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Scarabaeoidea Order Family Genus Species Author Year Exx. Region Coleoptera Ceratocanthidae Germarostes aphodioides Illiger 1800 71 nea Coleoptera Ceratocanthidae Germarostes globosus Say 1835 33 nea Coleoptera Ceratocanthidae Germarostes macleayi Perty 1830 1 pal Coleoptera Ceratocanthidae Germarostes plicatus Germar 1843 3 pal Coleoptera Ceratocanthidae Germarostes punctulatus Ohaus 1911 1 pal Coleoptera Ceratocanthidae Haroldostes senegalensis Castelnau 1840 1 pal Coleoptera Diphyllostomatidae Diphyllostoma fimbriata Fall 1901 3 nea Coleoptera Diphyllostomatidae Diphyllostoma linsleyi Fall 1932 6 nea Coleoptera Diphyllostomatidae Diphyllostoma nigricollis Fall 1912 34 nea Coleoptera Geotrupidae Anoplotrupes ballyi Jekel 1865 45 nea Coleoptera Geotrupidae Anoplotrupes hornii Blanchard 1888 13 nea Coleoptera Geotrupidae Anoplotrupes stercorosus Scriba 1791 77 pal Coleoptera Geotrupidae Athyreus excavatus Castelnau 1840 1 pal Coleoptera Geotrupidae Athyreus sp. 1 pal Coleoptera Geotrupidae Blackburnium insigne Lea 1916 2 aus Coleoptera Geotrupidae Blackburnium sloanei Blackburn 1889 1 aus Coleoptera Geotrupidae Bolbelasmmus horni Rivers 1886 1 nea Coleoptera Geotrupidae Bolbocerastes imperialis Cartwright 1953 9 nea Coleoptera Geotrupidae Bolbocerastes regalis Cartwright 1953 1 nea Coleoptera Geotrupidae Bolbocerastes serratus LeConte 1854 10 nea Coleoptera Geotrupidae Bolbocerosoma biplagiatum Dawson and McCollock 1924 4 nea Coleoptera Geotrupidae Bolbocerosoma bruneri Dawson and McCollock 1924 8 nea Coleoptera Geotrupidae Bolbocerosoma confusum Brown 1928 2 nea Coleoptera Geotrupidae Bolbocerosoma farctum Fabricius 1775 9 nea Coleoptera Geotrupidae Bolbocerosoma hamatum Brown 1929 3 Coleoptera Geotrupidae Bolbocerosoma pusillum Dawson and McCollock 1924 1 nea Coleoptera Geotrupidae Bolbocerosoma tumefactum Beauvois 1805 8 nea Coleoptera Geotrupidae Bolbochromus n.sp. 1 ori Coleoptera Geotrupidae Bolborhombus sallaei Bates 1887 9 nea Coleoptera Geotrupidae Bradycinetulus ferrugineus Beauvois 1805 1 nea Coleoptera Geotrupidae Bradycinetulus fossatus Haldeman 1853 5 nea Coleoptera Geotrupidae Ceratophyus polyceros Pallas 1771 2 pal Coleoptera Geotrupidae Cnemotrupes blackburnii Fabricius 1781 28 nea Coleoptera Geotrupidae Cnemotrupes semiopacus Jekel 1865 45 nea Coleoptera Geotrupidae Cnemotrupes splendidus Fabricius 1781 40 nea Coleoptera Geotrupidae Enoplotrupes sharpi Jordan and Rothschild 1893 2 pal Coleoptera Geotrupidae Eogeotrupes auratus Motschulsky 1857 2 Coleoptera Geotrupidae Epigeotrupes laevistriatus Motschulsky 1 pal Coleoptera Geotrupidae Eucanthus alutaceus Cartwright 1944 1 nea Coleoptera Geotrupidae Eucanthus greeni Robinson 6 nea Coleoptera Geotrupidae Eucanthus impressus Howden 1964 10 nea Coleoptera Geotrupidae Eucanthus lazarus Fabricius 1775 109 nea Coleoptera Geotrupidae Eucanthus lazarus 33 nea Coleoptera Geotrupidae Frickius variolosus Germain 1897 6 pal Coleoptera Geotrupidae Geohowdenius egeriei Germar 1824 14 nea Coleoptera Geotrupidae Geohowdenius opacus Haldeman 1853 7 nea Coleoptera Geotrupidae Geotrupes mutator Marsham 1802 12 pal Coleoptera Geotrupidae Geotrupes spiniger Marsham 1802 6 pal Coleoptera