Middle Jurassic Sanjianfang Formation, Turpan Basin) -A Preliminary Report
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Journal of Paleontology
Journal of Paleontology http://journals.cambridge.org/JPA Additional services for Journal of Paleontology: Email alerts: Click here Subscriptions: Click here Commercial reprints: Click here Terms of use : Click here New heterodontosaurid remains from the Cañadón Asfalto Formation: cursoriality and the functional importance of the pes in small heterodontosaurids Marcos G. Becerra, Diego Pol, Oliver W.M. Rauhut and Ignacio A. Cerda Journal of Paleontology / Volume 90 / Issue 03 / May 2016, pp 555 - 577 DOI: 10.1017/jpa.2016.24, Published online: 27 June 2016 Link to this article: http://journals.cambridge.org/abstract_S002233601600024X How to cite this article: Marcos G. Becerra, Diego Pol, Oliver W.M. Rauhut and Ignacio A. Cerda (2016). New heterodontosaurid remains from the Cañadón Asfalto Formation: cursoriality and the functional importance of the pes in small heterodontosaurids. Journal of Paleontology, 90, pp 555-577 doi:10.1017/jpa.2016.24 Request Permissions : Click here Downloaded from http://journals.cambridge.org/JPA, IP address: 190.172.49.57 on 16 Aug 2016 Journal of Paleontology, 90(3), 2016, p. 555–577 Copyright © 2016, The Paleontological Society 0022-3360/16/0088-0906 doi: 10.1017/jpa.2016.24 New heterodontosaurid remains from the Cañadón Asfalto Formation: cursoriality and the functional importance of the pes in small heterodontosaurids Marcos G. Becerra,1 Diego Pol,1 Oliver W.M. Rauhut,2 and Ignacio A. Cerda3 1CONICET- Museo Palaeontológico Egidio Feruglio, Fontana 140, Trelew, Chubut 9100, Argentina 〈[email protected]〉; 〈[email protected]〉 2SNSB, Bayerische Staatssammlung für Paläontologie und Geologie and Department of Earth and Environmental Sciences, LMU München, Richard-Wagner-Str. -
Eubrontes and Anomoepus Track
Sullivan, R.M. and Lucas, S.G., eds., 2016, Fossil Record 5. New Mexico Museum of Natural History and Science Bulletin 74. 345 EUBRONTES AND ANOMOEPUS TRACK ASSEMBLAGES FROM THE MIDDLE JURASSIC XIASHAXIMIAO FORMATION OF ZIZHONG COUNTY, SICHUAN, CHINA: REVIEW, ICHNOTAXONOMY AND NOTES ON PRESERVED TAIL TRACES LIDA XING1, MARTIN G. LOCKLEY2, GUANGZHAO PENG3, YONG YE3, JIANPING ZHANG1, MASAKI MATSUKAWA4, HENDRIK KLEIN5, RICHARD T. MCCREA6 and W. SCOTT PERSONS IV7 1School of the Earth Sciences and Resources, China University of Geosciences, Beijing 100083, China; -email: [email protected]; 2Dinosaur Trackers Research Group, University of Colorado Denver, P.O. Box 173364, Denver, CO 80217; 3 Zigong Dinosaur Museum, Zigong 643013, Sichuan, China; 4 Department of Environmental Sciences, Tokyo Gakugei University, Koganei, Tokyo 184-8501, Japan; 5 Saurierwelt Paläontologisches Museum Alte Richt 7, D-92318 Neumarkt, Germany; 6 Peace Region Palaeontology Research Centre, Box 1540, Tumbler Ridge, British Columbia V0C 2W0, Canada; 7 Department of Biological Sciences, University of Alberta 11455 Saskatchewan Drive, Edmonton, Alberta T6G 2E9, Canada Abstract—The Nianpanshan dinosaur tracksite, first studied in the 1980s, was designated as the type locality of the monospecific ichnogenus Jinlijingpus, and the source of another tridactyl track, Chuanchengpus, both presumably of theropod affinity. After the site was mapped in 2001, these two ichnotaxa were considered synonyms of Eubrontes and Anomoepus, respectively, the latter designation being the first identification of this ichnogenus in China. The assemblage indicates a typical Jurassic ichnofauna. The present study reinvestigates the site in the light of the purported new ichnospecies Chuanchengpus shenglingensis that was introduced in 2012. After re- evaluation of the morphological and extramorphological features, C. -
The Jurassic Fossil Wood Diversity from Western Liaoning, NE China
