Order Family Genus Species Author Year Exx. Region Coleoptera Ceratocanthidae Germarostes aphodioides Illiger 1800 71 nea Coleoptera Ceratocanthidae Germarostes globosus Say 1835 33 nea Coleoptera Ceratocanthidae Germarostes macleayi Perty 1830 1 pal Coleoptera Ceratocanthidae Germarostes plicatus Germar 1843 3 pal Coleoptera Ceratocanthidae Germarostes punctulatus Ohaus 1911 1 pal Coleoptera Ceratocanthidae Haroldostes senegalensis Castelnau 1840 1 pal Coleoptera Diphyllostomatidae Diphyllostoma fimbriata Fall 1901 3 nea Coleoptera Diphyllostomatidae Diphyllostoma linsleyi Fall 1932 6 nea Coleoptera Diphyllostomatidae Diphyllostoma nigricollis Fall 1912 34 nea Coleoptera Geotrupidae Anoplotrupes ballyi Jekel 1865 45 nea Coleoptera Geotrupidae Anoplotrupes hornii Blanchard 1888 13 nea Coleoptera Geotrupidae Anoplotrupes stercorosus Scriba 1791 77 pal Coleoptera Geotrupidae Athyreus excavatus Castelnau 1840 1 pal Coleoptera Geotrupidae Athyreus sp. 1 pal Coleoptera Geotrupidae Blackburnium insigne Lea 1916 2 aus Coleoptera Geotrupidae Blackburnium sloanei Blackburn 1889 1 aus Coleoptera Geotrupidae Bolbelasmmus horni Rivers 1886 1 nea Coleoptera Geotrupidae Bolbocerastes imperialis Cartwright 1953 9 nea Coleoptera Geotrupidae Bolbocerastes regalis Cartwright 1953 1 nea Coleoptera Geotrupidae Bolbocerastes serratus LeConte 1854 10 nea Coleoptera Geotrupidae Bolbocerosoma biplagiatum Dawson and McCollock 1924 4 nea Coleoptera Geotrupidae Bolbocerosoma bruneri Dawson and McCollock 1924 8 nea Coleoptera Geotrupidae Bolbocerosoma confusum Brown 1928 2 nea Coleoptera Geotrupidae Bolbocerosoma farctum Fabricius 1775 9 nea Coleoptera Geotrupidae Bolbocerosoma hamatum Brown 1929 3 Coleoptera Geotrupidae Bolbocerosoma pusillum Dawson and McCollock 1924 1 nea Coleoptera Geotrupidae Bolbocerosoma tumefactum Beauvois 1805 8 nea Coleoptera Geotrupidae Bolbochromus n.sp. 1 ori Coleoptera Geotrupidae Bolborhombus sallaei Bates 1887 9 nea Coleoptera Geotrupidae Bradycinetulus ferrugineus Beauvois 1805 1 nea Coleoptera Geotrupidae Bradycinetulus fossatus Haldeman 1853 5 nea Coleoptera Geotrupidae Ceratophyus polyceros Pallas 1771 2 pal Coleoptera Geotrupidae Cnemotrupes blackburnii Fabricius 1781 28 nea Coleoptera Geotrupidae Cnemotrupes semiopacus Jekel 1865 45 nea Coleoptera Geotrupidae Cnemotrupes splendidus Fabricius 1781 40 nea Coleoptera Geotrupidae Enoplotrupes sharpi Jordan and Rothschild 1893 2 pal Coleoptera Geotrupidae Eogeotrupes auratus Motschulsky 1857 2 Coleoptera Geotrupidae Epigeotrupes laevistriatus Motschulsky 1 pal Coleoptera Geotrupidae Eucanthus alutaceus Cartwright 1944 1 nea Coleoptera Geotrupidae Eucanthus greeni Robinson 6 nea Coleoptera Geotrupidae Eucanthus impressus Howden 1964 10 nea Coleoptera Geotrupidae Eucanthus lazarus Fabricius 1775 109 nea Coleoptera Geotrupidae Eucanthus lazarus 33 nea Coleoptera Geotrupidae Frickius variolosus Germain 1897 6 pal Coleoptera Geotrupidae Geohowdenius egeriei Germar 1824 14 nea Coleoptera Geotrupidae Geohowdenius opacus Haldeman 1853 7 nea Coleoptera Geotrupidae Geotrupes mutator Marsham 1802 12 pal Coleoptera Geotrupidae Geotrupes spiniger Marsham 1802 6 pal Coleoptera Geotrupidae Geotrupes stercorarius Linnaeus 1758 12 pal Coleoptera Geotrupidae Glyptogeotrupes impressus Gebler 1841 1 pal Coleoptera Geotrupidae Lethrus acutangulus Ballion 1871 2 pal Coleoptera Geotrupidae Lethrus apterus Laxm. 1770 2 Coleoptera Geotrupidae Lethrus apterus Fischer 1822 2 pal Coleoptera Geotrupidae Lethrus cephalotus Pallas 1771 4 neo Coleoptera Geotrupidae Lethrus inermis Reitter 1897 1 neo Coleoptera Geotrupidae Lethrus karelini Gebler 1845 2 neo Coleoptera Geotrupidae Lethrus koshantschikovi Semenov 1892 1 neo Coleoptera Geotrupidae Lethrus scoparius Fischer 1820 2 neo Coleoptera Geotrupidae Lethrus sulcipennis Kraatz 1883 2 neo Coleoptera Geotrupidae Mycotrupes gaigei Olsen and Hubbel 1954 1 nea Coleoptera Geotrupidae Mycotrupes retusus LeConte 1866 1 nea Coleoptera Geotrupidae Neoathyreus excavatus Laporte 1840 1 pal Coleoptera Geotrupidae Neoathyreus fissicornis Harold 1880 1 pal Coleoptera Geotrupidae Neoathyreus lanei Martinez 1952 2 pal Coleoptera Geotrupidae Odonteus binasutum Fairmaire 1861 2 pal Coleoptera Geotrupidae Odonteus capense Klug 1843 2 afr Coleoptera Geotrupidae Odonteus corniger Melsheimer 1844 4 nea Coleoptera Geotrupidae Odonteus darlingtoni Wallis 1928 4 nea Coleoptera Geotrupidae Odonteus distinguendum Fairmaire 1861 1 pal Coleoptera Geotrupidae Odonteus falli Wallis 1928 4 nea Coleoptera Geotrupidae Odonteus floridanus Wallis 1928 31 nea Coleoptera Geotrupidae Odonteus gallicus Mulsant 1842 3 neo Coleoptera Geotrupidae Odonteus laesicolle Fairmaire 1856 1 pal Coleoptera Geotrupidae Odonteus liebecki Wallis 1928 34 nea Coleoptera Geotrupidae Odonteus obesus LeConte 1859 20 nea Coleoptera Geotrupidae Odonteus simi Wallis 1928 3 nea Coleoptera Geotrupidae Odonteus sp. 1 nea Coleoptera Geotrupidae Odonteus sp. 1 ori Coleoptera Geotrupidae Odonteus striatopunctatum Castelnau 1840 1 Coleoptera Geotrupidae Odonteus thoracicornis Wallis 1928 2 nea Coleoptera Geotrupidae Peltotrupes profundus Howden 1952 5 nea Coleoptera Geotrupidae Sericotrupes niger Marsham 1802 2 pal Coleoptera Geotrupidae Taurocerastes patagonicus Phillipi 1866 2 pal Coleoptera Geotrupidae Thorectes geminatus Gene 1839 3 pal Coleoptera Geotrupidae Thorectes intermedius Costa 1827 9 pal Coleoptera Geotrupidae Thorectes marginatus Poiret 1787 2 pal Coleoptera Geotrupidae Thorectes rugatulus Jekel 1865 2 pal Coleoptera Geotrupidae Trypocopris pyrenaeus Charpentier 1825 2 pal Coleoptera Geotrupidae Trypocopris vernalis Linnaeus 1758 13 pal Coleoptera Geotrupidae Trypocopris vernalis Heer 1847 1 pal Coleoptera Geotrupidae Trypocopris vernalis Kuster 1853 4 pal Coleoptera Geotrupidae Typhoeus typhoeus Linn. 1758 8 pal Coleoptera Geotrupidae Typhoeus typhoeus Linnaeus 1758 8 pal Coleoptera Glaphyridae Amphicoma acrtos Faldermann 1835 1 pal Coleoptera Glaphyridae Amphicoma arctos Reitter 1890 1 pal Coleoptera Glaphyridae Amphicoma bombyliformis Pallas 1781 1 pal Coleoptera Glaphyridae Amphicoma bombylius Fabricius 1789 8 pal Coleoptera Glaphyridae Amphicoma fastuosa Reitter 1890 1 pal Coleoptera Glaphyridae Amphicoma hyrax Truqui 1848 1 pal Coleoptera Glaphyridae Amphicoma kuschakewitschi Ball 1871 1 pal Coleoptera Glaphyridae Amphicoma lasserrei Germar 1834 17 pal Coleoptera Glaphyridae Amphicoma malatia Truqui 1848 7 pal Coleoptera Glaphyridae Amphicoma papaveris Sturm 1843 1 pal Coleoptera Glaphyridae Amphicoma vittata Fabricius 1775 1 pal Coleoptera Glaphyridae Amphicoma vittata Reitter 1890 2 neo Coleoptera Glaphyridae Amphicoma vittatta Reitter 1890 1 neo Coleoptera Glaphyridae Amphicoma vulpes Brulle 1 neo Coleoptera Glaphyridae Amphicoma vulpes Fabricius 1792 2 pal Coleoptera Glaphyridae Cratoscelis villosa Blanchard 1850 7 neo Coleoptera Glaphyridae Cratoscelis vulpina Erichson 1835 2 neo Coleoptera Glaphyridae Glaphyrus festivus Menier 1836 1 pal Coleoptera Glaphyridae Glaphyrus maurus Linnaeus 1758 3 afr Coleoptera Glaphyridae Glaphyrus micans Faldermann 1835 1 pal Coleoptera Glaphyridae Lichanthe rathvoni LeConte 1863 13 nea Coleoptera Glaphyridae Lichanthe ursina LeConte 1861 4 nea Coleoptera Glaphyridae Lichanthe vulpina Hentz 1826 16 nea Coleoptera Glaphyridae Lichnia limbata Erichson 1851 4 neo Coleoptera Glaresidae Glaresis ecostata Fall 1907 1 nea Coleoptera Glaresidae Glaresis gordoni Warner 2 nea Coleoptera Glaresidae Glaresis phoenicia Fall 1907 1 nea Coleoptera Glaresidae Glaresis zvirgzdinsi Warner 2 nea Coleoptera Hybosoridae Coelodes ovalis Robinson Coleoptera Hybosoridae Hybosorus illigeri Reiche 1853 8 nea Coleoptera Hybosoridae Hybosorus spp. 18 Coleoptera Lucanidae Aegus minutus Gestro 1 Coleoptera Lucanidae Aesalus scarabaeoides Panzer 1794 1 neo Coleoptera Lucanidae Apterocyclus honoluluensis Waterhouse 1 Coleoptera Lucanidae Apterodorcus bacchus Westwood 4 Coleoptera Lucanidae Bomansodus impubis Parry 4 Coleoptera Lucanidae Brasilucanus alvaregei Vulcano and Pereira 1 Coleoptera Lucanidae Casignetus humboldti Gyllenhal 1 Coleoptera Lucanidae Ceratognathus curvidens Broun 1904 2 aus Coleoptera Lucanidae Ceruchus chrysomelinus Hochenwarth 1785 2 neo Coleoptera Lucanidae Ceruchus piceus Weber 1801 297 nea Coleoptera Lucanidae Ceruchus punctatus LeConte 1869 3 nea Coleoptera Lucanidae Ceruchus striatus LeConte 1859 9 nea Coleoptera Lucanidae Chalcodes carinatus Linnaeus 2 Coleoptera Lucanidae Chiasognathinus gaujoni Oberthuer 1 Coleoptera Lucanidae Chiasognathus granti Stephens 10 Coleoptera Lucanidae Cladognathus confucius Hope 1 Coleoptera Lucanidae Cladognathus confucius 1 Coleoptera Lucanidae Cladognathus giraffa Olivier 4 Coleoptera Lucanidae Cyclommatus frey-gessneri Ritsema 1 Coleoptera Lucanidae Cyclommatus tarandus Thuenberg 15 Coleoptera Lucanidae Dorcus brevis Say 4 Coleoptera Lucanidae Dorcus immundus Arrow 1 Coleoptera Lucanidae Dorcus parallelepipedus Linnaeus 21 Coleoptera Lucanidae Dorcus parallelepipedus Ganglbauer 3 Coleoptera Lucanidae Dorcus parallelus Say 82 Coleoptera Lucanidae Eurytrachellelus alcides Vollenhoven 1 Coleoptera Lucanidae Eurytrachellelus arfakianus Lansberge 2 Coleoptera Lucanidae Eurytrachellelus bucephalus Perty 5 Coleoptera Lucanidae Eurytrachellelus cribriceps Chevrolat 3 Coleoptera Lucanidae Eurytrachellelus egregius Moellenkamp 2 Coleoptera Lucanidae Eurytrachellelus gypaetus Castelnau 1 Coleoptera Lucanidae Eurytrachellelus intermedius Gestro 26 Coleoptera Lucanidae Eurytrachellelus reichei Hope 1 Coleoptera Lucanidae Eurytrachellelus rubrofemoratus Vollenhoven 4 Coleoptera Lucanidae Eurytrachellelus saiga Olivier 10 Coleoptera Lucanidae Eurytrachellelus striatipennis Motschulsky 7 Coleoptera Lucanidae Eurytrachellelus submolaris Westwood 1 Coleoptera Lucanidae Eurytrachellelus titanus Boisduval 6 Coleoptera Lucanidae Eurytrachellelus tityus Hope 1 Coleoptera Lucanidae Figulus foveicollis Boisduval 2 Coleoptera Lucanidae Gnaphaloryx opacus Burmeister 1 Coleoptera Lucanidae Hemisodorcus napalensis Hope 1 Coleoptera Lucanidae Hexarthrius deyrollei Parry 1 Coleoptera Lucanidae Hexarthrius parryi Hope 1 Coleoptera Lucanidae Lamprima adolphinae Gestro 2 Coleoptera Lucanidae Lamprima aurata Latreille 2 Coleoptera Lucanidae Lamprima latreille McLeay 1 Coleoptera Lucanidae Lamprima varians Germar 1 Coleoptera Lucanidae Leptinopterus ibex Billberg 2 Coleoptera Lucanidae Leptinopterus tibialis Eschscholtz 5 Coleoptera Lucanidae Lissapterus howittanus Westwood 1 Coleoptera Lucanidae Lissotes reticulatus Westwood 2 Coleoptera Lucanidae Lissotes spp. 2 Coleoptera Lucanidae Lucanus cervus Linnaeus 53 Coleoptera Lucanidae Lucanus elaphus Fabricius 7 Coleoptera Lucanidae Lucanus smithi Parry 3 Coleoptera Lucanidae Metopodontus bison Olivier 7 Coleoptera Lucanidae Metopodontus cinctus Montrouzier 1 Coleoptera Lucanidae Metopodontus foveatus Hope 1 Coleoptera Lucanidae Metopodontus occipitalis Westwood 1 Coleoptera Lucanidae Metopodontus savagei Hope 5 Coleoptera Lucanidae Metopodontus spp. 5 Coleoptera Lucanidae Neolucanus alces Fabricius 4 Coleoptera Lucanidae Neolucanus baladeva Hope 5 Coleoptera Lucanidae Neolucanus castanopterus Hope 5 Coleoptera Lucanidae Nicagus obscurus LeConte 1848 15 nea Coleoptera Lucanidae Nigidius bonneuili Boileau 2 Coleoptera Lucanidae Nigidius laevicollis Westwood 2 Coleoptera Lucanidae Nigidius medimniennsis Seguy 1 Coleoptera Lucanidae Nigidius spp. 3 Coleoptera Lucanidae Odontolabis bellicosus Castelnau 2 Coleoptera Lucanidae Odontolabis cuvera HOpe 6 Coleoptera Lucanidae Odontolabis dalmani Westwood 12 Coleoptera Lucanidae Odontolabis delesserti Guerin 1 Coleoptera Lucanidae Odontolabis gazella Fabricius 1 Coleoptera Lucanidae Odontolabis moellenkampi Fruhstorfer 1 Coleoptera Lucanidae Odontolabis siva Westwood 2 Coleoptera Lucanidae Odontolabis sommeri Parry 2 Coleoptera Lucanidae Odontolabis wollastoni Parry 7 Coleoptera Lucanidae Phalacrognathus muelleri Kriesche 71 Coleoptera Lucanidae Platyceroides aeneus VanDyke 4 Coleoptera Lucanidae Platyceroides agassii LeConte 3 Coleoptera Lucanidae Platyceroides laticollis Casey 2 Coleoptera Lucanidae Platyceropsis keeni Casey 6 Coleoptera Lucanidae Platycerus caraboides Linnaeus 11 Coleoptera Lucanidae Platycerus marginalis Casey 14 Coleoptera Lucanidae Platycerus marginalis Benesh 2 Coleoptera Lucanidae Platycerus oregonensis Westwood 28 Coleoptera Lucanidae Platycerus piceus Kirby 14 Coleoptera Lucanidae Platycerus spp. 15 Coleoptera Lucanidae Platycerus virescens Fabricius 54 Coleoptera Lucanidae Prismognathus angularis Waterhouse 1 Coleoptera Lucanidae Prismognathus subaeneus Motschulsky 2 Coleoptera Lucanidae Prosopocoelus dorsalis Erichson 2 Coleoptera Lucanidae Prosopocoelus hasterti Boileau 2 Coleoptera Lucanidae Prosopocoelus lorquini Deyrolle 1 Coleoptera Lucanidae Prosopocoelus natalensis Waterhouse 4 Coleoptera Lucanidae Prosopocoelus senegalensis Klug 2 Coleoptera Lucanidae Prosopocoelus serricornis Latreille 2 Coleoptera Lucanidae Prosopocoelus spp. 1 Coleoptera Lucanidae Psalidoremus inclinatus Motschulsky 14 Coleoptera Lucanidae Pseudolucanus capreolus Linnaeus 184 Coleoptera Lucanidae Pseudolucanus mazama LeConte 9 Coleoptera Lucanidae Pseudolucanus placidus Say 53 Coleoptera Lucanidae Pseudolucanus spp. 35 Coleoptera Lucanidae Pycnosiphorus caelatus Blanchard 4 Coleoptera Lucanidae Pycnosiphorus costatus Benesh 1 Coleoptera Lucanidae Pycnosiphorus fasciatus Germain 5 Coleoptera Lucanidae Pycnosiphorus lessoni Buquet 2 Coleoptera Lucanidae Pycnosiphorus mandibularis Solier 1 Coleoptera Lucanidae Pycnosiphorus virgatus Nagel 1 Coleoptera Lucanidae Sclerostomus cucullatus Blanchard 4 Coleoptera Lucanidae Sclerostomus tuberculatus Solier 2 Coleoptera Lucanidae Sinodendron cylindricum Linnaeus 1758 12 neo Coleoptera Lucanidae Sinodendron rugosum Mannerheim 1843 75 nea Coleoptera Lucanidae Sphenognathus higginsi Parry 1 Coleoptera Lucanidae Sphenognathus nobilis Parry 1 Coleoptera Lucanidae Streptocerus speciosus Fairaire 1 Coleoptera Lucanidae Streptocerus speciosus Fairmaire 1 Coleoptera Lucanidae undetermined spp. 64 Coleoptera Ochodaeidae Ochodaeus biarmatus LeConte 1868 25 nea Coleoptera Ochodaeidae Ochodaeus chrysomeloides Schrank 1781 2 pal Coleoptera Ochodaeidae Ochodaeus frontalis LeConte 1863 1 nea Coleoptera Ochodaeidae Ochodaeus gnatho Fall 1907 1 nea Coleoptera Ochodaeidae Ochodaeus inarmatus Schaeffer 1906 9 nea Coleoptera Ochodaeidae Ochodaeus kansanus Fall 1909 4 nea Coleoptera Ochodaeidae Ochodaeus n.sp. nr.biarmatus 2 nea Coleoptera Ochodaeidae Ochodaeus n.sp. nr.biarmatus ? 2 nea Coleoptera Ochodaeidae Ochodaeus pectoralis LeConte 1868 1 nea Coleoptera Ochodaeidae Ochodaeus peninsularis Horn 1895 1 nea Coleoptera Ochodaeidae Ochodaeus planifrons Schaeffer 1906 7 nea Coleoptera Ochodaeidae Ochodaeus praesidii Bates 1887 14 nea Coleoptera Ochodaeidae Ochodaeus repandus Fall 1909 3 nea Coleoptera Ochodaeidae Ochodaeus simplex LeConte 1854 5 nea Coleoptera Ochodaeidae Ochodaeus sparsus LeConte 1868 1 nea Coleoptera Ochodaeidae Ochodaeus spp. 3 Coleoptera Pleocomidae Pleocoma australis Fall 1911 1 nea Coleoptera Pleocomidae Pleocoma badia Fall 1917 1 nea Coleoptera Pleocomidae Pleocoma behrensi LeConte 1874 2 nea Coleoptera Pleocomidae Pleocoma crinita Linsley 1938 6 nea Coleoptera Pleocomidae Pleocoma dubitalis Davis 1935 2 nea Coleoptera Pleocomidae Pleocoma edwardsi LeConte 1874 1 nea Coleoptera Pleocomidae Pleocoma minor Linsley 1938 6 nea Coleoptera Pleocomidae Pleocoma oregonensis Leach 1933 2 nea Coleoptera Pleocomidae Pleocoma puncticollis Rivers 1889 1 nea Coleoptera Pleocomidae Pleocoma simi Davis 1934 2 nea Coleoptera Pleocomidae Pleocoma spp. 8 Coleoptera Pleocomidae Pleocoma staff Schaufuss 1870 1 nea Coleoptera Pleocomidae Pleocoma tularensis Leach 1933 1 nea Coleoptera Scarabaeidae Actinobolus radians Westwood 1841 1 neo Coleoptera Scarabaeidae Adoretus compressus Weber 1801 4 afr Coleoptera Scarabaeidae Adoretus flaveolus Fahraeus 1857 2 afr Coleoptera Scarabaeidae Adoretus nigrifrons Stephens 1809 1 afr Coleoptera Scarabaeidae Adoretus sinicus Burmeister 1855 3 ori Coleoptera Scarabaeidae Adoretus tenuimaculatus Waterhouse 1875 4 ori Coleoptera Scarabaeidae Aegialia arenaria Fabricius 1787 6 nea Coleoptera Scarabaeidae Aegialia arenaria Fabricius 1787 6 pal Coleoptera Scarabaeidae Aegialia blanchardi Horn 1887 2 nea Coleoptera Scarabaeidae Aegialia conferta Horn 1871 8 nea Coleoptera Scarabaeidae Aegialia convexa Fall 1932 3 nea Coleoptera Scarabaeidae Aegialia criddlei Brown 1931 3 nea Coleoptera Scarabaeidae Aegialia lacustris LeConte 1850 5 nea Coleoptera Scarabaeidae Aegialia latispina LeConte 1878 4 nea Coleoptera Scarabaeidae Aegialia opaca Brown 1931 1 nea Coleoptera Scarabaeidae Aegialia opifex Horn 1887 1 nea Coleoptera Scarabaeidae Aegialia punctata Brown 1931 6 nea Coleoptera Scarabaeidae Aegialia sabuleti Panzer 1797 2 pal Coleoptera Scarabaeidae Aegopsis curvicornis Burmeister 1847 1 neo Coleoptera Scarabaeidae Aegopsis sp. 2 neo Coleoptera Scarabaeidae Aethiessa floralis Fabricius 1787 31 afr Coleoptera Scarabaeidae Aethiessa floralis Herbst 1790 1 pal Coleoptera Scarabaeidae Aethiessa floralis Gory and Percheron 1833 1 afr Coleoptera Scarabaeidae Allomyrina