Experimental Archaeology, Macrofracture Analyses and Morphometric Techniques

Total Page:16

File Type:pdf, Size:1020Kb

Experimental Archaeology, Macrofracture Analyses and Morphometric Techniques INTERPRETATIVE TOOLS FOR STUDYING STONE AGE HUNTING TECHNOLOGIES: EXPERIMENTAL ARCHAEOLOGY, MACROFRACTURE ANALYSES AND MORPHOMETRIC TECHNIQUES. Justin Pargeter A dissertation submitted to the Faculty of Science, University of the Witwatersrand, in fulfilment of the requirements for the degree of Master of Science Johannesburg, 2011 i DECLARATION I declare that this dissertation is my own unaided work. It is submitted for the degree of Master of Science to the Faculty of Science at the University of the Witwatersrand, Johannesburg. It has not been submitted before for any other degree or examination at any other university. Signed Justin Harry Pargeter On this the 05 day of April 2011 ii ABSTRACT This dissertation contains an assessment and use of the macrofracture and morphometric methods for detecting Later Stone Age hunting weaponry. Two sets of replicated unretouched stone artefacts were trampled by cattle and humans to determine the formation of impact fractures under these, and knapping conditions. The results suggest that small frequencies (c. 3 %) of certain impact fracture types do occur on flakes subject to trampling and knapping forces. Macrofracture and morphometric data were recorded for stone artefacts (bladelets, backed artefacts and convergent pieces) from Robberg (c. 18 000 - 12 000 years ago) and Wilton (c. 8000 - 2000 years ago) Later Stone Age assemblages on the southern Cape coast. Impact fracture frequencies were similar in these two samples, but were significantly higher than in the trampling experiments. The morphometric data suggests, on average, congruence between Later Stone Age tools with impact fractures and experimental, archaeological and ethnohistoric spear and arrow tips. Based on these results it appears likely that Wilton backed artefacts, specifically segments, were used as arrowheads and it is unclear at present what weapon types were used during the Robberg phase although the use of spears seems probable. iii ACKNOWLEDGMENTS This dissertation is dedicated to all the hooves and feet that made the experiments possible. I would like to thank all my family and friends who gave support, advice and input into this project and I would especially like to thank my supervisors, post-graduate support staff and the PAST and NRF foundations for financial and academic assistance during this project. iv CONTENTS PAGE DECLARATION ............................................................................................................................................... II ABSTRACT ......................................................................................................................................................... III ACKNOWLEDGMENTS .................................................................................................................................... IV LIST OF FIGURES ............................................................................................................................................ VIII LIST OF TABLES ............................................................................................................................................... XI 1 CHAPTER 1: INTRODUCTION ......................................................................................................... 1 1.1 MACROFRACTURE ANALYSIS AND TRAMPLING/KNAPPING EXPERIMENTS .............................................. 1 1.2 ASSESSING HUNTING WEAPONRY USING MACROFRACTURE ANALYSIS ................................................... 2 1.3 DISSERTATION STRUCTURE .................................................................................................................... 4 2 CHAPTER 2: MACROFRACTURE METHOD BACKGROUND .................................................. 6 2.1 FRACTURE TYPES, NOMENCLATURE AND VARIABLES ............................................................................. 7 2.2 MACROFRACTURE EXPERIMENTS AND ARCHAEOLOGICAL USES OUTSIDE OF AFRICA .......................... 12 2.3 MACROFRACTURE EXPERIMENTS IN AFRICA AND THE MIDDLE EAST .................................................. 18 2.4 ARCHAEOLOGICAL APPLICATIONS OF THE MACROFRACTURE METHOD IN AFRICA ............................... 26 2.5 CHAPTER SUMMARY ............................................................................................................................ 28 3 CHAPTER 3: BACKGROUND TO MORPHOMETRIC ANALYSES ......................................... 29 3.1 CHAPTER SUMMARY ............................................................................................................................ 31 4 CHAPTER 4: BACKGROUND TO EXPERIMENTAL ARCHAEOLOGY ................................. 32 4.1 INTRODUCTION .................................................................................................................................... 32 4.2 THEORETICAL TENETS OF EXPERIMENTAL ARCHAEOLOGY ................................................................... 33 4.3 BUILDING ANALOGIES ALONG EXPERIMENTAL LINES ........................................................................... 34 4.4 THE APPLICABILITY OF EXPERIMENTS IN ARCHAEOLOGY .................................................................... 35 4.5 REFERRING TO THE ARCHAEOLOGICAL RECORD ................................................................................... 37 4.6 A TYPOLOGY OF EXPERIMENTS IN ARCHAEOLOGY ............................................................................... 38 4.7 CHAPTER SUMMARY ............................................................................................................................ 