giovannoni, PhD, Oregon State Elise Van Meerssche University Mates Giselher Gust, PhD, Hamburg University Courtney Barber III Mike Helm, PhD rIsK PredIctIOn InItIatIVe E. James Doyle David Karl, PhD, University of Hawaii Science Program Manager Quinton S. Lewis David Malmquist, PhD, Virginia Institute of Richard J. Murnane, PhD Mark T. Maloof Marine Science Richard C. Ogus John J Manock, PhD, University of North Communications Technician Carolina, Wilmington & Webmaster Chief Engineer Murray Moinester, PhD, University of Tel Charles F. King Blake Barber Aviv Richard E. Smith Jr. Norman B. Nelson, PhD, University of California, Santa Barbara

pg 22 Bermuda institute of ocean Sciences Annual Report 2010 FACulty & StAFF r/V atlantic explorer crew Bookkeepers OPeratIOns through Bernhard schulte Cheryl E.M. Iris Laboratory Operations Technician ship management Shirley M. Sousa Bruce Williams Officers Mel June Inocencio, 2nd Officer Accounts Payable Clerk Laboratory Attendant Manuel Par, 2nd Officer LeeAnn Fox-Moreno Georgianna White Juan Cabungcal, 4th Engineer Accommodations Reservations & Motormen InFOrmatIOn tecHnOLOGY House Manager Cristito Briones Chief Technology Officer Jane L. Burrows Juan Cabungcal James S. Marquez Alchamor Soliva Reservations Assistant/Receptionist IT Manager Heidi Smith Chefs K. Mark Jolley Ades Gregorio Housekeeper Bernardo Manalo Network and Systems Administrator Sharon Minors Dexter Ojan Alexander J. Hidalgo James G. Wicker Office Attendants Bosun Alexis E. Fox-Moreno Joseph Paitone T. Trevon S Hayling Ronaldo De Leon educatIOn Sharon Minors Director of Education Caurline J. Smith Seamen F. Gerald Plumley, PhD Kanyour L. Wales Jesus Ambrocio Georgianna White Ciriaco Mutas Jr. Education Officer Eric Parcon John Paul Skinner Room Attendants June F. Manalo Caurline J. Smith Education Coordinators Kanyour L. Wales Motormen Meaghan Meredith Cristito Briones Chloé Newcomb Hodgetts Juan Cabungcal dInInG Alchamor Soliva Librarian Head Chef, Kitchen Manager Chloé Newcomb Hodgetts George Crane

Small Boats and Diving Outreach Coordinator Chef Diving Safety Officer/Small Boats Dean E. Lea Helena Simoes Supervisor Alexander D. Chequer Senior Lecturer Kitchen Assistants Samantha de Putron, PhD Charlene Millet Captain R/V Stommel Georgianna White Alfred A. Smith Instructors Angela Mead, PhD maIntenance FInance Co-Instructors Maintenance Superintendent Treasurer and Controller Kaitlin Baird Wilfred J. Stovell Stephen M. Markman, CPA Chloé Newcomb Hodgetts Carpenter Human Resources Officer Teaching Assistants Chesley W. Foggo Sandra M. Spurling Lisa M. Bourassa Chloé Newcomb Hodgetts Electrician/Plumber Qualified Accountant Warren A. Smith Kelly McLaren Truck/Bus Driver Accountant Kenneth E. Trott Ruth M Heron-White