dichotomus Linnaeus 1771 5 ori Coleoptera Scarabaeidae Allorrhina carmelita Burmeister 1842 1 neo Coleoptera Scarabaeidae Allorrhina scabriuscula Swederus 1787 2 neo Coleoptera Scarabaeidae Amaladera euphorbiae Burmeister 1855 1 pal Coleoptera Scarabaeidae Amaurodes passerini Westwood 1844 1 afr Coleoptera Scarabaeidae Amazula suavis Burmeister 1847 1 afr Coleoptera Scarabaeidae Amithao decemguttatus Waterhouse 1876 1 neo Coleoptera Scarabaeidae Amphimallon assimilis Herbst 1790 1 pal Coleoptera Scarabaeidae Amphimallon ater Herbst 1790 2 pal Coleoptera Scarabaeidae Amphimallon ater Olivier 1789 1 pal Coleoptera Scarabaeidae Amphimallon caucasicus Gyllenhal 1817 1 pal Coleoptera Scarabaeidae Amphimallon glabripennis Ballion 1871 1 pal Coleoptera Scarabaeidae Amphimallon majalis Razoumowsky 1789 122 nea Coleoptera Scarabaeidae Amphimallon quercanus Burmeister 1855 3 pal Coleoptera Scarabaeidae Amphimallon seidlitzi Kraatz 1891 1 pal Coleoptera Scarabaeidae Amphimallon solstitialis Linnaeus 1758 17 pal Coleoptera Scarabaeidae Amphimallon solstitialis Gyllenhal 1817 4 pal Coleoptera Scarabaeidae Anatropis verticalis Horn 1879 11 nea Coleoptera Scarabaeidae Ancognatha manca LeConte 1866 43 nea Coleoptera Scarabaeidae Ancognatha spp. 29 neo Coleoptera Scarabaeidae Anisonyx hilaris Peringuey 1902 2 afr Coleoptera Scarabaeidae Anisoplia agnata Reitter 1889 1 pal Coleoptera Scarabaeidae Anisoplia agricola Poda 1761 5 pal Coleoptera Scarabaeidae Anisoplia agricola Goeze 1777 15 pal Coleoptera Scarabaeidae Anisoplia austriaca Herbst 1783 2 pal Coleoptera Scarabaeidae Anisoplia austriaca Herbst 1783 10 pal Coleoptera Scarabaeidae Anisoplia baetica Erichson 1847 1 pal Coleoptera Scarabaeidae Anisoplia bromicola Germar 1817 2 pal Coleoptera Scarabaeidae Anisoplia farraria Erichson 1847 1 pal Coleoptera Scarabaeidae Anisoplia flavipennis Brulle 1832 1 pal Coleoptera Scarabaeidae Anisoplia flavipennis Erichson 1847 13 pal Coleoptera Scarabaeidae Anisoplia leucaspis Laporte 1840 2 pal Coleoptera Scarabaeidae Anisoplia remota Reitter 1889 3 pal Coleoptera Scarabaeidae Anisoplia segetum Herbst 1783 16 pal Coleoptera Scarabaeidae Anisoplia syriaca Burmeister 1844 2 pal Coleoptera Scarabaeidae Anisoplia tempestiva Erichson 1847 2 pal Coleoptera Scarabaeidae Anisoplia tenebralis Burmeister 1844 2 pal Coleoptera Scarabaeidae Anochilia laevigata Gory and Percheron 1835 4 ori Coleoptera Scarabaeidae Anochilia spp. 4 ori Coleoptera Scarabaeidae Anomala arida Casey 1915 4 nea Coleoptera Scarabaeidae Anomala atriplicis Fabricius 1787 1 pal Coleoptera Scarabaeidae Anomala aulax Wiedemann 1832 1 ori Coleoptera Scarabaeidae Anomala binotata Gyllenhal 1817 nea Coleoptera Scarabaeidae Anomala camancha Wickham 1913 5 nea Coleoptera Scarabaeidae Anomala cavifrons LeConte 1868 50 nea Coleoptera Scarabaeidae Anomala cinta Bates 1888 3 neo Coleoptera Scarabaeidae Anomala cupricollis Chevrolat 2 neo Coleoptera Scarabaeidae Anomala delicata Casey 1915 3 nea Coleoptera Scarabaeidae Anomala digressa Casey 1915 5 nea Coleoptera Scarabaeidae Anomala donovani Blanchard 1850 2 neo Coleoptera Scarabaeidae Anomala dubia Scopoli 1763 1 pal Coleoptera Scarabaeidae Anomala dubia Scopoli 1763 8 pal Coleoptera Scarabaeidae Anomala dubia Scopoli 1763 9 pal Coleoptera Scarabaeidae Anomala dubia Scopoli 1763 14 pal Coleoptera Scarabaeidae Anomala dubia Fabricius 1775 1 pal Coleoptera Scarabaeidae Anomala dubia Fabricius 1775 7 pal Coleoptera Scarabaeidae Anomala flavilla Bates 1888 2 nea Coleoptera Scarabaeidae Anomala flavilla LeConte 1863 1 nea Coleoptera Scarabaeidae Anomala flavipennis Burmeister 1844 29 nea Coleoptera Scarabaeidae Anomala flavipennis LeConte 1854 24 nea Coleoptera Scarabaeidae Anomala flavipennis Casey 1915 3 nea Coleoptera Scarabaeidae Anomala foraminosa Bates 1888 2 nea Coleoptera Scarabaeidae Anomala foraminosa Bates 1888 2 neo Coleoptera Scarabaeidae Anomala forreri Bates 1888 1 neo Coleoptera Scarabaeidae Anomala fusciceps Fahraeus 1857 2 afr Coleoptera Scarabaeidae Anomala gemella Say 1835 3 neo Coleoptera Scarabaeidae Anomala histrionella Bates 1888 3 neo Coleoptera Scarabaeidae Anomala hoepfneri Bates 1888 1 neo Coleoptera Scarabaeidae Anomala immatura Bohemann 1860 2 afr Coleoptera Scarabaeidae Anomala inconstans Burmeister 1844 1 neo Coleoptera Scarabaeidae Anomala inconstans Burmeister 1844 2 neo Coleoptera Scarabaeidae Anomala innuba Fabricius 1787 11 nea Coleoptera Scarabaeidae Anomala lucicola Fabricius 1798 85 nea Coleoptera Scarabaeidae Anomala ludoviciana Schaeffer 1906 4 nea Coleoptera Scarabaeidae Anomala marginata Fabricius 1775 77 nea Coleoptera Scarabaeidae Anomala marginicollis Bates 1888 2 neo Coleoptera Scarabaeidae Anomala maritima Casey 1915 4 nea Coleoptera Scarabaeidae Anomala medorensis Casey 1915 1 nea Coleoptera Scarabaeidae Anomala mendica Casey 1915 2 nea Coleoptera Scarabaeidae Anomala minuta Burmeister 1844 5 nea Coleoptera Scarabaeidae Anomala nigropicta Casey 1915 210 nea Coleoptera Scarabaeidae Anomala oblivia Horn 1884 26 nea Coleoptera Scarabaeidae Anomala orientalis Waterhouse 1875 3 aus Coleoptera Scarabaeidae Anomala orientalis Waterhouse 1875 15 nea Coleoptera Scarabaeidae Anomala osmanlis Blanchard 1850 1 pal Coleoptera Scarabaeidae Anomala papagoana Casey 1915 26 nea Coleoptera Scarabaeidae Anomala parvula Burmeister 1844 1 nea Coleoptera Scarabaeidae Anomala pasadenae Casey 3 nea Coleoptera Scarabaeidae Anomala sagax Casey 1915 4 nea Coleoptera Scarabaeidae Anomala semilivida LeConte 1878 6 nea Coleoptera Scarabaeidae Anomala semitonsa Bates 1888 2 neo Coleoptera Scarabaeidae Anomala sp. nr.plurisulcata neo Coleoptera Scarabaeidae Anomala spp. 6 nea Coleoptera Scarabaeidae Anomala spp. 11 neo Coleoptera Scarabaeidae Anomala stigmotella Casey 1915 3 nea Coleoptera Scarabaeidae Anomala tenera Casey 1915 3 nea Coleoptera Scarabaeidae Anomala tenera Casey 1915 21 neo Coleoptera Scarabaeidae Anomala undulata Melsheimer 1844 45 nea Coleoptera Scarabaeidae Anomala vanpatteni Bates 1888 4 neo Coleoptera Scarabaeidae Anomala villosella Blanchard 1850 1 neo Coleoptera Scarabaeidae Anomala vitis Fabricius 1775 10 pal Coleoptera Scarabaeidae Anomiopsoides pereirai Menetries neo Coleoptera Scarabaeidae Anoplocheilus figuratus Bohmann 1857 2 afr Coleoptera Scarabaeidae Anoplodrepanus pecki Howden neo Coleoptera Scarabaeidae Anoplodrepanus reconditus Matthews 2 neo Coleoptera Scarabaeidae Anoplognathus macleayi Blackburn 1892 5 aus Coleoptera Scarabaeidae Anoplognathus montanus MacLeay 1873 5 aus Coleoptera Scarabaeidae Anoplognathus olivieri Dalman 1817 1 aus Coleoptera Scarabaeidae Anoplognathus pallidicollis Blanchard 1851 8 aus Coleoptera Scarabaeidae Anoplognathus sp. 2 Coleoptera Scarabaeidae Anoplognathus sp. 1 aus Coleoptera Scarabaeidae Anoplognathus sp. 3 aus Coleoptera Scarabaeidae Anoplognathus suturalis Boisduval 1835 3 aus Coleoptera Scarabaeidae Anoplognathus viriditarsis Leach 1815 1 aus Coleoptera Scarabaeidae Anoplosiagum sp. nr.rufum neo Coleoptera Scarabaeidae Anoxia asiatica Desbrocher 1871 2 pal Coleoptera Scarabaeidae Anoxia orientalis Krynicki 1832 1 pal Coleoptera Scarabaeidae Anoxia pilosa Fabricius 1792 12 pal Coleoptera Scarabaeidae Anoxia scutellaris Mulsant 1842 2 pal Coleoptera Scarabaeidae Anoxia villosa Fabricius 1781 3 pal Coleoptera Scarabaeidae Anticheira campisilvaticus Ohaus 1905 4 neo Coleoptera Scarabaeidae Aphodius aemulus Horn 1887 6 nea Coleoptera Scarabaeidae Aphodius albertanus Brown 1928 1 nea Coleoptera Scarabaeidae Aphodius aleutus Eschscholtz 1822 7 nea Coleoptera Scarabaeidae Aphodius aleutus Fall and Cockerell 1907 14 nea Coleoptera Scarabaeidae Aphodius aleutus Horn 1870 16 nea Coleoptera Scarabaeidae Aphodius alpinus Scopoli 1763 4 neo Coleoptera Scarabaeidae Aphodius alternatus Horn 1870 3 nea Coleoptera Scarabaeidae Aphodius alternatus Horn 1879 7 nea Coleoptera Scarabaeidae Aphodius angustus Hatch 1972 3 nea Coleoptera Scarabaeidae Aphodius anthracinus LeConte 1878 12 nea Coleoptera Scarabaeidae Aphodius aquilonarius Brown 1928 1 nea Coleoptera Scarabaeidae Aphodius ater Degeer 1774 7 neo Coleoptera Scarabaeidae Aphodius badipes Melsheimer 1844 5 nea Coleoptera Scarabaeidae Aphodius badipes Melshiemer 1844 5 nea Coleoptera Scarabaeidae Aphodius bicolor Say 1823 15 nea Coleoptera Scarabaeidae Aphodius bimaculatus Laxmann 1770 8 neo Coleoptera Scarabaeidae Aphodius bonvouloiri Harold 1860 1 neo Coleoptera Scarabaeidae Aphodius brasiliensis Castelnau 1840 1 pal Coleoptera Scarabaeidae Aphodius brimleyi Cartwright 1939 5 nea Coleoptera Scarabaeidae Aphodius browni Hinton 1934 1 nea Coleoptera Scarabaeidae Aphodius bryanti Brown 1929 1 nea Coleoptera Scarabaeidae Aphodius campestris Blatchley 1912 53 nea Coleoptera Scarabaeidae Aphodius carri Brown 1929 1 nea Coleoptera Scarabaeidae Aphodius cavidomus Brown 1927 4 nea Coleoptera Scarabaeidae Aphodius claudus Fall 1932 9 nea Coleoptera Scarabaeidae Aphodius coenosus Panzer 1798 6 neo Coleoptera Scarabaeidae Aphodius coloradensis Horn 1870 106 nea Coleoptera Scarabaeidae Aphodius comma Reitter 1892 7 neo Coleoptera Scarabaeidae Aphodius concavus Say 1823 7 nea Coleoptera Scarabaeidae Aphodius congregatus Mannerheim 1853 55 nea Coleoptera Scarabaeidae Aphodius consentaneus LeConte 1850 10 nea Coleoptera Scarabaeidae Aphodius consociatus Horn 1887 4 nea Coleoptera Scarabaeidae Aphodius consputus Creutzer 1799 6 neo Coleoptera Scarabaeidae Aphodius constans Duftschmid 1805 5 neo Coleoptera Scarabaeidae Aphodius contaminatus Herbst 1783 1 neo Coleoptera Scarabaeidae Aphodius corruptor Brown 1929 2 nea Coleoptera Scarabaeidae Aphodius crassuloides Fall and Cockerell 1907 15 nea Coleoptera Scarabaeidae Aphodius crassulus Horn 1870 1 nea Coleoptera Scarabaeidae Aphodius cribrarius Brulle 1836 5 neo Coleoptera Scarabaeidae Aphodius criddlei Brown 1928 1 nea Coleoptera Scarabaeidae Aphodius cuniculus Chevrolat 1864 9 nea Coleoptera Scarabaeidae Aphodius denticulatus Haldeman 1848 2 nea Coleoptera Scarabaeidae Aphodius dentigerulus Brown 1929 4 nea Coleoptera Scarabaeidae Aphodius depressus Kugelann 1792 1 neo Coleoptera Scarabaeidae Aphodius depressus Felderman 1835 6 neo Coleoptera Scarabaeidae Aphodius depressus Erichson 1848 11 neo Coleoptera Scarabaeidae Aphodius distinctus Muller 1776 6 neo Coleoptera Scarabaeidae Aphodius distinctus Muller 1776 169 nea Coleoptera Scarabaeidae Aphodius duplex LeConte 1878 1 nea Coleoptera Scarabaeidae Aphodius elegans Allibert 1847 1 neo Coleoptera Scarabaeidae Aphodius erraticus Linnaeus 1758 118 nea Coleoptera Scarabaeidae Aphodius erraticus Mulsant 1842 3 neo Coleoptera Scarabaeidae Aphodius erraticus Schmidt 1916 5 neo Coleoptera Scarabaeidae Aphodius erraticus Mulsant 1842 2 neo Coleoptera Scarabaeidae Aphodius erraticus Gebler 1809 8 neo Coleoptera Scarabaeidae Aphodius explanatus LeConte 1878 1 nea Coleoptera Scarabaeidae Aphodius femoralis Say 1823 4 nea Coleoptera Scarabaeidae Aphodius fimetarius Linnaeus 1758 469 nea Coleoptera Scarabaeidae Aphodius fimetarius Reitter 1892 5 neo Coleoptera Scarabaeidae Aphodius fossor Linnaeus 1758 121 nea Coleoptera Scarabaeidae Aphodius fucosus Schmidt 1922 13 nea Coleoptera Scarabaeidae Aphodius fuliginosus Harold 1863 1 pal Coleoptera Scarabaeidae Aphodius geomysi Cartwright 1939 1 nea Coleoptera Scarabaeidae Aphodius granarius Linnaeus 1767 401 nea Coleoptera Scarabaeidae Aphodius granarius Mulsant 1842 1 neo Coleoptera Scarabaeidae Aphodius grombezewskyi Koshant 1891 neo Coleoptera Scarabaeidae Aphodius guttatus Eschscholtz 1823 2 nea Coleoptera Scarabaeidae Aphodius haemorrhoidalis Linnaeus 1758 1 neo Coleoptera Scarabaeidae Aphodius haemorrhoidalis Linnaeus 1758 28 nea Coleoptera Scarabaeidae Aphodius haemorrhoidalis Linneaus 1758 1 neo Coleoptera Scarabaeidae Aphodius hamatus Say 1823 23 nea Coleoptera Scarabaeidae Aphodius hydrochaeris Fabricius 1798 2 neo Coleoptera Scarabaeidae Aphodius hyperboreus LeConte 1850 1 nea Coleoptera Scarabaeidae Aphodius ictericus Laicharting 1781 12 neo Coleoptera Scarabaeidae Aphodius immundus Creutzer 1799 7 neo Coleoptera Scarabaeidae Aphodius iowensis Wickham 1913 2 nea Coleoptera Scarabaeidae Aphodius kirni Cartwright 1944 6 nea Coleoptera Scarabaeidae Aphodius lacustris LeConte 1850 5 nea Coleoptera Scarabaeidae Aphodius larreae Horn 1887 4 nea Coleoptera Scarabaeidae Aphodius lentus Horn 1870 1 nea Coleoptera Scarabaeidae Aphodius lividus Olivier 1789 422 nea Coleoptera Scarabaeidae Aphodius lugens Creutzer 1799 12 neo Coleoptera Scarabaeidae Aphodius lunifer Solsky 1876 1 neo Coleoptera Scarabaeidae Aphodius luridus Fabricius 1775 4 Coleoptera Scarabaeidae Aphodius luridus Fabricius 1775 11 neo Coleoptera Scarabaeidae Aphodius luridus Muller 1776 5 neo Coleoptera Scarabaeidae Aphodius luteolus Horn 1887 18 nea Coleoptera Scarabaeidae Aphodius lutulentus Haldeman 1843 7 nea Coleoptera Scarabaeidae Aphodius luxatus Horn 1887 1 nea Coleoptera Scarabaeidae Aphodius melanostictus Schm. 1840 5 neo Coleoptera Scarabaeidae Aphodius melanostictus Schmidt 1840 5 neo Coleoptera Scarabaeidae Aphodius merdarius Fabricius 1775 11 neo Coleoptera Scarabaeidae Aphodius militaris LeConte 1858 1 nea Coleoptera Scarabaeidae Aphodius nemoralis Erichson 1848 1 neo Coleoptera Scarabaeidae Aphodius neotomae Fall 1907 12 nea Coleoptera Scarabaeidae Aphodius nigritus Fabricius 1801 3 nea Coleoptera Scarabaeidae Aphodius nigrivittus Solsky 1876 2 neo Coleoptera Scarabaeidae Aphodius obscuras Fabricius 1792 6 Coleoptera Scarabaeidae Aphodius obscurus Fabricius 1792 6 neo Coleoptera Scarabaeidae Aphodius oklahomensis Brown 1928 2 nea Coleoptera Scarabaeidae Aphodius omissus LeConte 1850 5 nea Coleoptera Scarabaeidae Aphodius omissus Horn 1 nea Coleoptera Scarabaeidae Aphodius opacus LeConte 1872 14 nea Coleoptera Scarabaeidae Aphodius opifex Horn 1887 1 nea Coleoptera Scarabaeidae Aphodius pardalis LeConte 1857 57 nea Coleoptera Scarabaeidae Aphodius paykulli Bedel 1911 4 neo Coleoptera Scarabaeidae Aphodius pectoralis LeConte 1857 2 nea Coleoptera Scarabaeidae Aphodius peculiosus Brown 1928 2 nea Coleoptera Scarabaeidae Aphodius phaeopterus LeConte 1878 24 nea Coleoptera Scarabaeidae Aphodius phaleroides Horn 1870 14 nea Coleoptera Scarabaeidae Aphodius piceatus Robinson 1946 1 nea Coleoptera Scarabaeidae Aphodius piceus Gyllenhal 1808 1 neo Coleoptera Scarabaeidae Aphodius pictus Sturm 1805 8 neo Coleoptera Scarabaeidae Aphodius pingellus Brown 1929 1 nea Coleoptera Scarabaeidae Aphodius pinguis Haldeman 1848 3 nea Coleoptera Scarabaeidae Aphodius prodromus Braham 1790 260 nea Coleoptera Scarabaeidae Aphodius prodromus Solsky 1876 2 neo Coleoptera Scarabaeidae Aphodius pseudabusus Cartwright 1957 7 nea Coleoptera Scarabaeidae Aphodius pubescens Sturm 1800 6 neo Coleoptera Scarabaeidae Aphodius punctissimus Brown 1928 2 nea Coleoptera Scarabaeidae Aphodius pusillus Herbst 1789 10 neo Coleoptera Scarabaeidae Aphodius quadriguttatus Herbst 1783 15 neo Coleoptera Scarabaeidae Aphodius quadriguttatus Mulsant 1842 1 neo Coleoptera Scarabaeidae Aphodius quadrimaculatus Linnaeus 1761 1 neo Coleoptera Scarabaeidae Aphodius rectus Motschulsky 1866 1 neo Coleoptera Scarabaeidae Aphodius rectus Reitter 1892 2 neo Coleoptera Scarabaeidae Aphodius rossi Cartwright 1944 2 nea Coleoptera Scarabaeidae Aphodius rubeolus Beauvois 1805 18 nea Coleoptera Scarabaeidae Aphodius rubripennis Horn 1870 24 nea Coleoptera Scarabaeidae Aphodius rudis LeConte 1878 1 nea Coleoptera Scarabaeidae Aphodius rudis LeConte 1878 3 nea Coleoptera Scarabaeidae Aphodius ruficlaris 1932 3 nea Coleoptera Scarabaeidae Aphodius ruficlarus Fall 3 nea Coleoptera Scarabaeidae Aphodius rufipes Linnaeus 1758 158 nea Coleoptera Scarabaeidae Aphodius rugatus Schmidt 1907 1 nea Coleoptera Scarabaeidae Aphodius rusicola Melsheimer 1845 148 nea Coleoptera Scarabaeidae Aphodius sallei 1863 29 nea Coleoptera Scarabaeidae Aphodius satellitus Herbst 1789 8 neo Coleoptera Scarabaeidae Aphodius scrofa Fabricius 1787 5 neo Coleoptera Scarabaeidae Aphodius scrutator Herbst 1798 2 neo Coleoptera Scarabaeidae Aphodius scuticollis Semenov 1898 1 neo Coleoptera Scarabaeidae Aphodius scybalarius Fabricius 1781 23 neo Coleoptera Scarabaeidae Aphodius sepultus Cartwright 1944 4 nea Coleoptera Scarabaeidae Aphodius serval Say 1835 6 nea Coleoptera Scarabaeidae Aphodius sigmoides VanDyke 1918 2 nea Coleoptera Scarabaeidae Aphodius snowi Brown 1929 19 nea Coleoptera Scarabaeidae Aphodius socialis Brown 1928 5 nea Coleoptera Scarabaeidae Aphodius sordidus Fabricius 1775 8 neo Coleoptera Scarabaeidae Aphodius sparsus LeConte 1878 11 nea Coleoptera Scarabaeidae Aphodius sphacelatus Sturm 1805 1 neo Coleoptera Scarabaeidae Aphodius stercorosus Melsheimer 1845 72 nea Coleoptera Scarabaeidae Aphodius sticticus Panzer 1798 6 neo Coleoptera Scarabaeidae Aphodius stuessyi Gordon 1 nea Coleoptera Scarabaeidae Aphodius sturmi Harold 1897 6 neo Coleoptera Scarabaeidae Aphodius subterraneus Linnaeus 1758 59 nea Coleoptera Scarabaeidae Aphodius talpoidesi Brown 1928 5 nea Coleoptera Scarabaeidae Aphodius tasmaniae Hope 1846 3 aus Coleoptera Scarabaeidae Aphodius tenellus Say 1823 1 nea Coleoptera Scarabaeidae Aphodius tenuistriatus Horn 1887 4 nea Coleoptera Scarabaeidae Aphodius terminalis Say 1823 15 nea Coleoptera Scarabaeidae Aphodius testaceiventris Fall 