40 5 CHAPTER 5: BACKGROUND TO THE FUNCTIONAL STUDY OF BLADELETS, BACKED ARTEFACTS AND CONVERGENT PIECES .............................................................................. 41 5.1 INTRODUCTION .................................................................................................................................... 41 5.2 BACKGROUND TO BLADELETS, BACKED ARTEFACTS AND CONVERGENT PIECES................................... 41 5.3 HUNTING WEAPONRY FORMS AND FUNCTIONS ..................................................................................... 50 5.4 ARCHAEOLOGICAL EVIDENCE FOR MECHANICAL AND NON-MECHANICAL HUNTING WEAPON TYPES..................................................................................................................................... 53 v 5.5 CHAPTER SUMMARY ............................................................................................................................ 56 6 CHAPTER 6: BACKGROUND TO ARCHAEOLOGICAL SITES AND SAMPLES ................. 57 6.1 INTRODUCTION .................................................................................................................................... 57 6.2 BACKGROUND TO NELSON BAY CAVE ................................................................................................. 57 6.3 BACKGROUND TO BYNESKRANSKOP 1 ................................................................................................. 66 6.4 BACKGROUND TO BLOMBOSFONTEIN NATURE RESERVE SITE 4 .......................................................... 74 6.5 CHAPTER SUMMARY ............................................................................................................................ 79 7 CHAPTER 7: EXPERIMENTAL METHODS ................................................................................. 80 7.1 INTRODUCTION AND AIMS .................................................................................................................... 80 7.2 BACKGROUND TO THE TRAMPLING EXPERIMENTS ................................................................................ 81 7.3 EXPERIMENTAL MATERIALS ................................................................................................................. 82 7.4 EXPERIMENTAL METHODS ................................................................................................................... 86 7.5 MACROFRACTURE ANALYSIS ............................................................................................................... 91 7.6 CHAPTER SUMMARY ............................................................................................................................ 91 8 CHAPTER 8: MACROFRACTURE AND MORPHOMETRIC METHODS ............................... 92 8.1 INTRODUCTION .................................................................................................................................... 92 8.2 APPLICATION OF THE MACROFRACTURE METHOD ................................................................................ 92 8.3 MORPHOMETRIC AND STATISTICAL METHODS EMPLOYED IN THIS STUDY ............................................ 94 8.4 CHAPTER SUMMARY ............................................................................................................................ 99 9 CHAPTER 9: RESULTS OF MACROFRACTURE ANALYSIS ON THE EXPERIMENTAL MATERIALS ................................................................................................................................... 100 9.1 INTRODUCTION .................................................................................................................................. 100 9.2 MACROFRACTURE RESULTS ON THE TRAMPLING AND KNAPPING EXPERIMENTAL FLAKES ................. 100 9.3 RESULTS AS PER DEPOSITIONAL DEPTH .............................................................................................. 107 9.4 RESULTS AS PER ROCK TYPE..............................................................................................................
Recommended publications
  • Standards for Ruminant Sanctuaries
    Global Federation of Animal Sanctuaries Standards For Ruminant Sanctuaries Version: April 2019 ©2012 Global Federation of Animal Sanctuaries Global Federation of Animal Sanctuaries – Standards for Ruminant Sanctuaries Table of Contents INTRODUCTION...................................................................................................................................... 1 GFAS PRINCIPLES ................................................................................................................................................... 1 ANIMALS COVERED BY THESE STANDARDS ............................................................................................................ 1 STANDARDS UPDATES ........................................................................................................................................... 2 RUMINANT STANDARDS ........................................................................................................................................ 2 RUMINANT HOUSING ........................................................................................................................... 2 H-1. Types of Space and Size ..................................................................................................................................... 2 H-2. Containment ...................................................................................................................................................... 5 H-3. Ground and Plantings ........................................................................................................................................
