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10 Annual Meeting of the

European Association of Vertebrate Palaeontologists

Royo-Torres, R., Gascó, F. and Alcalá, L., coord. (2012). 10th Annual Meeting of the European

Association of Vertebrate Palaeontologists. ¡Fundamental! 20: 1–290.
EDITOR: © Fundación Conjunto Paleontológico deTeruel – Dinópolis

COORDINATION: Rafael Royo-Torres, Francisco Gascó and Luis Alcalá. DISEÑOY MAQUETA: © EKIX Soluciones Gráficas DL:TE–72–2012 ISBN–13: 978–84–938173–4–3

PROJECT: CGL 2009 06194-E/BTE Ministerio de Ciencia e Innovación

Queda rigurosamente prohibida, sin la autorización escrita de los autores y del editor, bajo las sanciones establecidas en la ley, la reproducción total o parcial de esta obra por cualquier medio o procedimiento, comprendidos la reprografía y el tratamiento informático. T o dos los derechos reservados.

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the accurate geological study has contributed to understand the succession of dinosaur faunas from the latest Campanian to the end of the Maastrichtian in the Ibero-armorican Island.

The last dinosaurs of Europe: clade-specific

heterogeneity in the dinosaur record of the southern Pyrenees

Geological Setting

Àngel Galobart1, José Ignacio Canudo2, Oriol Oms3, BernatVila2,1, Penélope Cruzado-Caballero2, Violeta Riera3, Rodrigo Gaete4, Fabio M. DallaVecchia1, Josep Marmi1 and Albert G. Sellés1

TheArén Sandstone andTremp Formations represent the coastal and coastal to fully continental deposition, respectively, during the Late Cretaceous-Palaeocene interval in the southern Pyrenees. They record a marine regression that began near the CampanianMaastrichtian boundary. In the Tremp Formation four informal lithostratigraphic units have been distinguished (Rosell et al., 2001) (Fig. 1). Dinosaurs bearing sites occur in the two lower units: the

“grey unit” and in the “lower red unit”. The first one consists in grey

marls with abundant invertebrates that usually are interbedded with layers of coal seams, limestones with charophytes and sandstones beds. It deposited in lagoonal and coastal wetland settings with variable salinity that laterally evolve to a barrier-island. The age ranges from the earliest to the latest Maastrichtian.

1Institut Català de Paleontologia Miquel Crusafont, C/ Escola Industrial 23, 08201, Sabadell, Catalonia, Spain.

[email protected]; [email protected]; [email protected]; [email protected]

2Grupo Aragosaurus-IUCA, Universidad de Zaragoza, C/ Pedro Cerbuna 12, 50009, Zaragoza, Spain.

jicanudo@uniza r . es; bernat.vila@uniza r . es; penelope@uniza r . es

3Departament de Geologia, Universitat Autònoma de Barcelona, 08193, Cerdanyola del V a llès, Catalonia, Spain.

[email protected]; [email protected]

4Museu de la Conca Dellà,

C/ del Museu 4, 25650, Isona, Catalonia, Spain. [email protected]

The outcrops of the Aren and Tremp Formations in the southern Pyrenees, which are located in the Aragón and Catalunya communities, have yielded the most important collection of Late Cretaceous dinosaur remains of Europe. Bones, ichnites and eggs are found in a wide exposure that extends over one hundred

kilometres. The importance of these outcrops was first noted by

Talens (1955) and Lapparent (1958) in the middle 50s, and later corroborated by other scientists in the last decades (e.g., Casanovas

et al., 1993; Sanz et al., 1995). Recent fieldworks conducted by the

Universidad de Zaragoza and the Institut Català de Paleontologia have uncovered new dinosaur sites. These new data, together with

Figure 1. Chronostratigraphy of the Tremp and Aren formations in the

southern Pyrenees (modified from Riera et al., 2009). From east to west:

Vallcebre Syncline, Coll de Nargó Syncline, Tremp Syncline, and Àger Synclines.