Jiang et al. Journal of Palaeogeography (2019) 8:1 https://doi.org/10.1186/s42501-018-0018-y Journal of Palaeogeography RESEARCH Open Access The Jurassic fossil wood diversity from western Liaoning, NE China Zi-Kun Jiang1,2, Yong-Dong Wang2,3*, Ning Tian4,5, Ao-Wei Xie2,6, Wu Zhang7, Li-Qin Li2 and Min Huang1 Abstract Western Liaoning is a unique region in China that bears diverse types of Jurassic plants, including leaves, fern rhizomes, and wood, providing significant proxy for vegetation and palaeoenvironment reconstruction of the well-known Yanliao Flora in East Asia. In particular, the silicified wood is very abundant in the fossil Lagerstätte of the Jurassic Tiaojishan Formation in Beipiao, western Liaoning. Previous and recent systematic investigations documented a high diversity of the Jurassic wood assemblages. These assemblages are dominated by conifers, followed by cycads and ginkgoaleans. In total, about 30 species belonging to 21 genera of fossil wood have been recorded so far, which are represented by Cycadopsida, Ginkgopsida, Coniferopsida, and Gymnospermae incertae sedis. The evolutionary implications of several distinctive fossil wood taxa as well as palaeoclimate implications are summarized based on their anatomical structures and growth ring patterns. This work approaches the vegetation development and evolutionary significances of the wood taxa and their relatives, and provides clues for the further understanding of the diversity of the Jurassic Yanliao Flora in East Asia. Keywords: Fossil wood, Diversity, Evolution, Tiaojishan Formation, Jurassic 1 Introduction 2004;Wangetal.,2009). Among these localities, western Fossil floras are a significant record for the vegetation Liaoning is a well-known fossil Lagerstätte with diverse and for the palaeoenvironment reconstructions of the and well-preserved fossil plant foliages and wood (Zhang Mesozoic. -
New Heterodontosaurid Remains from the Cañadón Asfalto Formation: Cursoriality and the Functional Importance of the Pes in Small Heterodontosaurids
Journal of Paleontology, 90(3), 2016, p. 555–577 Copyright © 2016, The Paleontological Society 0022-3360/16/0088-0906 doi: 10.1017/jpa.2016.24 New heterodontosaurid remains from the Cañadón Asfalto Formation: cursoriality and the functional importance of the pes in small heterodontosaurids Marcos G. Becerra,1 Diego Pol,1 Oliver W.M. Rauhut,2 and Ignacio A. Cerda3 1CONICET- Museo Palaeontológico Egidio Feruglio, Fontana 140, Trelew, Chubut 9100, Argentina 〈[email protected]〉; 〈[email protected]〉 2SNSB, Bayerische Staatssammlung für Paläontologie und Geologie and Department of Earth and Environmental Sciences, LMU München, Richard-Wagner-Str. 10, Munich 80333, Germany 〈[email protected]〉 3CONICET- Instituto de Investigación en Paleobiología y Geología, Universidad Nacional de Río Negro, Museo Carlos Ameghino, Belgrano 1700, Paraje Pichi Ruca (predio Marabunta), Cipolletti, Río Negro, Argentina 〈[email protected]〉 Abstract.—New ornithischian remains reported here (MPEF-PV 3826) include two complete metatarsi with associated phalanges and caudal vertebrae, from the late Toarcian levels of the Cañadón Asfalto Formation. We conclude that these fossil remains represent a bipedal heterodontosaurid but lack diagnostic characters to identify them at the species level, although they probably represent remains of Manidens condorensis, known from the same locality. Histological features suggest a subadult ontogenetic stage for the individual. A cluster analysis based on pedal measurements identifies similarities of this specimen with heterodontosaurid taxa and the inclusion of the new material in a phylogenetic analysis with expanded character sampling on pedal remains confirms the described specimen as a heterodontosaurid. Finally, uncommon features of the digits (length proportions among nonungual phalanges of digit III, and claw features) are also quantitatively compared to several ornithischians, theropods, and birds, suggesting that this may represent a bipedal cursorial heterodontosaurid with gracile and grasping feet and long digits. -