    [Show full text]
  • Boselaphus Tragocamelus</I>
    University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln USGS Staff -- Published Research US Geological Survey 2008 Boselaphus tragocamelus (Artiodactyla: Bovidae) David M. Leslie Jr. U.S. Geological Survey, [email protected] Follow this and additional works at: https://digitalcommons.unl.edu/usgsstaffpub Leslie, David M. Jr., "Boselaphus tragocamelus (Artiodactyla: Bovidae)" (2008). USGS Staff -- Published Research. 723. https://digitalcommons.unl.edu/usgsstaffpub/723 This Article is brought to you for free and open access by the US Geological Survey at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in USGS Staff -- Published Research by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. MAMMALIAN SPECIES 813:1–16 Boselaphus tragocamelus (Artiodactyla: Bovidae) DAVID M. LESLIE,JR. United States Geological Survey, Oklahoma Cooperative Fish and Wildlife Research Unit and Department of Natural Resource Ecology and Management, Oklahoma State University, Stillwater, OK 74078-3051, USA; [email protected] Abstract: Boselaphus tragocamelus (Pallas, 1766) is a bovid commonly called the nilgai or blue bull and is Asia’s largest antelope. A sexually dimorphic ungulate of large stature and unique coloration, it is the only species in the genus Boselaphus. It is endemic to peninsular India and small parts of Pakistan and Nepal, has been extirpated from Bangladesh, and has been introduced in the United States (Texas), Mexico, South Africa, and Italy. It prefers open grassland and savannas and locally is a significant agricultural pest in India. It is not of special conservation concern and is well represented in zoos and private collections throughout the world. DOI: 10.1644/813.1.
    [Show full text]
  • Biodiversity in Sub-Saharan Africa and Its Islands Conservation, Management and Sustainable Use
    Biodiversity in Sub-Saharan Africa and its Islands Conservation, Management and Sustainable Use Occasional Papers of the IUCN Species Survival Commission No. 6 IUCN - The World Conservation Union IUCN Species Survival Commission Role of the SSC The Species Survival Commission (SSC) is IUCN's primary source of the 4. To provide advice, information, and expertise to the Secretariat of the scientific and technical information required for the maintenance of biologi- Convention on International Trade in Endangered Species of Wild Fauna cal diversity through the conservation of endangered and vulnerable species and Flora (CITES) and other international agreements affecting conser- of fauna and flora, whilst recommending and promoting measures for their vation of species or biological diversity. conservation, and for the management of other species of conservation con- cern. Its objective is to mobilize action to prevent the extinction of species, 5. To carry out specific tasks on behalf of the Union, including: sub-species and discrete populations of fauna and flora, thereby not only maintaining biological diversity but improving the status of endangered and • coordination of a programme of activities for the conservation of bio- vulnerable species. logical diversity within the framework of the IUCN Conservation Programme. Objectives of the SSC • promotion of the maintenance of biological diversity by monitoring 1. To participate in the further development, promotion and implementation the status of species and populations of conservation concern. of the World Conservation Strategy; to advise on the development of IUCN's Conservation Programme; to support the implementation of the • development and review of conservation action plans and priorities Programme' and to assist in the development, screening, and monitoring for species and their populations.