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The “lower red unit” is mainly made of red lutites with sandstones, lacustrine limestones and palaeosols. These facies are

interpreted as floodplain alluvial and fluvial deposits with braided

and meandering channels and palustrine-lacustrine limestones interbedded (Rosell et al., 2001). Its age is late Maastrichtian (Riera et al., 2009; Oms et al., 2007; Oms and Canudo, 2004). teeth (Torices et al., 2004; Riera et al., 2009). Peculiar eggs (Sankofa pyrenaica) from a small theropod are known from the upper Arén Formation (López-Martínez and Vicens, 2012) and Prismatoolithidae-type eggshells are known from a few sites of the “lower red unit” (Galobart, 2006).

Ornithopods

Dinosaurian fossil record

Hadrosauroids are the most abundant dinosaurs in the Arén and Tremp Formations. About sixty localities with hadrosauroid

bone remains and tracks have been identified, but no egg-sites

have been reported to date (Dalla Vecchia et al., 2011). Nearly all sites are located in the “lower red unit” whereas only a few are in the upper part of the underlying “grey unit”. The sampled record indicates high hadrosaurid diversity in the upper Maastrichtian with at least three lambeosaurines - Pararhabdodon isonensis,

Arenysaurus ardevoli and Blasisaurus canudoi – an euhadrosaurid

and an indeterminate ‘hadrosaurine’ (Prieto-Márquez et al., 2006; Pereda Suberbiola et al., 2009; Cruzado-Caballero et al., 2010a, Cruzado-Caballero et al., 2010b, Casanovas et al., 1999).

We completed a continuous stratigraphic record for the

Maastrichtian of the southern Pyrenees that provides a physical framework in which the dinosaur sites can be located.

Sauropods

Sauropod evidence include tracks and trackways from two basal levels (Fumanya and Orcau-2; Vila et al., 2005; Vila et al., 2011a) in the “grey unit” and a few sites with eggs and clutches (Font del Bullidor, Biscarri and Basturs-1, 2; Vila et al., 2010a; López-Martínez et al., 2000; Sanz et al., 1995) at the top of theArén Formation and the “grey unit” (Tremp Formation). Bone remains are scant and they occasionally occur in the upper part of the “grey

unit” and in the “lower red unit”, probably reflecting differences

in rock availability between the two units. In the “lower red unit”, the eggs and clutches are abundant (e.g. Pinyes and Els Terrers sites; Vila et al., 2010b; Vila et al., 2011b) being scarcer upwards.

Tracks are rare and they are only found near the K-Pg boundary. In

terms of sauropod diversity, at least four distinct titanosaurs forms can be distinguished in the upper part of the Tremp Formation, late Maastrichtian in age (Vila et al., 2009).

Ankylosaurs

To date, nodosaurid remains are probably the most infrequent dinosaur fossils in the south-Pyrenean basins. They include a few teeth and some isolated postcranial material (López-Martínez et al., 2000; Riera et al., 2009; Escaso et al., 2010). Neither eggs

nor tracks have been identified yet. They are always found in the

lower Maastrichtian deposits (lowermost part of the “grey unit” in the eastern Tremp Syncline), while they were never reported from the “lower red unit”.

Theropods

Conclusions

Theropod remains are rare and distributed in a few localities throughout the upper Arén Formation and Cretaceous part of Tremp Formation. They include teeth, eggshells, eggs and rarely

bones. Five different theropod taxa have been identified, at least,

on the basis of dental morphology: Coelurosauria indet., cf.

Dromaeosauridae indet., cf. Richardoestesia, cf. Euronychodon

and an indeterminate, large-sized taxon with plesiomorphic
The southern Pyrenees contain a rich paleontological record to understand the terrestrial environmental changes and biota successions

of the last five million years of the Mesozoic. Over 300 sites with dinosaur remains have been identified to date, containing titanosaur

sauropods, theropods, hadrosauroid ornithopods and nodosaurid ankylosaurians, as well as many eggs sites and thousand of tracks.

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On the basis of this fossil record, a well dated succession of dinosaurs is established. This is featured by the following

statements: first, there is clear heterogeneity in the stratigraphic

distribution of the dinosaur taxa in the southern Pyrenees. Second, dinosaur sites are less frequent in the ‘grey unit’, so most of the fossils are from the upper Maastrichtian “lower red unit” of the Tremp Formation. Third, changes in dinosaur associations along

the chronostratigraphic succession may reflect time-related events

(e.g., extinctions and colonizations) or biases (rock availability, sampling, environment, and ecology).
Latest Cretaceous of Arén (Huesca, Spain). Canadian Journal

of Earth Sciences, 47 (12): 1507-1517.