1932 1 nea Coleoptera Scarabaeidae Aphodius thermicola Sturm 1800 9 neo Coleoptera Scarabaeidae Aphodius thomomsi Brown 1928 4 nea Coleoptera Scarabaeidae Aphodius tomentosus Muller 1776 3 neo Coleoptera Scarabaeidae Aphodius troglodytes Hubbard 1894 1 nea Coleoptera Scarabaeidae Aphodius urostigma Harold 1862 2 neo Coleoptera Scarabaeidae Aphodius vandykei Barrett 1931 8 nea Coleoptera Scarabaeidae Aphodius varians Duftschmid 1805 12 neo Coleoptera Scarabaeidae Aphodius vittatus Say 1825 89 nea Coleoptera Scarabaeidae Aphodius walshi Horn 1870 10 nea Coleoptera Scarabaeidae Aphodius wickhami Brown 1928 1 nea Coleoptera Scarabaeidae Aphonides dunnianus Rivers 1889 4 nea Coleoptera Scarabaeidae Aphonus brevicruris Cartwright 1944 2 nea Coleoptera Scarabaeidae Aphonus castaneus Melsheimer 1846 6 nea Coleoptera Scarabaeidae Aphonus sp. 9 nea Coleoptera Scarabaeidae Aphonus tridentatus Say 1823 10 nea Coleoptera Scarabaeidae Aphotaenius carolinus VanDyke 1928 6 nea Coleoptera Scarabaeidae Archophileurus cribrosus LeConte 1835 5 nea Coleoptera Scarabaeidae Argyrophegges kolbei Kraatz 1895 1 afr Coleoptera Scarabaeidae Aspidolea fulginea Burmeister 1847 11 neo Coleoptera Scarabaeidae Aspidolea singularis Bates 1888 18 neo Coleoptera Scarabaeidae Aspidolea sp. 1 neo Coleoptera Scarabaeidae Ataenius abditus Haldeman 1848 36 nea Coleoptera Scarabaeidae Ataenius abditus Brown 1932 7 nea Coleoptera Scarabaeidae Ataenius alternatus Melsheimer 1844 183 nea Coleoptera Scarabaeidae Ataenius alternatus Melshiemer 1844 183 nea Coleoptera Scarabaeidae Ataenius apicalis Hinton 1937 10 nea Coleoptera Scarabaeidae Ataenius brevis Fall 1930 2 nea Coleoptera Scarabaeidae Ataenius californicus Horn 1887 7 nea Coleoptera Scarabaeidae Ataenius cognatus LeConte 1858 46 nea Coleoptera Scarabaeidae Ataenius confertus Fall 1909 6 nea Coleoptera Scarabaeidae Ataenius cylindrus Horn 1871 34 nea Coleoptera Scarabaeidae Ataenius desertus Horn 1871 32 nea Coleoptera Scarabaeidae Ataenius duncani Cartwright 1974 1 nea Coleoptera Scarabaeidae Ataenius fattigi Cartwight 1948 23 nea Coleoptera Scarabaeidae Ataenius gracilis Melsheimer 1844 269 nea Coleoptera Scarabaeidae Ataenius haroldi Steinheil 1872 5 nea Coleoptera Scarabaeidae Ataenius hirsutus Horn 1871 37 nea Coleoptera Scarabaeidae Ataenius imbricatoides Schmidt 1909 1 neo Coleoptera Scarabaeidae Ataenius imbricatus Melsheimer 1844 8 neo Coleoptera Scarabaeidae Ataenius imbricatus Melsheimer 1844 91 nea Coleoptera Scarabaeidae Ataenius inquisitus Horn 1887 6 nea Coleoptera Scarabaeidae Ataenius miamii Cartwright 1934 7 nea Coleoptera Scarabaeidae Ataenius morator Harold 1869 neo Coleoptera Scarabaeidae Ataenius n.spp. 25 nea Coleoptera Scarabaeidae Ataenius nugator Harold 1880 3 neo Coleoptera Scarabaeidae Ataenius ovatulus Horn 1871 3 nea Coleoptera Scarabaeidae Ataenius picinus Harold 1867 1 nea Coleoptera Scarabaeidae Ataenius picinus Harold 1867 17 neo Coleoptera Scarabaeidae Ataenius picipes Fleutiaux and Salle 1889 33 neo Coleoptera Scarabaeidae Ataenius platensis Blanchard 1843 44 nea Coleoptera Scarabaeidae Ataenius puncticollis LeConte 1868 61 nea Coleoptera Scarabaeidae Ataenius salutator Fabricius),? 1 neo Coleoptera Scarabaeidae Ataenius salutator Fabricius),??? 1 neo Coleoptera Scarabaeidae Ataenius saxatilis Cartwright 1944 5 nea Coleoptera Scarabaeidae Ataenius scutellaris Harold 1867 1 neo Coleoptera Scarabaeidae Ataenius semipilosus VanDyke 1928 18 nea Coleoptera Scarabaeidae Ataenius sp. 1 neo Coleoptera Scarabaeidae Ataenius sp. 2 neo Coleoptera Scarabaeidae Ataenius spp. 12 Coleoptera Scarabaeidae Ataenius spretulus Haldemann 1848 35 nea Coleoptera Scarabaeidae Ataenius stephani Cartwright 1974 1 nea Coleoptera Scarabaeidae Ataenius stercorator Fabricius 1775 2 neo Coleoptera Scarabaeidae Ataenius strigatus Say 1823 207 nea Coleoptera Scarabaeidae Ataenius strigicauda Bates 1887 2 nea Coleoptera Scarabaeidae Ataenius terminalis Chevrolat 1864 4 neo Coleoptera Scarabaeidae Ataenius texanus Harold 1874 1 neo Coleoptera Scarabaeidae Ataenius texanus Harold 1874 28 nea Coleoptera Scarabaeidae Ataenius tuberculatus Schmidt 1911 1 neo Coleoptera Scarabaeidae Ateuchus histeroides Weber 1801 36 nea Coleoptera Scarabaeidae Ateuchus illaesum Harold 1868 2 neo Coleoptera Scarabaeidae Ateuchus lecontei Harold 1875 25 nea Coleoptera Scarabaeidae Ateuchus robustum Harold 1868 11 neo Coleoptera Scarabaeidae Ateuchus subquadratum Harold 1868 4 neo Coleoptera Scarabaeidae Ateuchus texanus Robinson 1948 1 nea Coleoptera Scarabaeidae Atrichelaphinis tigrina Olivier 2 afr Coleoptera Scarabaeidae Augosoma centaurus Fabricius 1775 18 afr Coleoptera Scarabaeidae Aulacopalpus ciliatus Solier 1851 neo Coleoptera Scarabaeidae Aulacopalpus variabilis Philippi 1861 1 neo Coleoptera Scarabaeidae Blackbolbus echinicollis Howden 1 aus Coleoptera Scarabaeidae Blackbolbus hollowayorum Howden 1 aus Coleoptera Scarabaeidae Blackbolbus inopinus Howden 1 aus Coleoptera Scarabaeidae Blaesia atra Burmeister 1842 4 neo Coleoptera Scarabaeidae Blitopertha campestris Latreille 1804 11 pal Coleoptera Scarabaeidae Blitopertha conspurcata Harold 1878 2 pal Coleoptera Scarabaeidae Blitopertha lineolata Fischer 1823 1 pal Coleoptera Scarabaeidae Blitopertha orientalis Waterhouse 1875 10 ori Coleoptera Scarabaeidae Bolbites onitoides Harold 1868 5 neo Coleoptera Scarabaeidae Bolborhachium anneae Howden 1 aus Coleoptera Scarabaeidae Bolborhachium concavum Howden 1 aus Coleoptera Scarabaeidae Bolborhachium deceptum Howden 1 aus Coleoptera Scarabaeidae Bolborhachium inclinatum Howden 1 aus Coleoptera Scarabaeidae Bolborhachium keithi Howden 1 aus Coleoptera Scarabaeidae Bolborhachium nanum Howden 1 aus Coleoptera Scarabaeidae Boreocanthon ebenus Say 1823 17 nea Coleoptera Scarabaeidae Boreocanthon lecontei Harold 1870 2 nea Coleoptera Scarabaeidae Boreocanthon melanus Robinson 1948 1 nea Coleoptera Scarabaeidae Boreocanthon praticola LeConte 1859 13 nea Coleoptera Scarabaeidae Boreocanthon probus Germar 1824 19 nea Coleoptera Scarabaeidae Boreocanthon puncticollis LeConte 1866 5 nea Coleoptera Scarabaeidae Boreocanthon simplex LeConte 1857 73 nea Coleoptera Scarabaeidae Bothynus dasypleurus Germar 1824 4 neo Coleoptera Scarabaeidae Bothynus entellus Serville 1825 4 neo Coleoptera Scarabaeidae Bothynus laevicollis Bates 1888 1 neo Coleoptera Scarabaeidae Bothynus selanderi Cartwright 1959 2 neo Coleoptera Scarabaeidae Bothynus sp. 2 neo Coleoptera Scarabaeidae Bothynus spp. 7 neo Coleoptera Scarabaeidae Bothynus spp. 18 neo Coleoptera Scarabaeidae Brahmina dilaticolis Ballion 1870 1 pal Coleoptera Scarabaeidae Bubas bison Linnaeus 1767 6 pal Coleoptera Scarabaeidae Bubas bubalus Olivier 1811 1 pal Coleoptera Scarabaeidae Caccobius mundus Menetries 1838 1 pal Coleoptera Scarabaeidae Caccobius schreberi Linnaeus 1767 16 pal Coleoptera Scarabaeidae Caelorrhina relucens Bates 1881 1 afr Coleoptera Scarabaeidae Caelorrhina superba Gerstaecker 1883 4 afr Coleoptera Scarabaeidae Calomacraspis concinna Blanchard 1850 2 neo Coleoptera Scarabaeidae Calomacraspis splendens Burmeister 1844 1 neo Coleoptera Scarabaeidae Calonota inconstans Brookes 1 aus Coleoptera Scarabaeidae Calonota sobrina Sharp 1876 4 aus Coleoptera Scarabaeidae Cantharsius exiguus Boheman 1860 6 afr Coleoptera Scarabaeidae Cantharsius exiguus Bohemann 1860 6 afr Coleoptera Scarabaeidae Canthidium aeneolum Harold 1867 1 neo Coleoptera Scarabaeidae Canthidium ardens Bates 1887 1 neo Coleoptera Scarabaeidae Canthidium aurifex Bates 1887 1 neo Coleoptera Scarabaeidae Canthidium breve Germar 1824 6 neo Coleoptera Scarabaeidae Canthidium haroldi P.deBorre 1886 4 neo Coleoptera Scarabaeidae Canthidium leucopterum Howden and Young 1981 1 neo Coleoptera Scarabaeidae Canthidium lucidum Harold 1867 1 neo Coleoptera Scarabaeidae Canthidium planovultum Howden and Young 1981 2 neo Coleoptera Scarabaeidae Canthidium sp. 2 neo Coleoptera Scarabaeidae Canthidium tenebrosum Howden and Young 1981 2 neo Coleoptera Scarabaeidae Canthon affinis Fabricius 1801 1 neo Coleoptera Scarabaeidae Canthon angularis Harold 1868 2 neo Coleoptera Scarabaeidae Canthon angustatus Harold 1867 2 neo Coleoptera Scarabaeidae Canthon bispinatus Robinson 1941 1 nea Coleoptera Scarabaeidae Canthon callosus Harold 1868 3 neo Coleoptera Scarabaeidae Canthon chalcites Halderman 1843 25 nea Coleoptera Scarabaeidae Canthon chevrolati Harold 1868 1 neo Coleoptera Scarabaeidae Canthon coeruleicollis Blanchard 1843 2 neo Coleoptera Scarabaeidae Canthon conformis Harold 1868 6 neo Coleoptera Scarabaeidae Canthon curvipes Harold 1868 4 neo Coleoptera Scarabaeidae Canthon curvodilatum Schaum 3 neo Coleoptera Scarabaeidae Canthon cyanellus LeConte 1859 3 neo Coleoptera Scarabaeidae Canthon cyanellus LeConte 1859 10 nea Coleoptera Scarabaeidae Canthon depressipennis LeConte 1859 4 nea Coleoptera Scarabaeidae Canthon dives Harold 1868 2 neo Coleoptera Scarabaeidae Canthon fastuosus Harold 1868 4 neo Coleoptera Scarabaeidae Canthon forreri Bates 1887 1 neo Coleoptera Scarabaeidae Canthon fulgidus Redtenbacher 1867 4 neo Coleoptera Scarabaeidae Canthon humectus Say 1832 26 neo Coleoptera Scarabaeidae Canthon humectus Say 1832 44 nea Coleoptera Scarabaeidae Canthon imitator Brown 42 nea Coleoptera Scarabaeidae Canthon lamprimus Bates 1887 1 neo Coleoptera Scarabaeidae Canthon lividus Blanchard 1843 2 neo Coleoptera Scarabaeidae Canthon moniliatus Bates 1887 1 neo Coleoptera Scarabaeidae Canthon mutabilis Lucas 1857 1 neo Coleoptera Scarabaeidae Canthon nigricornis Say 1823 31 nea Coleoptera Scarabaeidae Canthon pilularius LeConte 1758 124 nea Coleoptera Scarabaeidae Canthon quinquemaculatus Castelnau 1840 1 neo Coleoptera Scarabaeidae Canthon rugosus Blanchard 1868 7 neo Coleoptera Scarabaeidae Canthon septemmaculatus Latreille 1807 13 neo Coleoptera Scarabaeidae Canthon smaragdulus Fabricius 1781 5 neo Coleoptera Scarabaeidae Canthon speculifer Castelnau 1840 2 neo Coleoptera Scarabaeidae Canthon spp. 8 nea Coleoptera Scarabaeidae Canthon triangularis Drury 1770 19 neo Coleoptera Scarabaeidae Canthon tristis Harold 1862 7 neo Coleoptera Scarabaeidae Canthon vigilans LeConte 1858 21 nea Coleoptera Scarabaeidae Canthon violaceus Olivier 1789 4 neo Coleoptera Scarabaeidae Canthon viridis Beauvois 1805 4 nea Coleoptera Scarabaeidae Cartwrightia intertribalis Islas 1958 11 neo Coleoptera Scarabaeidae Catharsius molossus Linnaeus 1758 9 ori Coleoptera Scarabaeidae Catharsius pithecius Fabricius 1775 8 ori Coleoptera Scarabaeidae Catharsius sp. 1 ori Coleoptera Scarabaeidae Celidota stephens Gory and Percheron 1833 2 afr Coleoptera Scarabaeidae Cetonia aurata Linnaeus 1761 52 pal Coleoptera Scarabaeidae Cetonia aurata Reitter 1896 1 pal Coleoptera Scarabaeidae Cetonia aurata Fieber 1831 3 pal Coleoptera Scarabaeidae Cetonia aurata Reitter 1896 9 pal Coleoptera Scarabaeidae Cetonia aurata Reitter 1896 3 pal Coleoptera Scarabaeidae Cetonia aurata Reitter 1896 2 pal Coleoptera Scarabaeidae Cetonia aurata Reitter 1898 4 pal Coleoptera Scarabaeidae Cetonia aurata Reitter 1896 1 pal Coleoptera Scarabaeidae Cetonia chrysosoma Reitter 1896 1 pal Coleoptera Scarabaeidae Cetonia magnifica Ballion 1870 4 pal Coleoptera Scarabaeidae Cetonia pilifera Motschulsky 1860 1 pal Coleoptera Scarabaeidae Cetonia roelofsi Harold 1880 5 pal Coleoptera Scarabaeidae Cetonia viridiopaca Motschulsky 1860 1 pal Coleoptera Scarabaeidae Chalcocopris hesperus Olivier 1789 1 neo Coleoptera Scarabaeidae Chalcocrates sp. 1 aus Coleoptera Scarabaeidae Chalcoplethis velutipes Arrow 1900 1 neo Coleoptera Scarabaeidae Chalcosoma atlas Linnaeus 1758 5 ori Coleoptera Scarabaeidae Chalcosoma atlas Erichson 1834 2 ori Coleoptera Scarabaeidae Chalcosoma sp. 2 ori Coleoptera Scarabaeidae Chalepides barbatus Fabricius 1787 16 neo Coleoptera Scarabaeidae Chalepides spp. 3 neo Coleoptera Scarabaeidae Chasmatopterus hirtulus Illiger 1803 1 pal Coleoptera Scarabaeidae Chasmatopterus vilosulus Illiger 1803 1 pal Coleoptera Scarabaeidae Cheiroplatys borealis Carne 1957 1 aus Coleoptera Scarabaeidae Chelorrhina polyphemus Fabricius 1781 4 afr Coleoptera Scarabaeidae Chelorrhina savagei Harris 1844 3 afr Coleoptera Scarabaeidae Chioneosoma gorilla Brenske 1886 1 pal Coleoptera Scarabaeidae Chioneosoma mucosum Semen 1 pal Coleoptera Scarabaeidae Chioneosoma porosum Fischer 1824 1 pal Coleoptera Scarabaeidae Chioneosoma reitteri Semen 1887 1 pal Coleoptera Scarabaeidae Chiron cylindrus Fabricius 1798 1 pal Coleoptera Scarabaeidae Chiron grandis Gory 1829 14 afr Coleoptera Scarabaeidae Chiron puncticollis Harold 1862 2 afr Coleoptera Scarabaeidae Chironitis furcifer Rossi 1792 5 pal Coleoptera Scarabaeidae Chironitis hungaricus Herbst 1789 3 pal Coleoptera Scarabaeidae Chironitis irroratus Rossi 1792 1 pal Coleoptera Scarabaeidae Chlorocala africana Drury 1773 1 afr Coleoptera Scarabaeidae Chlorocala africana Harold 1878 1 afr Coleoptera Scarabaeidae Chlorocala sp. 1 afr Coleoptera Scarabaeidae Chlorocala viridicyanea Pallas 1805 1 afr Coleoptera Scarabaeidae Chlorochiton suturalis Fabricius 1775 3 aus Coleoptera Scarabaeidae Chnaunanthus chapini Saylor 1937 5 nea Coleoptera Scarabaeidae Chnaunanthus flavipennis Horn 1867 1 nea Coleoptera Scarabaeidae Chrysina amoena Sturm 1843 1 neo Coleoptera Scarabaeidae Chrysina macropus Francillon 1795 11 neo Coleoptera Scarabaeidae Chrysophora chrysochlora Latreille 1811 8 neo Coleoptera Scarabaeidae Clinteria spilota Gory and Percheron 1833 1 pal Coleoptera Scarabaeidae Clinteroides permutans Burmeister 1847 2 afr Coleoptera Scarabaeidae Cnemida lacerata Germar 1824 1 neo Coleoptera Scarabaeidae Cnemida retusa Fabricius 1801 10 neo Coleoptera Scarabaeidae Coelosis bicornis Leske 1779 28 neo Coleoptera Scarabaeidae Coelosis biloba Linnaeus 1767 10 neo Coleoptera Scarabaeidae Coelosis sp.1 1 neo Coleoptera Scarabaeidae Coelosis sp.2 2 neo Coleoptera Scarabaeidae Coelosis sp.3 1 neo Coleoptera Scarabaeidae Coenoyncha fusca McClay 1943 2 nea Coleoptera Scarabaeidae Coenoyncha testacea Cazier 1937 1 nea Coleoptera Scarabaeidae Copris arizonensis Schaeffer 1906 13 nea Coleoptera Scarabaeidae Copris boucardi Harold 1869 1 neo Coleoptera Scarabaeidae Copris capensis Waterhouse 1891 2 afr Coleoptera Scarabaeidae Copris costaricensis Gahan 1894 1 neo Coleoptera Scarabaeidae Copris eudamas Olivier??? 1 neo Coleoptera Scarabaeidae Copris fidius Olivier 1789 1 ori Coleoptera Scarabaeidae Copris fricator Fabricius 1787 44 nea Coleoptera Scarabaeidae Copris hispanus Linnaeus 1764 4 pal Coleoptera Scarabaeidae Copris howdeni Matthews and Halffter, 1 nea Coleoptera Scarabaeidae Copris incertus Say 1835 1 neo Coleoptera Scarabaeidae Copris incertus Say 1835 2 neo Coleoptera Scarabaeidae Copris inemarginatus Blatchley 1918 1 nea Coleoptera Scarabaeidae Copris jacchus Fabricius 1775 1 afr Coleoptera Scarabaeidae Copris klugi Harold 1869 4 neo Coleoptera Scarabaeidae Copris klugi Harold 1869 12 neo Coleoptera Scarabaeidae Copris laeviceps Harold 1869 1 neo Coleoptera Scarabaeidae Copris lecontei Matthews 5 neo Coleoptera Scarabaeidae Copris lugubris Boheman 1858 6 neo Coleoptera Scarabaeidae Copris lugubris Bohemann 1858 6 neo Coleoptera Scarabaeidae Copris lunaris Linnaeus 1758 19 pal Coleoptera Scarabaeidae Copris macer Peringuey 1900 1 afr Coleoptera Scarabaeidae Copris minutus Drury 1773 76 nea Coleoptera Scarabaeidae Copris moechus LeConte 1854 1 neo Coleoptera Scarabaeidae Copris nevinsoni Waterhouse 1891 3 ori Coleoptera Scarabaeidae Copris ochus Motschulsky 1860 2 ori Coleoptera Scarabaeidae Copris remotus LeConte 1866 3 nea Coleoptera Scarabaeidae Copris repertus Walker 1858 4 ori Coleoptera Scarabaeidae Copris sacontala Redtenbacher 1848 1 ori Coleoptera Scarabaeidae Copris tullius Olivier 1789 47 nea Coleoptera Scarabaeidae Coproecus hemisphaericus Guerin 1830 1 aus Coleoptera Scarabaeidae Coptomia prasina Burmeister 1842 1 afr Coleoptera Scarabaeidae Coscinocephalus cribrifrons Schaeffer 1915 17 nea Coleoptera Scarabaeidae Coscinocephalus cribrifrons Schaeffer 1915 53 pal Coleoptera Scarabaeidae Cotalpa consobrina Horn 1871 13 neo Coleoptera Scarabaeidae Cotalpa consobrina Horn 1871 30 nea Coleoptera Scarabaeidae Cotalpa flavida Horn 1878 2 nea Coleoptera Scarabaeidae Cotalpa lanigera Linnaeus 1764 65 nea Coleoptera Scarabaeidae Cotalpa lanigera Casey 1915 7 nea Coleoptera Scarabaeidae Cotalpa subcrinita Wickham 1905 5 nea Coleoptera Scarabaeidae Cotinis lebas Gory and Percheron 1833 1 neo Coleoptera Scarabaeidae Cotinis mutabilis Gory and Percheron 1833 8 neo Coleoptera Scarabaeidae Cotinis mutabilis Gory and Percheron 1833 244 nea Coleoptera Scarabaeidae Cotinis nitida Linnaeus 1764 90 nea Coleoptera Scarabaeidae Cotinis pauperula Burmeister 1847 1 neo Coleoptera Scarabaeidae Cotinis sp. neo Coleoptera Scarabaeidae Cremastocheilus academicus Krikken 1982 3 neo Coleoptera Scarabaeidae Cremastocheilus angularis LeConte 1857 2 nea Coleoptera Scarabaeidae Cremastocheilus armatus Walker 1866 3 nea Coleoptera Scarabaeidae Cremastocheilus armatus Casey 1915 6 nea Coleoptera Scarabaeidae Cremastocheilus canaliculatus Kirby 1827 12 nea Coleoptera Scarabaeidae Cremastocheilus castanea Knoch 1801 51 nea Coleoptera Scarabaeidae Cremastocheilus constricticollis Cazier 1940 1 nea Coleoptera Scarabaeidae Cremastocheilus