    [Show full text]
  • Research Description for Shyam Thapa
    Research description for Shyam Thapa Doubling tiger numbers means doubling the prey-base: but how? The Terai Arc is landscape lying between the Ganges Plain and the foothills of the Himalayas. Biogeographically it is the most northern part of the Oriental Kingdom, and zoologically it is characterised by a mixture of Tropical species, like the Indian Rhinoceros, the Indian Tiger, the Asian Elephant, the Gavials or Hornbills and Palearctic species, such as the many migratory bird species but also mammals such as Wolves and Wild Boar. Botanically, the Oriental Floral Kingdom is a mixture of Pan-Tropical species, and shares Ethiopian species with Africa and Indo-Malayan species with the tropical Far-East. The Terai, which stretches from the area bordering Assam and Myanmar to the Punjab, was till recently nearly one hundred percent forested. Malaria eradication programmes in the 1960s enabled settlement by non-malaria resistant people, and the Terai quickly filled up with settlers from the foothills of the Himalayas to the North and from the Gangetic Plain to the South. This landnam resulted in a very fast deforestation of this stretch of land of some 1200 km E-W and 60 km N-S and now, in 2017, only little of the original vegetation still exists in a few isolated remnants which got legal protection as National Parks, both in India and Nepal. In these ever-shrinking pieces of forest an ever-shrinking population of tigers occurred. At the start of the 21st century, the world-wide number of tiger remaining in the wild approached 2000 in total in the so-called ‘range states’ (e.g., Russia, Nepal, China, India, Malaysia, Indonesia).
    [Show full text]
  • The Paris Bloubok (Hippotragus Leucophaeus (Pallas, 1766) [Bovidae]) and Its Provenance
    DIRECTEUR DE LA PUBLICATION : Bruno David Président du Muséum national d’Histoire naturelle RÉDACTRICE EN CHEF / EDITOR-IN-CHIEF : Laure Desutter-Grandcolas ASSISTANTS DE RÉDACTION / ASSISTANT EDITORS : Anne Mabille ([email protected]) MISE EN PAGE / PAGE LAYOUT : Anne Mabille COMITÉ SCIENTIFIQUE / SCIENTIFIC BOARD : James Carpenter (AMNH, New York, États-Unis) Maria Marta Cigliano (Museo de La Plata, La Plata, Argentine) Henrik Enghoff (NHMD, Copenhague, Danemark) Rafael Marquez (CSIC, Madrid, Espagne) Peter Ng (University of Singapore) Norman I. Platnick (AMNH, New York, États-Unis) Jean-Yves Rasplus (INRA, Montferrier-sur-Lez, France) Jean-François Silvain (IRD, Gif-sur-Yvette, France) Wanda M. Weiner (Polish Academy of Sciences, Cracovie, Pologne) John Wenzel (The Ohio State University, Columbus, États-Unis) COUVERTURE / COVER : Illustration of Levaillant’s Bloubok. University of Leiden Library. Courtesy of the Library. Zoosystema est indexé dans / Zoosystema is indexed in: – Science Citation Index Expanded (SciSearch®) – ISI Alerting Services® – Current Contents® / Agriculture, Biology, and Environmental Sciences® – Scopus® Zoosystema est distribué en version électronique par / Zoosystema is distributed electronically by: – BioOne® (http://www.bioone.org) Les articles ainsi que les nouveautés nomenclaturales publiés dans Zoosystema sont référencés par / Articles and nomenclatural novelties published in Zoosystema are referenced by: – ZooBank® (http://zoobank.org) Zoosystema est une revue en flux continu publiée par les Publications scientifiques du Muséum, Paris / Zoosystema is a fast track journal published by the Museum Science Press, Paris Les Publications scientifiques du Muséum publient aussi / The Museum Science Press also publish: Adansonia, Geodiversitas, Anthropozoologica, European Journal of Taxonomy, Naturae, Cryptogamie sous-sections Algologie, Bryologie, Mycologie. Diffusion – Publications scientifiques Muséum national d’Histoire naturelle CP 41 – 57 rue Cuvier F-75231 Paris cedex 05 (France) Tél.