CRUZADO-CABALLERO, P., RUIZ-OMEÑACA, J.I. and
CANUDO, J.I. 2010b. Evidencias de la coexistencia de hadrosaurinos y lambeosaurinos en el Maastrichtiano superior de la Península Ibérica (Arén, Huesca, España). Ameghiniana, 47(2): 153-164.
DALLA VECCHIA, F.M., GAETE, R., RIERA, V., OMS, O.,
PRIETO-MÁRQUEZ, A., VILA, A., GARCIA SELLéS, A. and GALOBART, A. 2011. The hadrosaurid record in the Maastrichtian of the Eastern Tremp Syncline (Northern Spain).

In: Hadrosaur Symposium at the Royal Tyrrell Museum of

Palaeontology. Abstract V o lume: 38-44.

The abundance and diversity of dinosaur fossils in the upper
Maastrichtian of the Pyrenees has a great potential to improve our knowledge of the extinction pattern of continental vertebrates at the Cretaceous – Palaeogene boundary in Europe.
ESCASO, F., PéREZ-GARCÍA, A., ORTEGA, F. and SANZ, J.L.
2010. Ankylosaurian evidence from the Upper Cretaceous of South Central Pyrennees (Lleida, Spain): a reappraisal. In: 8

EAVP Meeting, Aix-en-Provence 2010. Abstract V o lume: 33.

GALOBART, A. 2006. Importancia del registro español de huevos

de dinosaurio. In: Los dinosaurios en el siglo XXI, Nuevas respuestas al inagotable enigma de los dinosaurios, (J.L. Sanz,

Ed). Colección Metatemas, Tusquets Editores, Barcelona, 99, 98-119 p.

Acknowledgements

We would like to thank the Projects of the Subdirección

general de Proyectos de Investigació, Ministerio de Economía y Competitividad: CGL2011-30069-C02-01/02 and CGL2010- 16447.BV acknowledges support from the Ministerio de Ciencia e Innovación (Subprograma Juan de la Cierva (MICINN-JDC) 2011).
LAPPARENT, A.F. 1958. Découverte d’un gisement d’oeufs de
Dinosauriens dans le Crétacé supérieur du bassin de Tremp

(Province de Lérida, Espagne). Comptes Rendus de l ’ A cadémie

des Sciences de Paris: 1879-1880.

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  • The Upper Maastrichtian Dinosaur Fossil Record from the Southern

    The Upper Maastrichtian Dinosaur Fossil Record from the Southern

    Cretaceous Research 57 (2016) 540e551 Contents lists available at ScienceDirect Cretaceous Research journal homepage: www.elsevier.com/locate/CretRes The upper Maastrichtian dinosaur fossil record from the southern Pyrenees and its contribution to the topic of the CretaceousePalaeogene mass extinction event * Jose I. Canudo a, , Oriol Oms b, Bernat Vila a, Angel Galobart c, g, Víctor Fondevilla b, Eduardo Puertolas-Pascual a, Albert G. Selles c, Penelope Cruzado-Caballero d, Jaume Dinares-Turell e, Enric Vicens b, Diego Castanera a, Julio Company f, Laura Burrel b, Rita Estrada b, Josep Marmi c, Alejandro Blanco c a AragosauruseIUCA, Paleontología, Facultad de Ciencias, Universidad de Zaragoza, 50009, Zaragoza, Spain b Universitat Autonoma de Barcelona, Facultat de Ciencies (Geologia), 08193, Bellaterra, Barcelona, Spain c Institut Catala de Paleontologia Miquel Crusafont, C/ Escola Industrial 23, 08201, Sabadell, Spain d CONICET, Instituto de Investigacion en Paleobiología y Geología, Universidad Nacional de Río Negro, Av. Roca 1242, General Roca, 8332, Río Negro, Argentina e Istituto Nazionale di Geofisica e Vulcanologia, Via di Vigna Murata 605, 00143, Roma, Italy f Departamento de Ingeniería del Terreno, Universidad Politecnica de Valencia, Camino de Vera s/n, 46022, Valencia, Spain g Museu de la Conca Della, C/ del Museu 4, 25650, Isona i Conca Della, Lleida, Spain article info abstract Article history: In the present paper, the fossil record of the archosaurs (dinosaurs, crocodylomorphs and pterosaurs) of Received 3 June 2015 the southern Pyrenees before the CretaceousePalaeogene (KePg) transition is revised. On the basis of Received in revised form this fossil record, a well-dated succession of dinosaurs and other archosaurs is established within polarity 23 June 2015 magnetochrons C30 and C29r.
  • The Tetrapod Fossil Record from the Uppermost

