Tema 23: Clado Malvids (1): Familia Malvaceae

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Tema 23: Clado Malvids (1): Familia Malvaceae Tema 23: Clado Malvids (1): Familia Malvaceae Prof. Francisco J. García Breijo Unidad Docente de Botánica Dep. Ecosistemas Agroforestales Escuela Técnica Superior del Medio Rural y Enología Universidad Politécnica de Valencia Diapositiva nº: 1 Taxonomía -1 al clado Eudicotiledóneas (A.P.G. III, 2009) Clado Eudicots nucleares Clado Rósidas Clado Eurósidas II (Malvids) . Orden Malvales (Juss. Ex Bercht. & J.Presl, 1820) . Orden Brassicales (Bromhead, 1838) . Orden Sapindales (Juss. Ex Bercht. & J.Presl, 1820) a la subclase Dilleniidae (Takhtajan, 1997; Cronquist, 1981). Familia Malváceas ©: Francisco José García Breijo Diapositiva 2 Unidad Docente de Botánica. ETSMRE, UPV Taxonomía -2 Pertenece al: Orden Malvales (Jussieu ex Berchtold & J. Presl, 1820) (A.P.G. III, 2009; Thorne, 1992; Dahlgren, 1985; Cronquist, 1981) Superorden Malvanae, orden Malvales (Takhtajan, 1997). Familia Malváceas ©: Francisco José García Breijo Diapositiva 3 Unidad Docente de Botánica. ETSMRE, UPV Generalidades Malvaceae (Jussieu, 1789). Familia cosmopolita de hierbas, raramente arbustos o árboles. Usos económicos: cultivos para obtención de fibras (algodón, yute de China) y ornamentales (hibiscos, malva locas, ...) Familia Malváceas ©: Francisco José García Breijo Diapositiva 4 Unidad Docente de Botánica. ETSMRE, UPV Distribución Caracteres diagnósticos -1 HÁBITO: Hierbas, arbustos o árboles. Mesófitas o xerófitas. HOJAS: Pecioladas, alternas, simples y con estípulas pequeñas y caducas. Lámina simple o palmatilobadas. Estomas en ambas caras; anomocíticos. Tricomas complejos y glandulares. REPRODUCCIÓN y POLINIZACIÓN: Plantas hermafroditas (o dióicas). Polinización entomófila. Familia Malváceas ©: Francisco José García Breijo Diapositiva 6 Unidad Docente de Botánica. ETSMRE, UPV Caracteres diagnósticos -2 INFLORESCENCIAS: Flores conspicuas, solitarias o en inflorescencias cimosas (cincinios) Flores regulares con nectarios. Cíclicas. CÁLIZ: 5 sépalos, libres o soldados, a menudo con calículo (epicáliz). COROLA: 5 pétalos, libres pero individualmente unidos a la columna de estambres. Pétalos contornados o imbricados. Familia Malváceas ©: Francisco José García Breijo Diapositiva 7 Unidad Docente de Botánica. ETSMRE, UPV Caracteres diagnósticos -3 ANDROCEO: Muchos estambres (5-)15-100, centrífugos, con los filamentos unidos formando un tubo que envuelve al pistilo (monoadelfos); anteras uniloculares. GINECEO: Carpelos de (1-)5(-100), soldados; ovario súpero con (1-)5(-100) lóculos y 1 o varios óvulos axilares; 1 estilo, más o menos dividido. FRUTO: en cápsula o en esquizocarpo. Familia Malváceas ©: Francisco José García Breijo Diapositiva 8 Unidad Docente de Botánica. ETSMRE, UPV Caracteres diagnósticos -4 Fisiología: Plantas C3. Bioquímica. Cianogénicas o no. Proantocianidinas presentes: cianidina y delfinidina. Flavonoles presentes: quercetina, kenferol. Proantocianidinas presentes: cianidina y delfinidina. Arbutina y ácido elágico ausentes. Alcaloides, saponinas e iridoides ausentes. Familia Malváceas ©: Francisco José García Breijo Diapositiva 9 Unidad Docente de