pg 23 Bermuda institute of ocean Sciences Annual Report 2010 FACulty & StAFF

InstItutIOnaL adVancement Chief Development Officer Arthur G. Wasserman

Personal Assistant to the BIOS, UK Associates Joanna Moore

Development Officers Katherine J. Lantagne Keesha Simon-Walker

cOmmunIcatIOns & marKetInG Public Relations Officer Tiffany Wardman

Community Outreach and Events Officer Vanessa Shorto

pg 24 Bermuda institute of ocean Sciences Annual Report 2010 HonoR Roll of donoRS the 107-year history of INDIVIDUALS Mr. and Mrs. James B. Butterfield, JP achievement at the Bermuda The following individuals made gifts to BIOS George and Sue Cubbon institute of ocean Sciences (BioS) for operating support in 2010. Rod and Martha Ferguson Ms. Diana H. Frazier would not be possible without the Mr. John Gaugain support of the donors who are part THE CHAIRMAN'S CIRCLE Sir David and Lady Gibbons of the BioS community. year after $100,000 or more Ms. Linda K. Glover year, BioS Annual Fund donors have Mr. and Mrs. Robert E. Cawthorn Peter and Lennie Gotcher provided truly significant support Mr. Peter B. Green Ms. Jamie Gutteridge for marine research and education. THE OCEAN PROTECTORS Walwyn and Betsey Hughes $50,000 – $99,999 Dr. Anthony H. Knap Mr. and Mrs. Brian Duperreault Dr. Georgia E. Lesh-Laurie Mr. and Mrs. Ross Perot Jack and Virginia Lightbourn Mr. and Mrs. Sjur H. Linberg Alan and Rochelle McGowan THE INTERNATIONAL PARTNERS Mr. and Mrs. Raymond E. Moore $10,000 – $24,999 Mr. Paolo M. Pellegrini Mr. and Mrs. Philip V. Bancroft Juan and Nancy Prado Mr. and Mrs. Michael A. Butt Mrs. Fred M. Reiss Mrs. Traudl Engelhorn-Vechiatto Dr. and Mrs. William Russel Mr. and Mrs. Handel Evans John and Ruth Sainsbury Richard M. and Helen T. Fraser John and Rosemary Talbot Mr. Michael Hamer and Ms. Clare Bob and Anne Tucker Warburton Mr. and Mrs. George A. Wardman Mr. James Maguire Mr. Arthur Wasserman Mr. Chris Maybury Mr. Anthony Williams Mr. and Mrs. Michael McGavick Brian and Nancy O'Hara Mr. and Mrs. Ian Strecker THE ASSOCIATES PROGRAM

THE VOYAGERS THE EXPLORERS $500 – $999 $5,000 – $9,999 Anonymous Mr. and Mrs. Michael A. Butt Mr. Casper Young and Ms. Jennifer Ayres Mr. and Mrs. Neill Currie Mr. George B. Bennett John and Rosemary Galbraith Mr. and Mrs. J. Stewart Bryan, III Janet Outerbridge Mr. Graham B. Collis Lady Antoinette Pearman M.M. Cone - J.H. Cone Dr. F. Gerald Plumley Bill and Anne Cooke Jonathan and Britt Reiss Mr. John R. Cox John and Marilyn Dickinson Mr. and Mrs. James F.H. Hannam THE ARGONAUTS Mr. J C H Johnson $1,000 – $4,999 Dr Charles F. Kennel Anonymous Dr. Charlotte M. Lord Mr. Foster Bam Dr. and Mrs. Fred T. Mackenzie Hon. Robert V. and Grace Barritt Mr. and Mrs. Larry Marsland Mr. and Mrs. Jeremiah M. Bogert Edith A. Maynard David and Valerie Brown Dr. John J. McDermott In memory of Mrs. Dudley G. Butterfield, Brenda McLean CMG Gretl Nunnemacher

pg 25 Bermuda institute of ocean Sciences Annual Report 2010 HonoR Roll of donoRS