    The Tetrapod Fossil Record from the Uppermost

    geosciences Review The Tetrapod Fossil Record from the Uppermost Maastrichtian of the Ibero-Armorican Island: An Integrative Review Based on the Outcrops of the Western Tremp Syncline (Aragón, Huesca Province, NE Spain) Manuel Pérez-Pueyo 1,* , Penélope Cruzado-Caballero 1,2,3,4 , Miguel Moreno-Azanza 1,5,6 , Bernat Vila 7, Diego Castanera 1,7 , José Manuel Gasca 1 , Eduardo Puértolas-Pascual 1,5,6, Beatriz Bádenas 1 and José Ignacio Canudo 1 1 Grupo Aragosaurus-IUCA, Facultad de Ciencias, Universidad de Zaragoza, C/Pedro Cerbuna, 12, 50009 Zaragoza, Aragón, Spain; [email protected] (P.C.-C.); [email protected] (M.M.-A.); [email protected] (D.C.); [email protected] (J.M.G.); [email protected] (E.P.-P.); [email protected] (B.B.); [email protected] (J.I.C.) 2 Área de Paleontología, Departamento de Biología Animal, Edafología y Geología, Universidad de La Laguna, Citation: Pérez-Pueyo, M.; 38200 San Cristóbal de La Laguna, Santa Cruz de Tenerife, Spain 3 Cruzado-Caballero, P.; Instituto de Investigación en Paleobiología y Geología (IIPG), Universidad Nacional de Río Negro, Moreno-Azanza, M.; Vila, B.; 8500 Río Negro, Argentina 4 IIPG, UNRN, Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Castanera, D.; Gasca, J.M.; 2300 Buenos Aires, Argentina Puértolas-Pascual, E.; Bádenas, B.; 5 GEOBIOTEC, Department of Earth Sciences, NOVA School of Science and Technology, Campus de Caparica, Canudo, J.I. The Tetrapod Fossil 2829-516 Caparica, Portugal Record from the Uppermost 6 Espaço Nova Paleo, Museu de Lourinhã, Rua João Luis de Moura 95, 2530-158 Lourinhã, Portugal Maastrichtian of the Ibero-Armorican 7 Institut Català de Paleontologia Miquel Crusafont, Edifici Z, C/de les Columnes s/n, Campus de la Island: An Integrative Review Based Universitat Autònoma de Barcelona, Cerdanyola del Vallès, 08193 Barcelona, Spain; [email protected] on the Outcrops of the Western Tremp * Correspondence: [email protected] Syncline (Aragón, Huesca Province, NE Spain).
  • Palaeobiodiversity of Crocodylomorphs from the Lourinhã Formation Based on the Tooth Record: Insights Into the Palaeoecology of the Late Jurassic of Portugal

    Palaeobiodiversity of Crocodylomorphs from the Lourinhã Formation Based on the Tooth Record: Insights Into the Palaeoecology of the Late Jurassic of Portugal