    [Show full text]
  • California Geophytesgeophytes
    $12.00 (Free to Members) VOL. 44, NO.3 • DECEMBER 2016 FREMONTIAFREMONTIA JOURNAL OF THE CALIFORNIA NATIVE PLANT SOCIETY SPECIAL ISSUE: VOL. 44, NO. 3, DECEMBER 2016 FREMONTIA CALIFORNIACALIFORNIA GEOPHYTESGEOPHYTES V44_3_cover.pmd 1 2/20/17, 5:26 AM CALIFORNIA NATIVE PLANT SOCIETY CNPS, 2707 K Street, Suite 1; Sacramento, CA 95816-5130 FREMONTIA Phone: (916) 447-2677 Fax: (916) 447-2727 Web site: www.cnps.org Email: [email protected] VOL. 44, NO. 3, DECEMBER 2016 MEMBERSHIP Copyright © 2016 Members receive many benefits, including subscriptions to Fremontia and California Native Plant Society the CNPS Bulletin. Membership form is on inside back cover. Mariposa Lily . $1,500 Family or Group . $75 Benefactor . $600 International or Library . $75 M. Kat Anderson, Guest Editor Patron . $300 Individual . $45 Michael Kauffmann, Editor Plant Lover . $100 Student/Retired/Limited Income . $25 CORPORATE/ORGANIZATIONAL Beth Hansen-Winter, Designer 10+ Employees . $2,500 4-6 Employees . $500 7-10 Employees . $1,000 1-3 Employees . $150 california Native STAFF & CONTRACTORS Plant Society Dan Gluesenkamp: Executive Director Marin: Charlotte Torgovitsky Chris Brown: Admin Assistant Milo Baker: Leia Giambastiani, Sarah Protecting California’s Native Flora Jennifer Buck-Diaz: Vegetation Ecologist Gordon Since 1965 Catherine Curley: Assistant Botanist Mojave Desert: Timothy Thomas Joslyn Curtis, Assistant Veg. Ecologist Monterey Bay: Christopher Hauser The views expressed by authors do not Julie Evens: Vegetation Program Dir. Mount Lassen: Woody Elliot necessarily
    [Show full text]
  • Digestive Strategies in Ruminants and Nonruminants S.E. Van Wieren
    DIGESTIVE STRATEGIES IN RUMINANTS AND NONRUMINANTS S.E. VANWIERE N 0000 0714 3726 Promotoren: Dr. ir. S. Tamminga, buitengewoon hoogleraar op het vakgebied van de veevoeding in het bijzonder de voeding van herkauwers Dr. H.H.T. Prins, hoogleraar in het natuurbeheer in de tropen en oecologie van vertebraten WMoPûi , ZI 9 S.E. van Wieren DIGESTIVE STRATEGIES IN RUMINANTS AND NONRUMINANTS Proefschrift ter verkrijging van de graad van doctor op gezag van de rector magnificus van de Landbouwuniversiteit Wageningen, dr. C.M. Karssen, in het openbaar te verdedigen op dinsdag 3 december 1996 des namiddags te vier uur in de Aula. aiqo?<? T CIP-DATA KONINKLIJKE BIBLIOTHEEK, DEN HAAG Van Wieren, S.E. Digestive strategies in ruminants and nonruminants / S.E. van Wieren. - Thesis Landbouw Universiteit Wageningen. - With réf. - With summary in Dutch. ISBN 90-5485-611-4 Subject headings: digestion / ruminants /nonruminants / feeding ecology cr.-::i n; . •Y.:r Cover and illustrations: Esther van Nie & Marjolein Wiersma STELLINGEN I De veelvuldig aangehangen gedachte dat paarden, wat betreft de dagelijkse opname van metaboliseerbare energie uit laagwaardig voedsel, superieur zijn aan runderen, wordt niet door alle onderzoek ondersteund en blijkt ook niet altijd uit praktijkervaringen. P. Duncan et al. (1990). Oecologia 84:411-418. R. Meydam. Evaluatie begrazing Meyendel. 1996. II Wilde zwijnen die op een dieet van uitsluitend mast leven, krijgen onherroepelijk eiwitgebrek. III Het concept van duurzame ontwikkeling leidt in de praktijk niet zozeer tot beperkingen aan de groei, maar meer tot de groei van de beperkingen. B. Willers. (1994). Conservation Biology (8):1146-1148 . IV De typisch nederlandse gedachte dat de mens verrijkend heeft gewerkt op de natuur, is een gevaarlijke misvatting.