Botánica. ETSMRE, UPV Caracteres diagnósticos -5 FÓRMULA FLORAL: ⊕ K(5) [C5 A(∞)] G (5- ∞) Familia Malváceas ©: Francisco José García Breijo Diapositiva 10 Unidad Docente de Botánica. ETSMRE, UPV Caracteres diagnósticos -6 DIAGRAMA FLORAL Familia Malváceas ©: Francisco José García Breijo Diapositiva 11 Unidad Docente de Botánica. ETSMRE, UPV Cápsula abierta de Rama con hojas simples y flores de Hoja palmeada de Lavatera Cápsula de Lagunaria Gossypium sp. Lagunaria patersonii (Andrews) G.Don assurgentiflora Kellog patersonii (Andrews) G.Don Pétalos Gineceo (el estilo pasa a través de la columna de estambres) Columna de Esquizocarpo de estambres (los Lavatera filamentos están soldados: assurgentiflora Kellog monoadelfos) Anteras uniloculares y Sépalos granos de polen Flor de Lavatera Flor de Lagunaria assurgentiflora Kellog patersonii (Andrews) G.Don La flor de las Malváceas Pétalos Estigma ramificado Estambres monoadelfos Sépalos Ovario con varios óvulos Receptáculo Calículo Dibujo de una sección longitudinal de una flor de Pedúnculo floral Gossypium hirsutum L. (algodón) Estilo Tubo estaminal monoadelfo Pétalo Lugar de unión de los pétalos al tubo estaminal Sépalo Ovario súpero con placentación axial Epicáliz Sección Longitudinal de una flor de Hibiscus sp. La flor de las Malváceas Estigma ramificado Estilo Estambres monoadelfos Fotografía de los estambres monoadelfos y de los estigmas ramificados en trompetilla típicos de la flor de Hibiscus sp. Géneros y especies -1 243 géneros y 4225 especies (APG III). Géneros y especies importantes por sus fibras: Género Gossypium: G. hirsutum L., G. barbadense L. y G. herbaceum L.(especies productoras de algodón). Género Abutilon: Especie Abutilon avicennae Gaertn (Yute de China) Familia Malváceas ©: Francisco José García Breijo Diapositiva 16 Unidad Docente de Botánica. ETSMRE, UPV Fruto maduro con 5 partes, dentro de las cuales se encuentran muchas semillas densamente cubiertas con pelos epidérmicos. Cada semilla se encuentra recubiertas de abundantes pelos epidérmicos (algodón). Dibujo de una planta de algodón (Gossypium hirsutum L.) mostrando las hojas, una flor madura y varios frutos en cápsula. Fibras de algodón Calículo Lámina que muestra los caracteres morfológicos del (7) Gossypium barbadense L. (5) (A) Rama florífera. (4) (1) Androceo, gineceo y calículo. (2) Detalle antera. (3) ST de la antera. (A) (4) SL del ovario. (5) ST del ovario (6) Fruto (cápsula). (6) (7) Semilla (8) SL de una semilla (1) (3) (2) (8) Géneros y especies -2 Géneros y especies importantes por su uso en ornamentación: Género Malva Género Hibiscus Género Lavatera Género Althaea Género Abutilon Género Sida Familia Malváceas ©: Francisco José García Breijo Diapositiva 29 Unidad Docente de Botánica. ETSMRE, UPV Género Malva Género de unas 25 especies de plantas herbáceas. Este género se encuentra localizado en las zonas templadas, subtropical y tropical de África, Asia y Europa. Las especies de Malva las usan como fuente de alimentación las larvas de algunas especies de Lepidópteros. Especies representativas: Malva sylvestris L. Familia Malváceas ©: Francisco José García Breijo Diapositiva 30 Unidad Docente de Botánica. ETSMRE, UPV (5) (3) (2) (A) Rama florífera de Malva