On the Arenig and Llandeilo Groups
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Evolutionary Patterns of Trilobites Across the End Ordovician Mass Extinction
Evolutionary Patterns of Trilobites Across the End Ordovician Mass Extinction by Curtis R. Congreve B.S., University of Rochester, 2006 M.S., University of Kansas, 2008 Submitted to the Department of Geology and the Faculty of the Graduate School of The University of Kansas in partial fulfillment on the requirements for the degree of Doctor of Philosophy 2012 Advisory Committee: ______________________________ Bruce Lieberman, Chair ______________________________ Paul Selden ______________________________ David Fowle ______________________________ Ed Wiley ______________________________ Xingong Li Defense Date: December 12, 2012 ii The Dissertation Committee for Curtis R. Congreve certifies that this is the approved Version of the following thesis: Evolutionary Patterns of Trilobites Across the End Ordovician Mass Extinction Advisory Committee: ______________________________ Bruce Lieberman, Chair ______________________________ Paul Selden ______________________________ David Fowle ______________________________ Ed Wiley ______________________________ Xingong Li Accepted: April 18, 2013 iii Abstract: The end Ordovician mass extinction is the second largest extinction event in the history or life and it is classically interpreted as being caused by a sudden and unstable icehouse during otherwise greenhouse conditions. The extinction occurred in two pulses, with a brief rise of a recovery fauna (Hirnantia fauna) between pulses. The extinction patterns of trilobites are studied in this thesis in order to better understand selectivity of the -
001-012 Primeras Páginas
PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. -
The Ordovician Trilobite Genus Robergia Wiman, 1905 and Some Other Species Hitherto Included
The Ordovician trilobite genus Robergia Wiman, 1905 and some other species hitherto included FRANK NIKOLAISEN Nikolaisen, F.: The Ordovician trilobite genus Robergia Wiman, 1905 and some other species hitherto included. Norsk Geologisk Tidsskrift, Vol. 71, pp. 37-62. Oslo 1991. ISSN 0029-196X The genus Robergia Wiman, 1905 is revised together with type and additional material including R. striatel/a sp. nov. from the upper Llandeilo/Iower Caradoc of Jiimtland, Sweden. This species is similar to both older and younger forms from North Arnerica, and 'European' and 'American' stocks are recognized. Pugilator gen. nov. is erected for late Ordovician species with short palpebral lobes and a large glabellar tongue, and Arator gen. nov. for species from Kazakhstan, U.S.S.R. , with a narrow glabellar tongue and a falcate , prominent anterior border. Type material of R. breviceps Raymond, 1925 is illustrated for the first time and the species assigned to Eorobergia. Frank Nikolaisen, Paleontologisk Museum, Sars gate l, N-{)562 Oslo 5, Norway. On a visit to the Palaeontological Institute at the Uni ammonium chloride before photographing, using the versity of Uppsala in 1968 I received on loan from the late technique describ,ed by Whittington (1956). Professor Per Thorslund some excellent remopleuridid Magnesian oxychloride replicas of figured type sped material from Jamtland, including that of an undescribed mens from Kazakhstan have been made from 'Anto species of Robergia. It was agreed that this should be germetic' rubber paste casts using the method described included with Norwegian material I was revising, but by Stitt (1982). Illustrated cranidia are oriented with the this was not done (Nikolaisen 1983). -
Paleozoic and Quaternary Geology of the St. Louis