    [Show full text]
  • The Cape Floristic Region, Within Which
    1 heCape Floristic Region, within which in the Overberg wheat belt, so that landowners you are farming, is the smallest, yet can see how we are all working together towards richest Plant Kingdom in the world. a shared vision, through partnering with farmers It is comprised of Mountain Fynbos to ensure the long-term survival of this Critically T(nutrient-poor soils), Coastal Strandveld (nutrient Endangered habitat and all its dependent rich, alkaline soils) and Renosterveld (nutrient- wildlife. The remnants and watercourses that rich, clay-derived soils). Fynbos and Strandveld hold the ecological functioning of the Rûens tend to be dominated by the showier Proteas, wheat belt together are currently threatened by Ericas and Restios, while Renosterveld is generally mismanagement. In contrast, several coastal and dominated by grasses and shrubs belonging to montane reserves exist in the areas surrounding the daisy family. Because Renosterveld is found the wheat belt. The remnants of natural habitats on more fertile soils, it has been converted and watercourses in the Rûens can be considered extensively to croplands, to such a degree that the ‘life blood’ of this region. Landowners are less than 5% remains. Lowland Renosterveld, starting to realise that a paradigm shift in mind- often referred to as uitvalgrond (‘wasteland’) by sets and management is required if we are to keep farmers, has been overlooked, misunderstood this landscape alive and functioning. The Overberg and largely neglected. However, Renosterveld is Lowlands Conservation Trust is committed to considered the richest bulb habitat on Earth and is facilitating these changes, through building renowned for its incredible spring ower displays relationships with landowners and assisting with - many popular horticultural bulbs were bred out management on the ground.
    [Show full text]
  • BIODIVERSITY MANAGEMENT PLAN for the BONTEBOK Damaliscus Pygargus Pygargus in SOUTH AFRICA
    SCHEDULE BIODIVERSITY MANAGEMENT PLAN FOR THE BONTEBOK Damaliscus pygargus pygargus IN SOUTH AFRICA Jointly developed by SANParks and CapeNature Carly Cowell¹ and Coral Birss² ¹Cape Research Centre, South African National Parks ²Scientific Services, CapeNature 1 | P a g e Citation Cowell, C.R. and Birss, C. 2017. Biodiversity Management Plan for The Bontebok (Damaliscus pygargus pygargus) in South Africa. Jointly developed by South African National Parks, CapeNature and the National Department of Environmental Affairs. Version 1.0 2 | P a g e FOREWORD - PROF CHRISTO FABRICIUS Soon after the first Europeans arrived in South Africa they encountered two spectacularly coloured antelope types that closely resembled one another. They called them bontebok and Blesbok but, as one does with siblings that look alike, regularly mistook one for the other; the first of many mix- ups in the bontebok’s recent history. This on-going confusion was an important contributor to the near-extinction of bontebok with its narrower habitat requirements and smaller numbers. The unfortunate animal was, after being almost hunted to extinction, practically hybridized to extinction through human-influenced interbreeding. But it was also humans, the cause of the problem, who brought bontebok back from the brink when a few farmers in the Bredasdorp area started to actively conserve them. These bold actions, aimed at breeding the species, first gave rise to the establishment of bontebok National Park and later led to the proclamation of De Hoop Nature Reserve. The fortunate (albeit unintended) consequence of this was the preservation of equally threatened ecosystems: Fynbos and Renosterveld. Its fascinating history and strong cultural and ecological association with the Fynbos biome make the bontebok an ideal flagship for ecosystem conservation.
    [Show full text]
  • Western Cape Province, South Africa Mammals
    Western Cape State of Biodiversity 2000 State of Biodiversity: Western Cape Province, South Africa Mammals Peter H. Lloyd Scientific Services Western Cape Nature Conservation Board Private Bag 5014 STELLENBOSCH 7599 Introduction (1986) which was most heavily relied upon. The result of this exercise was a list of all the mammals known to occur The Western Cape Province (W.C.P.) of South Africa, one within the southern African subregion (a region defined of nine provinces forming the country, is generally loosely as that south of a line joining the Kunene and regarded in South African terms as being “poor” with Zambesi Rivers), at currently recognized (warts and all!) respect to the number of its mammals when compared to subspecific level. For reference purposes typical the other eight provinces. This misconception has its subspecies (i.e. those subspecies whose trinomial is the origin in the fact that the Western Cape, essentially a same as the specific epithet; in other words where the winter rainfall region, lies adjacent to one of the richest species and subspecies names are the same) were included spots in the world in terms of mammalian diversity, even if that typical subspecies did not occur within the namely the summer rainfall region of the rest of South subregion. For the purposes of this paper, with minor Africa. In fact only when compared with the rest of our exceptions, only species are discussed. own country can the Western Cape be regarded as having The next step was to identify from W.C.N.C.B. records, a “poor” mammofauna in terms of biodiversity.