crinitis LeConte 1874 1 nea Coleoptera Scarabaeidae Cremastocheilus crinitus LeConte 1874 3 nea Coleoptera Scarabaeidae Cremastocheilus harrisi Kirby 1827 15 nea Coleoptera Scarabaeidae Cremastocheilus hirsutus VanDyke 1918 1 nea Coleoptera Scarabaeidae Cremastocheilus hirsutus VanDyke 1918 2 nea Coleoptera Scarabaeidae Cremastocheilus incisus Casey 1915 1 nea Coleoptera Scarabaeidae Cremastocheilus knochi LeConte 1853 5 nea Coleoptera Scarabaeidae Cremastocheilus lengi Cazier 1940 1 nea Coleoptera Scarabaeidae Cremastocheilus maritimus Casey 1915 2 nea Coleoptera Scarabaeidae Cremastocheilus mentalis Cazier 1940 1 nea Coleoptera Scarabaeidae Cremastocheilus mexicanus Schaum 1841 2 nea Coleoptera Scarabaeidae Cremastocheilus mexicanus Schaum 1841 5 nea Coleoptera Scarabaeidae Cremastocheilus nitens LeConte 1853 3 nea Coleoptera Scarabaeidae Cremastocheilus planatus LeConte 1863 1 nea Coleoptera Scarabaeidae Cremastocheilus planipes Horn 1885 3 nea Coleoptera Scarabaeidae Cremastocheilus schaumi LeConte 1853 3 nea Coleoptera Scarabaeidae Cremastocheilus squamulosus LeConte 1858 1 nea Coleoptera Scarabaeidae Cremastocheilus tomentosus Warner 1985 2 nea Coleoptera Scarabaeidae Cremastocheilus tridens Casey 1915 2 nea Coleoptera Scarabaeidae Cremastocheilus variolosus Kirby 1826 3 nea Coleoptera Scarabaeidae Ctenotis obesa Burmeister 1855 2 neo Coleoptera Scarabaeidae Cyclidinus lugubris Westwood 1874 2 afr Coleoptera Scarabaeidae Cyclidius sp. 1 neo Coleoptera Scarabaeidae Cyclocephala amazonica Fabricius 1781 1 neo Coleoptera Scarabaeidae Cyclocephala borealis Arrow 1911 47 nea Coleoptera Scarabaeidae Cyclocephala brittoni Endrodi 1964 1 neo Coleoptera Scarabaeidae Cyclocephala carbonaria Arrow 1911 1 neo Coleoptera Scarabaeidae Cyclocephala castanea Olivier 1789 3 neo Coleoptera Scarabaeidae Cyclocephala castaniella Bates 1888 1 neo Coleoptera Scarabaeidae Cyclocephala cubana Chapin 1932 2 neo Coleoptera Scarabaeidae Cyclocephala freudei Endrodi 1963 2 neo Coleoptera Scarabaeidae Cyclocephala helavai Endrodi 1975 1 neo Coleoptera Scarabaeidae Cyclocephala herteli Endrodi 1964 2 neo Coleoptera Scarabaeidae Cyclocephala hirta LeConte 1861 17 nea Coleoptera Scarabaeidae Cyclocephala longula LeConte 1863 36 nea Coleoptera Scarabaeidae Cyclocephala lunulata Burmeister 1847 26 neo Coleoptera Scarabaeidae Cyclocephala lurida Bland 1863 107 nea Coleoptera Scarabaeidae Cyclocephala mafaffa Burmeister 1847 10 neo Coleoptera Scarabaeidae Cyclocephala melanocephala Fabricius 1775 5 neo Coleoptera Scarabaeidae Cyclocephala melanocephala Fabricius 1775 156 nea Coleoptera Scarabaeidae Cyclocephala modesta Burmeister 1847 2 neo Coleoptera Scarabaeidae Cyclocephala nigricollis Burmeister 1847 4 nea Coleoptera Scarabaeidae Cyclocephala olivieri Arrow 1911 5 neo Coleoptera Scarabaeidae Cyclocephala ovulum Bates 1888 89 pal Coleoptera Scarabaeidae Cyclocephala parallela Casey 1915 31 nea Coleoptera Scarabaeidae Cyclocephala pasadenae Casey 1915 77 nea Coleoptera Scarabaeidae Cyclocephala prolongata Arrow 1902 1 neo Coleoptera Scarabaeidae Cyclocephala puberela LeConte 1873 3 nea Coleoptera Scarabaeidae Cyclocephala putrida Burmeister 1847 2 neo Coleoptera Scarabaeidae Cyclocephala seditiosa LeConte 1863 12 nea Coleoptera Scarabaeidae Cyclocephala signaticollis Burmeister 1847 6 neo Coleoptera Scarabaeidae Cyclocephala sinaloe Howden and Endrodi 1966 1 neo Coleoptera Scarabaeidae Cyclocephala spp. 144 Coleoptera Scarabaeidae Cyclocephala stictica Burmeister 1847 1 neo Coleoptera Scarabaeidae Cyclocephala testacea Burmeister 1847 5 neo Coleoptera Scarabaeidae Cyclocephala tetrica Burmeister 1847 6 neo Coleoptera Scarabaeidae Cyclocephala villosa Blanchard 1846 2 nea Coleoptera Scarabaeidae Cyphochilus farinosus Waterhouse 1867 1 pal Coleoptera Scarabaeidae Cyprolais hornimani Bates 1877 2 afr Coleoptera Scarabaeidae Cyriopertha glabra Semenov 1890 1 pal Coleoptera Scarabaeidae Cyriopertha glabra Reitter 1890 1 pal Coleoptera Scarabaeidae Dasyvalgus formosanis Moser 1915 2 ori Coleoptera Scarabaeidae Dasyvalgus spp. 14 ori Coleoptera Scarabaeidae Deltochilum amazomicum Bates 1887 2 neo Coleoptera Scarabaeidae Deltochilum brasiliense Castelnau 1840 1 neo Coleoptera Scarabaeidae Deltochilum denitpes Eschscholtz 1822 1 neo Coleoptera Scarabaeidae Deltochilum furcatum Castelnau 1840 1 neo Coleoptera Scarabaeidae Deltochilum gibbosum Fabricius 1775 19 nea Coleoptera Scarabaeidae Deltochilum icarus Olivier 1789 2 neo Coleoptera Scarabaeidae Deltochilum rubripenne Gory 1831 1 neo Coleoptera Scarabaeidae Deltochilum scabriusculum Bates 1887 1 neo Coleoptera Scarabaeidae Deltochilum tesselatum Bates 1870 1 neo Coleoptera Scarabaeidae Democrates burmeisteri Reiche 1852 1 neo Coleoptera Scarabaeidae Dendropemon convexus Harold 1869 1 Coleoptera Scarabaeidae Dialytellus dialytoides Fall 1907 11 nea Coleoptera Scarabaeidae Dialytellus humeralis LeConte 1878 2 nea Coleoptera Scarabaeidae Dialytes criddlei Brown,??? 2 nea Coleoptera Scarabaeidae Dialytes n.sp. 3 neo Coleoptera Scarabaeidae Dialytes striatulus Say 1825 19 nea Coleoptera Scarabaeidae Dialytes trunctatus Melsheimer 1844 23 nea Coleoptera Scarabaeidae Dialytes ulkei Horn 1875 4 nea Coleoptera Scarabaeidae Diaphonia mniszechi Janson 1873 2 aus Coleoptera Scarabaeidae Dichelonyx albicollis Burmeister 1855 52 nea Coleoptera Scarabaeidae Dichelonyx backii Barrett 1933 15 nea Coleoptera Scarabaeidae Dichelonyx canadensis Horn 1876 6 nea Coleoptera Scarabaeidae Dichelonyx crotchi Horn 1876 5 nea Coleoptera Scarabaeidae Dichelonyx decolorata Fall 1901 11 nea Coleoptera Scarabaeidae Dichelonyx diluta Fall 1901 16 nea Coleoptera Scarabaeidae Dichelonyx elongata Fabricius 1792 165 nea Coleoptera Scarabaeidae Dichelonyx elongatula Schonherr 1817 12 nea Coleoptera Scarabaeidae Dichelonyx fulgida LeConte 1856 45 nea Coleoptera Scarabaeidae Dichelonyx fuscula LeConte 1856 6 nea Coleoptera Scarabaeidae Dichelonyx lateralis Fall 1901 1 nea Coleoptera Scarabaeidae Dichelonyx linearis Gyllenhal 1817 3 nea Coleoptera Scarabaeidae Dichelonyx muscula Fall 1901 9 nea Coleoptera Scarabaeidae Dichelonyx nana Fall 1901 1 nea Coleoptera Scarabaeidae Dichelonyx pusilla LeConte 1856 11 nea Coleoptera Scarabaeidae Dichelonyx sp. 5 nea Coleoptera Scarabaeidae Dichelonyx subvittata LeConte 1856 47 nea Coleoptera Scarabaeidae Dichelonyx sulcata LeConte 1856 2 nea Coleoptera Scarabaeidae Dichelonyx sulcata Saylor 1933 9 nea Coleoptera Scarabaeidae Dichelonyx testacea Kirby 1837 5 nea Coleoptera Scarabaeidae Dichelonyx truncata LeConte 1856 26 nea Coleoptera Scarabaeidae Dichelonyx vaga Fall 1901 5 nea Coleoptera Scarabaeidae Dichelonyx valida LeConte 1856 3 nea Coleoptera Scarabaeidae Dichelonyx vicina Fall 1901 10 nea Coleoptera Scarabaeidae Dichelonyx vicina Hopping 1931 1 nea Coleoptera Scarabaeidae Dicheros burmeisteri Schoch 1898 2 ori Coleoptera Scarabaeidae Dichotomius anaglypticus Mannerheim 1829 5 neo Coleoptera Scarabaeidae Dichotomius ascanius Harold 1869 4 neo Coleoptera Scarabaeidae Dichotomius carolinus Linnaeus 1 neo Coleoptera Scarabaeidae Dichotomius carolinus Linnaeus 40 nea Coleoptera Scarabaeidae Dichotomius carolinus Say 21 nea Coleoptera Scarabaeidae Dichotomius carolinus Say 1835 2 neo Coleoptera Scarabaeidae Dichotomius colonicus Say 1835 1 nea Coleoptera Scarabaeidae Dichotomius fimbriatus Harold 1869 5 neo Coleoptera Scarabaeidae Dichotomius glaucus Harold 1869 12 neo Coleoptera Scarabaeidae Dichotomius longiceps Taschb. 1870 4 neo Coleoptera Scarabaeidae Dichotomius mormon Ljungh. 1799 5 neo Coleoptera Scarabaeidae Dichotomius nisus Olivier 1789 10 neo Coleoptera Scarabaeidae Dichotomius semiaeneus Germar 1824 4 neo Coleoptera Scarabaeidae Dichotomius sexdentatus Luderwaldt 1 neo Coleoptera Scarabaeidae Dichotomius torulosus Eschscholtz 1822 2 neo Coleoptera Scarabaeidae Dicranocephalus bowringi Pascoe 3 ori Coleoptera Scarabaeidae Dicronorhina derbyana Westwood 1843 2 afr Coleoptera Scarabaeidae Dicronorhina oberthueri Deyrolle 1876 3 afr Coleoptera Scarabaeidae Didactylia knausi Fall 1905 5 nea Coleoptera Scarabaeidae Didactylia sp. 1 neo Coleoptera Scarabaeidae Didactylia sp. 3 neo Coleoptera Scarabaeidae Diloboderus abderus Sturm 1826 12 neo Coleoptera Scarabaeidae Dilochrosis balteata Vollenhoven 1871 2 aus Coleoptera Scarabaeidae Dinacoma caseyi Blaisdell 1930 5 nea Coleoptera Scarabaeidae Dipelicus optatus Sharp 1875 1 aus Coleoptera Scarabaeidae Diphucephala colaspidoides Gyllenhal 1817 1 aus Coleoptera Scarabaeidae Diplognatha gagates Forster 1771 14 afr Coleoptera Scarabaeidae Diplognatha gagates MacLeay 1838 4 afr Coleoptera Scarabaeidae Diplognatha subaenea Duviv 1891 1 afr Coleoptera Scarabaeidae Diplotaxis abnormis Fall 1909 2 nea Coleoptera Scarabaeidae Diplotaxis aequalis Cazier 1940 2 nea Coleoptera Scarabaeidae Diplotaxis ambigua Fall 1909 2 nea Coleoptera Scarabaeidae Diplotaxis angularis LeConte 1856 59 nea Coleoptera Scarabaeidae Diplotaxis angustula Moser 1918 4 neo Coleoptera Scarabaeidae Diplotaxis arizonica Schaeffer 1907 4 nea Coleoptera Scarabaeidae Diplotaxis atlantis Fall 1909 18 nea Coleoptera Scarabaeidae Diplotaxis atramentaria Bates 1888 5 neo Coleoptera Scarabaeidae Diplotaxis barbarae Vaurie and Cazier 1955 1 neo Coleoptera Scarabaeidae Diplotaxis belfragei Fall 1909 115 nea Coleoptera Scarabaeidae Diplotaxis bidentata LeConte 1856 4 nea Coleoptera Scarabaeidae Diplotaxis boops Bates 1888 5 nea Coleoptera Scarabaeidae Diplotaxis brevicollis LeConte 1856 6 nea Coleoptera Scarabaeidae Diplotaxis brevicollis LeConte 1856 57 nea Coleoptera Scarabaeidae Diplotaxis brevicornis Cazier 1940 1 nea Coleoptera Scarabaeidae Diplotaxis brevidens LeConte 1856 1 nea Coleoptera Scarabaeidae Diplotaxis carbonata LeConte 1856 4 nea Coleoptera Scarabaeidae Diplotaxis chiricahuae Fall 1909 11 nea Coleoptera Scarabaeidae Diplotaxis chiricahuae Fall 1909 32 nea Coleoptera Scarabaeidae Diplotaxis conformis Fall 1909 1 nea Coleoptera Scarabaeidae Diplotaxis conformis Fall 1909 5 nea Coleoptera Scarabaeidae Diplotaxis costulata Fall 1909 1 nea Coleoptera Scarabaeidae Diplotaxis cribratella Bates 1889 3 neo Coleoptera Scarabaeidae Diplotaxis cribulosa LeConte 1856 239 nea Coleoptera Scarabaeidae Diplotaxis cribulosa LeConte 1856 227 nea Coleoptera Scarabaeidae Diplotaxis dentella Fall 1909 6 nea Coleoptera Scarabaeidae Diplotaxis dubia LeConte 1856 2 nea Coleoptera Scarabaeidae Diplotaxis fimbriata Fall 1909 5 nea Coleoptera Scarabaeidae Diplotaxis frondicola Say 1825 28 nea Coleoptera Scarabaeidae Diplotaxis fulva LeConte 1856 1 nea Coleoptera Scarabaeidae Diplotaxis harperi Blanchard 1850 20 nea Coleoptera Scarabaeidae Diplotaxis haydeni LeConte 1856 7 nea Coleoptera Scarabaeidae Diplotaxis hebes Bates 1888 1 neo Coleoptera Scarabaeidae Diplotaxis hirsuta Vaurie 1958 4 neo Coleoptera Scarabaeidae Diplotaxis illustris Fall 1909 1 nea Coleoptera Scarabaeidae Diplotaxis ingenua Fall 1909 25 nea Coleoptera Scarabaeidae Diplotaxis jamaicensis Cazier 1952 6 neo Coleoptera Scarabaeidae Diplotaxis knausi Schaeffer 1907 12 nea Coleoptera Scarabaeidae Diplotaxis knausii Schaeffer 1907 2 nea Coleoptera Scarabaeidae Diplotaxis languida LeConte 1878 12 nea Coleoptera Scarabaeidae Diplotaxis latispina Vaurie 1960 6 neo Coleoptera Scarabaeidae Diplotaxis liberta Germar 1824 11 nea Coleoptera Scarabaeidae Diplotaxis magna Bates 1888 4 nea Coleoptera Scarabaeidae Diplotaxis magna Bates 1888 7 nea Coleoptera Scarabaeidae Diplotaxis marginicollis Fall 1909 14 nea Coleoptera Scarabaeidae Diplotaxis marginicollis Fall 1909 144 nea Coleoptera Scarabaeidae Diplotaxis maura Fall 1909 6 nea Coleoptera Scarabaeidae Diplotaxis mentalis Fall 1909 14 nea Coleoptera Scarabaeidae Diplotaxis mimosae Fall 101 nea Coleoptera Scarabaeidae Diplotaxis mimosae Fall 1909 4 nea Coleoptera Scarabaeidae Diplotaxis moerens LeConte 1856 9 nea Coleoptera Scarabaeidae Diplotaxis muricata Schaeffer 1907 1 nea Coleoptera Scarabaeidae Diplotaxis mus Fall 1932 1 neo Coleoptera Scarabaeidae Diplotaxis obscura LeConte 1856 24 nea Coleoptera Scarabaeidae Diplotaxis pacata LeConte 1856 34 nea Coleoptera Scarabaeidae Diplotaxis parallela Fall 1909 3 nea Coleoptera Scarabaeidae Diplotaxis parvicollis Fall 1909 1 nea Coleoptera Scarabaeidae Diplotaxis pinalica Fall 1932 7 nea Coleoptera Scarabaeidae Diplotaxis planidens Fall 1909 1 nea Coleoptera Scarabaeidae Diplotaxis poropyge Bates 1887 2 neo Coleoptera Scarabaeidae Diplotaxis punctata LeConte 1856 3 nea Coleoptera Scarabaeidae Diplotaxis punctaturugosa Blanchard 1850 22 nea Coleoptera Scarabaeidae Diplotaxis rosae Vaurie and Cazier 1955 1 neo Coleoptera Scarabaeidae Diplotaxis rudis LeConte 1859 1 nea Coleoptera Scarabaeidae Diplotaxis rufa Linell 1896 1 nea Coleoptera Scarabaeidae Diplotaxis rufiola Fall 1909 8 nea Coleoptera Scarabaeidae Diplotaxis sierrae Fall 1909 1 nea Coleoptera Scarabaeidae Diplotaxis simplex Blanchard 1850 4 neo Coleoptera Scarabaeidae Diplotaxis sordida Say 1825 111 nea Coleoptera Scarabaeidae Diplotaxis sp. pubereagroup 1 neo Coleoptera Scarabaeidae Diplotaxis sp. trapeziferagroup 5 neo Coleoptera Scarabaeidae Diplotaxis spp. 16 nea Coleoptera Scarabaeidae Diplotaxis statura Cazier 2 nea Coleoptera Scarabaeidae Diplotaxis subangulata Blanchard 1850 1 nea Coleoptera Scarabaeidae Diplotaxis subangulata LeConte 1856 93 nea Coleoptera Scarabaeidae Diplotaxis subangulata Schaeffer 1907 6 neo Coleoptera Scarabaeidae Diplotaxis subcostata Blanchard 1850 44 nea Coleoptera Scarabaeidae Diplotaxis sulcatulus Fall 1909 6 nea Coleoptera Scarabaeidae Diplotaxis tarsalis Schaeffer 1907 2 nea Coleoptera Scarabaeidae Diplotaxis texana LeConte 1856 11 nea Coleoptera Scarabaeidae Diplotaxis thoracica Fall 1909 12 nea Coleoptera Scarabaeidae Diplotaxis trapezifera Bates 1887 71 neo Coleoptera Scarabaeidae Diplotaxis trementina Fall 1909 2 nea Coleoptera Scarabaeidae Diplotaxis tristis Kirby 1837 93 nea Coleoptera Scarabaeidae Diplotaxis truncatula LeConte 1856 63 nea Coleoptera Scarabaeidae Dischista cincta DeGeer 1778 1 afr Coleoptera Scarabaeidae Dymusia cyanea Olivier 1789 1 afr Coleoptera Scarabaeidae Dynastes granti Horn 46 nea Coleoptera Scarabaeidae Dynastes hercules Linnaeus 1758 9 neo Coleoptera Scarabaeidae Dynastes hercules Verril 1906 1 neo Coleoptera Scarabaeidae Dynastes hercules Scopoli 1769 1 neo Coleoptera Scarabaeidae Dynastes hyllus Chevrolat 1843 2 neo Coleoptera Scarabaeidae Dynastes neptunus Quensel 1817 2 neo Coleoptera Scarabaeidae Dynastes tityus Linnaeus 1763 68 nea Coleoptera Scarabaeidae Dyscinetus dubius Olivier 1789 149 neo Coleoptera Scarabaeidae Dyscinetus gagates Burmeister 1847 2 neo Coleoptera Scarabaeidae Dyscinetus morator Fabricius 1798 150 nea Coleoptera Scarabaeidae Dyscinetus picipes Burmeister 1847 17 neo Coleoptera Scarabaeidae Dyscinetus picipes Burmeister 1847 24 nea Coleoptera Scarabaeidae Dyscinetus rugifrons Burmeister 1847 6 neo Coleoptera Scarabaeidae Dyscinetus sp. 3 neo Coleoptera Scarabaeidae Dyscinetus spp. 72 nea Coleoptera Scarabaeidae Dyscinetus spp. 176 Coleoptera Scarabaeidae Endroedyantus paradoxus Scholtz and Howden 1 afr Coleoptera Scarabaeidae Enema pan Fabricius 1775 11 neo Coleoptera Scarabaeidae Enema pan Fabricius 1787 6 neo Coleoptera Scarabaeidae Enema sp. 10 neo Coleoptera Scarabaeidae Eophileurus chinensis Falderman 1835 1 pal Coleoptera Scarabaeidae Epadoretus reitteri Semenov 1890 1 pal Coleoptera Scarabaeidae Epectinaspis mexicanus Burmeister 1844 8 neo Coleoptera Scarabaeidae Epectinaspis opacicollis Bates 1888 1 neo Coleoptera Scarabaeidae Epirinus silvestris Cambefort 2 afr Coleoptera Scarabaeidae Epirinus validus Peringuey 1901 1 afr Coleoptera Scarabaeidae Eremazus unistriatus Mulsant 1854 1 pal Coleoptera Scarabaeidae Erioscelis emarginata Mannerheim 1829 33 neo Coleoptera Scarabaeidae Eucanthus impressus Howden 1 nea Coleoptera Scarabaeidae Euchirus dupontianus Burmeister 1 ori Coleoptera Scarabaeidae Euchirus longimanus Linnaeus 1 ori Coleoptera Scarabaeidae Euchirus macleayi Hope 4 ori Coleoptera Scarabaeidae Euchirus parryi Gray 2 ori Coleoptera Scarabaeidae Euchirus sp. 2 ori Coleoptera Scarabaeidae Euchloropus laetus Fabricius 1801 7 ori Coleoptera Scarabaeidae Eucranium arachnoides Brulle 1834 1 neo Coleoptera Scarabaeidae Eudicella gralli Buquet 1836 3 afr Coleoptera Scarabaeidae Eudicella morgani White 1839 2 afr Coleoptera Scarabaeidae Euheptaulacus carinatus Germar 1824 2 pal Coleoptera Scarabaeidae Euheptaulacus sus Herbst 1783 4 pal Coleoptera Scarabaeidae Euheptaulacus villosus Gyllenhal 1808 1 pal Coleoptera Scarabaeidae Euparia castanea Lepeletier and Serville 1828 14 nea Coleoptera Scarabaeidae Euparia spp. 3 neo Coleoptera Scarabaeidae Eupatorus hardwickii Hope 1831 4 ori Coleoptera Scarabaeidae Euphoria basalis Gory and Percheron 1833 36 neo Coleoptera Scarabaeidae Euphoria biguttata Gory and Percheron 1833 3 neo Coleoptera Scarabaeidae Euphoria candezei Janson 1875 1 neo Coleoptera Scarabaeidae Euphoria canescens Gory and Percheron 1833 2 neo Coleoptera Scarabaeidae Euphoria dimidiata Gory and Percheron 1833 2 neo Coleoptera Scarabaeidae Euphoria fulgida Fabricius 1775 49 nea Coleoptera Scarabaeidae Euphoria fulgida Casey 1915 13 nea Coleoptera Scarabaeidae Euphoria fulgida Fall 1905 44 nea Coleoptera Scarabaeidae Euphoria fulgida Fall 1905 8 nea Coleoptera Scarabaeidae Euphoria herbacea Olivier 1789 10 nea Coleoptera Scarabaeidae Euphoria histrionica Thomas 1878 32 nea Coleoptera Scarabaeidae Euphoria inda Linnaeus 1760 101 nea Coleoptera Scarabaeidae Euphoria inda Klages 1894 Coleoptera Scarabaeidae Euphoria kernii Haldeman 1852 95 nea Coleoptera Scarabaeidae Euphoria lineoligera Blanchard 1850 2 neo Coleoptera Scarabaeidae Euphoria pulchella Gory and Percheron 1833 17 neo Coleoptera Scarabaeidae Euphoria sepulchralis Fabricius 1801 227 nea Coleoptera Scarabaeidae Euphoria sepulchralis Casey 1915 17 nea Coleoptera Scarabaeidae Euphoria sepulchralis Gory and Percheron 1833 4 neo Coleoptera Scarabaeidae Euphoria sepulchralis Gory and Percheron 1833 12 nea Coleoptera Scarabaeidae Euphoria sp. 2 nea Coleoptera Scarabaeidae Euphoria subtomentosa Mannerheim 1837 3 neo Coleoptera Scarabaeidae Euphoriopsis hirtipes Horn 1880 2 nea Coleoptera Scarabaeidae Eupoecila australasiae Donovan 1805 1 aus Coleoptera Scarabaeidae Europtron confusum Marseul? 