Paleozoic and Quaternary Geology of the St. Louis Metro East Area of Western Illinois 63 rd Annual Tri-State Geological Field Conference October 6-8, 2000 Rodney D. Norby and Zakaria Lasemi, Editors ILLINOIS STATE GEOLOGICAL SURVEY Champaign, Illinois ISGS Guidebook 32 Cover photo Aerial view of Casper Stolle Quarry at the upper right, Falling Springs Quarry at the left, and the adjacent rural residential subdivision at the right, all situated at the northernmost extent of the karst terrain in St. Clair County, Illinois. Photograph taken April 7, 1988, U.S. Geological Survey, National Aerial Photography Program (NAPP1). ILLINOIS DEPARTMENT OF NATURAL RESOURCES © printed with soybean ink on recycled paper Printed by authority of the State of Illinois/2000/700 Paleozoic and Quaternary Geology of the St. Louis Metro East Area of Western Illinois Rodney D. Norby and Zakaria Lasemi, Editors and Field Conference Chairmen F. Brett Denny, Joseph A. Devera, David A. Grimley, Zakaria Lasemi, Donald G. Mikulic, Rodney D. Norby, and C. Pius Weibel Illinois State Geological Survey Joanne Kluessendorf University of Illinois rd 63 Annual Tri-State Geological Field Conference Sponsored by the Illinois State Geological Survey October 6-8, 2000 WISCONSIN ISGS Guidebook 32 Department of Natural Resources George H. Ryan, Governor William W. Shilts, Chief Illinois State Geological Survey 615 East Peabody Drive Champaign, IL 61820-6964 (217)333-4747 http://www.isgs.uiuc.edu Digitized by the Internet Archive in 2012 with funding from University of Illinois Urbana-Champaign http://archive.org/details/paleozoicquatern32tris 51 CONTENTS Middle Mississippian Carbonates in the St. Louis Metro East Area: Stratigraphy and Economic Significance Zakaria Lasemi and Rodney D. -
Late Silurian Trilobite Palaeobiology And
LATE SILURIAN TRILOBITE PALAEOBIOLOGY AND BIODIVERSITY by ANDREW JAMES STOREY A thesis submitted to the University of Birmingham for the degree of DOCTOR OF PHILOSOPHY School of Geography, Earth and Environmental Sciences University of Birmingham February 2012 University of Birmingham Research Archive e-theses repository This unpublished thesis/dissertation is copyright of the author and/or third parties. The intellectual property rights of the author or third parties in respect of this work are as defined by The Copyright Designs and Patents Act 1988 or as modified by any successor legislation. Any use made of information contained in this thesis/dissertation must be in accordance with that legislation and must be properly acknowledged. Further distribution or reproduction in any format is prohibited without the permission of the copyright holder. ABSTRACT Trilobites from the Ludlow and Přídolí of England and Wales are described. A total of 15 families; 36 genera and 53 species are documented herein, including a new genus and seventeen new species; fourteen of which remain under open nomenclature. Most of the trilobites in the British late Silurian are restricted to the shelf, and predominantly occur in the Elton, Bringewood, Leintwardine, and Whitcliffe groups of Wales and the Welsh Borderland. The Elton to Whitcliffe groups represent a shallowing upwards sequence overall; each is characterised by a distinct lithofacies and fauna. The trilobites and brachiopods of the Coldwell Formation of the Lake District Basin are documented, and are comparable with faunas in the Swedish Colonus Shale and the Mottled Mudstones of North Wales. Ludlow trilobite associations, containing commonly co-occurring trilobite taxa, are defined for each palaeoenvironment. -
Phylogenetic Analyses of Trilobites from the Cambrian and Ordovician Radiations
Phylogenetic analyses of trilobites from the Cambrian and Ordovician radiations by Copyright 2014 Ian Wesley Gapp Submitted to the graduate degree program in Geology and the Graduate Faculty of the University of Kansas in partial fulfillment of the requirements for the degree of Doctor of Philosophy. ________________________________ Chairperson: Bruce S. Lieberman ________________________________ Alison Olcott Marshall ________________________________ Jennifer A. Roberts ________________________________ Paul A. Selden ________________________________ Edward O. Wiley Date Defended: January 27, 2014 The Dissertation Committee for Ian Wesley Gapp certifies that this is the approved version of the following dissertation: Phylogenetic analyses of trilobites from