    [Show full text]
  • New Antelope Fossils from Awash, Ethiopia, and Phylogenetic Analysis of Hippotragini (Bovidae, Mammalia)
    Palaeont. afr., 31,55-72 (1994) NEW ANTELOPE FOSSILS FROM AWASH, ETHIOPIA, AND PHYLOGENETIC ANALYSIS OF HIPPOTRAGINI (BOVIDAE, MAMMALIA) by ~E. S. Vrba1 and J. Gatesy2 lDepartment of Geology and Geophysics, Yale University, P.O. Box 6666, New Haven, CT 06511, U.S.A. 2Department of Invertebrates, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024, U.S .A. ABSTRACT New hippotragine antelope fossils from the Middle Awash, Ethiopia, include a new species Praedamalis howelli from Maka, an early species of Oryx from Matabaietu, and Hippotragus gigas from Gamedah. Our comparisons of these fossils with all other known fossil and Recent Hippotragini includes a cladistic analysis. The results suggest that the two major lineages in the monophyletic Hippotragini are Hippotragus including taxa from the Siwaliks previously ascribed to Sivatragus, and a Praedamalis - Oryx sister-group. Some evolutionary and biogeographic implications of the new fossils from the Middle A wash deposits are discussed. KEY WORDS: Hippotragini, Awash, Praedamalis INTRODUCTION introduced in Table 1, which also contains first and last The new fossils form part of a large assemblage of appearance data for all hippotragines. Plio-Pleistocene fossil vertebrates, including In a previous survey of Makapansgat Limeworks representatives of Hominidae, that has recently been Hippotragini (Vrba, 1987) it was suggested that recovered from sedimentary deposits in eastern Wellsiana torticornuta Vrba 1987 and Brabovus Ethiopia's Middle Awash Valley (White et al., 1994). nanincisivus Gentry 1987 might be hippotragine. The Awash faunas are of particular interest because Based on these tentative suggestions we included these their evolution was affected by pervasi ve tectonism and taxa in the cladistic analysis.
    [Show full text]
  • Extinct in the Wild Poster.Indd
    A Few African Species Extinctin the Wild Flickr.com John Gerrard Keulemans/Wikipedia.com John Gerrard Danny Barron/Flickr.com Danny Giant Tortoise Egyptian Barbary Sheep Seychelles Parakeet (Extinct) Cylindraspis Ammotragus lervia ornata Psittacula wardi Mauritius, Seychelles Eygpt Seychelles Giant tortoises were considered extremely The native range of the Egyptian Barbary sheep The Seychelles parakeet was endemic to Mahé valuable by early mariners for food as they could was the arid hills east of Cairo, Egypt, and the and Silhouette, two islands in the Seychelles John and Karen Hollingsworth, US Fish and Wildlife Service Wildlife and John and Karen Hollingsworth, US Fish Flickr.com G. Reclos/MCH survive for months in captivity without food and rugged terrain bordering both sides of the group. This small, primarily green parrot was Barbary Lion Pinstripe Dambo West African Black Rhino (Extinct) water. Their fl esh and oil was considered a cure Nile River in southern Egypt. It is thought that already rare when it was fi rst described by for scurvy. Sadly, thousands were wastefully the Egyptian Barbary sheep probably became Europeans in the 1860s. The Seychelles parakeet Panthera leo leo Paretroplus menarambo Diceros bicornis longipes harvested, with many specimens being left to extinct in the wild in the 1970s or 1980s. The was fi nally driven to extinction in 1906, largely North Africa Madagascar Central West Africa rot after their valuable liver and oil had been species does survive, however, in captive due to the clearing of its forest habitat for removed. Most sub-species became extinct in breeding programmes. coconut plantations and eradication eff orts to The Barbary (also called Atlas or Nubian) lion The pinstripe dambo was endemic to a Among two of Africa’s most threatened rhinoceros the early years of the 18th century.
    [Show full text]