Zircon U-Pb SHRIMP Dating of the Yixian Formation in Sihetun, Northeast China
Cretaceous Research 28 (2007) 177e182 www.elsevier.com/locate/CretRes New evidence for Cretaceous age of the feathered dinosaurs of Liaoning: zircon U-Pb SHRIMP dating of the Yixian Formation in Sihetun, northeast China Wei Yang a, Shuguang Li a,*, Baoyu Jiang b a Department of Earth and Space Sciences, University of Science and Technology of China, Hefei 230026, China b Department of Earth Sciences, Nanjing University, Nanjing 210093, China Accepted 30 May 2006 Available online 25 January 2007 Abstract We present the first report of U-Pb SHRIMP dating of zircons from three tuffs interbedded within the ‘‘feathered dinosaur’’-bearing deposits in western Liaoning, China. One is a sample from the Bed 6 tuff (LX-SHT-12) of the Yixian Formation in Sihetun, for which our zircon U-Pb SHRIMP analyses gave a Cretaceous age (124.7 Æ 2.7 Ma), in agreement with a previously published sanidine 40Ar/39Ar age (124.60 Æ 0.25 Ma). The other two are from the Bed 1 tuff (LX-HBJ-1) and Bed 8 tuff (LX-HBJ-6) of the Yixian Formation in Huangbanjigou; the former gave an age of 124.9 Æ 1.7 Ma, the latter an age of 122.8 Æ 1.6 Ma. The three consistent ages indicate that the Yixian Formation was deposited in the Early Cretaceous within a very short time period (ca. 2 Ma). Ó 2006 Published by Elsevier Ltd. Keywords: Zircon U-Pb SHRIMP dating; Yixian Formation; Feathered dinosaurs; Liaoning; Jehol Biota 1. Introduction still the subject of disagreement. Previously K-Ar and Rb-Sr dat- ing results for the volcanic rocks of the formation have yielded Recently, a wide variety of spectacular fossils, including the Jurassic ages of 137 Æ 7 Ma and 142.5 Æ 4Ma(Wang and Diao, ‘‘feathered dinosaurs’’ Sinosauropteryx (Chen et al., 1998), Pro- 1984). -
At Carowinds
at Carowinds EDUCATOR’S GUIDE CLASSROOM LESSON PLANS & FIELD TRIP ACTIVITIES Table of Contents at Carowinds Introduction The Field Trip ................................... 2 The Educator’s Guide ....................... 3 Field Trip Activity .................................. 4 Lesson Plans Lesson 1: Form and Function ........... 6 Lesson 2: Dinosaur Detectives ....... 10 Lesson 3: Mesozoic Math .............. 14 Lesson 4: Fossil Stories.................. 22 Games & Puzzles Crossword Puzzles ......................... 29 Logic Puzzles ................................. 32 Word Searches ............................... 37 Answer Keys ...................................... 39 Additional Resources © 2012 Dinosaurs Unearthed Recommended Reading ................. 44 All rights reserved. Except for educational fair use, no portion of this guide may be reproduced, stored in a retrieval system, or transmitted in any form or by any Dinosaur Data ................................ 45 means—electronic, mechanical, photocopy, recording, or any other without Discovering Dinosaurs .................... 52 explicit prior permission from Dinosaurs Unearthed. Multiple copies may only be made by or for the teacher for class use. Glossary .............................................. 54 Content co-created by TurnKey Education, Inc. and Dinosaurs Unearthed, 2012 Standards www.turnkeyeducation.net www.dinosaursunearthed.com Curriculum Standards .................... 59 Introduction The Field Trip From the time of the first exhibition unveiled in 1854 at the Crystal -
A Phylogenetic Analysis of the Basal Ornithischia (Reptilia, Dinosauria)