the Cambrian and Ordovician radiations ________________________________ Chairperson: Bruce S. Lieberman Date approved: January 27, 2014 ii Abstract Trilobites are an ideal group used for phylogenetic studies due to their complex morphology and abundance in the Paleozoic fossil record. Because of this, trilobite phylogenetics has become an important component of understanding macroevolutionary patterns during key evolutionary radiations. Two radiations that this dissertation investigates are the Cambrian and Ordovician radiations. While both occur during the early Paleozoic, they present very different patterns. The Cambrian radiation was the sudden appearance of all major metazoan phyla, whereas in the Ordovician, those new beauplans were already in place and there was instead a great increase in diversity among those groups. Presented here are phylogenetic analyses of trilobites from those radiations to further our understanding of the evolutionary patterns occurring at those times. iii Acknowledgements It has been an incredible six and a half years at The University of Kansas, here in Lawrence. I have met many fantastic people during my time here and want to be sure that several people get the recognition that they deserve. -
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https://doi.org/10.24199/j.mmv.1985.46.03 20 July 1985 TREMADOC TR1LOBITES OF THE DIGGER ISLAND FORMATION, WARATAH BAY, VICTORIA By P. A. Jell Museum of Victoria, 285-321 Russell Street, Melbourne, Victoria 3000 Abstract Trilobites of the Digger Island Formation at Digger Island, 1 .5 km south of Walkerville on Waratah equivalent to the Bay . South Gippsland are described and assigned an early Tremadoc age approximately Kaineila meridionalis zone of Argentina. It is impossible to correlate directly with any known Australian and sequence but indirectly it is considered older than La 1 .5 zone of the Victorian graptolite sequence ap- proximately contemporaneous with the Oneolodus bkuspaius-Drepanodus simplex zone of western Queensland. Four new genera, Natmus (Hystricuridae), Barachyhipposiderus (Harpedidae), and Landyia and lictorispina (Pilekiidae) are erected with eleven new species, N. viclns, N. tuberus, B. logimus, L. elizabelhae, V. holmesorum, Neougnosms eckurdti, Onychopyge parkerae, Pseudokainellu diggerensis, Austraioharpes singleioni, A. expansus, and Proioplionwrops lindneri. New taxa left in open nomenclature are referred 10 Pilekia. Tessalacauda, and the Hystricuridae. The Argentinian species Micragnosius hoeki (Kobayashi, 1939), Slmmardia erquemis Kobayashi, 1937, and Leiostegium douglasi Harrington, 1937 are identified, Introduction from samples taken some distance along strike trilobite locality; they come from near Digger Island is a small stack approximately from the Lindner fig. 3) (D. J. Ken- 75 m in diameter, isolated from ihe mainland locality 2 of (1953, using above half-tide, and situated approximately nedy pets, coram.). Webby et al. (1981) Leiostegium made a direct correla- 1.5 km south of Walkerville on the western Kaineila and Kainella-Leiostegium zone shore of Waratah Bay, South Gippsland; it lion between the of Ross (1951) and Hintze consists of brown, largely decalcified mud- (i.e. -
The Brachymetopid Trilobite Radnoria in the Silurian (Wenlock) of New York State and Arctic Canada
2087 The brachymetopid trilobite Radnoria in the Silurian (Wenlock) of New York State and Arctic Canada Jonathan M. Adrain and Denis K. Tetreault Abstract: Radnoria bretti n. sp., from the Wenlock (Sheinwoodian) Rochester Formation of western New York State, is the best preserved and most completely known member of its genus. It provides the first definitive information on Radnoria’s hourglass-shaped rostral plate, the first known hypostome of a Silurian member of the genus, demonstrates that Radnoria engaged in sphaeroidal enrollment, and reveals that early holaspid individuals had tubercles on the posterior thoracic axes and pygidial axial rings that were effaced with maturity. Three new species from the Wenlock