Carl E. Painter Susan Holland Mr. and Mrs. John A. Capstick Mr. and Mrs. Leslie R. Patterson Scott Hunter and Janet Kemp Mr. Andrew W. Card Mr. and Mrs. John D. Pereira Mrs. C.H. Ford Hutchings Ms. Kathleen Casey Walter and Kate Scott Carl and Suzi Johnson Maria C. Charest Mr. and Mrs. Walter A. Scott Bill and Janet Lang Dr. Rita R. Colwell Mark Smith and Mariette Savoie Donald and Janine Lines Stephen and Carolyn Copeland Jan and Barbara Spiering Mr. and Mrs. David Lonsdale Tony and Elizabeth Cordeiro Mr. and Mrs. Richard D. Spurling Ms. Natalie C. Luthi Colin and Peggy Couper Mr. and Mrs. John Stockbridge Drs. David Malmquist and Deborah Mr. and Mrs. David W. Cox Ben and Charlotte Sullivan Steinberg Mrs. H.C.D. Cox Mrs. Dorothy Trimingham Mr. Stephen Markman Dr. Nancy Owens Cunningham Tiffany Wardman Dr. and Mrs. H. Roger Marlene Melton Owen and Pam Darrell Dr. David B. Wingate, OBE Mr. A.E. Ming, Jr. Roger and Lee Davidson Mrs. Keith R.Winter Mrs. Howard Mitchell Connie Dey Mrs. Joy Young-Tucker Captain and Mrs. J. Willard Moore Tommy and Elaine Dickinson Cummings and Katherine Zuill Dr. Duncan and Janice Moran Ross and Dianna Doe Terry and Bebe Moulton Terry and Mary Draycott Dr. and Mrs. Donald A. Munson Anne Engelhardt THE MARINERS Peter and Colleen Nelson Dr. and Mrs. John W. Farrington $250 – $499 Sheila Nicoll and Steve Lake Dr. and Mrs. Faripour Forouhar Mr. and Mrs. Mickey Adderley Ms. Joan M. Petracca D.M.D Dr. Joanne E. Fortune Mrs. Tia D. Andrew Mrs. Sylvia Savery Tina Fraser-Epps Anonymous Mr. and Mrs. Sam A. Stephens Harold Frith Ed and Pat Argenta Dr. Elijah W. Stommel Dr. Robert B. Gagosian Mr. and Mrs. Per Arneberg Mr. and Mrs. J. Clive Thatcher Grant and Kathy Gibbons Dr. and Mrs. Edwin Boger, Sr. Mr. and Mrs. William D. Thomson Mr. Crayton Greene Mr. and Mrs. William J. Brewer James and Calista Todd Mr. and Mrs Tony Hay Karen and Boyce Budd Sharon Vesey Mr. and Mrs. Stephen 'Straws' Haycock Dr. and Mrs. James N. Butler Mr. and Mrs. Joseph E. Wakefield Keith and Norrie Hollis Ted and Loretta Cassidy Mr. and Mrs. Colin R. Young Buell and Margaret Hollister Mr. and Mrs. Wayne Chapman Jonathan Ingham Jack and Pat Chiappa William and Margery Jahoda Hazel Clifford THE ADVENTURERS Mrs. Patricia M. Thorne-Kemp Dr. Richard H. Colby $100 – $249 Mr. Richard Klein John and Judy Collis Elma Anfossi Fred and Suzanne Lepple Mrs. Jean Conyers Anonymous R. McLeod Lightbourn Tom and Heather Conyers Dr. and Mrs. Ali Arouzi Mrs. Joy Lusher Margot M. Cooper Quinton and Cleta Astwood Mr and Mrs Andrew Marsh Alex and Sheelagh Cooper Colin and Moyra Benbow Gervase and Elsie Marson Mr. and Mrs. Dudley R. Cottingham John and Hope Berg Janet E. McCulloch Colin and Peggy Couper John and Jane Bielby Mr. and Mrs. Newton P.S. Merrill Mrs. J. Henry Dallas Jean Bird Ms. Vikki Mesich Peter and Sally Darling W.A. and Susan Black Mr. and Mrs. Leo N. Mills II Ian and Felicité Davidson Malcolm A. Borg Ms. Judith Montgomery-Moore Mr. and Mrs. Nicholas B. Dill Mr. Kevin Bossenmeyer and Mrs. Teri Mary Mullen and Michael Hoffman Mr. and Mrs. Michael J. Drew Wielenga Robert Nunnemacher Elizabeth A.C. Durrant Mrs. Hope B. Bowker Bernard and Lily Oatley Mr. and Mrs. Richard Ferrett Mr. Anthony Boxall Eric and Muriel Parker Keith and Suzette Fisher Kenneth and Nancy Brink Mrs. George M. Parker Peter and Elizabeth Gebhard Dr. and Mrs. Albert L. Brooks Mr. and Mrs. Richard S.L. Pearman Mr. and Mrs. Tredick Gorham David S. Brown Dr. and Mrs. Brian W. Peckett Patrick and Bridget Hagan Norman and Anne Brown Mrs. Audrey Nunnemacher Pertl Henry and Patricia Hayward Warren Brown Mr. and Mrs. Kenneth Pestone Jim and Diana Hindess Mr. David K. Bulloch Mr. and Mrs. Joseph Pilaro

pg 26 Bermuda institute of ocean Sciences Annual Report 2010 HonoR Roll of donoRS