A PHYLOGENETIC ANALYSIS OF THE BASAL ORNITHISCHIA (REPTILIA, DINOSAURIA) Marc Richard Spencer A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements of the degree of MASTER OF SCIENCE December 2007 Committee: Margaret M. Yacobucci, Advisor Don C. Steinker Daniel M. Pavuk © 2007 Marc Richard Spencer All Rights Reserved iii ABSTRACT Margaret M. Yacobucci, Advisor The placement of Lesothosaurus diagnosticus and the Heterodontosauridae within the Ornithischia has been problematic. Historically, Lesothosaurus has been regarded as a basal ornithischian dinosaur, the sister taxon to the Genasauria. Recent phylogenetic analyses, however, have placed Lesothosaurus as a more derived ornithischian within the Genasauria. The Fabrosauridae, of which Lesothosaurus was considered a member, has never been phylogenetically corroborated and has been considered a paraphyletic assemblage. Prior to recent phylogenetic analyses, the problematic Heterodontosauridae was placed within the Ornithopoda as the sister taxon to the Euornithopoda. The heterodontosaurids have also been considered as the basal member of the Cerapoda (Ornithopoda + Marginocephalia), the sister taxon to the Marginocephalia, and as the sister taxon to the Genasauria. To reevaluate the placement of these taxa, along with other basal ornithischians and more derived subclades, a phylogenetic analysis of 19 taxonomic units, including two outgroup taxa, was performed. Analysis of 97 characters and their associated character states culled, modified, and/or rescored from published literature based on published descriptions, produced four most parsimonious trees. Consistency and retention indices were calculated and a bootstrap analysis was performed to determine the relative support for the resultant phylogeny. The Ornithischia was recovered with Pisanosaurus as its basalmost member. -
The First Record of Anomoepus Tracks from the Middle Jurassic of Henan Province, Central China
Historical Biology An International Journal of Paleobiology ISSN: 0891-2963 (Print) 1029-2381 (Online) Journal homepage: http://www.tandfonline.com/loi/ghbi20 The first record of Anomoepus tracks from the Middle Jurassic of Henan Province, Central China Lida Xing, Nasrollah Abbassi, Martin G. Lockley, Hendrik Klein, Songhai Jia, Richard T. McCrea & W. Scott Persons IV To cite this article: Lida Xing, Nasrollah Abbassi, Martin G. Lockley, Hendrik Klein, Songhai Jia, Richard T. McCrea & W. Scott Persons IV (2016): The first record of Anomoepus tracks from the Middle Jurassic of Henan Province, Central China, Historical Biology, DOI: 10.1080/08912963.2016.1149480 To link to this article: http://dx.doi.org/10.1080/08912963.2016.1149480 Published online: 22 Feb 2016. Submit your article to this journal View related articles View Crossmark data Full Terms & Conditions of access and use can be found at http://www.tandfonline.com/action/journalInformation?journalCode=ghbi20 Download by: [Lida Xing] Date: 22 February 2016, At: 05:53 HISTORICAL BIOLOGY, 2016 http://dx.doi.org/10.1080/08912963.2016.1149480 The first record of Anomoepus tracks from the Middle Jurassic of Henan Province, Central China Lida Xinga, Nasrollah Abbassib, Martin G. Lockleyc, Hendrik Kleind, Songhai Jiae, Richard T. McCreaf and W. Scott PersonsIVg aSchool of the Earth Sciences and Resources, China University of Geosciences, Beijing, China; bFaculty of Sciences, Department of Geology, University of Zanjan, Zanjan, Iran; cDinosaur Tracks Museum, University of Colorado Denver, Denver, CO, USA; dSaurierwelt Paläontologisches Museum, Neumarkt, Germany; eHenan Geological Museum, Henan Province, China; fPeace Region Palaeontology Research Centre, British Columbia, Canada; gDepartment of Biological Sciences, University of Alberta, Edmonton, Canada ABSTRACT ARTICLE HISTORY Small, gracile mostly tridactyl tracks from the Middle Jurassic of Henan Province represent the first example Received 30 December 2015 of the ichnogenus Anomoepus from this region. -
Monster Polar Shift, Shifts Back: Paleoclimate and CA-ID-TIMS Evidence from Northern China