of the Cape Phillips Formation of Nunavut are known from sparse material and are reported in open nomenclature. Together, the species greatly increase knowledge of Laurentian Silurian brachymetopids, which have until now been known from a single cranidium from the Wenlock of Arkansas. Résumé : Radnoria bretti n. sp., de la Formation de Rochester du Wenlock (Sheinwoodien) de l’Ouest de l’État de New York, est le membre le mieux préservé et dont la description est la plus complète de ce genre. Il fournit les premières données définitives sur la plaque rostrale en forme de sablier de Radnoria, le premier hypostome connu d’un membre d’âge silurien de ce genre, démontre que Radnoria était capable d’enroulement sphéroïdal, et révèle que les individus holaspides précoces présentaient des tubercules sur leurs axes thoraciques postérieurs et des anneaux axiaux pygidiaux qui s’effaçaient chez les individus à maturité. Trois nouvelles espèces provenant de matériel dispersé du Wenlock de la Formation de Cape Phillips, au Nunavut, sont connues et signalées en nomenclature ouverte. -
Sborník Národního Museav PRAZE ACTA MUSEI NATIONALIS PRAGAE Vol
SBORNíK NÁRODNíHO MUSEAv PRAZE ACTA MUSEI NATIONALIS PRAGAE Vol. III. B (1947) No 1. Geologia et Paleontologia No. !1. FERDINAND PRANTL a ALOIS PRIBYL: Roztřídění některých českých Cheiruridů. (Trilobitae.) Classification of some Bohemian Cheiruridae. (Trilobitae.) (Předloženo 4. března 1947.) Tato práce je věnována novému systematickému roztřídění českých cheiruridních trilobitů z podčeledi Cheirurinae (HAWLE & CORDA), emend. RAYMOND, 1913 a Cyrtometopinae ÚPIK, 1937. J. BARRANDE (1846; 1850; 1852 a 1872) zahrnoval většinu sem náležejících druhů do jediného rodu - Cheirurus BEYRICH, 1845. Rody Actinopeltis HAWLE & CORDA, 1847 a Eccoptochile HAWLE & CORDA, 1847 pokládal při tom za totožné s rodem výše zmíněným. Později přeřadilO. NOVÁK (1890) některé české druhy k SALTEROVU podrodu Crotalocephalus SALTER, 1853. V novější době oddělil C. D. BARTON dva BARRANDOVY druhy (Chei rurus comes a Ch. vinculum) do zvláštních rodů Ceraurinus BARTON, 1913 a Lehua BARTON, 1915. R. Růžr.čKA (1926; 1934) popsal z českého ordoviku dvě nové cyrtometopinní formy pod rodovým jménem Cyrto metopus ANGELIN, 1854, které však T. KOBAYASHI (1934; 1935) přeřadil do svého nového rodu Parapilekic~ KOBAYASHI, 1934. Jinak se však v po slední době nikdo, s výjimkou starších prací REEDOVÝCH (1896; 1898) a práce C. D. BARTONA (1915), otázkou systematické příslušnosti českých cheirurinních a cyrtometopinních trilobitů nezabýval. Proto si všímáme i celkové klasifikace čeledi Cheiru'ridae (HAWLE & CORDA), emend. RAYMOND, 1913. Rozdělujeme ji do pěti podčeledí, a to: Cheirurinae RAYMOND, 1913; Cyrtometopinae ÚPIK, 1937; Dei phoninae RAYMOND, 1913; Sphaeroxochinae ÚPIK, 1937 a Areiinae nov. subfam. Rod Staurocerphalus BARRANDE, 1846, který byl až dosud kladen rovněž do čeledi Cheiruridae, přeřazujeme současně do če l edi Encri nuridae ANGELIN, 1854 a tvoříme pro něj novou podčeleď Staurocepha linae nov. -
Early Silurian Phacopide Trilobites from Central Victoria, Australia
Memoirs of Museum Victoria 63(2): 215–255 (2006) ISSN 1447-2546 (Print) 1447-2554 (On-line) http://www.museum.vic.gov.au/memoirs/index.asp Early Silurian phacopide trilobites from central Victoria, Australia ANDREW C. SANDFORD1 AND DAV I D J. HOLLOWAY2 1PO BOX 2682, CHELTENHAM , VICTORIA 3192, AUSTRALIA ([email protected] ). 2 MUSEUM VICTORIA, PO BOX 666, MELBOURNE, VICTORIA 3001, AUSTRALIA ([email protected]). Abstract Sandford, A.C. and Holloway, D.J. 2006. Early Silurian phacopide trilobites from central Victoria, Australia. Memoirs of Museum Victoria 63(2): 215–255. A diverse component of trilobite faunas in lower Silurian marine strata of central Victoria are members of the suborder Phacopina, here assigned to 19 species within the Phacopidae, Dalmanitidae and Acastidae. New genera are represented by the blind acastid Berylacaste berylae gen. et sp. nov. and the phacopid Ivops wallanensis gen. et sp. nov. Other new species are