Mrs. Bill M. Pimm Mr. and Mrs. Peter Holmes THE CORPORATE INVESTOR'S Mr. and Mrs. Robert Rans Mrs. Margaret King PROGRAM Mr. and Mrs. John Ely Riegel Ms. Suzanne Koren The following companies made gifts to BIOS Dr. Priscilla F. Roslansky Louis and Cricket Laun for operating support. Mr. and Mrs. Laurence P. Rubinow Mr. Nicholas Leporacci Martin and Jane Salwen Mr. and Mrs. David Lines THE PRESIDENT'S CIRCLE Winifred C. Savill Ms. Jennifer M. Mancell Jonathan and Gwyneth Sharp Mr. and Mrs. Robert E. Mello THE ROUNDTABLE Kathryn Siggins Dr. John M. Miller $25,000 – $49,999 Dr. June Lindstedt Siva Claire Perry Flagstone Reinsurance Limited Mr. Charles Slatkin Dr. Harold R. Pestana Audrey C. Smith Mrs. Albert A. Powell Mr. and Mrs. C.H. Smith, Jr. Patricia Powell THE INTERNATIONAL PARTNERS Peter and Pamela Smith Baroness E. Raben-Levetzau $10,000 – $24,999 Mr. and Mrs. Reginald A.R. Smith Miss Elizabeth E. Roosevelt Bermuda Container Line Ltd. John and Evelyn Steele Dr. Klaus Ruetzler Mrs. Ivan Swan Ms. Joan Senghas Dr. Robert H. Warren Mr. Rich Seymour THE EXPLORERS Joyce Watlington David and Susan Sims $5,000 – $9,999 Mr. and Mrs. Bruce F. Wilkie Ann Smith Gordon, MBE, JP Bacardi International Limited Mrs. Nea P. Willits Mr. and Mrs. Elliot Sylvan BF&M Insurance Group Dr. and Mrs. Joseph H. Woolf Sylvia B. Tompkins Hannover Life Re Edward and Yvonne Wright Dr. Robert L. Vadas, Sr. New England Biolabs, Inc. Mr. and Mrs. Owen Walsh Oil Management Services Ltd. Mr and Mrs Geordie Wardman Renaissance Reinsurance Ltd. THE RESEARCHERS Ronica Watlington Royal Dutch Shell Companies in $50 – $99 Mr. and Mrs. W. Graham Welton Bermuda Dr. C.J. and Sharon Abraham Mr. and Mrs. William S. Zuill, Sr. Anonymous Mr. and Mrs. Gerald B. Ardis THE ARGONAUTS Ms. Barbara Atwood $1,000 – $4,999 Professor Harold L. Atwood A.P. Moller (Bermuda) Limited Mr. and Mrs. Edward Babor Chevron International Limited Sydney and Sheila Bamber Conyers, Dill & Pearman Peter and Joan Banks Gibbons Management Services Limited Mr. David Barlet International Gas Transportation Mr. and Mrs. Robin M. Blackburne Company Limited Dr. Vincent Brach Orbis Investment Management Ltd. Robert and Marcia Brixius Renaissance Reinsurance Ltd. Duffy and Laurie Brookes Robertson's Drug Store Than and Nancy Butterfield Standard Steamship Owners' P & I Assn. Mr. and Mrs. J. David Chew (Bermuda) Ltd. Herman and Barbara Cooper The Bermuda Telephone Company, Ltd. Margot Cox Carleen Currie Mr. and Mrs. Byrd W. Davenport, Jr. Barbara Elkin Mr. and Mrs. Chris Estwanik Ms. Emily Ferguson Dr. Katherine Fleischmann Dr. Gloria J. Frederick Professor Robert N. Ginsburg Dr. and Mrs. Seymour Handler Mr. Daniel Harrop Dr. Robert B. Hill pg 27 Bermuda institute of ocean Sciences Annual Report 2010 HonoR Roll of donoRS