Geotrupidae Geotrupes stercorarius Linnaeus 1758 12 pal Coleoptera Geotrupidae Glyptogeotrupes impressus Gebler 1841 1 pal Coleoptera Geotrupidae Lethrus acutangulus Ballion 1871 2 pal Coleoptera Geotrupidae Lethrus apterus Laxm. 1770 2 Coleoptera Geotrupidae Lethrus apterus Fischer 1822 2 pal Coleoptera Geotrupidae Lethrus cephalotus Pallas 1771 4 neo Coleoptera Geotrupidae Lethrus inermis Reitter 1897 1 neo Coleoptera Geotrupidae Lethrus karelini Gebler 1845 2 neo Coleoptera Geotrupidae Lethrus koshantschikovi Semenov 1892 1 neo Coleoptera Geotrupidae Lethrus scoparius Fischer 1820 2 neo Coleoptera Geotrupidae Lethrus sulcipennis Kraatz 1883 2 neo Coleoptera Geotrupidae Mycotrupes gaigei Olsen and Hubbel 1954 1 nea Coleoptera Geotrupidae Mycotrupes retusus LeConte 1866 1 nea Coleoptera Geotrupidae Neoathyreus excavatus Laporte 1840 1 pal Coleoptera Geotrupidae Neoathyreus fissicornis Harold 1880 1 pal Coleoptera Geotrupidae Neoathyreus lanei Martinez 1952 2 pal Coleoptera Geotrupidae Odonteus binasutum Fairmaire 1861 2 pal Coleoptera Geotrupidae Odonteus capense Klug 1843 2 afr Coleoptera Geotrupidae Odonteus corniger Melsheimer 1844 4 nea Coleoptera Geotrupidae Odonteus darlingtoni Wallis 1928 4 nea Coleoptera Geotrupidae Odonteus distinguendum Fairmaire 1861 1 pal Coleoptera Geotrupidae Odonteus falli Wallis 1928 4 nea Coleoptera Geotrupidae Odonteus floridanus Wallis 1928 31 nea Coleoptera Geotrupidae Odonteus gallicus Mulsant 1842 3 neo Coleoptera Geotrupidae Odonteus laesicolle Fairmaire 1856 1 pal Coleoptera Geotrupidae Odonteus liebecki Wallis 1928 34 nea Coleoptera Geotrupidae Odonteus obesus LeConte 1859 20 nea Coleoptera Geotrupidae Odonteus simi Wallis 1928 3 nea Coleoptera Geotrupidae Odonteus sp. 1 nea Coleoptera Geotrupidae Odonteus sp. 1 ori Coleoptera Geotrupidae Odonteus striatopunctatum Castelnau 1840 1 Coleoptera Geotrupidae Odonteus thoracicornis Wallis 1928 2 nea Coleoptera Geotrupidae Peltotrupes profundus Howden 1952 5 nea Coleoptera Geotrupidae Sericotrupes niger Marsham 1802 2 pal Coleoptera Geotrupidae Taurocerastes patagonicus Phillipi 1866 2 pal Coleoptera Geotrupidae Thorectes geminatus Gene 1839 3 pal Coleoptera Geotrupidae Thorectes intermedius Costa 1827 9 pal Coleoptera Geotrupidae Thorectes marginatus Poiret 1787 2 pal Coleoptera Geotrupidae Thorectes rugatulus Jekel 1865 2 pal Coleoptera Geotrupidae Trypocopris pyrenaeus Charpentier 1825 2 pal Coleoptera Geotrupidae Trypocopris vernalis Linnaeus 1758 13 pal Coleoptera Geotrupidae Trypocopris vernalis Heer 1847 1 pal Coleoptera Geotrupidae Trypocopris vernalis Kuster 1853 4 pal Coleoptera Geotrupidae Typhoeus typhoeus Linn. 1758 8 pal Coleoptera Geotrupidae Typhoeus typhoeus Linnaeus 1758 8 pal Coleoptera Glaphyridae Amphicoma acrtos Faldermann 1835 1 pal Coleoptera Glaphyridae Amphicoma arctos Reitter 1890 1 pal Coleoptera Glaphyridae Amphicoma bombyliformis Pallas 1781 1 pal Coleoptera Glaphyridae Amphicoma bombylius Fabricius 1789 8 pal Coleoptera Glaphyridae Amphicoma fastuosa Reitter 1890 1 pal Coleoptera Glaphyridae Amphicoma hyrax Truqui 1848 1 pal Coleoptera Glaphyridae Amphicoma kuschakewitschi Ball 1871 1 pal Coleoptera Glaphyridae Amphicoma lasserrei Germar 1834 17 pal Coleoptera