    applyparastyle “fig//caption/p[1]” parastyle “FigCapt” Zoological Journal of the Linnean Society, 2019, XX, 1–35. With 17 figures. Downloaded from https://academic.oup.com/zoolinnean/advance-article-abstract/doi/10.1093/zoolinnean/zlz112/5648910 by Boston University user on 02 December 2019 Palaeobiodiversity of crocodylomorphs from the Lourinhã Formation based on the tooth record: insights into the palaeoecology of the Late Jurassic of Portugal ALEXANDRE R. D. GUILLAUME1,2,*, , MIGUEL MORENO-AZANZA1,2, EDUARDO PUÉRTOLAS-PASCUAL1,2, and OCTÁVIO MATEUS1,2, 1GeoBioTec, Faculdade de Ciências e Tecnologia da Universidade Nova de Lisboa, Caparica, Portugal 2Museu da Lourinhã, Lourinhã. Portugal Received 7 March 2019; revised 19 August 2019; accepted for publication 28 August 2019 Crocodylomorphs were a diverse clade in the Late Jurassic of Portugal, with six taxa reported to date. Here we describe 126 isolated teeth recovered by screen-washing of sediments from Valmitão (Lourinhã, Portugal, late Kimmeridgian–Tithonian), a vertebrate microfossil assemblage in which at least five distinct crocodylomorph taxa are represented. Ten morphotypes are described and attributed to five clades (Lusitanisuchus, Atoposauridae, Goniopholididae, Bernissartiidae and an undetermined mesoeucrocodylian). Four different ecomorphotypes are here proposed according to ecological niches and feeding behaviours: these correspond to a diet based on arthropods and small vertebrates (Lusitanisuchus and Atoposauridae), a generalist diet (Goniopholididae), a durophagous diet (Bernissartiidae) and a carnivorous diet. Lusitanisuchus mitracostatus material from Guimarota is here redescribed to achieve a better illustration and comparison with the new material. This assemblage shares similar ecomorphotypes with other Mesozoic west-central European localities, where a diversity of crocodylomorphs lived together, avoiding direct ecological competition through niche partitioning.
  • Late Cretaceous (Maastrichtian) Chondrichthyes and Osteichthyes from Northeastern Iberia

    Late Cretaceous (Maastrichtian) Chondrichthyes and Osteichthyes from Northeastern Iberia

    Palaeogeography, Palaeoclimatology, Palaeoecology 465 (2017) 278–294 Contents lists available at ScienceDirect Palaeogeography, Palaeoclimatology, Palaeoecology journal homepage: www.elsevier.com/locate/palaeo Late Cretaceous (Maastrichtian) Chondrichthyes and Osteichthyes from northeastern Iberia Alejandro Blanco a,⁎,MártonSzabób,c, Àngel Blanco-Lapaz d, Josep Marmi a a Institut Català de Paleontologia Miquel Crusafont, Universitat Autònoma de Barcelona, C/Escola Industrial 23, 08201 Sabadell, Cataluña, Spain b Hungarian Natural History Museum, Department of Paleontology and Geology, Ludovika tér 2, Budapest 1083, Hungary c MTA-ELTE Lendület Dinosaur Research Group, Pázmány Péter Sétány 1/C, Budapest 1117, Hungary d Institut für Naturwissenschaftliche Archäologie, Universität Tübingen, Rümelinstr. 23, 72070 Tübingen, Germany article info abstract Article history: Intensive sampling for vertebrate microfossils has yielded abundant fish remains in the Maastrichtian units of the Received 21 May 2016 Tremp Formation (southern Pyrenees, Catalonia, Spain). Samples were taken from eight new sites representing Received in revised form 24 October 2016 different palaeoenvironments including coastal wetlands and floodplains, in order to assess the fish diversity and Accepted 25 October 2016 to gain a better understanding of the last dinosaur-dominated ecosystems of northeastern Spain. The results sug- Available online 5 November 2016 gest that a diverse ichthyofauna inhabited these transitional to inland fluvial settings throughout the Maastrichtian, comprising both marine and freshwater taxa. Three different chondrichthyans, eight basal Keywords: Lepisosteiformes neopterygians and at least seven teleostean species were found, the latter being more diverse than in other Phyllodontidae Maastrichtian localities in Europe. Fossil evidence from the studied late Maastrichtian assemblages suggests Osteoglossiformes that teleosteans were present in all the trophic guilds.
  • Peat-Forming Plants in the Maastrichtian Coals of the Eastern Pyrenees