EGU2020-20827 https://doi.org/10.5194/egusphere-egu2020-20827 EGU General Assembly 2020 © Author(s) 2021. This work is distributed under the Creative Commons Attribution 4.0 License. Monster polar shift, shifts back: paleoclimate and CA-ID-TIMS evidence from northern China Paul Olsen1, Jingeng Sha2, Scott Maclennan3, Sean Kinney1, Yanan Fang2, Clara Chang1, Theo Kuhn1, Roger Fu4, Dennis Kent1, and Blair Schone5 1Columbia University, Lamont-Doherty Earth Observatory, Palisades, New York, United States of America ([email protected]) 2State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute for Geology and Paleontology and Center for Excellence in Life and Paleoenvironment, 210008 Nanjing, China 3Department of Earth and Environmental Sciences, University of Rochester, Rochester, New York, United States of America 4Department of Earth and Planetary Sciences, Harvard University, Cambridge, Massachusetts, United States of America 5Department of Geosciences, Princeton University, Princeton, New Jersey, United States of America The two great lacustrine fossil Konservat-Lagerstätten of northeastern China producing feathered dinosaurs, the Jurassic Yanliao Biota and the Jehol Biota, were deposited during relatively humid times and are separated by a major redbed interval, typified by the Tuchengzi Formation deposited under a much more arid climate (1). We present new zircon CA-TIMS U-Pb ages for the peaks of the Yanliao [~160 Ma] and the Jehol biotas [Yixian Fm ~125 Ma] constraining a shift in that region from a higher-latitude temperate zone to a lower-latitude semiarid zone consistent with a ~30° arc distance shift true polar wander shift (1, 2, 3). The Yanliao Biota and the Jehol Biota are preserved in remarkably similar facies almost lacking signs of desiccation, while the Tuchengzi Formation has abundant evidence for desiccation and even eolian dune sands. -
Ichnos: an International Journal for Plant and Animal Traces Sauropod
This article was downloaded by: [University of Alberta] On: 02 June 2014, At: 06:41 Publisher: Taylor & Francis Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: Mortimer House, 37-41 Mortimer Street, London W1T 3JH, UK Ichnos: An International Journal for Plant and Animal Traces Publication details, including instructions for authors and subscription information: http://www.tandfonline.com/loi/gich20 Sauropod and Small Theropod Tracks from the Lower Jurassic Ziliujing Formation of Zigong City, Sichuan, China, with an Overview of Triassic–Jurassic Dinosaur Fossils and Footprints of the Sichuan Basin Lida Xinga, Guangzhao Pengb, Yong Yeb, Martin G. Lockleyc, Hendrik Kleind, W. Scott Persons IVe, Jianping Zhanga, Chunkang Shub & Baoqiao Haob a School of the Earth Sciences and Resources, China University of Geosciences, Beijing, China b Zigong Dinosaur Museum, Zigong, Sichuan, China c Dinosaur Trackers Research Group, University of Colorado, Denver, Colorado, USA d Saurierwelt Paläontologisches Museum, Neumarkt, Germany e Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada Published online: 28 May 2014. To cite this article: Lida Xing, Guangzhao Peng, Yong Ye, Martin G. Lockley, Hendrik Klein, W. Scott Persons IV, Jianping Zhang, Chunkang Shu & Baoqiao Hao (2014) Sauropod and Small Theropod Tracks from the Lower Jurassic Ziliujing Formation of Zigong City, Sichuan, China, with an Overview of Triassic–Jurassic Dinosaur Fossils and Footprints of the Sichuan Basin, Ichnos: An International Journal for Plant and Animal Traces, 21:2, 119-130, DOI: 10.1080/10420940.2014.909352 To link to this article: http://dx.doi.org/10.1080/10420940.2014.909352 PLEASE SCROLL DOWN FOR ARTICLE Taylor & Francis makes every effort to ensure the accuracy of all the information (the “Content”) contained in the publications on our platform. -