1878 1 pal Coleoptera Scarabaeidae Eutheola bidentata Burmeister 1847 66 neo Coleoptera Scarabaeidae Eutheola humilis LeConte 1856 23 nea Coleoptera Scarabaeidae Eutheola sp. 2 pal Coleoptera Scarabaeidae Eutheola sp. 18 nea Coleoptera Scarabaeidae Eutheola spp. 64 neo Coleoptera Scarabaeidae Eutheola subglabra Schaeffer 1909 6 nea Coleoptera Scarabaeidae Exolontha mandarinen Fairmaire 1 ori Coleoptera Scarabaeidae Gametis jucunda Falderman 1835 16 ori Coleoptera Scarabaeidae Gametis jucunda Gory and Percheron 1833 5 ori Coleoptera Scarabaeidae Gametis jucunda Hope 1831 3 ori Coleoptera Scarabaeidae Gametis jucunda Reitter 1896 2 ori Coleoptera Scarabaeidae Gametis jucunda Reitter 1898 1 ori Coleoptera Scarabaeidae Gametis versicolor Arrow 1910 2 ori Coleoptera Scarabaeidae Gametis versicolor Arrow 1910 1 ori Coleoptera Scarabaeidae Gametis versicolor Arrow 1910 1 ori Coleoptera Scarabaeidae Gametis versicolor Arrow 1910 2 ori Coleoptera Scarabaeidae Gametis versicolor Arrow 1910 1 ori Coleoptera Scarabaeidae Geniates bornelli Camerano 1894 2 neo Coleoptera Scarabaeidae Geniates spinolae Burmeister 1844 9 neo Coleoptera Scarabaeidae Genuchinus ineptus Horn 1885 2 nea Coleoptera Scarabaeidae Geopsammodius hydropicus Horn 1887 14 nea Coleoptera Scarabaeidae Geotrogus gabalus Fairmaire 1860 2 pal Coleoptera Scarabaeidae Glycyphana cuculus Burmeister 1842 8 ori Coleoptera Scarabaeidae Glycyphana fulvistemma Motschulsky 1860 5 pal Coleoptera Scarabaeidae Glycyphana magnifica Moser 1906 4 ori Coleoptera Scarabaeidae Glycyphana roquesi Bourgin 1916 2 ori Coleoptera Scarabaeidae Glycyphana rubripes Moser 1906 2 ori Coleoptera Scarabaeidae Glycyphana rubroscutellaris Mohnike 1873 1 ori Coleoptera Scarabaeidae Glycyphana sp. 1 afr Coleoptera Scarabaeidae Glycyphana sp. nr. mollendorfi 1 ori Coleoptera Scarabaeidae Glycyphana spp. 19 ori Coleoptera Scarabaeidae Glycyphana stolata Fabricius 2 aus Coleoptera Scarabaeidae Glycyphana tibialis Moser 1913 24 ori Coleoptera Scarabaeidae Gnathocera trivittata Swederus 1789 2 afr Coleoptera Scarabaeidae Gnathocera trivittata Swartz 1817 3 afr Coleoptera Scarabaeidae Gnorimella maculosa Knoch 16 nea Coleoptera Scarabaeidae Gnorimus nobilis Linnaeus 1758 9 pal Coleoptera Scarabaeidae Gnorimus octopunctatus Linnaeus 1758 1 pal Coleoptera Scarabaeidae Gnorimus octopunctatus Linnaeus 1758 3 pal Coleoptera Scarabaeidae Gnorimus subopacus Motschulsky 1860 1 pal Coleoptera Scarabaeidae Goliathus cacicus Voet 1779 4 afr Coleoptera Scarabaeidae Goliathus goliatus Drury 1770 10 afr Coleoptera Scarabaeidae Goliathus orientalis Moser 1909 1 afr Coleoptera Scarabaeidae Goliathus orientalis Moser 1909 2 afr Coleoptera Scarabaeidae Goliathus regius Klug 1835 4 afr Coleoptera Scarabaeidae Goliathus sp. 3 afr Coleoptera Scarabaeidae Golofa eacus Burmeister 1847 1 neo Coleoptera Scarabaeidae Golofa pelegon Burmeister 1847 2 neo Coleoptera Scarabaeidae Golofa pizarro Hope 1837 3 neo Coleoptera Scarabaeidae Golofa porteri Hope 1837 2 neo Coleoptera Scarabaeidae Golofa spp. 18 neo Coleoptera Scarabaeidae Gromphas aeruginosa Perty 1830 1 neo Coleoptera Scarabaeidae Gromphas lacordairei Brulle 1834 6 neo Coleoptera Scarabaeidae Gronocharus autumnalis Schaeffer 8 nea Coleoptera Scarabaeidae Gronocharus multispinosus Howden 1967 4 nea Coleoptera Scarabaeidae Guatemalica marginicollis Burmeister 1842 3 neo Coleoptera Scarabaeidae Gymnetis chalcipes Gory and Percheron 1833 1 neo Coleoptera Scarabaeidae Gymnetis cinerea Gory and Percheron 1833 6 neo Coleoptera Scarabaeidae Gymnetis cretacea LeConte 1863 2 nea Coleoptera Scarabaeidae Gymnetis holosericea Voet 1779 Coleoptera Scarabaeidae Gymnetis insculpta Kirby 1818 1 neo Coleoptera Scarabaeidae Gymnetis irregularis Gory and Percheron 1833 1 neo Coleoptera Scarabaeidae Gymnetis maculosa Olivier 1789 1 neo Coleoptera Scarabaeidae Gymnetis pantherina Burmeister 1842 1 neo Coleoptera Scarabaeidae Gymnetis rufolatris Illiger 1800 1 neo Coleoptera Scarabaeidae Gymnetis sallei Schaum 1849 5 nea Coleoptera Scarabaeidae Gymnetis sp. 1 neo Coleoptera Scarabaeidae Gymnetis sp. nr sallei 1 neo Coleoptera Scarabaeidae Gymnetis tigrina Gory and Percheron 1833 5 neo Coleoptera Scarabaeidae Gymnetis undulata Vigors 1825 2 neo Coleoptera Scarabaeidae Gymnopleurus aciculatus Gebler 1841 2 pal Coleoptera Scarabaeidae Gymnopleurus coriarius Mulsant 1842 1 pal Coleoptera Scarabaeidae Gymnopleurus coriarius Mulsant 1842 2 pal Coleoptera Scarabaeidae Gymnopleurus coriarius Fischer 1821 1 pal Coleoptera Scarabaeidae Gymnopleurus cyaneus Fabricius 1798 1 ori Coleoptera Scarabaeidae Gymnopleurus geoffroyi Fuessely 1775 4 pal Coleoptera Scarabaeidae Gymnopleurus miliaris Fabricius 1775 1 ori Coleoptera Scarabaeidae Gymnopleurus mopsus Pallas 1781 2 pal Coleoptera Scarabaeidae Gymnopleurus mopsus Pallas 1781 4 pal Coleoptera Scarabaeidae Gymnopleurus sinuatus Olivier 1789 1 ori Coleoptera Scarabaeidae Gymnopleurus smaragdifer Walker 1858 2 ori Coleoptera Scarabaeidae Gymnopleurus sturmi MacLeay 1821 9 pal Coleoptera Scarabaeidae Gymnopleurus sturmi Duft 1805 1 pal Coleoptera Scarabaeidae Gymnopleurus virens Erichson 1843 1 afr Coleoptera Scarabaeidae Gymnopyge hirsuta Cazier 1939 5 nea Coleoptera Scarabaeidae Gymnopyge hopliaformis Linell 1896 16 nea Coleoptera Scarabaeidae Hadrosticta viridiflua Kraatz 1892 1 neo Coleoptera Scarabaeidae Haplidia graeca Kraatz 1882 1 pal Coleoptera Scarabaeidae Haplidia transversa Fabricius 1801 2 pal Coleoptera Scarabaeidae Heliocopris bucephalus Fabricius 1775 6 ori Coleoptera Scarabaeidae Heliocopris dominus Bates 1868 1 ori Coleoptera Scarabaeidae Heliocopris gigas Olivier 1798 4 afr Coleoptera Scarabaeidae Heliocopris hamadryas Fabricius 1775 1 afr Coleoptera Scarabaeidae Heliocopris japetus Klug 1855 1 afr Coleoptera Scarabaeidae Heliocopris neptunis Boheman 1857 1 afr Coleoptera Scarabaeidae Heliocopris neptunis Bohemann 1857 1 afr Coleoptera Scarabaeidae Hemictenius nigrociliatus Reitter 1897 1 pal Coleoptera Scarabaeidae Hemictenius opacipes Reitter 1901 1 pal Coleoptera Scarabaeidae Hemictenius simplicitarsis Reitter 1897 1 pal Coleoptera Scarabaeidae Heptophylla picea Motschultsky 1857 1 ori Coleoptera Scarabaeidae Heterogomphus chevrolati Burmeister 1847 15 neo Coleoptera Scarabaeidae Heterogomphus dilaticollis Burmeister 1847 2 neo Coleoptera Scarabaeidae Heterogomphus pauson Perty 1830 5 neo Coleoptera Scarabaeidae Heterogomphus schonherri Burmeister 1847 5 neo Coleoptera Scarabaeidae Heteronychus arator Fabricius 1775 4 aus Coleoptera Scarabaeidae Heteronychus sp. 21 afr Coleoptera Scarabaeidae Heterophana lineatta Burmeister 1842 2 afr Coleoptera Scarabaeidae Heterorhina dohni Lansberge 1883 3 ori Coleoptera Scarabaeidae Heterorhina imperatrix Mohnike 1871 1 ori Coleoptera Scarabaeidae Heterorhina punctatissima Westwood 1842 4 ori Coleoptera Scarabaeidae Holotrichia diomphalia Bates 1888 1 pal Coleoptera Scarabaeidae Holotrichia morosa Waterhouse 1875 8 ori Coleoptera Scarabaeidae Homalopia marginata Fuessley 1775 1 pal Coleoptera Scarabaeidae Homalopia ruricola Fabricius 1775 4 pal Coleoptera Scarabaeidae Homalopia spiraeae Pallas 1773 4 pal Coleoptera Scarabaeidae Homonyx sp. 2 neo Coleoptera Scarabaeidae Homovalgus seticollis Beauvois 1805 24 nea Coleoptera Scarabaeidae Hoplia angulata Reitter 1902 1 pal Coleoptera Scarabaeidae Hoplia aulica Linnaeus 1766 2 pal Coleoptera Scarabaeidae Hoplia aulica Fabricius 1801 1 pal Coleoptera Scarabaeidae Hoplia aureola Pallas 1781 2 ori Coleoptera Scarabaeidae Hoplia callipyge LeConte 1856 63 nea Coleoptera Scarabaeidae Hoplia caucasica Kolenati 1846 1 pal Coleoptera Scarabaeidae Hoplia coerulea Drury 1773 31 neo Coleoptera Scarabaeidae Hoplia dispar LeConte 1880 11 nea Coleoptera Scarabaeidae Hoplia equina LeConte 1880 6 nea Coleoptera Scarabaeidae Hoplia farinosa Linnaeus 1761 1 pal Coleoptera Scarabaeidae Hoplia graminicola Fabricius 1792 2 pal Coleoptera Scarabaeidae Hoplia modesta Haldeman 1843 21 nea Coleoptera Scarabaeidae Hoplia parvula Krynicki 1832 1 pal Coleoptera Scarabaeidae Hoplia paupera Krynicki 1832 1 pal Coleoptera Scarabaeidae Hoplia philanthus Fussley 1775 14 pal Coleoptera Scarabaeidae Hoplia practicola Duftschmid 1805 2 pal Coleoptera Scarabaeidae Hoplia rufipes Motschulsky 1860 1 ori Coleoptera Scarabaeidae Hoplia sackenii LeConte 1880 5 nea Coleoptera Scarabaeidae Hoplia sp. 1 neo Coleoptera Scarabaeidae Hoplia sp. 3 pal Coleoptera Scarabaeidae Hoplia sulphurea Dufour 1843 8 pal Coleoptera Scarabaeidae Hoplia trifasciata Say 1823 177 nea Coleoptera Scarabaeidae Hoplia trivialis Harold 1869 3 nea Coleoptera Scarabaeidae Hoplopyga brasiliensis Gory and Percheron 1833 26 neo Coleoptera Scarabaeidae Hoplopyga liturata Olivier 1789 1 neo Coleoptera Scarabaeidae Hoplopyga miliaris Gory and Percheron 1833 3 neo Coleoptera Scarabaeidae Hoplopyga ocellata Gory and Percheron 1833 1 neo Coleoptera Scarabaeidae Hoplopyga sp.1 1 neo Coleoptera Scarabaeidae Hoplopyga sp.2 3 neo Coleoptera Scarabaeidae Hylamorpha elegans Burmeister 1844 6 neo Coleoptera Scarabaeidae Hylamorpha elegans Phillipi ? 4 neo Coleoptera Scarabaeidae Hymenoplia cinerascens Rosenhauer 1856 1 pal Coleoptera Scarabaeidae Hymenoplia rugulosa Mulsant 1842 2 pal Coleoptera Scarabaeidae Hypothyce mixta Howden 1968 2 nea Coleoptera Scarabaeidae Idutywa collaris Burmeister 1844 2 afr Coleoptera Scarabaeidae Inca clathratus Olivier 1792 4 neo Coleoptera Scarabaeidae Incala sp. 4 afr Coleoptera Scarabaeidae Ischiopsopha esmeralda Wallace 1868 2 pal Coleoptera Scarabaeidae Ischiopsopha jamesi Waterhouse 1876 1 pal Coleoptera Scarabaeidae Ischiopsopha lucivorax Kraatz 1890 3 pal Coleoptera Scarabaeidae Ischiopsopha rugata Blanchard 1842 1 pal Coleoptera Scarabaeidae Ischiopsopha sp. 2 pal Coleoptera Scarabaeidae Ischiopsopha sp. 5 aus Coleoptera Scarabaeidae Ischiopsopha virens Blanchard 1853 2 pal Coleoptera Scarabaeidae Ischiopsopha wallacei Thomas 1857 1 aus Coleoptera Scarabaeidae Ischnoscelis dohrni Westwood 1855 1 neo Coleoptera Scarabaeidae Isonychus arizonensis Howden 9 nea Coleoptera Scarabaeidae Isonychus ocellatus Burmeister 1855 9 neo Coleoptera Scarabaeidae Lachnota henningi Fischer 1836 1 pal Coleoptera Scarabaeidae Leiopsammodius malkini Cartwright 1946 3 nea Coleoptera Scarabaeidae Leptohoplia testaceipennis Saylor 1935 2 nea Coleoptera Scarabaeidae Leptothyrea perroudi Schaum 1844 1 afr Coleoptera Scarabaeidae Leucoserica arenicola Solsky 1876 1 pal Coleoptera Scarabaeidae Leucothyreus kirbyanus Macleay 1819 3 neo Coleoptera Scarabaeidae Ligyrus cuniculus Fabricius 1801 46 neo Coleoptera Scarabaeidae Ligyrus ebenus De Geer 1774 2 neo Coleoptera Scarabaeidae Ligyrus fossor Latreille 1833 2 neo Coleoptera Scarabaeidae Ligyrus gibbosus DeGeer 1774 244 nea Coleoptera Scarabaeidae Ligyrus gibbosus LeConte 1847 9 nea Coleoptera Scarabaeidae Ligyrus gibbosus LeConte 1856 116 nea Coleoptera Scarabaeidae Ligyrus gyas Erichson 1848 15 neo Coleoptera Scarabaeidae Ligyrus neglectus LeConte 1847 4 neo Coleoptera Scarabaeidae Ligyrus neglectus LeConte 1847 34 nea Coleoptera Scarabaeidae Ligyrus relictus Say 1825 37 nea Coleoptera Scarabaeidae Ligyrus sallaei Bates 1888 6 neo Coleoptera Scarabaeidae Ligyrus sallaei Bates 1888 7 nea Coleoptera Scarabaeidae Ligyrus sp. 1 nea Coleoptera Scarabaeidae Ligyrus sp. 2 neo Coleoptera Scarabaeidae Ligyrus spp. 190 Coleoptera Scarabaeidae Ligyrus subtropicus Blatchley 1922 4 nea Coleoptera Scarabaeidae Ligyrus villosus Burmeister 1847 10 neo Coleoptera Scarabaeidae Liocola brevitarsis Lewis 1879 22 afr Coleoptera Scarabaeidae Liocola brevitarsis Kraatz 1889 1 ori Coleoptera Scarabaeidae Liocola catharica Bates 1890 1 ori Coleoptera Scarabaeidae Liocola lugubris Herbst 1786 1 ori Coleoptera Scarabaeidae Liocola lugubris Fabricius 1792 3 pal Coleoptera Scarabaeidae Liocola speculifera Swartz 1817 4 ori Coleoptera Scarabaeidae Liogenys palpalis Eschscholtz 1822 2 neo Coleoptera Scarabaeidae Lomanoxoides bitubericolis Schmidt 1909 1 neo Coleoptera Scarabaeidae Lomaptera cinnamomea Thomas 1878 1 pal Coleoptera Scarabaeidae Lomaptera helleri Moser 1908 2 pal Coleoptera Scarabaeidae Lomaptera salvadorii Gestro 1876 1 aus Coleoptera Scarabaeidae Lomaptera satanus Schaum 1912 2 pal Coleoptera Scarabaeidae Lomaptera simbagensis Schaum 1 pal Coleoptera Scarabaeidae Lomaptera soror Kraatz 1890 3 afr Coleoptera Scarabaeidae Lomaptera wahnesi Moser 1906 1 pal Coleoptera Scarabaeidae Macraspis chalcea Burmeister 1844 8 neo Coleoptera Scarabaeidae Macraspis cincta MacLeay 1819 4 neo Coleoptera Scarabaeidae Macraspis dichroa Waterhouse 1881 6 neo Coleoptera Scarabaeidae Macraspis festiva Burmeister 1844 6 neo Coleoptera Scarabaeidae Macraspis lucida Olivier 1789 5 neo Coleoptera Scarabaeidae Macraspis tetradactyla Linnaeus 1771 4 ori Coleoptera Scarabaeidae Macrodactylus angustatus Beauvois 1805 5 nea Coleoptera Scarabaeidae Macrodactylus fulvescens Bates 1887 6 neo Coleoptera Scarabaeidae Macrodactylus mexicanus Burmeister 1855 7 neo Coleoptera Scarabaeidae Macrodactylus nigricornis? Moser? 1 neo Coleoptera Scarabaeidae Macrodactylus subspinosus Fabricius 1775 108 nea Coleoptera Scarabaeidae Macrodactylus thoracicus Kirsch 1885 3 neo Coleoptera Scarabaeidae Macrodactylus uniformis Horn 1876 36 nea Coleoptera Scarabaeidae Macrodactylus uniformis Horn 1876 46 nea Coleoptera Scarabaeidae Macronota nigritus Waterhouse 1841 1 ori Coleoptera Scarabaeidae Macronota regia Fabricius 1801 4 aus Coleoptera Scarabaeidae Macronota sp. 4 ori Coleoptera Scarabaeidae Macronota spp. 19 ori Coleoptera Scarabaeidae Maladera castanea Arrow 1913 52 nea Coleoptera Scarabaeidae Maladera holosericea Scopoli 1772 12 pal Coleoptera Scarabaeidae Maladera orientalis Motschulsky 1857 1 pal Coleoptera Scarabaeidae Maladera renardi Ballion 1870 1 neo Coleoptera Scarabaeidae Martinezia excavaticollis Blanchard 1843 2 nea Coleoptera Scarabaeidae Mausoleopsis amabilis Schaum 1844 1 afr Coleoptera Scarabaeidae Megaceras jason Fabricius 1775 2 neo Coleoptera Scarabaeidae Megaceras morpheus Burmeister 1847 7 neo Coleoptera Scarabaeidae Megaceras philoctetes Olivier 1789 1 neo Coleoptera Scarabaeidae Megaceras spp. 6 neo Coleoptera Scarabaeidae Megalorhina harrisi Westwood 1847 3 afr Coleoptera Scarabaeidae Megasoma actaeon Linnaeus 1758 21 neo Coleoptera Scarabaeidae Megasoma anubis Chevrolat 1836 2 neo Coleoptera Scarabaeidae Megasoma elephas Fabricius 1775 23 neo Coleoptera Scarabaeidae Megasoma gyas Herbst 1785 1 neo Coleoptera Scarabaeidae Megasoma mars Reiche 1852 1 neo Coleoptera Scarabaeidae Megasoma thersites LeConte 1861 1 neo Coleoptera Scarabaeidae Megathopa argentina Gillet 1911 1 neo Coleoptera Scarabaeidae Megathopa bicolor Guerin 1839 8 neo Coleoptera Scarabaeidae Megathopa villosa Eschscholtz 1839 6 neo Coleoptera Scarabaeidae Melinesthes umbonata Gory and Percheron 1833 2 afr Coleoptera Scarabaeidae Melolontha hippocastani Fabricius 1801 7 pal Coleoptera Scarabaeidae Melolontha hippocastani Comolli 1837 2 pal Coleoptera Scarabaeidae Melolontha melolontha Linnaeus 1758 36 pal Coleoptera Scarabaeidae Melolontha pectoralis Germar 1824 1 pal Coleoptera Scarabaeidae Metanastes vulgivagus Olliff 1889 1 aus Coleoptera Scarabaeidae Mimela chinensis Kirby 1825 1 ori Coleoptera Scarabaeidae Mimela junii Duftschmidt 1805 2 pal Coleoptera Scarabaeidae Mimela splendens Gyllenhal 1817 1 ori Coleoptera Scarabaeidae Mycterophallus validipes Heller 1894 2 aus Coleoptera Scarabaeidae Mycterophallus xanthopus Boisduval 1835 1 ori Coleoptera Scarabaeidae Myodermum alutaceum Afzeli 1817 1 afr Coleoptera Scarabaeidae Neodon laevis Burmeister 1847 1 aus Coleoptera Scarabaeidae Neodon pecuarius Reiche 1860 1 aus Coleoptera Scarabaeidae Neolucanus castanopterus Hope 2 pal Coleoptera Scarabaeidae Neopsammodius interuptus Say 1835 3 nea Coleoptera Scarabaeidae Neopsammodius quinqueplicatus Horn 1871 24 nea Coleoptera Scarabaeidae Neopsammodius werneri Cartwright 1955 1 nea Coleoptera Scarabaeidae Neopsammodius werneri Cartwright 1955 1 neo Coleoptera Scarabaeidae Neopsammodius werneri Cartwright 1955 5 neo Coleoptera Scarabaeidae Neorhyssemus mimicus Gordon and Cartwright 1980 1 neo Coleoptera Scarabaeidae Neptunides stanleyi Janson 1889 1 afr Coleoptera Scarabaeidae Odontoloma endroedyi