the phacopids Acernaspis georgei and Ananaspis kenleyi, and the dalmanitids Preodontochile springfi eldensis and Struveria? plinthourgos. Two previously described species, Ananaspis typhlagogus Öpik, 1953 and Dalmanites athamas Öpik, 1953 [= ʻDalmanitina (Eudolatites)ʼ aborigenum Öpik, 1953], are revised. Other species are assigned to Bessazoon, Phacopidella? and indeterminate phacopid and dalmanitid genera. These trilobites are a signifi cant representation of early Silurian Phacopina on the world scale. Keywords Trilobita, Silurian, Phacopidae, Dalmanitidae, Acastidae, Victoria, Australia, systematics, -
On the Silurian Rocks of the English Lake District
H. HICKS ON CAMBRIAN AND SILURIAN CLASSIFICATION. 105 The Arenig Group, which comes in next in the Table, is divided into an Upper and a Lower series, both characterised by a distinct assemblage of fossils, The Lower series occurs chiefly in the neighbourhood of St. David's, and is of about 1,000 feet in thick ness. The fossils consist of numerous graptolites, especially dendroid forms, and trilobites of the genera Ogygia, Trinucleue, and .k:glina. All the forms are thoroughly Silurian in character, and the series marks a definite line for the base of the Silurian groups. The Upper series comprise the well known Skiddaw Slates. Upon the Arenig comes the Llandeilo Group, the lowest rocks recognised by Sir R. Murchison in the typical Llandeilo district. This group in the Table is divided also into an Upper and Lower series, marked out by distinct faunas. The Bala or Caradoc Group now comes in with its exceedingly rich fauna, and it forms in the Table the upper member of our Lower Silurian, and also of Professor Sedgwick's Upper Cambrian. Sir R. Murchison added to the Lower Silurian the Lower Llandovery rocks, but Sir Charles Lyell in his" Student's Manual of Geology," 1871, preferred "to regard them as the base of the Upper Silurian, to which group they are linked by more than twice as many species as to the Lower Silurian." The evidence of Professor Hall and others, in America, tends to prove, however, that ultimately it will be necessary to make a Middle Silurian division, and though the group is scarcely important enough, as hitherto known in this country, to warrant such a division, yet should a spot be found where the present unconformity is filled up, a series may be discovered, like that in the American continent, sufficiently large to enable it to be done. -
A Pictorial Guide to the Orders of Trilobites
A PICTORIAL GUIDE TO THE ORDERS OF TRILOBITES by Samuel M. Gon III, Ph.D. ORDER ORDER PROETIDA ASAPHIDA Superfamily Superfamily Superfamily Superfamily Superfamily Superfamily Superfamily Superfamily Superfamily Aulacopleuroidea Bathyuroidea Proetoidea Anomocaroidea Asaphoidea Trinucleioidea Dikelocephaloidea Cyclopygoidea Remopleuridoidea Aulacopleuridae Bathyuridae Proetidae Anomocarellidae Asaphidae Trinucleidae Dikelocephalidae Cyclopygidae Remopleurididae Aulacopleura Goniotelus Cyphoproetus Glyphaspis Ogygiocaris Cryptolithus Dikelocephalus Cyclopyge Remopleurides ORDER ORDER ORDER ORDER HARPETIDA CORYNEXOCHIDA LICHIDA ODONTOPLEURIDA Suborder Suborder Suborder Suborder Superfamily Superfamily Superfamily Harpetina Corynexochina Illaenina Leiostegiina Lichoidea Dameselloidea Odontopleuroidea Harpetidae Dorypygidae Illaenidae Kaolishaniidae Lichidae Damesellidae Odontopleuridae Harpes Olenoides Bumastus Mansuyia Arctinurus Drepanura Kettneraspis ORDER ORDER PHACOPIDA PTYCHOPARIIDA Suborder Suborder Suborder Suborder Suborder Calymenina Cheirurina Phacopina Ptychopariina Olenina Calymenidae Cheiruridae Phacopidae Acastidae Tricrepicephalidae Ptychopariidae Ellipsocephalidae Olenidae Calymene Ceraurus Phacops Kayserops Tricrepicephalus Modocia Ellipsocephalus Olenus ORDER ORDER AGNOSTIDA REDLICHIIDA Suborder Suborder Suborder Suborder Agnostina Eodiscina Olenellina Redlichiina Agnostidae Condylopygidae Eodiscidae Olenellidae Fallotaspididae Redlichiidae Paradoxididae Xystriduridae Emuellidae Agnostus Pleuroctenium Pagetia Paedeumias