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Biosis: Biological Systems (2021) 2(2), 271-282 https://doi.org/10.37819/biosis.002.02.0102 ORIGINAL RESEARCH Theropod tracks from the Jurassic–Cretaceous boundary, Tuchengzi Formation, Chengde, China: Review and new observations Lida Xing a, b, Martin G. Lockley c a State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Beijing 100083, China. Corresponding author. Lida Xing: [email protected] b School of the Earth Sciences and Resources, China University of Geosciences, Beijing 100083, China; c Dinosaur Trackers Research Group, University of Colorado Denver, PO Box 173364, Denver, CO 80217, USA. © The Authors, 2021 ABSTRACT ARTICLE HISTORY Previously known theropod dinosaur footprints preserved as natural Received: 10-02-2021 casts in the Tuchengzi Formation, on a rock wall beside the railway in Revised: 18-03-2021 Nanshuangmiao Village, Shangbancheng Town, Chengde City, were Accepted: 16-04-2021 originally assigned to ichnogenus Anchisauripus and tentatively attributed to oviraptosaurs. The assemblage was restudied in more detail KEYWORDS by examining the entire assemblage of 55 tracks associated with two Theropod tracks horizons. The size range of the 27 measured tracks suggests a more Anchisauripus diverse grallatorid–eubrontid assemblage and potentially greater Grallatorid diversity of theropod trackmakers. The label Anchisauripus, which has Eubrontid fallen into disuse in some recent literature, implies trackmakers of Oviraptosaurs medium shape and size in the grallatorid–eubrontid morphological spectrum. However, given the presence of other theropod ichnotaxa in the Jurassic to Early Cretaceous strata of the Tuchengzi Formation and time equivalent units we suggest that explicit reference to the Grallator- Anchisauripus-Eubrontes (GAE) plexus, or simply the term Grallator- Eubrontes plexus be confined to Lower Jurassic assemblages as originally defined and intended. -
Body-Size Evolution in the Dinosauria
8 Body-Size Evolution in the Dinosauria Matthew T. Carrano Introduction The evolution of body size and its influence on organismal biology have received scientific attention since the earliest decades of evolutionary study (e.g., Cope, 1887, 1896; Thompson, 1917). Both paleontologists and neontologists have attempted to determine correlations between body size and numerous aspects of life history, with the ultimate goal of docu- menting both the predictive and causal connections involved (LaBarbera, 1986, 1989). These studies have generated an appreciation for the thor- oughgoing interrelationships between body size and nearly every sig- nificant facet of organismal biology, including metabolism (Lindstedt & Calder, 1981; Schmidt-Nielsen, 1984; McNab, 1989), population ecology (Damuth, 1981; Juanes, 1986; Gittleman & Purvis, 1998), locomotion (Mc- Mahon, 1975; Biewener, 1989; Alexander, 1996), and reproduction (Alex- ander, 1996). An enduring focus of these studies has been Cope’s Rule, the notion that body size tends to increase over time within lineages (Kurtén, 1953; Stanley, 1973; Polly, 1998). Such an observation has been made regarding many different clades but has been examined specifically in only a few (MacFadden, 1986; Arnold et al., 1995; Jablonski, 1996, 1997; Trammer & Kaim, 1997, 1999; Alroy, 1998). The discordant results of such analyses have underscored two points: (1) Cope’s Rule does not apply universally to all groups; and (2) even when present, size increases in different clades may reflect very different underlying processes. Thus, the question, “does Cope’s Rule exist?” is better parsed into two questions: “to which groups does Cope’s Rule apply?” and “what process is responsible for it in each?” Several recent works (McShea, 1994, 2000; Jablonski, 1997; Alroy, 1998, 2000a, 2000b) have begun to address these more specific questions, attempting to quantify patterns of body-size evolution in a phylogenetic (rather than strictly temporal) context, as well as developing methods for interpreting the resultant patterns.