Howden and Scholtz 1 afr Coleoptera Scarabaeidae Odontoloma peckorum Howden and Scholtz 4 afr Coleoptera Scarabaeidae Odontopsammodius armaticeps Fall 1932 2 nea Coleoptera Scarabaeidae Odontopsammodius bidens Horn 1871 9 nea Coleoptera Scarabaeidae Odontria australis Given 4 aus Coleoptera Scarabaeidae Odontria halii Broun 3 aus Coleoptera Scarabaeidae Odontria nitidula Broun 3 aus Coleoptera Scarabaeidae Odontria smithii Broun 1893 3 aus Coleoptera Scarabaeidae Odontria striata White 1846 3 aus Coleoptera Scarabaeidae Odontria varicolorata Given 4 aus Coleoptera Scarabaeidae Odontria xanthosticta White 1846 1 aus Coleoptera Scarabaeidae Odontria zealandica White 1846 4 aus Coleoptera Scarabaeidae Oncerus floralis LeConte 1856 5 nea Coleoptera Scarabaeidae Oniticellus festivus Steven 1809 1 pal Coleoptera Scarabaeidae Oniticellus fulvus Goeze 1777 15 pal Coleoptera Scarabaeidae Oniticellus fulvus Fabricius 1781 4 pal Coleoptera Scarabaeidae Oniticellus militarius Castelnau 1840 4 afr Coleoptera Scarabaeidae Oniticellus nasicornis Reiche 1849 4 afr Coleoptera Scarabaeidae Oniticellus pallens Olivier 1789 1 pal Coleoptera Scarabaeidae Oniticellus pallipes Fabricius 1781 9 pal Coleoptera Scarabaeidae Oniticellus sp. Horn 1882 2 nea Coleoptera Scarabaeidae Onitis aerarius Harold 1878 2 afr Coleoptera Scarabaeidae Onitis crenatus Reiche 1847 2 pal Coleoptera Scarabaeidae Onitis falcatus Wulfen 1786 1 ori Coleoptera Scarabaeidae Onitis humerosus Pallas 1771 1 pal Coleoptera Scarabaeidae Onitis ion Olivier 1789 1 pal Coleoptera Scarabaeidae Onitis sphinx Fabricius 1775 2 afr Coleoptera Scarabaeidae Onitis virens Lansberge 1875 1 pal Coleoptera Scarabaeidae Ontherus appendiculatus Mannerheim 1829 10 neo Coleoptera Scarabaeidae Ontherus sulcator Fabricius 1775 2 neo Coleoptera Scarabaeidae Onthobium howdeni Paulian 2 aus Coleoptera Scarabaeidae Onthobium lerati Paulian 2 aus Coleoptera Scarabaeidae Onthobium montrouzieri Paulian 2 aus Coleoptera Scarabaeidae Onthophagus alluvius Howden and Cartwright 1 nea Coleoptera Scarabaeidae Onthophagus amyntas Olivier 1789 4 pal Coleoptera Scarabaeidae Onthophagus anthracinus Harold 1873 7 nea Coleoptera Scarabaeidae Onthophagus ater Waterhouse 1875 2 ori Coleoptera Scarabaeidae Onthophagus atramentarius Menetries 1832 1 pal Coleoptera Scarabaeidae Onthophagus bonasus Linnaeus 1775 9 ori Coleoptera Scarabaeidae Onthophagus brevifrons Horn 1881 1 nea Coleoptera Scarabaeidae Onthophagus browni Howden and Cartwright 8 nea Coleoptera Scarabaeidae Onthophagus castanea Serville 1825 14 nea Coleoptera Scarabaeidae Onthophagus chevrolati Harold 1869 2 neo Coleoptera Scarabaeidae Onthophagus cochisus Brown 1927 nea Coleoptera Scarabaeidae Onthophagus coenobita Herbst 1783 4 pal Coleoptera Scarabaeidae Onthophagus concinnus Castelnau 1840 1 nea Coleoptera Scarabaeidae Onthophagus concinnus LaPorte 1840 7 nea Coleoptera Scarabaeidae Onthophagus crocatus Mulsant,??? 1 pal Coleoptera Scarabaeidae Onthophagus crocatus Mulsant? 1 pal Coleoptera Scarabaeidae Onthophagus depressus Harold 1871 13 nea Coleoptera Scarabaeidae Onthophagus finschi Harold 1877 2 ori Coleoptera Scarabaeidae Onthophagus fissicornis Steven 1809 1 pal Coleoptera Scarabaeidae Onthophagus flagrans Reitter 1893 1 pal Coleoptera Scarabaeidae Onthophagus fracticornis Preyssler 1790 26 pal Coleoptera Scarabaeidae Onthophagus furcatus Fabricius 1781 20 pal Coleoptera Scarabaeidae Onthophagus gazella Fabricius 27 neo Coleoptera Scarabaeidae Onthophagus gazella Fabricius 1787 3 afr Coleoptera Scarabaeidae Onthophagus gazella Fabricius),? 17 nea Coleoptera Scarabaeidae Onthophagus gazella Fabricius),?? 17 nea Coleoptera Scarabaeidae Onthophagus germanus Linnaeus 1767 3 pal Coleoptera Scarabaeidae Onthophagus gibbosus Reitter 1891 3 pal Coleoptera Scarabaeidae Onthophagus gibbulus Pallas 1781 2 pal Coleoptera Scarabaeidae Onthophagus granulatus Boheman 1858 3 aus Coleoptera Scarabaeidae Onthophagus granulatus Bohemann 1858 3 aus Coleoptera Scarabaeidae Onthophagus grossepunctatus Muller 1906 6 pal Coleoptera Scarabaeidae Onthophagus haroldi Ball 1871 2 pal Coleoptera Scarabaeidae Onthophagus hecate Panzer 1794 100 nea Coleoptera Scarabaeidae Onthophagus hecate Brown 1929 11 nea Coleoptera Scarabaeidae Onthophagus hoepfneri Harold 1869 1 nea Coleoptera Scarabaeidae Onthophagus hopfneri Harold 1869 5 neo Coleoptera Scarabaeidae Onthophagus knausi Brown 1927 1 nea Coleoptera Scarabaeidae Onthophagus knulli Howden and Cartwright 1963 2 nea Coleoptera Scarabaeidae Onthophagus koshantschikoffi Reitter 1891 2 pal Coleoptera Scarabaeidae Onthophagus landolti Harold 1880 5 nea Coleoptera Scarabaeidae Onthophagus lecontei Harold 1871 5 neo Coleoptera Scarabaeidae Onthophagus lemur Fabricius 1781 11 pal Coleoptera Scarabaeidae Onthophagus marginalis Gebler 1817 1 pal Coleoptera Scarabaeidae Onthophagus marginatus Castelnau 1840 14 neo Coleoptera Scarabaeidae Onthophagus marginicollis Harold 1880 2 neo Coleoptera Scarabaeidae Onthophagus medorensis Brown 1929 12 nea Coleoptera Scarabaeidae Onthophagus mexicanus Bates 1887 3 neo Coleoptera Scarabaeidae Onthophagus nebulosus Reiche 1864 1 afr Coleoptera Scarabaeidae Onthophagus nuchicornis Linnaeus 1758 12 pal Coleoptera Scarabaeidae Onthophagus nuchicornis Linnaeus 1758 69 nea Coleoptera Scarabaeidae Onthophagus oklahomensis Brown 1927 11 nea Coleoptera Scarabaeidae Onthophagus orpheus Panzer 1794 15 nea Coleoptera Scarabaeidae Onthophagus orpheus Fabricius 1801 6 nea Coleoptera Scarabaeidae Onthophagus ovatus Linnaeus 1767 26 pal Coleoptera Scarabaeidae Onthophagus pennsylvanicus Harold 1871 42 nea Coleoptera Scarabaeidae Onthophagus polyphemi Hubbard 1894 1 nea Coleoptera Scarabaeidae Onthophagus polyphemi Howden and Cartwright 1963 3 nea Coleoptera Scarabaeidae Onthophagus pygargus Motschuskly 1845 1 pal Coleoptera Scarabaeidae Onthophagus rangifer Klug 1855 2 afr Coleoptera Scarabaeidae Onthophagus ruficapillus Brulle 1832 6 pal Coleoptera Scarabaeidae Onthophagus sibiricus Harold 1877 1 pal Coleoptera Scarabaeidae Onthophagus speculifer Solsky 1876 1 pal Coleoptera Scarabaeidae Onthophagus spp. 22 neo Coleoptera Scarabaeidae Onthophagus striatulus Beauvois 1809 1 nea Coleoptera Scarabaeidae Onthophagus striatulus Beauvois 1809 2 nea Coleoptera Scarabaeidae Onthophagus striatulus Beauvois 1809 78 nea Coleoptera Scarabaeidae Onthophagus subaeneus Beauvois 1811 8 nea Coleoptera Scarabaeidae Onthophagus subtropicus Howden and Cartwright 1963 1 nea Coleoptera Scarabaeidae Onthophagus tapirus Sharp 1877 16 neo Coleoptera Scarabaeidae Onthophagus taurus Schreber 32 nea Coleoptera Scarabaeidae Onthophagus taurus Schreber 1759 14 pal Coleoptera Scarabaeidae Onthophagus tuberculifrons Harold 1873 23 nea Coleoptera Scarabaeidae Onthophagus vacca Linnaeus 1767 15 pal Coleoptera Scarabaeidae Onthophagus velutinus Horn 35 nea Coleoptera Scarabaeidae Onthophagus velutinus Horn 1875 1 nea Coleoptera Scarabaeidae Onthophagus verticicornis Laich 1781 8 pal Coleoptera Scarabaeidae Onthophagus vitulus Fabricius 1776 2 pal Coleoptera Scarabaeidae Oplostomus fuligineus Olivier 1789 4 afr Coleoptera Scarabaeidae Orizabus clunalis LeConte 1856 280 nea Coleoptera Scarabaeidae Oruscatus opalescens Bates 1870 5 neo Coleoptera Scarabaeidae Oryctes boas Fabricius 1775 3 afr Coleoptera Scarabaeidae Oryctes gigas Castelnau 1840 8 afr Coleoptera Scarabaeidae Oryctes gnu Mohnike 1874 4 ori Coleoptera Scarabaeidae Oryctes nasicornis Linnaeus 1758 12 pal Coleoptera Scarabaeidae Oryctes nasicornis Illiger 1803 6 pal Coleoptera Scarabaeidae Oryctes rhinoceros Linnaeus 1758 5 afr Coleoptera Scarabaeidae Oryctes rhinoceros Linnaeus 1758 12 ori Coleoptera Scarabaeidae Oryctes rhinoceros Linnaeus 1758 12 pal Coleoptera Scarabaeidae Oryctes sp. 1 neo Coleoptera Scarabaeidae Oryctes sp. 7 afr Coleoptera Scarabaeidae Oryctes spp. 12 Coleoptera Scarabaeidae Oryctoderus latitarsis Boisduval 1835 3 aus Coleoptera Scarabaeidae Oryctomorphus bimaculatus Guerin 1830 8 neo Coleoptera Scarabaeidae Oryctomorphus maculicollis Guerin 1838 4 neo Coleoptera Scarabaeidae Osmoderma eremicola Knoch 1801 64 nea Coleoptera Scarabaeidae Osmoderma eremita Scopoli 1763 5 pal Coleoptera Scarabaeidae Osmoderma scabra Beauvois 1805 86 nea Coleoptera Scarabaeidae Oxygrylius ruginasus LeConte 1856 262 nea Coleoptera Scarabaeidae Oxyligyrus nasutus Burmeister 1847 1 neo Coleoptera Scarabaeidae Oxyomus silvestris Scopoli 1763 4 nea Coleoptera Scarabaeidae Oxysternon conspicillatum Weber 1801 13 neo Coleoptera Scarabaeidae Oxysternon festivum Linnaeus 1767 106 neo Coleoptera Scarabaeidae Oxysternon palaemon Castelnau 1840 11 neo Coleoptera Scarabaeidae Oxythyrea amethystina MacLeay 1838 4 afr Coleoptera Scarabaeidae Oxythyrea cinctella Schaum 1841 23 pal Coleoptera Scarabaeidae Oxythyrea funesta Poda 1761 61 pal Coleoptera Scarabaeidae Oxythyrea funesta Linnaeus 1766 2 pal Coleoptera Scarabaeidae Oxythyrea haemorrhoridalis Fabricius 1775 3 afr Coleoptera Scarabaeidae Oxythyrea hildebrandti Kraatz 1880 3 afr Coleoptera Scarabaeidae Oxythyrea marginalis Swartz 1817 4 afr Coleoptera Scarabaeidae Oxythyrea pantherina Gory and Percheron 1833 1 pal Coleoptera Scarabaeidae Oxythyrea plebeja Kolbe 1895 1 afr Coleoptera Scarabaeidae Oxythyrea rubra Gory and Percheron 1833 1 afr Coleoptera Scarabaeidae Oxythyrea spp. 11 afr Coleoptera Scarabaeidae Oxythyrea testaceoguttata Blanchard 1850 4 afr Coleoptera Scarabaeidae Oxythyrea tripolitano Reitter 1891 6 pal Coleoptera Scarabaeidae Pachnoda consentanea Schaum. 1844 2 afr Coleoptera Scarabaeidae Pachnoda interrupta Olivier 1789 2 afr Coleoptera Scarabaeidae Pachnoda marginata Drury 1773 2 afr Coleoptera Scarabaeidae Pachnoda marginella Fabricius 1775 1 afr Coleoptera Scarabaeidae Pachnoda rufa DeGeer 1778 15 afr Coleoptera Scarabaeidae Pachnoda savigny Gory and Percheron 1833 2 afr Coleoptera Scarabaeidae Pachnoda sinuata Gory and Percheron 1833 6 afr Coleoptera Scarabaeidae Pachnoda sp. 1 afr Coleoptera Scarabaeidae Pachnoda sp. 3 afr Coleoptera Scarabaeidae Pachnoda sp. 8 afr Coleoptera Scarabaeidae Pachnoda spp. 9 afr Coleoptera Scarabaeidae Pachnoda stehelini Schaum 1841 1 afr Coleoptera Scarabaeidae Pachnoda tridentata Olivier 1789 1 afr Coleoptera Scarabaeidae Pachnodella impressa Goldfuss 1805 2 afr Coleoptera Scarabaeidae Pachydema bucharica Reitter 1898 1 pal Coleoptera Scarabaeidae Pachydema hirticollis Fabricius 1787 5 pal Coleoptera Scarabaeidae Pachylomerus femoralis Kirby 1828 2 afr Coleoptera Scarabaeidae Panglaphyra duboulayi Thomas 1879 4 afr Coleoptera Scarabaeidae Panotrogus myschenkowi Ballion 1871 1 pal Coleoptera Scarabaeidae Parabrysopolis arizonae Ohaus? 1915 5 nea Coleoptera Scarabaeidae Parabrysopolis chihuahuae Bates 1888 3 nea Coleoptera Scarabaeidae Paracotalpa puncticollis LeConte 1863 5 nea Coleoptera Scarabaeidae Paracotalpa ursina Horn 1867 17 nea Coleoptera Scarabaeidae Paragnorimus sambucas Howden 1970 1 neo Coleoptera Scarabaeidae Paragymnetis hebraica Drapiez 1820 1 neo Coleoptera Scarabaeidae Paragymnetis spp. 3 neo Coleoptera Scarabaeidae Parastasia brevipes LeConte 1856 9 nea Coleoptera Scarabaeidae Parataenius simulator Harold 1868 5 nea Coleoptera Scarabaeidae Parataenius simulator Harold 1868 116 nea Coleoptera Scarabaeidae Parathyce fieldi Fall 1908 1 nea Coleoptera Scarabaeidae Parathyce harfordi Casey 1889 3 nea Coleoptera Scarabaeidae Parathyce palpalis Horn 1880 9 nea Coleoptera Scarabaeidae Pedinorrhina mediana Westwood 1841 1 afr Coleoptera Scarabaeidae Pelidnota chalcothorax Perty 1834 3 neo Coleoptera Scarabaeidae Pelidnota fulva Blanchard 1850 2 neo Coleoptera Scarabaeidae Pelidnota laevissima Burmeister 1855 1 neo Coleoptera Scarabaeidae Pelidnota ludovici Ohaus 1905 11 neo Coleoptera Scarabaeidae Pelidnota lugubris LeConte 1874 16 nea Coleoptera Scarabaeidae Pelidnota lutea Olivier 1789 7 nea Coleoptera Scarabaeidae Pelidnota lutea Casey 1915 2 nea Coleoptera Scarabaeidae Pelidnota lutea Casey 1915 18 nea Coleoptera Scarabaeidae Pelidnota nitescens Vigors 1825 2 neo Coleoptera Scarabaeidae Pelidnota notata Blanchard 1850 3 neo Coleoptera Scarabaeidae Pelidnota prasima Burmeister 1844 1 neo Coleoptera Scarabaeidae Pelidnota prolixa Sharp 1877 1 neo Coleoptera Scarabaeidae Pelidnota pulchella Kirby 1818 1 neo Coleoptera Scarabaeidae Pelidnota punctata Linnaeus 1758 176 nea Coleoptera Scarabaeidae Pelidnota spp. 22 nea Coleoptera Scarabaeidae Pelidnota spp. 117 neo Coleoptera Scarabaeidae Pelidnota virescens Burmeister 1844 32 neo Coleoptera Scarabaeidae Pentodon bidens Pallas 1771 3 pal Coleoptera Scarabaeidae Pentodon bidens Villers 1789 7 pal Coleoptera Scarabaeidae Pentodon idiota Herbst 1789 6 pal Coleoptera Scarabaeidae Pericoptus punctatus White 1846 3 aus Coleoptera Scarabaeidae Pericoptus truncatus Fabricius 1787 2 aus Coleoptera Scarabaeidae Phaeadoretus comptus Menetries 1849 1 afr Coleoptera Scarabaeidae Phaenomeris magnifica Hope 1835 2 pal Coleoptera Scarabaeidae Phanaeus amethystinus Harold 1871 3 neo Coleoptera Scarabaeidae Phanaeus amithaon Harold 1875 11 nea Coleoptera Scarabaeidae Phanaeus batesi Harold 1868 8 neo Coleoptera Scarabaeidae Phanaeus bellicosus Olivier 1789 2 neo Coleoptera Scarabaeidae Phanaeus bispinus Bates 1887 1 neo Coleoptera Scarabaeidae Phanaeus bonariensis Gory 1844 2 neo Coleoptera Scarabaeidae Phanaeus cerberus Harold 1869 1 neo Coleoptera Scarabaeidae Phanaeus corythus Harold 1863 1 neo Coleoptera Scarabaeidae Phanaeus damon Castelnau 1840 7 neo Coleoptera Scarabaeidae Phanaeus daphnis Harold 1863 23 neo Coleoptera Scarabaeidae Phanaeus dardanus MacLeay 1819 1 neo Coleoptera Scarabaeidae Phanaeus difformis LeConte 1847 19 nea Coleoptera Scarabaeidae Phanaeus ensifera Germar 1812 9 neo Coleoptera Scarabaeidae Phanaeus faunus Fabricius 1775 3 neo Coleoptera Scarabaeidae Phanaeus furcosus Bates 1887 7 neo Coleoptera Scarabaeidae Phanaeus hermes Harold 1868 2 neo Coleoptera Scarabaeidae Phanaeus howdeni Arnaud 1983 1 neo Coleoptera Scarabaeidae Phanaeus howdeni Arnaud 1984 1 neo Coleoptera Scarabaeidae Phanaeus igneus MacLeay 1819 41 nea Coleoptera Scarabaeidae Phanaeus imperator Chevrolet 1844 6 neo Coleoptera Scarabaeidae Phanaeus jasius Olivier 1789 27 neo Coleoptera Scarabaeidae Phanaeus jasius MacLeay 1819 3 neo Coleoptera Scarabaeidae Phanaeus kirbyi Vigors 1825 neo Coleoptera Scarabaeidae Phanaeus laeviceps Harold 1869 1 neo Coleoptera Scarabaeidae Phanaeus lancifer Linnaeus 1767 11 neo Coleoptera Scarabaeidae Phanaeus leander Waterhouse 1891 84 neo Coleoptera Scarabaeidae Phanaeus melampus Harold 1863 1 neo Coleoptera Scarabaeidae Phanaeus meleagris Blanchard 1843 3 neo Coleoptera Scarabaeidae Phanaeus mexicanus Harold 1863 6 neo Coleoptera Scarabaeidae Phanaeus milon Blanchard 1843 2 neo Coleoptera Scarabaeidae Phanaeus palaeno Blanchard 1843 1 neo Coleoptera Scarabaeidae Phanaeus prasinus Harold 1868 6 neo Coleoptera Scarabaeidae Phanaeus pyrois Bates 1887 1 neo Coleoptera Scarabaeidae Phanaeus quadridens Say 1835 21 nea Coleoptera Scarabaeidae Phanaeus sallei Harold 1863 1 neo Coleoptera Scarabaeidae Phanaeus sapphirinus Sturm 1826 8 neo Coleoptera Scarabaeidae Phanaeus septentrionalis Pess??? 1 neo Coleoptera Scarabaeidae Phanaeus splendidulus Fabricius 1781 43 neo Coleoptera Scarabaeidae Phanaeus spp. 21 neo Coleoptera Scarabaeidae Phanaeus sulcatus Drury 1770 4 neo Coleoptera Scarabaeidae Phanaeus torrens Olsontieff 1924 1 nea Coleoptera Scarabaeidae Phanaeus triangularis Edmonds 1994 1 nea Coleoptera Scarabaeidae Phanaeus velutinus Murray 1856 2 neo Coleoptera Scarabaeidae Phanaeus vindex MacLeay 1819 118 nea Coleoptera Scarabaeidae Pharaonus lederi Reitter 1888 1 pal Coleoptera Scarabaeidae Pharaonus semenowi Reitter 1887 1 pal Coleoptera Scarabaeidae Phileurus didymus Linnaeus 1758 2 neo Coleoptera Scarabaeidae Phileurus illatus LeConte 1854 26 nea Coleoptera Scarabaeidae Phileurus quadrituberculatus Beauvois 1806 1 neo Coleoptera Scarabaeidae Phileurus sp. 4 nea Coleoptera Scarabaeidae Phileurus sp. 4 1 neo Coleoptera Scarabaeidae Phileurus sp.1 3 neo Coleoptera Scarabaeidae Phileurus sp.2 1 neo Coleoptera Scarabaeidae Phileurus sp.3 1 neo Coleoptera Scarabaeidae Phileurus sp.5 1 neo Coleoptera Scarabaeidae Phileurus spp. 10 neo Coleoptera Scarabaeidae Phileurus truncatus Beauvois 1807 1 neo Coleoptera Scarabaeidae Phileurus truncatus Beauvois 1807 25 nea Coleoptera Scarabaeidae Phileurus valgus Olivier 1789 10 neo Coleoptera Scarabaeidae Phileurus valgus Bates 1888 1 neo Coleoptera Scarabaeidae Phileurus valgus Casey 6 nea Coleoptera Scarabaeidae Phileurus valgus Olivier 1789 16 nea Coleoptera Scarabaeidae Philoscaptus bonariensis Burmeister 1847 7 neo Coleoptera Scarabaeidae Phobetus centralis Casey 1909 1 nea Coleoptera Scarabaeidae Phobetus comatus Barrett 1856 1 nea Coleoptera Scarabaeidae Phobetus mojavus Barrett 1933 11 nea Coleoptera Scarabaeidae Phobetus palpalis Saylor 1936 1 nea Coleoptera Scarabaeidae Phobetus spp. 2 nea Coleoptera Scarabaeidae Phyllognathus excavatus Forster 1771 7 pal Coleoptera Scarabaeidae Phyllopertha horticola Linnaeus 1758 26 pal Coleoptera Scarabaeidae Phyllopertha massageta Kirsch 1881 2 pal Coleoptera Scarabaeidae Phyllophaga acinosa Arrow 1920 2 neo Coleoptera Scarabaeidae Phyllophaga aemula Horn 1887 7 nea Coleoptera Scarabaeidae