THE ASSOCIATES PROGRAM The Arthur & Annette Bolton Charitable RESEARCH AND OTHER GRANTS Foundation The ACE Foundation - Bermuda THE VOYAGERS Bank of Bermuda Foundation ExxonMobil Foundation $500 – $999 Centre on Philanthropy Hamilton Charitable Foundation Atlantic Marine LP Ernest E. Stempel Foundation PADI Foundation Attorney's Liability Assurance Society Fidelity Charitable Gift Fund Partnership for Observation of the Ltd. Foster-Davis Foundation, Inc. Global Oceans (POGO) Bermuda Air Conditioning Ltd. The G. Unger Vetlesen Foundation Mr. and Mrs. William H. Williams John Barritt & Son, Ltd. Johnston Family Foundation Lockheed Martin Sippican, Inc. The MacPherson Fund, Inc. RISK PREDICTION INITIATIVE Merck Sharp & Dohme International Ltd. The McCance Foundation Trust Amlin Plc Thomas Miller (Bermuda) Ltd. Michael and Susan Newburger Aspen Insurance Holdings Limited Vallis & Hayward Ltd. Foundation AXIS Specialty Limited Mostyn Foundation, Inc. Nephila Capital Ltd. Network for Good PartnerRe Ltd. THE MARINERS Pfizer Foundation Risk Management Solutions, Inc. $250 – $499 Robert Wood Johnson Foundation State Farm Fire and Casualty Company Anfossi Management Ltd. Walter and Mary Scott Foundation WeatherPredict Arch Reinsurance Ltd. (Bermuda) Senior's Learning XL Re Ltd. Burrows, Lightbourn Ltd. Vita Foundation Ernst & Young The Wendell & Margaret Smith ROGER PERRY SCHOLARSHIP Foundation ENDOWMENT FUND The William Ewing Foundation Isobel Perry THE ADVENTURERS Mary and Redwood Wright $100 – $249 ACE Installers RESTRICTED GIFTS AND GRANTS THE SUNDERMAN FUND P.M. Limited The following companies, foundations and Martin and Jane Salwen Rosedon Hotel individuals made gifts to BIOS for specific John and Evelyn Steele The Total Group of Companies programs or projects. Windcrest Management Limited CANADIAN ASSOCIATES OF BIOS THE ANTI-AGING PROJECT Nick & Lynn Ross Charitable Foundation Canadian Associates of BIOS St. Marys Cement Inc. (Canada) MATCHING GIFTS Hannover Life Re Mr. James Beal We are grateful to the following Mr. and Mrs. Raymond E. Moore Dr. Neil Campbell corporations and private foundations that Mr. and Mrs. Michael Davies have provided matching support for gifts R/V HSBC ATLANTIC EXPLORER Mr. William Iams & Mrs. Marie Iams made to BIOS by their employees, retirees Bank of Bermuda Foundation Ms. Patricia Pocklington or directors. HSBC Mr. Edward Truant Mr. Karl Wildi The ACE Foundation - Bermuda EDUCATION Arch Reinsurance Ltd. (Bermuda) Dr. and Mrs. Kenneth H. Brink UK ASSOCIATES OF BIOS Bank of Bermuda Foundation Canadian Associates of BIOS The Fishmonger's Company Flagstone Reinsurance Limited LCDR Donald S. Delikat Mr. Clive Askew General Atlantic Group ExxonMobil Foundation Michael Bradfield Pfizer Foundation The Agouron Institute Maldwin Drummond (Chairman) Renaissance Reinsurance Ltd. The Charrock Foundation Chris Day Robert Wood Johnson Foundation U.K. Associates of BIOS Professor Mike Depledge XL Foundation Diana Dunrossil THE HEIN BESSELAAR FUND Charles Howeson Toni Besselaar Professor Richard Lampitt SUPPORT FROM PRIVATE Raymond Sykes FOUNDATIONS NUNNEMACHER NATURE RESERVE Sir Crispin Tickell Abacus Trust Limited Dr. and Mrs. Robert B. McComb Professor John Woods The ACE Foundation - Bermuda

pg 28 Bermuda institute of ocean Sciences Annual Report 2010 HonoR Roll oF donoRS