sylvestris (1) Flor en SL. (2) Pétalo. (3) Androceo (est. monoadelfos). (12) (4) Detalle de las anteras. (11) (5) Detalle del polen. (6) Gineceo. (1) (4) (10) (7) Detalle del ovario en ST (8) Detalle del fruto (esquizocarpo). (A) (9) Detalle de un merocarpo. (9) (10) ST de un merocarpo. (11) SL de un merocarpo. (12) Semilla. (8) (6) Lámina que muestra los caracteres morfológicos de la malva silvestre (7) (Malva sylvestris L.) Género Lavatera Género de unas 25 especies de plantas. Nativo de la región del Mediterráneo, este y centro de Asia, y Australia. Las especies de Lavatera las usan como fuente de alimentación las larvas de algunas especies de Lepidópteros. Especies representativas: Lavatera cretica L. Lavatera arborea L. Lavatera bicolor L. Familia Malváceas ©: Francisco José García Breijo Diapositiva 33 Unidad Docente de Botánica. ETSMRE, UPV (A) Rama florífera (1) Flor. (2) Esquizocarpo y calículo (3) Merocarpos (1) (A) (2) (3) Lámina que muestra el aspecto de la Lavatera arborea L. Género Hibiscus Son los hibiscos, llamados cayena en latinoamérica, forman un amplio género de alrededor de 220 especies, típicas de ambientes cálidos, en regiones tropicales y subtropicales. Especies representativas: Hibiscus rosa-sinensis L. (Rosa china o cayena). Hibiscus cannabinus L. (Kenaf) Familia Malváceas ©: Francisco José García Breijo Diapositiva 37 Unidad Docente de Botánica. ETSMRE, UPV (2) (1) (A) Rama florífera (1) Flor (2) Gineceo y androceo monoadelfo. (3) Esquizocarpo protegido por los sépalos (4) SL del esquizocarpo protegido por los sépalos. (A) (5) Merocarpos Lámina que muestra los caracteres morfológicos más importantes de Hibiscus alcea L. (3) (5) (4) Fotografías de flores de Hibisco (Hibiscus rosa-sinensis L.) mostrando las características más sobresalientes de las malváceas. Estambres monoadelfos Calículo Sépalos Estigmas ramificados (5) en forma de trompetilla Pétalos Género Althaea Género de 6-12 especies de plantas herbáceas. Las especies de Althaea las usan como fuente de alimentación las larvas de algunas especies de Lepidópteros. Especies representativas: Althaea officinalis L. (Malvavisco). Familia Malváceas ©: Francisco José García Breijo Diapositiva 43 Unidad Docente de Botánica. ETSMRE, UPV (2) (6) (5) (A) Rama florífera (1) Gineceo y androceo en SL. (2) Androceo monoadelfo. (3) Detalle del gineceo. (4) Vista superior del ovario. (3) (5) Detalle de las anteras. (6) Grano de polen. (4) (7) Esquizocarpo rodeado de sépalos (8) Detalle de un merocarpo y semilla. (9) Secciones de los merocarpos. (1) Lámina de Althaea officinalis L. (7) (9) (8) (A) (6) (1) (A) Rama florífera (1) Gineceo y androceo en SL. (2) (5) (2) Androceo monoadelfo. (3) Detalle de las anteras. (4) (4) Detalle del polen. (5) Gineceo. (3) (6) ST del ovario. (9) (7) Esquizocarpo cubierto de sépalos (8) Esquizocarpo (9) Detalle de merocarpos. (11) (10) SL ST de un merocarpo. (11) Semillas. (10) Lámina de Althaea rosea L. (8) (7) (A) Género Abutilon Gran género de alrededor de 150 especies de plantas de flores. Algunas especies de Sida las usan como fuente de alimentación las larvas de algunas especies de Lepidópteros. Especies representativas: Abutilon
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