Phyllophaga aequalis LeConte 1857 4 nea Coleoptera Scarabaeidae Phyllophaga analis Burmeister 1855 1 neo Coleoptera Scarabaeidae Phyllophaga anxia LeConte 1850 118 nea Coleoptera Scarabaeidae Phyllophaga arcta Horn 1887 2 nea Coleoptera Scarabaeidae Phyllophaga arizona Saylor 1940 1 nea Coleoptera Scarabaeidae Phyllophaga arkansana Schaeffer 1906 9 nea Coleoptera Scarabaeidae Phyllophaga balia Say 1825 43 nea Coleoptera Scarabaeidae Phyllophaga barda Horn 1887 10 nea Coleoptera Scarabaeidae Phyllophaga batillifer Bates 1 neo Coleoptera Scarabaeidae Phyllophaga beckeri Moser 20 nea Coleoptera Scarabaeidae Phyllophaga bellamyi Warner and Moron 1992 1 nea Coleoptera Scarabaeidae Phyllophaga bilobatata Saylor 1939 55 nea Coleoptera Scarabaeidae Phyllophaga bilobatata Saylor 1939 165 nea Coleoptera Scarabaeidae Phyllophaga bipartita Horn 1887 30 nea Coleoptera Scarabaeidae Phyllophaga brevidens Bates 1 neo Coleoptera Scarabaeidae Phyllophaga bruneri Chapin 27 nea Coleoptera Scarabaeidae Phyllophaga calceata LeConte 1856 10 nea Coleoptera Scarabaeidae Phyllophaga cephalicus Horn 4 nea Coleoptera Scarabaeidae Phyllophaga chapini Saylor 1940 9 nea Coleoptera Scarabaeidae Phyllophaga chiriquina Bates 9 neo Coleoptera Scarabaeidae Phyllophaga citri Smyth 1917 7 neo Coleoptera Scarabaeidae Phyllophaga clemens Horn 1887 4 nea Coleoptera Scarabaeidae Phyllophaga clypeata Horn 1887 1 nea Coleoptera Scarabaeidae Phyllophaga congrua LeConte 1856 8 nea Coleoptera Scarabaeidae Phyllophaga corrosa LeConte 1856 4 nea Coleoptera Scarabaeidae Phyllophaga crassissima Blanchard 1850 36 nea Coleoptera Scarabaeidae Phyllophaga crenulata Froelich 1792 71 nea Coleoptera Scarabaeidae Phyllophaga cribrosa LeConte 1853 8 nea Coleoptera Scarabaeidae Phyllophaga cribrosa LeConte 1853 49 nea Coleoptera Scarabaeidae Phyllophaga crinata Smith 1889 6 nea Coleoptera Scarabaeidae Phyllophaga crinita Burmeister 1855 19 nea Coleoptera Scarabaeidae Phyllophaga cushmani Say 1940 1 nea Coleoptera Scarabaeidae Phyllophaga debilis LeConte 1856 31 nea Coleoptera Scarabaeidae Phyllophaga delata Horn 1887 26 nea Coleoptera Scarabaeidae Phyllophaga densepunctata Moser 6 neo Coleoptera Scarabaeidae Phyllophaga diffinis Blanchard 1850 4 nea Coleoptera Scarabaeidae Phyllophaga dispar Burmeister 1855 8 nea Coleoptera Scarabaeidae Phyllophaga disparilis Horn 1878 3 nea Coleoptera Scarabaeidae Phyllophaga disparilis Horn 1878 7 nea Coleoptera Scarabaeidae Phyllophaga divertens Bates 2 nea Coleoptera Scarabaeidae Phyllophaga drakei Kirby 41 nea Coleoptera Scarabaeidae Phyllophaga duncani Barrett 1933 1 nea Coleoptera Scarabaeidae Phyllophaga duncani Barrett 1933 42 nea Coleoptera Scarabaeidae Phyllophaga elizoria Saylor 1937 33 nea Coleoptera Scarabaeidae Phyllophaga ephilida Say 1825 37 nea Coleoptera Scarabaeidae Phyllophaga ephilida Reinhardt 2 nea Coleoptera Scarabaeidae Phyllophaga epigaea Wickham 1903 3 nea Coleoptera Scarabaeidae Phyllophaga errans LeConte 1860 4 nea Coleoptera Scarabaeidae Phyllophaga falsa LeConte 1856 12 nea Coleoptera Scarabaeidae Phyllophaga farcta LeConte 1856 12 nea Coleoptera Scarabaeidae Phyllophaga fervida Fabricius 1781 18 nea Coleoptera Scarabaeidae Phyllophaga floridana Robinson 1838 3 nea Coleoptera Scarabaeidae Phyllophaga forbesi Glasgow 1916 9 nea Coleoptera Scarabaeidae Phyllophaga forsteri Burmeister 1850 39 nea Coleoptera Scarabaeidae Phyllophaga foxii Davis 1920 2 nea Coleoptera Scarabaeidae Phyllophaga fragilipennis Blanchard 1850 neo Coleoptera Scarabaeidae Phyllophaga fraterna Harris 1842 35 nea Coleoptera Scarabaeidae Phyllophaga fucata Horn 1887 5 nea Coleoptera Scarabaeidae Phyllophaga fulviventris Moser 2 neo Coleoptera Scarabaeidae Phyllophaga fusca Froelich 1792 86 nea Coleoptera Scarabaeidae Phyllophaga futilis LeConte 1850 58 nea Coleoptera Scarabaeidae Phyllophaga gaigei Sanderson 1948 Coleoptera Scarabaeidae Phyllophaga georgiana Horn 1 nea Coleoptera Scarabaeidae Phyllophaga glabberima Blanchard 1850 73 nea Coleoptera Scarabaeidae Phyllophaga glabricula LeConte 1856 4 nea Coleoptera Scarabaeidae Phyllophaga gracilis Burmeister 1855 28 nea Coleoptera Scarabaeidae Phyllophaga gracilis Glasg. 1925 12 nea Coleoptera Scarabaeidae Phyllophaga granti Saylor 1940 2 nea Coleoptera Scarabaeidae Phyllophaga granti Saylor 1940 8 nea Coleoptera Scarabaeidae Phyllophaga hirsuta Knoch 1801 17 nea Coleoptera Scarabaeidae Phyllophaga hirticula Knoch 1801 89 nea Coleoptera Scarabaeidae Phyllophaga hirtiventris Horn 1887 9 nea Coleoptera Scarabaeidae Phyllophaga hoogsraali Saylor 2 neo Coleoptera Scarabaeidae Phyllophaga hornii Smith 1889 8 nea Coleoptera Scarabaeidae Phyllophaga huachuca Saylor 1940 24 nea Coleoptera Scarabaeidae Phyllophaga huachuca Saylor 1940 32 nea Coleoptera Scarabaeidae Phyllophaga idonea Sanderson 1948 8 nea Coleoptera Scarabaeidae Phyllophaga ignava Horn 1887 28 nea Coleoptera Scarabaeidae Phyllophaga ilicis Knoch 1801 2 nea Coleoptera Scarabaeidae Phyllophaga ilicis Knoch 1801 45 nea Coleoptera Scarabaeidae Phyllophaga impar Davis 1920 2 nea Coleoptera Scarabaeidae Phyllophaga implicata Horn 1887 17 nea Coleoptera Scarabaeidae Phyllophaga incuria Sanderson 1942 1 nea Coleoptera Scarabaeidae Phyllophaga inepta Horn 1887 2 nea Coleoptera Scarabaeidae Phyllophaga infidelis Horn 1887 16 nea Coleoptera Scarabaeidae Phyllophaga inflexa Barrett 1935 1 nea Coleoptera Scarabaeidae Phyllophaga inversa Horn 1887 15 nea Coleoptera Scarabaeidae Phyllophaga jonesi Sanderson 2 nea Coleoptera Scarabaeidae Phyllophaga juvenilis Fall 1932 1 nea Coleoptera Scarabaeidae Phyllophaga karlsioei Linell 1898 2 nea Coleoptera Scarabaeidae Phyllophaga kentuckiana Richter 1937 2 nea Coleoptera Scarabaeidae Phyllophaga knochii Gyllenhal 1817 7 nea Coleoptera Scarabaeidae Phyllophaga koehleriana Saylor 1940 4 nea Coleoptera Scarabaeidae Phyllophaga lanceolata Say 1825 6 nea Coleoptera Scarabaeidae Phyllophaga lanceolata Say 1825 27 nea Coleoptera Scarabaeidae Phyllophaga latidens Schaeffer 1906 6 nea Coleoptera Scarabaeidae Phyllophaga latidens Schaeffer 1906 25 nea Coleoptera Scarabaeidae Phyllophaga latifrons LeConte 1856 64 nea Coleoptera Scarabaeidae Phyllophaga lenis Horn 1887 1 nea Coleoptera Scarabaeidae Phyllophaga lenis Horn 1887 19 nea Coleoptera Scarabaeidae Phyllophaga lobata Fall 1908 6 nea Coleoptera Scarabaeidae Phyllophaga lodingi Sanderson 1939 2 nea Coleoptera Scarabaeidae Phyllophaga longispina Smith 1889 23 nea Coleoptera Scarabaeidae Phyllophaga longitarsa Say 1824 24 nea Coleoptera Scarabaeidae Phyllophaga lota Luginbill 1928 2 nea Coleoptera Scarabaeidae Phyllophaga luctuosa Horn 1887 18 nea Coleoptera Scarabaeidae Phyllophaga luginbilli Saylor 1941 1 nea Coleoptera Scarabaeidae Phyllophaga marginalis LeConte 1856 37 nea Coleoptera Scarabaeidae Phyllophaga micans Knoch 1801 32 nea Coleoptera Scarabaeidae Phyllophaga micans Langston 1924 3 nea Coleoptera Scarabaeidae Phyllophaga misteca Bates 1888 2 neo Coleoptera Scarabaeidae Phyllophaga mucorea LeConte 1856 44 nea Coleoptera Scarabaeidae Phyllophaga neomexicana? Saylor 1940 8 nea Coleoptera Scarabaeidae Phyllophaga nitida LeConte 1856 10 nea Coleoptera Scarabaeidae Phyllophaga obsoleta Blanchard 1850 2 neo Coleoptera Scarabaeidae Phyllophaga opacicollis Horn 1878 2 nea Coleoptera Scarabaeidae Phyllophaga opacita Reinhard 1939 2 nea Coleoptera Scarabaeidae Phyllophaga orosina Moser 1 neo Coleoptera Scarabaeidae Phyllophaga pallida Horn 1885 1 nea Coleoptera Scarabaeidae Phyllophaga parvidens LeConte 1856 8 nea Coleoptera Scarabaeidae Phyllophaga parvisetis Bates 1888 1 neo Coleoptera Scarabaeidae Phyllophaga paternoi Glasgow 1925 1 nea Coleoptera Scarabaeidae Phyllophaga plena Fall 1932 5 nea Coleoptera Scarabaeidae Phyllophaga plenus Fall 1932 3 nea Coleoptera Scarabaeidae Phyllophaga pleroma Reinhard 1940 4 nea Coleoptera Scarabaeidae Phyllophaga postrema Horn 1887 1 nea Coleoptera Scarabaeidae Phyllophaga praetermissa Horn 1887 8 nea Coleoptera Scarabaeidae Phyllophaga profunda Blanchard 1850 9 nea Coleoptera Scarabaeidae Phyllophaga prolixa Bates 1888 2 neo Coleoptera Scarabaeidae Phyllophaga prunina LeConte 1856 17 nea Coleoptera Scarabaeidae Phyllophaga prununculina Burmeister 1855 40 nea Coleoptera Scarabaeidae Phyllophaga pusillidens Fall 1937 10 nea Coleoptera Scarabaeidae Phyllophaga pygidialis Schaeffer 1906 19 nea Coleoptera Scarabaeidae Phyllophaga quercus Knoch 1801 26 nea Coleoptera Scarabaeidae Phyllophaga ravida Blanchard 1850 2 nea Coleoptera Scarabaeidae Phyllophaga rolbakeri Saylor 2 neo Coleoptera Scarabaeidae Phyllophaga rorulenta Burmeister 1855 11 neo Coleoptera Scarabaeidae Phyllophaga rossi Saylor 1939 2 nea Coleoptera Scarabaeidae Phyllophaga rubiginosa LeConte 1856 11 nea Coleoptera Scarabaeidae Phyllophaga rubricosa Reinhard 1939 4 nea Coleoptera Scarabaeidae Phyllophaga rugosa Melsheimer 1846 38 nea Coleoptera Scarabaeidae Phyllophaga schaefferi Saylor 1937 7 nea Coleoptera Scarabaeidae Phyllophaga scoparia LeConte 1856 21 nea Coleoptera Scarabaeidae Phyllophaga sequoiana Saylor 1936 4 nea Coleoptera Scarabaeidae Phyllophaga servicata Blanchard 5 neo Coleoptera Scarabaeidae Phyllophaga setifera Burmeister 1885 1 nea Coleoptera Scarabaeidae Phyllophaga sonora Saylor 1939 11 nea Coleoptera Scarabaeidae Phyllophaga sp. 1 Coleoptera Scarabaeidae Phyllophaga sp. graciliscomplex 2 nea Coleoptera Scarabaeidae Phyllophaga sp. nrtimidus 3 neo Coleoptera Scarabaeidae Phyllophaga sp. setidorsisgroup 8 neo Coleoptera Scarabaeidae Phyllophaga sp.sonora/pallida 13 nea Coleoptera Scarabaeidae Phyllophaga spp. testaceipennisgroup 4 neo Coleoptera Scarabaeidae Phyllophaga straminea Bates 2 neo Coleoptera Scarabaeidae Phyllophaga submucida LeConte 1856 37 nea Coleoptera Scarabaeidae Phyllophaga subpruinosa Casey 1884 5 nea Coleoptera Scarabaeidae Phyllophaga subtonsa LeConte 1856 4 nea Coleoptera Scarabaeidae Phyllophaga suttonana Reinhardt 1 nea Coleoptera Scarabaeidae Phyllophaga sylvatica Sanderson 1942 5 nea Coleoptera Scarabaeidae Phyllophaga tarsalis Schaeffer 1908 2 nea Coleoptera Scarabaeidae Phyllophaga tarsalis Schaeffer 1908 12 nea Coleoptera Scarabaeidae Phyllophaga tarsalis Schaeffer 1908 268 nea Coleoptera Scarabaeidae Phyllophaga taxodii Langston 1924 6 nea Coleoptera Scarabaeidae Phyllophaga tecta Cartwright 1944 2 nea Coleoptera Scarabaeidae Phyllophaga temora Saylor 2 neo Coleoptera Scarabaeidae Phyllophaga testaceipennis Moser 1921 1 neo Coleoptera Scarabaeidae Phyllophaga texensis Saylor 1940 1 nea Coleoptera Scarabaeidae Phyllophaga timida Horn 1878 2 nea Coleoptera Scarabaeidae Phyllophaga timida Horn 1878 4 nea Coleoptera Scarabaeidae Phyllophaga timida Horn 1878 8 nea Coleoptera Scarabaeidae Phyllophaga timidus Horn 8 nea Coleoptera Scarabaeidae Phyllophaga torta LeConte 1856 44 nea Coleoptera Scarabaeidae Phyllophaga transversicollis Moser 1 neo Coleoptera Scarabaeidae Phyllophaga trichodes Bates 1890 2 neo Coleoptera Scarabaeidae Phyllophaga tristis Fabricius 1781 1 nea Coleoptera Scarabaeidae Phyllophaga tristis Fabricius 1781 77 nea Coleoptera Scarabaeidae Phyllophaga tristis Reinhard 1939 2 nea Coleoptera Scarabaeidae Phyllophaga trochanter Saylor 1940 1 nea Coleoptera Scarabaeidae Phyllophaga trochanter Saylor 1940 11 nea Coleoptera Scarabaeidae Phyllophaga trochanter Saylor 1940 24 nea Coleoptera Scarabaeidae Phyllophaga tusa Horn 1887 1 nea Coleoptera Scarabaeidae Phyllophaga ulkei Smith 1888 10 nea Coleoptera Scarabaeidae Phyllophaga uniformis Blanchard 1850 13 nea Coleoptera Scarabaeidae Phyllophaga vanalleni Schaeffer 1927 10 nea Coleoptera Scarabaeidae Phyllophaga vandykei Saylor 1 neo Coleoptera Scarabaeidae Phyllophaga vehemens Horn 1887 6 nea Coleoptera Scarabaeidae Phyllophaga vetula Horn 1887 24 nea Coleoptera Scarabaeidae Phyllophaga vetula Horn 1887 130 nea Coleoptera Scarabaeidae Phyllophaga vexata Horn 1885 2 neo Coleoptera Scarabaeidae Phyllophaga villifrons LeConte 1856 8 nea Coleoptera Scarabaeidae Phyllophaga wickhami Saylor 1940 8 nea Coleoptera Scarabaeidae Phyllophaga youngi Cartwright 1935 4 nea Coleoptera Scarabaeidae Phyllophaga zeteki Saylor 2 neo Coleoptera Scarabaeidae Pimelopus dubius Blackburn 1885 1 aus Coleoptera Scarabaeidae Plaesiorrhina recurva Wiedemeyer 1821 1 afr Coleoptera Scarabaeidae Plaesiorrhina recurva Wiedemeyer 1821 4 afr Coleoptera Scarabaeidae Plaesiorrhina watkinsiana Lewis 1879 2 afr Coleoptera Scarabaeidae Platychelus dimidiatus Burmeister 1844 2 afr Coleoptera Scarabaeidae Platygenia barbata Afzeli 1817 1 afr Coleoptera Scarabaeidae Platytomus atlanticus Cartwright 1948 3 nea Coleoptera Scarabaeidae Platytomus caelicollis Cartwright 1948 2 nea Coleoptera Scarabaeidae Platytomus longulus Cartwright 1948 35 nea Coleoptera Scarabaeidae Platytomus micros Bates 1887 5 nea Coleoptera Scarabaeidae Platytomus notialis Cartwright 1948 2 nea Coleoptera Scarabaeidae Plectris aliena Chapin 2 nea Coleoptera Scarabaeidae Plectrodes pubescens Horn 1867 1 nea Coleoptera Scarabaeidae Plectrone nigrocoerulea Waterhouse 1841 2 afr Coleoptera Scarabaeidae Pleurophorus apicipennis Reitter 1897 2 pal Coleoptera Scarabaeidae Pleurophorus caesus Creutzer 1796 2 pal Coleoptera Scarabaeidae Pleurophorus caesus Creutzer 1896 31 nea Coleoptera Scarabaeidae Pleurophorus variolosus Kolenati 1846 1 pal Coleoptera Scarabaeidae Plusiotis beyeri Skinner 1905 156 nea Coleoptera Scarabaeidae Plusiotis gloriosa LeConte 1854 223 nea Coleoptera Scarabaeidae Plusiotis lecontei Horn 1882 149 nea Coleoptera Scarabaeidae Plusiotis resplendens Boucard 1875 1 neo Coleoptera Scarabaeidae Plusiotis woodi Horn 1883 27 nea Coleoptera Scarabaeidae Podischnus agenor Olivier 1789 7 neo Coleoptera Scarabaeidae Podolasia bottimeri Howden 1958 1 nea Coleoptera Scarabaeidae Poecilopharis femorata Waterhouse 1894 1 aus Coleoptera Scarabaeidae Poecilopharis woodfordi Waterhouse 1887 1 aus Coleoptera Scarabaeidae Poecilophila hebraea Olivier 1789 4 afr Coleoptera Scarabaeidae Poecilophila maculatissima Bohmann 1860 1 afr Coleoptera Scarabaeidae Poecilophilides rusticola Burmeister 1842 1 ori Coleoptera Scarabaeidae Polybaphes aequinoctialis Olivier 1789 3 afr Coleoptera Scarabaeidae Polybaphes balteata DeGeer 1778 1 afr Coleoptera Scarabaeidae Polybaphes scalaris Gory and Percheron 1833 1 pal Coleoptera Scarabaeidae Polybaphes sp. 1 afr Coleoptera Scarabaeidae Polyphylla 1 Young 1 nea Coleoptera Scarabaeidae Polyphylla alba Reitter 1903 1 pal Coleoptera Scarabaeidae Polyphylla arguta Casey 1914 1 nea Coleoptera Scarabaeidae Polyphylla autumnalis Schaeffer 1927 8 nea Coleoptera Scarabaeidae Polyphylla cavifrons LeConte 1854 1 nea Coleoptera Scarabaeidae Polyphylla comes Casey 1914 7 nea Coleoptera Scarabaeidae Polyphylla crinita LeConte 1856 7 nea Coleoptera Scarabaeidae Polyphylla decimlineata Say 1824 58 nea Coleoptera Scarabaeidae Polyphylla devestiva Young 1966 1 nea Coleoptera Scarabaeidae Polyphylla diffracta Casey 1891 78 nea Coleoptera Scarabaeidae Polyphylla fullo Linnaeus 1728 nea Coleoptera Scarabaeidae Polyphylla fullo Linnaeus 1728 15 pal Coleoptera Scarabaeidae Polyphylla gracilis Horn 1881 4 nea Coleoptera Scarabaeidae Polyphylla hammondi LeConte 1856 172 nea Coleoptera Scarabaeidae Polyphylla nigra Casey 1914 5 nea Coleoptera Scarabaeidae Polyphylla occidentalis Linnaeus 1767 8 nea Coleoptera Scarabaeidae Polyphylla sobrina Casey 1914 4 nea Coleoptera Scarabaeidae Polyphylla tridentata Reitter 1890 1 pal Coleoptera Scarabaeidae Polyphylla variolosa Hentz 1830 125 nea Coleoptera Scarabaeidae Polystigma punctata Donovan 1805 2 aus Coleoptera Scarabaeidae Popillia atrocoerulea Bates 1888 17 ori Coleoptera Scarabaeidae Popillia atrocoerulea Kraatz 1829 1 pal Coleoptera Scarabaeidae Popillia bipunctata Fabricius 1781 2 afr Coleoptera Scarabaeidae Popillia cyanea Hope 1831 11 ori Coleoptera Scarabaeidae Popillia formosana Arrow 1913 11 ori Coleoptera Scarabaeidae Popillia histeroidea Gyllenhal 1817 7 ori Coleoptera Scarabaeidae Popillia indigonacea Motschulsky 1853 11 ori Coleoptera Scarabaeidae Popillia japonica Newman 1838 272 nea Coleoptera Scarabaeidae Popillia mongolica Arrow 1913 2 ori Coleoptera Scarabaeidae Popillia mutans Newman 1838 16 ori Coleoptera Scarabaeidae Popillia pustulata Fairmaire 1887 14 ori Coleoptera Scarabaeidae Popillia quadriguttata Fabricius 1787 8 ori Coleoptera Scarabaeidae Protaetia acuminata Fabricius 1775 4 ori Coleoptera Scarabaeidae Protaetia aerata Erichson 1834 1 ori Coleoptera Scarabaeidae Protaetia aeruginosa Drury 1770 5 ori Coleoptera Scarabaeidae Protaetia aeruginosa Scopoli 1786 1 pal Coleoptera Scarabaeidae Protaetia affinis Andersch 1797 4 pal Coleoptera Scarabaeidae Protaetia affinis Reitter 1891 1 pal Coleoptera Scarabaeidae Protaetia afflicta Gory and Percheron 1833 3 pal Coleoptera Scarabaeidae Protaetia angustata Germar 1817 6 pal Coleoptera Scarabaeidae Protaetia angustata Reitter 1902 1 pal Coleoptera Scarabaeidae Protaetia anovittata Chevrolat 1841 1 ori Coleoptera Scarabaeidae Protaetia bakeri Moser 1917 1 ori Coleoptera Scarabaeidae Protaetia brevitaris Kraatz 1883 1 ori Coleoptera Scarabaeidae Protaetia brevitarsis Lewis 1879 22 afr Coleoptera Scarabaeidae Protaetia catharica Bates 1890 1 ori Coleoptera Scarabaeidae Protaetia culta Waterhouse 1879 2 ori Coleoptera Scarabaeidae Protaetia cuprea Fabricius 1775 7 pal