GIFTS IN KIND Mr. and Mrs. Thomas N. Wadson St. John’s Ambulance Tiffany Wardman Tops Ltd. INDIVIDUALS John B. Watson and Susan Walton Turkish Imports Grand Bazaar Mr. Tom Anderson Mrs. William F. Way Undercover Tent Rentals Ms. Magdalena Andres Mr. Tripp West Mr. and Mrs. Marcello Ausenda Mr. George White Johnny Barnes Bob and Christine Whiting VOLUNTEERS Miss Joy Barnum Ms. Eileen Zerba Greg Burrows Toni Besselaar Mr. Li Zhai Tia Burrows Mr. Brian W. Billings Cummings and Katherine Zuill Zak Burrows Ms. Heather Bouman Sean Correia Mr. and Mrs. Barry Brewer ORGANIZATIONS Travis De Sousa Kenneth and Nancy Brink Argus Optical Brian Fox Mr. Dwayne Caines Astwood Dickinson Dellesha Fox Mrs. Beecher Chapin Bermuda Air Conditioning Ltd. Alex Gerenda Mr. and Mrs. William Collieson Bacardi International Limited Kaelah Hines Mr. and Mrs. J. Cyril Cooper BELCO Alice Jolley Mr. and Mrs. William J. Cox Bermuda Golf Academy and Driving Mai Knap Allan and Judith Davidson Range Jim McLaren Mr. Maldwin Drummond Bermuda Natural History Museum Anthony Mocklow Mr. and Mrs. Peter C. Durhager Bermuda Regiment Capt. John Moore Mr. John Ellison Bermuda Zoological Society Alexis Moreno Mrs. Kenneth Finsness BUEI KJ Moreno Ms. Logan Franklin Burrows, Lightbourn Ltd. Kathleen Munson Ms. Jamie Gauk Butterfield and Vallis Liz Nystrom Mr. Stephen Franklin C Travel Sandra Outerbridge Ms. Linda K. Glover Capital G Helle Patterson Mrs. Adrianna Goodfellow CCMP - Bigelow Laboratory for Ocean Lily Peters Miss. Clare Gould Sciences Donald Richardson Mr. and Mrs. Peter Head The Charrock Foundation Rickai Robinson Dr. John Heidelburg Coral Beach & Tennis Club Crystal Schultz Dr. and Mrs. Michael W. Lomas Department of Youth, Sport & Recreation Joshua Simoes Miss. Heather MacDonald Fairmont Hamilton Princess Hotel Robbie Smith Mr. and Mrs. Kenneth C. MacGillivray, Jr. Gosling Brothers Limited Nancy Stevens Mr. and Mrs. John F. Mariani Graphix Signs Matt Tiahlo Mr. Christopher Marson Greenway Landscaping Vince Tweed Miss Meghan Meredith Henry VIIIth Restaurant Jonathan Watts Mr. James Mulderig The Irish Linen Shop Ltd. Kathleen Watts Ms. Cindy Mitchell John Barritt & Son, Ltd. Helga Wingate Mr. Trevor Nicholls John Barritt and Son Mr. Trevor Platt Kaleidoscope Arts Foundation the time period covered for recognition of Ms. Margaret Potts Keep Bermuda Beautiful (KBB) donor support to BioS in this report is the Mrs. Lisa Quin Kirk's Jewellers calendar year of 2010. if we inadvertently Mr. and Mrs. Ralph E. Richardson MarketPlace ommitted your name from the Honor Roll of donors or the Associates program, please Mr. Hamish Shorto MEF Enterprises accept our appoligies and let us know so that Ms. Emily Shroyer Melanie Burrows we may rectify the error. you may do so by Dr. Shubha Sathyendranath The Meyer Group of Companies contacting Keesha Simon-Walker at Ms. Nancy Smythe Noah’s Ark [email protected] or by calling Mrs. Anita Stockbridge Parks Department 441-297-1880 ext. 114. Mr. Ben Tompkins Partnership for Observation of the Global Ed and Bobbie Trippe Oceans (POGO) Mr. Christopher Trott Pulp and Circumstance Mr. and Mrs. Alex Venn Rayclan Sir John and Lady Vereker The Reefs Beach Club Diane Von Furstenberg Riddells’ Bay Golf Club Peter Vorley and Marion Watlington Southampton Princess Hotel

pg 29 Bermuda institute of ocean Sciences Annual Report 2010 17 Biological Station • St. George’s • GE 01 Bermuda • 441.297.1880 • www.bios.edu