Glaphyridae Amphicoma malatia Truqui 1848 7 pal Coleoptera Glaphyridae Amphicoma papaveris Sturm 1843 1 pal Coleoptera Glaphyridae Amphicoma vittata Fabricius 1775 1 pal Coleoptera Glaphyridae Amphicoma vittata Reitter 1890 2 neo Coleoptera Glaphyridae Amphicoma vittatta Reitter 1890 1 neo Coleoptera Glaphyridae Amphicoma vulpes Brulle 1 neo Coleoptera Glaphyridae Amphicoma vulpes Fabricius 1792 2 pal Coleoptera Glaphyridae Cratoscelis villosa Blanchard 1850 7 neo Coleoptera Glaphyridae Cratoscelis vulpina Erichson 1835 2 neo Coleoptera Glaphyridae Glaphyrus festivus Menier 1836 1 pal Coleoptera Glaphyridae Glaphyrus maurus Linnaeus 1758 3 afr Coleoptera Glaphyridae Glaphyrus micans Faldermann 1835 1 pal Coleoptera Glaphyridae Lichanthe rathvoni LeConte 1863 13 nea Coleoptera Glaphyridae Lichanthe ursina LeConte 1861 4 nea Coleoptera Glaphyridae Lichanthe vulpina Hentz 1826 16 nea Coleoptera Glaphyridae Lichnia limbata Erichson 1851 4 neo Coleoptera Glaresidae Glaresis ecostata Fall 1907 1 nea Coleoptera Glaresidae Glaresis gordoni Warner 2 nea Coleoptera Glaresidae Glaresis phoenicia Fall 1907 1 nea Coleoptera Glaresidae Glaresis zvirgzdinsi Warner 2 nea Coleoptera Hybosoridae Coelodes ovalis Robinson Coleoptera Hybosoridae Hybosorus illigeri Reiche 1853 8 nea Coleoptera Hybosoridae Hybosorus spp. 18 Coleoptera Lucanidae Aegus minutus Gestro 1 Coleoptera Lucanidae Aesalus scarabaeoides Panzer 1794 1 neo Coleoptera Lucanidae Apterocyclus honoluluensis Waterhouse 1 Coleoptera Lucanidae Apterodorcus bacchus Westwood 4 Coleoptera Lucanidae Bomansodus impubis Parry 4 Coleoptera Lucanidae Brasilucanus alvaregei Vulcano and Pereira 1 Coleoptera Lucanidae Casignetus humboldti Gyllenhal 1 Coleoptera Lucanidae Ceratognathus curvidens Broun 1904 2 aus Coleoptera Lucanidae Ceruchus chrysomelinus Hochenwarth 1785 2 neo Coleoptera Lucanidae Ceruchus piceus Weber 1801 297 nea Coleoptera Lucanidae Ceruchus punctatus LeConte 1869 3 nea Coleoptera Lucanidae Ceruchus striatus LeConte 1859 9 nea Coleoptera Lucanidae Chalcodes carinatus Linnaeus 2 Coleoptera Lucanidae Chiasognathinus gaujoni Oberthuer 1 Coleoptera Lucanidae Chiasognathus granti Stephens 10 Coleoptera Lucanidae Cladognathus confucius Hope 1 Coleoptera Lucanidae Cladognathus confucius 1 Coleoptera Lucanidae Cladognathus giraffa Olivier 4 Coleoptera Lucanidae Cyclommatus frey-gessneri Ritsema 1 Coleoptera Lucanidae Cyclommatus tarandus Thuenberg 15 Coleoptera Lucanidae Dorcus brevis Say 4 Coleoptera Lucanidae Dorcus immundus Arrow 1 Coleoptera Lucanidae Dorcus parallelepipedus Linnaeus 21 Coleoptera Lucanidae Dorcus parallelepipedus Ganglbauer 3 Coleoptera Lucanidae Dorcus parallelus Say 82 Coleoptera Lucanidae Eurytrachellelus alcides Vollenhoven 1 Coleoptera Lucanidae Eurytrachellelus arfakianus Lansberge 2 Coleoptera Lucanidae Eurytrachellelus bucephalus Perty 5 Coleoptera Lucanidae Eurytrachellelus cribriceps Chevrolat 3 Coleoptera Lucanidae Eurytrachellelus egregius Moellenkamp 2 Coleoptera Lucanidae Eurytrachellelus gypaetus Castelnau 1 Coleoptera Lucanidae Eurytrachellelus intermedius Gestro 26 Coleoptera Lucanidae Eurytrachellelus reichei Hope 1 Coleoptera Lucanidae Eurytrachellelus
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