Coleoptera Scarabaeidae Protaetia cuprea Andersch 1797 4 pal Coleoptera Scarabaeidae Protaetia cuprea Herbst 1790 8 pal Coleoptera Scarabaeidae Protaetia cuprea Gory and Percheron 1833 3 pal Coleoptera Scarabaeidae Protaetia cuprea Ball 1870 1 pal Coleoptera Scarabaeidae Protaetia cuprea Paykull 1799 13 pal Coleoptera Scarabaeidae Protaetia cuprea Andersch 1797 1 pal Coleoptera Scarabaeidae Protaetia cuprea Andersch 1797 6 pal Coleoptera Scarabaeidae Protaetia cuprea Reitter 1898 1 pal Coleoptera Scarabaeidae Protaetia cuprea Falderman 1835 1 pal Coleoptera Scarabaeidae Protaetia ducalis Mohnike 1873 1 ori Coleoptera Scarabaeidae Protaetia ferruginea Gory and Percheron 1833 7 ori Coleoptera Scarabaeidae Protaetia funebris Gory and Percheron 1833 1 pal Coleoptera Scarabaeidae Protaetia funesta Menetries 1836 2 pal Coleoptera Scarabaeidae Protaetia fusca Herbst 1790 97 ori Coleoptera Scarabaeidae Protaetia hungarica Herbst 1790 13 pal Coleoptera Scarabaeidae Protaetia hungarica Menetries 1832 1 afr Coleoptera Scarabaeidae Protaetia hungarica Reitter 1901 1 afr Coleoptera Scarabaeidae Protaetia incerta Mohnike 1873 1 ori Coleoptera Scarabaeidae Protaetia judith Reiche 1871 1 pal Coleoptera Scarabaeidae Protaetia karelini Zoubkoff 1829 2 afr Coleoptera Scarabaeidae Protaetia lugubris Herbst 1786 1 ori Coleoptera Scarabaeidae Protaetia lugubris Fabricius 1792 3 pal Coleoptera Scarabaeidae Protaetia morio Fabricius 1781 5 pal Coleoptera Scarabaeidae Protaetia oblonga Gory and Percheron 1833 2 pal Coleoptera Scarabaeidae Protaetia obscura Andersch 1797 1 pal Coleoptera Scarabaeidae Protaetia opaca Fabricius 1781 3 pal Coleoptera Scarabaeidae Protaetia papuana Moser 1914 5 ori Coleoptera Scarabaeidae Protaetia phillippensis Fabricius 1775 7 ori Coleoptera Scarabaeidae Protaetia sp. 1 ori Coleoptera Scarabaeidae Protaetia sp. 2 pal Coleoptera Scarabaeidae Protaetia sp. 5 ori Coleoptera Scarabaeidae Protaetia speciosa Gory and Percheron 1833 1 ori Coleoptera Scarabaeidae Protaetia speculifera Swartz 1817 4 ori Coleoptera Scarabaeidae Protaetia vidua Waltl 1838 4 pal Coleoptera Scarabaeidae Psammodius asper Fabricius 1775 6 pal Coleoptera Scarabaeidae Psammodius spp. 10 pal Coleoptera Scarabaeidae Pseudadoretus phthisicus Dohrn 1882 1 pal Coleoptera Scarabaeidae Pseudataenius socialis Horn 1871 9 nea Coleoptera Scarabaeidae Pseudinca admixtus Hope 1842 1 afr Coleoptera Scarabaeidae Pseudocanthon perplexus LeConte 1847 38 nea Coleoptera Scarabaeidae Pseudochalcothea pomacea Bates 1889 1 ori Coleoptera Scarabaeidae Pseudocotalpa andrewsi Hardy 1971 1 nea Coleoptera Scarabaeidae Pseudocotalpa giulianii Hardy 1974 2 nea Coleoptera Scarabaeidae Pseudoprotaetia stolata Olivier 1789 8 afr Coleoptera Scarabaeidae Pseudotrematodes frivaldszkyi Menetries 1836 8 pal Coleoptera Scarabaeidae Ptenomela gratiosa Sharp 1877 4 neo Coleoptera Scarabaeidae Pucaya pulchra Arrow 1911 1 neo Coleoptera Scarabaeidae Pycnoschema sp. 1 afr Coleoptera Scarabaeidae Pyronota festiva Fabricius 5 aus Coleoptera Scarabaeidae Reineria margaritacea Mohnike 1873 4 ori Coleoptera Scarabaeidae Reineria tigrina Mohnike 1873 1 ori Coleoptera Scarabaeidae Rhabdotis semipunctata Fabricius 1787 7 afr Coleoptera Scarabaeidae Rhizoplatys mucronatus Beauvois 1805 1 afr Coleoptera Scarabaeidae Rhizotrogus aequinoctialis Herbst 1790 6 pal Coleoptera Scarabaeidae Rhizotrogus aestivus Olivier 1789 11 pal Coleoptera Scarabaeidae Rhizotrogus maculicollis Villa 1833 1 pal Coleoptera Scarabaeidae Rhizotrogus marginipes Perris 1870 1 pal Coleoptera Scarabaeidae Rhizotrogus tusculus Buquet 1840 1 pal Coleoptera Scarabaeidae Rhizotrogus vernus Germar 1823 4 pal Coleoptera Scarabaeidae Rhomborhina japonica Hope 1841 1 ori Coleoptera Scarabaeidae Rhomborhina sp. 5 ori Coleoptera Scarabaeidae Rhomborhina unicolor Motschulsky 1861 5 ori Coleoptera Scarabaeidae Rhyparus risbeci Paulian,??? 20 aus Coleoptera Scarabaeidae Rhyssemodes orientalis Mulsant and Godart 1874 2 pal Coleoptera Scarabaeidae Rhyssemus californicus Horn 1871 3 nea Coleoptera Scarabaeidae Rhyssemus germanus Linnaeus 1767 3 pal Coleoptera Scarabaeidae Rhyssemus puncticollis Brown 1929 3 nea Coleoptera Scarabaeidae Rhyssemus scaber Haldeman 1848 5 nea Coleoptera Scarabaeidae Rhyssemus sonatus LeConte 1881 1 nea Coleoptera Scarabaeidae Rhyssemus spp. 2 Coleoptera Scarabaeidae Rutela dorcyi Olivier 1789 1 neo Coleoptera Scarabaeidae Rutela formosa Burmeister 1844 1 neo Coleoptera Scarabaeidae Rutela laeta Weber 1801 278 neo Coleoptera Scarabaeidae Rutela lineola Linnaeus 1767 78 neo Coleoptera Scarabaeidae Saprosites sp. 1 neo Coleoptera Scarabaeidae Scapanes australis Boisduval 1832 9 aus Coleoptera Scarabaeidae Scaptobius pentarthrius Westwood 1874 1 afr Coleoptera Scarabaeidae Scarabaeus cicatricosus Lucas 1846 1 pal Coleoptera Scarabaeidae Scarabaeus fritschi Harold 1868 1 afr Coleoptera Scarabaeidae Scarabaeus infernalis Klug 1855 1 afr Coleoptera Scarabaeidae Scarabaeus intricatus Fabricius 1801 2 afr Coleoptera Scarabaeidae Scarabaeus laticollis Linnaeus 1767 2 pal Coleoptera Scarabaeidae Scarabaeus pius Illiger 1803 4 pal Coleoptera Scarabaeidae Scarabaeus puncticollis Latreille 1819 5 pal Coleoptera Scarabaeidae Scarabaeus sacer Linnaeus 1758 13 pal Coleoptera Scarabaeidae Scarabaeus semipunctatus Fabricius 1792 8 pal Coleoptera Scarabaeidae Scarabaeus variolosus Fabricius 1787 9 pal Coleoptera Scarabaeidae Scatimus ovatus Harold 1862 Coleoptera Scarabaeidae Sclerostomus costatus H. and W. 1 neo Coleoptera Scarabaeidae Serica acicula Dawson 1932 1 nea Coleoptera Scarabaeidae Serica alternata LeConte 1856 85 nea Coleoptera Scarabaeidae Serica alternata Dawson 1933 1 nea Coleoptera Scarabaeidae Serica alternata Dawson 1933 2 nea Coleoptera Scarabaeidae Serica anthracina LeConte 1856 21 nea Coleoptera Scarabaeidae Serica apatela Dawson 1922 1 nea Coleoptera Scarabaeidae Serica aspera Dawson 1922 116 nea Coleoptera Scarabaeidae Serica atracapilla Kirby 1837 44 nea Coleoptera Scarabaeidae Serica brunnea Linnaeus 1758 18 pal Coleoptera Scarabaeidae Serica campestris Dawson 1919 2 nea Coleoptera Scarabaeidae Serica carolina Dawson 1920 3 nea Coleoptera Scarabaeidae Serica concinna Dawson 1947 3 nea Coleoptera Scarabaeidae Serica curvata LeConte 1856 3 nea Coleoptera Scarabaeidae Serica diablo Dawson 1967 3 nea Coleoptera Scarabaeidae Serica elusa Dawson 1919 7 nea Coleoptera Scarabaeidae Serica falli Dawson 1932 2 nea Coleoptera Scarabaeidae Serica fimbriata LeConte 1856 10 nea Coleoptera Scarabaeidae Serica floridana Dawson 1967 1 nea Coleoptera Scarabaeidae Serica frosti Dawson 1967 1 nea Coleoptera Scarabaeidae Serica frosti Dawson 1967 18 nea Coleoptera Scarabaeidae Serica frosti Dawson 1967 26 nea Coleoptera Scarabaeidae Serica georgiana Leng 1911 45 nea Coleoptera Scarabaeidae Serica georigana Dawson 1921 94 nea Coleoptera Scarabaeidae Serica holosericea Scopoli 1772 6 pal Coleoptera Scarabaeidae Serica intermixta Blatchley 1910 46 nea Coleoptera Scarabaeidae Serica iricolor Say 1824 57 nea Coleoptera Scarabaeidae Serica ligulata Dawson 1932 4 nea Coleoptera Scarabaeidae Serica mixta LeConte 1856 2 nea Coleoptera Scarabaeidae Serica opposita Dawson 1921 9 nea Coleoptera Scarabaeidae Serica parallela Casey 1884 27 nea Coleoptera Scarabaeidae Serica peregrina Chapin 1938 27 nea Coleoptera Scarabaeidae Serica perigonis Dawson 1920 2 nea Coleoptera Scarabaeidae Serica porcula Casey 1884 135 nea Coleoptera Scarabaeidae Serica prava Dawson 1933 1 nea Coleoptera Scarabaeidae Serica pusilla Dawson 1922 2 nea Coleoptera Scarabaeidae Serica pusilla Dawson 1922 8 nea Coleoptera Scarabaeidae Serica satrapa Dawson 1947 4 nea Coleoptera Scarabaeidae Serica sculptilus Dawson 1922 4 nea Coleoptera Scarabaeidae Serica seratina LeConte 1856 3 nea Coleoptera Scarabaeidae Serica sericea Illiger 1802 85 nea Coleoptera Scarabaeidae Serica similis LeConte 1895 1 nea Coleoptera Scarabaeidae Serica solita Dawson 1922 4 nea Coleoptera Scarabaeidae Serica sponsa Dawson 1919 6 nea Coleoptera Scarabaeidae Serica spp. 53 nea Coleoptera Scarabaeidae Serica tristis LeConte 1850 67 nea Coleoptera Scarabaeidae Serica trociformis Burmeister 1844 33 nea Coleoptera Scarabaeidae Serica trociformis Dawson 1932 18 nea Coleoptera Scarabaeidae Serica trogiformis Uhler 1855 10 nea Coleoptera Scarabaeidae Serica ventura Dawson 1932 1 nea Coleoptera Scarabaeidae Serica vespertina Gyllenhal 1817 47 nea Coleoptera Scarabaeidae Sericospilus advena Sharp 4 aus Coleoptera Scarabaeidae Sisyphus quadricollis Gory 1833 2 afr Coleoptera Scarabaeidae Sisyphus schaefferi Linnaeus 1758 5 pal Coleoptera Scarabaeidae Sisyraphora tomentosa Gory and Percheron 1833 4 afr Coleoptera Scarabaeidae Sphenognathus bellicosus Boileau 1 neo Coleoptera Scarabaeidae Spilophorus lugubris Fabricius 1775 2 afr Coleoptera Scarabaeidae Spilophorus plagosus Bohmann 1857 1 afr Coleoptera Scarabaeidae Spilophorus sp. 1 afr Coleoptera Scarabaeidae Spodistes mniszechi Thomas 1860 1 neo Coleoptera Scarabaeidae Stalagmosoma albella Pallas 1781 1 afr Coleoptera Scarabaeidae Stenocrates laborator Fabricius 1775 28 neo Coleoptera Scarabaeidae Stenocrates sp. 16 neo Coleoptera Scarabaeidae Stenocrates spp. 10 neo Coleoptera Scarabaeidae Stephanorrhina guttata Olivier 1789 25 afr Coleoptera Scarabaeidae Stephanucha areata Fabricius 1775 6 nea Coleoptera Scarabaeidae Stephanucha pilipennis Kraatz 1883 3 nea Coleoptera Scarabaeidae Stephanucha thoracica Casey 1915 5 nea Coleoptera Scarabaeidae Strategus aleous Linnaeus 11 neo Coleoptera Scarabaeidae Strategus aloeus Linnaeus 1758 53 nea Coleoptera Scarabaeidae Strategus aloeus Linnaeus 1758 39 nea Coleoptera Scarabaeidae Strategus aloeus Linnaeus 1758 87 neo Coleoptera Scarabaeidae Strategus antaeus Drury 1773 4 nea Coleoptera Scarabaeidae Strategus antaeus Drury 1773 113 nea Coleoptera Scarabaeidae Strategus centaurus Kolbe 1906 4 neo Coleoptera Scarabaeidae Strategus cessus LeConte 1866 35 nea Coleoptera Scarabaeidae Strategus mormon Burmeister 1847 1 nea Coleoptera Scarabaeidae Strategus splendens Beauvois 1809 27 nea Coleoptera Scarabaeidae Strategus spp. 6 Coleoptera Scarabaeidae Strategus spp. 38 nea Coleoptera Scarabaeidae Strategus spp. 66 neo Coleoptera Scarabaeidae Strategus validus Fabricius 1775 4 neo Coleoptera Scarabaeidae Strigidia liturella Kirby 1 neo Coleoptera Scarabaeidae Strigoderma arboricola Fabricius 1792 55 nea Coleoptera Scarabaeidae Strigoderma protea Burmeister 1844 21 neo Coleoptera Scarabaeidae Strigoderma pygmaea Fabricius 1798 69 nea Coleoptera Scarabaeidae Strigoderma pygmaea Ohaus 1905 2 nea Coleoptera Scarabaeidae Strigoderma pygmaea Brown 1925 2 nea Coleoptera Scarabaeidae Strigoderma sallaei Bates 1888 3 neo Coleoptera Scarabaeidae Strigoderma sulcipennis Burmeister 1844 20 neo Coleoptera Scarabaeidae Strigoderma texana Casey 1915 nea Coleoptera Scarabaeidae Strigoderma viridicollis Schaeffer 1907 2 nea Coleoptera Scarabaeidae Syrictes syrichthus Fabricius 1775 3 afr Coleoptera Scarabaeidae Tanyproctus reichei Rambur 1843 1 pal Coleoptera Scarabaeidae Taurhina longiceps Kolbe 1892 16 afr Coleoptera Scarabaeidae Taurocopris cadmus Harold 1868 1 neo Coleoptera Scarabaeidae Taurocopris mimas Linnaeus 1781 8 neo Coleoptera Scarabaeidae Telaugis aenescens Burmeister 1844 1 neo Coleoptera Scarabaeidae Tephraea dichroa Schaum 1844 1 afr Coleoptera Scarabaeidae Tesarius caelatus LeConte 1857 59 nea Coleoptera Scarabaeidae Tesarius mcclayi Cartwright 1955 13 nea Coleoptera Scarabaeidae Tesarius oregonensis Cartwright 1955 17 nea Coleoptera Scarabaeidae Thronistes rouxi Burmeister 1847 2 neo Coleoptera Scarabaeidae Thronistes spp. 3 neo Coleoptera Scarabaeidae Thyce squamicollis LeConte 1856 3 nea Coleoptera Scarabaeidae Thyridium semicinctum Bates 1888 3 neo Coleoptera Scarabaeidae Tmesorrhina iris Fabricius 1781 1 afr Coleoptera Scarabaeidae Torynorrhina hyacinthina Hope 1841 7 ori Coleoptera Scarabaeidae Trichillum pilosus Robinson 1 neo Coleoptera Scarabaeidae Trichiopsammobius brasiliensis Petrovitz 1963 1 neo Coleoptera Scarabaeidae Trichiorhyssemus riparius Horn 1871 36 nea Coleoptera Scarabaeidae Trichius abdominalis Menetries 1832 4 pal Coleoptera Scarabaeidae Trichius affinis Gory and Percheron 1833 188 nea Coleoptera Scarabaeidae Trichius assimilis Kirby 1837 132 nea Coleoptera Scarabaeidae Trichius bibens Fabricius 1775 9 nea Coleoptera Scarabaeidae Trichius fasciatus Linnaeus 1758 15 pal Coleoptera Scarabaeidae Trichius fasciatus Mulsant 1842 1 pal Coleoptera Scarabaeidae Trichius lunulatus Fabricius 1775 28 nea Coleoptera Scarabaeidae Trichius piger Fabricius 1775 77 nea Coleoptera Scarabaeidae Trichius rufobrunneus Casey 1914 54 nea Coleoptera Scarabaeidae Trichius sexualis Bedel 1906 5 pal Coleoptera Scarabaeidae Trichius sp. 7 pal Coleoptera Scarabaeidae Trichius spp. 124 nea Coleoptera Scarabaeidae Trichius succinctus Pallas 1781 2 pal Coleoptera Scarabaeidae Trichius texanus Horn 1876 11 nea Coleoptera Scarabaeidae Trichius viridans Kirby 1837 4 nea Coleoptera Scarabaeidae Trichius zonatus Germar 1794 9 pal Coleoptera Scarabaeidae Trichius zonatus Heer 1841 3 pal Coleoptera Scarabaeidae Trichogomphus excavatus Mohnike 1874 4 ori Coleoptera Scarabaeidae Trichogomphus mongol Arrow 1908 5 ori Coleoptera Scarabaeidae Tricholepis lactea Gory 1 ori Coleoptera Scarabaeidae Trichostetha signatus Fabricius 1 afr Coleoptera Scarabaeidae Trigonopeltastes delta Forster 1771 89 nea Coleoptera Scarabaeidae Trigonopeltastes floridana Casey 1909 5 nea Coleoptera Scarabaeidae Trigonopeltastes sallaei Bates 1889 1 neo Coleoptera Scarabaeidae Trigonopeltastes sinaloensis Howden 1968 2 neo Coleoptera Scarabaeidae Trigonophorus dilutus Bourgoin 1914 9 ori Coleoptera Scarabaeidae Trigonophorus gracilipes Westwood 1845 1 ori Coleoptera Scarabaeidae Trigonophorus nepalensis Hope 1831 1 ori Coleoptera Scarabaeidae Trigonophorus varians Bourgoin 1914 1 ori Coleoptera Scarabaeidae Triodonta cinctipennis Lucas 1849 9 afr Coleoptera Scarabaeidae Triodonta morio Fabricius 1798 1 pal Coleoptera Scarabaeidae Triodonta nitidula Rossi 1790 4 pal Coleoptera Scarabaeidae Trioplus cylindricus Mannerheim 1829 1 neo Coleoptera Scarabaeidae Trissodon glaber Arrow 1 aus Coleoptera Scarabaeidae Tropinota hirta Poda 1761 45 pal Coleoptera Scarabaeidae Tropinota hirta Linnaeus 1766 3 pal Coleoptera Scarabaeidae Tropinota squalida Scopoli 1783 26 pal Coleoptera Scarabaeidae Tropinota vestita Say 1825 12 pal Coleoptera Scarabaeidae Valgus californicus Horn 1870 6 nea Coleoptera Scarabaeidae Valgus canaliculatus Olivier 1789 29 nea Coleoptera Scarabaeidae Valgus hemipterus Linnaeus 1758 39 pal Coleoptera Scarabaeidae Valgus sp. 11 nea Coleoptera Scarabaeidae Xyloryctes jamaicensis Drury 1773 149 nea Coleoptera Scarabaeidae Xyloryctes spp. 64 neo Coleoptera Scarabaeidae Xyloryctes telephus Burmeister 1847 8 neo Coleoptera Scarabaeidae Xylotrupes gideon Linnaeus 1767 142 ori Coleoptera Trogidae Omorgus asper LeConte 1854 8 nea Coleoptera Trogidae Omorgus inflatus Loomis 1922 4 nea Coleoptera Trogidae Omorgus monachus Herbst 1790 22 nea Coleoptera Trogidae Omorgus nodosus Robinson 1940 6 nea Coleoptera Trogidae Omorgus persuberosus Vaurie 1962 16 pal Coleoptera Trogidae Omorgus punctatus Germar 1824 28 nea Coleoptera Trogidae Omorgus punctatus LeConte 1854 7 nea Coleoptera Trogidae Omorgus scabrosus Beauvois 1805 19 nea Coleoptera Trogidae Omorgus scutellaris Say 1823 8 nea Coleoptera Trogidae Omorgus suberosus Fabricius 1775 191 nea Coleoptera Trogidae Omorgus tesselatus LeConte 1854 4 nea Coleoptera Trogidae Omorgus tytus Robinson 1941 6 nea Coleoptera Trogidae Omorgus umbonatus LeConte 1854 1 nea Coleoptera Trogidae Polynoncus aricensis Gutierrez 1950 1 pal Coleoptera Trogidae Polynoncus brasiliensis Vaurie 1962 2 pal Coleoptera Trogidae Polynoncus bullatus Curtis 1845 1 pal Coleoptera Trogidae Polynoncus chilensis Harold 1872 1 pal Coleoptera Trogidae Polynoncus pilularius Germar 1824 2 pal Coleoptera Trogidae Trox aequalis Say 1831 9 nea Coleoptera Trogidae Trox atrox LeConte 1854 4 nea Coleoptera Trogidae Trox cadaverinus Illiger 1801 1 neo Coleoptera Trogidae Trox capillaris Say 1823 35 nea Coleoptera Trogidae Trox fabricii Reiche 1853 2 neo Coleoptera Trogidae Trox foveicollis Harold 1872 11 nea Coleoptera Trogidae Trox foveicollis Robinson 1940 4 nea Coleoptera Trogidae Trox gemmulatus Horn 1874 5 nea Coleoptera Trogidae Trox hamatus Robinson 1940 8 nea Coleoptera Trogidae Trox hispidus Pontoppidan 1763 5 neo Coleoptera Trogidae Trox insularis Chevrolat 1864 1 pal Coleoptera Trogidae Trox laticollis LeConte 1854 9 nea Coleoptera Trogidae Trox plicatus Robinson 1940 54 nea Coleoptera Trogidae Trox sabulosus Linnaeus 1758 3 neo Coleoptera Trogidae Trox scaber Linnaeus 1767 3 Coleoptera Trogidae Trox sonorae LeConte 1854 116 nea Coleoptera Trogidae Trox sordidus LeConte 1854 29 nea Coleoptera Trogidae Trox spinulosus Robinson 1940 26 nea Coleoptera Trogidae Trox striatus Melsheimer 1846 3 nea Coleoptera Trogidae Trox sulcatus Thunberg 1787 2 afr Coleoptera Trogidae Trox terrestris Say 1825 24 nea Coleoptera Trogidae Trox tuberculatus Degeer 1774 15 nea Coleoptera Trogidae Trox unistriatus Beauvois 1805 70 nea Coleoptera Trogidae Trox variolatus Melsheimer 1847 24 nea Coleoptera Trogidae undetermined spp. 6