    Peat-Forming Plants in the Maastrichtian Coals of the Eastern Pyrenees

    Geologica Acta, Vol.10, Nº 2, June 2012, 189-207 DOI: 10.1344/105.000001711 Available online at www.geologica-acta.com Peat-forming plants in the Maastrichtian coals of the Eastern Pyrenees 1 1 * 2 3 4 S. VILLALBA-BREVA C. MARTÍN-CLOSAS J. MARMI B. GOMEZ M.T. FERNÁNDEZ-MARRÓN 1 Departament d’Estratigrafia, Paleontologia i Geociències Marines, Facultat de Geologia, Universitat de Barcelona (UB) 08028, Barcelona, Catalonia, Spain 2 Institut Català de Paleontologia C/ Escola Industrial 23, 08201, Sabadell, Catalonia, Spain 3 CNRS-UMR 5125, Université Lyon 1 69622, Villeurbanne cedex, France 4 UEI de Paleontología. Instituto de Geociencias (CSIC-UCM) 28040, Madrid, Spain * Corresponding author E-mail: [email protected] ABSTRACT The Lower Maastrichtian of Fumanya and neighbouring localities of the Vallcebre syncline (Eastern Pyrenees, Catalonia, Spain) provide the first taphonomic evidence for the hypothesis that cheirolepidiacean conifers were significant precursors of Maastrichtian Pyrenean coal. Most Frenelopsis-rich lignite beds do not bear rootlet marks, suggesting that the original peat was detrital. Sedimentological and taphonomic evidence indicates deposition on the margins of a lagoon after the transport of the cheirolepidiacean remains by flotation. The same parautochthonous assemblage includes complete impressions of Sabalites longirhachis leaves and large impressions of logs attributed to the same palm trees. Other parautochthonous or allochthonous plant megaremains include extremely rare cycadalean and monocot leaves and abundant minute angiosperm seeds. Rootlet marks associated with thin lignite beds occur at the top of some charophyte limestones. The charophyte association, dominated by in situ accumulation of Peckichara and Microchara gyrogonites, suggests that these limestones were deposited in shallow, freshwater lakes and that the corresponding peat mires were limnic rather than paralic in nature.
  • CORE View Metadata, Citation and Similar Papers at Core.Ac.Uk

    CORE View Metadata, Citation and Similar Papers at Core.Ac.Uk

    View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by RERO DOC Digital Library Abstract Book-INTERNATIONAL SYMPOSIUM ON DINOSAURS AND OTHER VERTEBRATES PALAEOICHNOLOGY (2005) FUMANYA, BARCELONA 2 Abstract Book-INTERNATIONAL SYMPOSIUM ON DINOSAURS AND OTHER VERTEBRATES PALAEOICHNOLOGY (2005) FUMANYA, BARCELONA Organizing Committee Jean Le Loeuff, Musée des Dinosaures, Espéraza Bernat Vila, Consorci Ruta Minera, PhD student Josep Marmi, Consorci Ruta Minera Àngel Galobart, Institut de Paleontologia M. Crusafont de Sabadell Oriol Oms, Dep. Geologia (Estratigrafia), Fac. de Ciencies Univ. Autònoma de Barcelona Organizing Institutions Consorci Ruta Minera Dinosauria by means of European Project INTERREG-IIIA funds Collaborations Museu de les Mines de Cercs Cercs Town council Diputació de Barcelona Universitat Autònoma de Barcelona Fundación Patrimonio Paleontológico de La Rioja Museo Geominero (I.GM.E.) Obra Social “Fundació La Caixa” Scientific Committee Eric BUFFETAUT - Université Paris, France F.M. DALLA VECCHIA - Museo Paleontologico Cittadino, Monfalcone, Italy Àngel GALOBART- Institut de Paleontologia de Sabadell, Barcelona José Carlos GARCÍA-RAMOS - Universidad de Oviedo, España Jean LE LOEUFF - Musée des Dinosaures, Espéraza, France Martin G. LOCKLEY - University of Colorado, Denver, USA José Joaquin MORATALLA- Museo Geominero, (IGME), Madrid, España Oriol OMS- Universitat Autònoma de Barcelona Félix PÉREZ-LORENTE - Universidad de La Rioja , España Vanda Faria dos SANTOS - Museo Nacional de Historia